treatments-xml/data/CB/5E/87/CB5E87F7FA75FFE0FF69FDEF850EC515.xml
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<document id="B605D8FE74E97802A1F0102A06E33493" ID-DOI="10.11646/zootaxa.5296.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984382" ID-ZooBank="2EE722D2-463B-463F-8EA3-E9A099E85154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685428400234" checkinUser="plazi" docAuthor="Schawaller, Wolfgang &amp; Bellersheim, Aron" docDate="2023" docId="CB5E87F7FA75FFE0FF69FDEF850EC515" docLanguage="en" docName="zootaxa.5296.4.3.pdf" docOrigin="Zootaxa 5296 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laena brunkei Schawaller &amp; Bellersheim 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="541" masterDocId="3767FF8FFA74FFE1FFFEFFCC8013C040" masterDocTitle="New species and records of Laena Dejean (Coleoptera: Tenebrionidae: Lagriinae) from Vietnam, southeastern China, and Korea" masterLastPageNumber="550" masterPageNumber="540" pageNumber="541" updateTime="1685429945776" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="29DF12281A42E6E7FBCF103C82E26901">New species and records of Laena Dejean (Coleoptera: Tenebrionidae: Lagriinae) from Vietnam, southeastern China, and Korea</mods:title>
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<mods:namePart id="11E3881DD7E6DDFC96CD332C0EA14894">Schawaller, Wolfgang</mods:namePart>
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<mods:affiliation id="CB285CD09E830FA32D8F02698D64D2AC">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany &amp; schawaller. ehrenamt @ smns-bw. de; https: // orcid. org / 0000 - 0003 - 1482 - 7386</mods:affiliation>
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<mods:namePart id="82408D4190ABB09E8F57E4187AD4902C">Bellersheim, Aron</mods:namePart>
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<mods:affiliation id="B882C1AD63BD7C09B8DCD88A016EEE1C">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany &amp; bellersheim. aron @ smns-bw. de; https: // orcid. org / 0000 - 0002 - 5607 - 328 X</mods:affiliation>
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<taxonomicName id="84F74D62FA75FFE0FF69FDEF8159C27D" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[151,330,546,573]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="541" phylum="Arthropoda" rank="species" species="brunkei" status="sp. n.">
<emphasis id="7183EAF3FA75FFE0FF69FDEF8159C27D" bold="true" box="[151,330,546,573]" italics="true" pageId="1" pageNumber="541">Laena brunkei</emphasis>
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<paragraph id="434836E1FA75FFE0FF69FD8B810AC221" blockId="1.[151,403,546,609]" box="[151,281,583,609]" pageId="1" pageNumber="541">
(
<figureCitation id="DBCC2A64FA75FFE0FF61FD8B80FBC221" box="[159,232,583,609]" captionStart="FIGURES 17" captionStartId="2.[152,270,1801,1825]" captionTargetBox="[192,1388,207,1748]" captionTargetId="figure-17@2.[151,1436,179,1775]" captionTargetPageId="2" captionText="FIGURES 17. Dorsal view of Laena species from Vietnam and southeastern China. 1. L. faillei sp. n., ♀ paratype, Vietnam, SMNS. 2. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 3. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 4. L. hajeki sp. n., ♁ paratype, Guangdong, SMNS. 5. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 6. L. logshengica sp. n., ♁ paratype, Guangxi, NMPC. 7. L. honba sp. n., ♁ holotype, Vietnam, SMNS. Scale line: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984384" httpUri="https://zenodo.org/record/7984384/files/figure.png" pageId="1" pageNumber="541">Figs 2</figureCitation>
,
<figureCitation id="DBCC2A64FA75FFE0FF0AFD8B8102C221" box="[244,273,583,609]" captionStart="FIGURES 1120" captionStartId="5.[152,270,1254,1278]" captionTargetBox="[152,1435,185,1226]" captionTargetId="figure-20@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURES 1120. Aedeagi of Laena species from Vietnam and southeastern China. 11. L. tramtonica sp. n., ♁ holotype, Vietnam, NHMW. 12. L. tamdaoensis Masumoto, 1996, non-type, Vietnam, NHMW. 13. L. fanjingshanana Ren &amp; Hua, 2006, ♁ non-type, Guangxi, SMNS. 14. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 15. L. faillei sp. n., ♁ holotype, Vietnam, SMNS. 16. L. logshengica sp. n., ♁ holotype, Guangxi, NMPC. 17. L. honba sp. n., ♁ holotype, Vietnam, SMNS. 18. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 19. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 20. L. hajeki sp. n., ♁ holotype, Guangdong, NMPC. Scale line: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984388" httpUri="https://zenodo.org/record/7984388/files/figure.png" pageId="1" pageNumber="541">18</figureCitation>
