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<document id="A1233FAEFB2729F94B3260723B6F0D82" ID-DOI="10.11646/zootaxa.5296.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984382" ID-ZooBank="2EE722D2-463B-463F-8EA3-E9A099E85154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685428400234" checkinUser="plazi" docAuthor="Schawaller, Wolfgang &amp; Bellersheim, Aron" docDate="2023" docId="CB5E87F7FA73FFE9FF69FA0C846DC15D" docLanguage="en" docName="zootaxa.5296.4.3.pdf" docOrigin="Zootaxa 5296 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laena lianhua Schawaller &amp; Bellersheim 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="548" masterDocId="3767FF8FFA74FFE1FFFEFFCC8013C040" masterDocTitle="New species and records of Laena Dejean (Coleoptera: Tenebrionidae: Lagriinae) from Vietnam, southeastern China, and Korea" masterLastPageNumber