)
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<emphasis id="7183EAF3FA75FFE0FF69FD438156C2E8" bold="true" box="[151,325,654,681]" pageId="1" pageNumber="541">Type material.</emphasis>
<materialsCitation id="F39F3CBCFA75FFE0FEB2FD438269C28D" collectingDate="2019-05-17" collectingDateMax="2019-05-18" collectingDateMin="2019-05-17" collectionCode="NHMW" collectorName="Pia Ouac NP &amp; A. J. Brunke &amp; H. Schillhammer" country="Vietnam" elevation="1800" location="Cao Bang Prov." pageId="1" pageNumber="541" specimenCount="1" stateProvince="Cao Bang" typeStatus="holotype">
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<typeStatus id="9C4C8843FA75FFE0FEB2FD4381AEC2E9" box="[332,445,655,681]" pageId="1" pageNumber="541">Holotype</typeStatus>
:
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♁, N
<collectingCountry id="3BE07671FA75FFE0FDF8FD43827BC2E9" box="[518,616,655,681]" name="Vietnam" pageId="1" pageNumber="541">Vietnam</collectingCountry>
,
<collectingRegion id="8133F803FA75FFE0FD8DFD438335C2E9" box="[627,806,655,681]" country="Vietnam" name="Cao Bang" pageId="1" pageNumber="541">Cao Bang Prov.</collectingRegion>
,
<collectorName id="EE025337FA75FFE0FCCFFD4383D2C2E9" box="[817,961,655,681]" pageId="1" pageNumber="541">Pia Ouac NP</collectorName>
, summit road,
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<elevation id="C8DAD1D2FA75FFE0FB98FD4384AEC2E9" box="[1126,1213,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="541" unit="m" value="1800.0">1800 m</elevation>
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,
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</date>
, leg.
<collectorName id="EE025337FA75FFE0FF69FD7F8134C28D" box="[151,295,691,717]" pageId="1" pageNumber="541">A. J. Brunke</collectorName>
&amp;
<collectorName id="EE025337FA75FFE0FEB0FD7F8202C28D" box="[334,529,691,717]" pageId="1" pageNumber="541">H. Schillhammer</collectorName>
,
<collectionCode id="25E6AE24FA75FFE0FDE3FD7F8269C28D" box="[541,634,691,717]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="541" type="Museum">NHMW</collectionCode>
</materialsCitation>
.
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<typeStatus id="9C4C8843FA75FFE0FD7BFD7F82ECC28D" box="[645,767,691,717]" pageId="1" pageNumber="541">Paratypes</typeStatus>
.
</emphasis>
Same data as for holotype,
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,
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<collectionCode id="25E6AE24FA75FFE0FAD3FD7F8568C28D" box="[1325,1403,691,717]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="541">SMNS</collectionCode>
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.—
<materialsCitation id="F39F3CBCFA75FFE0FF69FD1B8467C2B1" box="[151,1140,726,753]" collectingDate="2014-06-05" collectingDateMax="2014-06-11" collectingDateMin="2014-06-05" collectionCode="SMNS" collectorName="C. Reuter" country="China" elevation="1900" location="Fanjing Shan" pageId="1" pageNumber="541" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<collectingCountry id="3BE07671FA75FFE0FF69FD1B80CEC2B1" box="[151,221,727,753]" name="China" pageId="1" pageNumber="541">China</collectingCountry>
,
<collectingRegion id="8133F803FA75FFE0FF16FD1B8158C2B1" box="[232,331,727,753]" country="China" name="Guizhou" pageId="1" pageNumber="541">Guizhou</collectingRegion>
,
<location id="4628603AFA75FFE0FEA9FD1B81FDC2B1" LSID="urn:lsid:plazi:treatment:CB5E87F7FA75FFE0FF69FDEF850EC515:4628603AFA75FFE0FEA9FD1B81FDC2B1" box="[343,494,727,753]" country="China" name="Fanjing Shan" pageId="1" pageNumber="541" stateProvince="Guizhou">Fanjing Shan</location>
,
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, leg.
<collectorName id="EE025337FA75FFE0FC85FD1B83F4C2B1" box="[891,999,727,753]" pageId="1" pageNumber="541">C. Reuter</collectorName>
,
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.
</paragraph>
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<paragraph id="434836E1FA75FFE0FF39FD368113C419" blockId="1.[151,1437,654,1365]" pageId="1" pageNumber="541">
<emphasis id="7183EAF3FA75FFE0FF39FD36814AC354" bold="true" box="[199,345,762,788]" pageId="1" pageNumber="541">Description.</emphasis>
Body length
<quantity id="840F9B04FA75FFE0FE07FD378296C355" box="[505,645,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.7" metricValueMin="3.7" pageId="1" pageNumber="541" unit="mm" value="4.2" valueMax="4.7" valueMin="3.7">3.74.7 mm</quantity>
, dorsal surface light brown without metallic reflection. Eyes weakly prominent. Antennomere 1 unusually enlarged. Pronotum round, widest across before middle, posterior margin similar wide as the anterior one; with large and confluent punctures, distance 0.51 puncture diameters, some punctures with short erect setae; surface convex and shining, disc without impressions; lateral margins not beaded; posterior corners rounded; prothoracic hypomera with similar punctures as on pronotum, but without visible setation. Elytra ovate with irregular punctation, not arranged in regular rows and intervals, punctures large as on pronotum but not confluent, punctures adjacent to suture forming two irregular rows; some punctures with short erect setae; lateral intervals without setiferous pores; surface shining. Apex of elytra not prolonged. All femora without teeth or sharp angles. Tibiae of both sexes simple, without peculiarities. Apicale of aedeagus triangular with rounded apex (
<figureCitation id="DBCC2A64FA75FFE0FF61FBF380E0C419" box="[159,243,1087,1113]" captionStart="FIGURES 1120" captionStartId="5.[152,270,1254,1278]" captionTargetBox="[152,1435,185,1226]" captionTargetId="figure-20@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURES 1120. Aedeagi of Laena species from Vietnam and southeastern China. 11. L. tramtonica sp. n., ♁ holotype, Vietnam, NHMW. 12. L. tamdaoensis Masumoto, 1996, non-type, Vietnam, NHMW. 13. L. fanjingshanana Ren &amp; Hua, 2006, ♁ non-type, Guangxi, SMNS. 14. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 15. L. faillei sp. n., ♁ holotype, Vietnam, SMNS. 16. L. logshengica sp. n., ♁ holotype, Guangxi, NMPC. 17. L. honba sp. n., ♁ holotype, Vietnam, SMNS. 18. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 19. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 20. L. hajeki sp. n., ♁ holotype, Guangdong, NMPC. Scale line: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984388" httpUri="https://zenodo.org/record/7984388/files/figure.png" pageId="1" pageNumber="541">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA75FFE0FF39FBAE859BC571" pageId="1" pageNumber="541" type="diagnosis">
<paragraph id="434836E1FA75FFE0FF39FBAE859BC571" blockId="1.[151,1437,654,1365]" pageId="1" pageNumber="541">
<emphasis id="7183EAF3FA75FFE0FF39FBAE8151C43C" bold="true" box="[199,322,1122,1148]" pageId="1" pageNumber="541">Diagnosis.</emphasis>
<taxonomicName id="84F74D62FA75FFE0FEB9FBAF81ABC43D" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[327,440,1123,1149]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="541" phylum="Arthropoda" rank="species" species="brunkei" status="sp. n.">
<emphasis id="7183EAF3FA75FFE0FEB9FBAF81ABC43D" box="[327,440,1123,1149]" italics="true" pageId="1" pageNumber="541">L. brunkei</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA75FFE0FE43FBAF81E8C43D" bold="true" box="[445,507,1123,1149]" pageId="1" pageNumber="541">
<taxonomicNameLabel id="6AB05788FA75FFE0FE43FBAF81E8C43D" box="[445,507,1123,1149]" pageId="1" pageNumber="541" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
shares with
<taxonomicName id="84F74D62FA75FFE0FD7CFBAF83F6C43D" authority="Schawaller, 1998" authorityName="Schawaller" authorityYear="1998" box="[642,997,1123,1149]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="541" phylum="Arthropoda" rank="species" species="burckhardti">
<emphasis id="7183EAF3FA75FFE0FD7CFBAF8333C43D" box="[642,800,1123,1149]" italics="true" pageId="1" pageNumber="541">L. burckhardti</emphasis>
<bibRefCitation id="27664B10FA75FFE0FCDBFBAF83F6C43D" author="Schawaller, W." box="[805,997,1123,1149]" pageId="1" pageNumber="541" pagination="375 - 382" refId="ref6232" refString="Schawaller, W. (1998) The genus Laena Latreille (Coleoptera: Tenebrionidae) in Thailand, with descriptions of new species. Revue suisse de Zoologie, 105, 375 - 382. https: // doi. org / 10.5962 / bhl. part. 80043" type="journal article" year="1998">Schawaller, 1998</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="3BE07671FA75FFE0FBDBFBAF849AC43D" box="[1061,1161,1123,1149]" name="Thailand" pageId="1" pageNumber="541">Thailand</collectingCountry>
and
<taxonomicName id="84F74D62FA75FFE0FB42FBAF80DCC4E1" authority="Schawaller, 2014" authorityName="Schawaller" authorityYear="2014" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="541" phylum="Arthropoda" rank="species" species="kresli">
<emphasis id="7183EAF3FA75FFE0FB42FBAF8506C43D" box="[1212,1301,1123,1149]" italics="true" pageId="1" pageNumber="541">L. kresli</emphasis>
<bibRefCitation id="27664B10FA75FFE0FAE4FBAF80DCC4E1" author="Schawaller, W." pageId="1" pageNumber="541" pagination="287 - 298" refId="ref6328" refString="Schawaller, W. (2014) New species of the genus Laena Dejean (Coleoptera: Tenebrionidae) from southeastern Asia (2). Annales Zoologici, 64, 287 - 298. https: // doi. org / 10.3161 / 000345414 x 682490" type="journal article" year="2014">Schawaller, 2014</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="3BE07671FA75FFE0FEE4FB4B8143C4E1" box="[282,336,1159,1185]" name="Laos" pageId="1" pageNumber="541">Laos</collectingCountry>
(known only from the
<typeStatus id="9C4C8843FA75FFE0FD9DFB4B8280C4E1" box="[611,659,1159,1185]" pageId="1" pageNumber="541">type</typeStatus>
loacalties) the small body size, the unarmed femora and the elytra with irregular punctation without separation into punctural rows and intervals. Additional species with irregularly punctured elytra are so far unknown in southeastern Asia.
<taxonomicName id="84F74D62FA75FFE0FCD7FB03838DC4A9" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[809,926,1231,1257]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="541" phylum="Arthropoda" rank="species" species="brunkei" status="sp. n.">
<emphasis id="7183EAF3FA75FFE0FCD7FB03838DC4A9" box="[809,926,1231,1257]" italics="true" pageId="1" pageNumber="541">L. brunkei</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA75FFE0FC5BFB0383F6C4A9" bold="true" box="[933,997,1231,1257]" pageId="1" pageNumber="541">
<taxonomicNameLabel id="6AB05788FA75FFE0FC5BFB0383F6C4A9" box="[933,997,1231,1257]" pageId="1" pageNumber="541" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be separated from both by lacking elytral setiferous pores, and by rough and confluent pronotal punctures. The aedeagi of these three species are similar, compare the figures in
<bibRefCitation id="27664B10FA75FFE0FE0AFADB82AFC571" author="Schawaller, W." box="[500,700,1303,1329]" pageId="1" pageNumber="541" pagination="375 - 382" refId="ref6232" refString="Schawaller, W. (1998) The genus Laena Latreille (Coleoptera: Tenebrionidae) in Thailand, with descriptions of new species. Revue suisse de Zoologie, 105, 375 - 382. https: // doi. org / 10.5962 / bhl. part. 80043" type="journal article" year="1998">Schawaller (1998</bibRefCitation>
,
<bibRefCitation id="27664B10FA75FFE0FD39FADB8317C571" author="Schawaller, W." box="[711,772,1303,1329]" pageId="1" pageNumber="541" pagination="287 - 298" refId="ref6328" refString="Schawaller, W. (2014) New species of the genus Laena Dejean (Coleoptera: Tenebrionidae) from southeastern Asia (2). Annales Zoologici, 64, 287 - 298. https: // doi. org / 10.3161 / 000345414 x 682490" type="journal article" year="2014">2014</bibRefCitation>
). See also figures and diagnosis of
<taxonomicName id="84F74D62FA75FFE0FB73FADB84E3C571" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[1165,1264,1303,1329]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="541" phylum="Arthropoda" rank="species" species="honba" status="sp. n.">
<emphasis id="7183EAF3FA75FFE0FB73FADB84E3C571" box="[1165,1264,1303,1329]" italics="true" pageId="1" pageNumber="541">L. honba</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA75FFE0FB09FADB8524C571" bold="true" box="[1271,1335,1303,1329]" pageId="1" pageNumber="541">
<taxonomicNameLabel id="6AB05788FA75FFE0FB09FADB8524C571" box="[1271,1335,1303,1329]" pageId="1" pageNumber="541" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
below.
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA75FFE0FF39FAF7850EC515" box="[199,1309,1339,1365]" pageId="1" pageNumber="541" type="etymology">
<paragraph id="434836E1FA75FFE0FF39FAF7850EC515" blockId="1.[151,1437,654,1365]" box="[199,1309,1339,1365]" pageId="1" pageNumber="541">
<emphasis id="7183EAF3FA75FFE0FF39FAF7815DC515" bold="true" box="[199,334,1339,1365]" pageId="1" pageNumber="541">Etymology.</emphasis>
Named in honour of Adam J. Brunke (Ottawa), one of the collectors of the
<typeStatus id="9C4C8843FA75FFE0FB60FAF784DDC515" box="[1182,1230,1339,1365]" pageId="1" pageNumber="541">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>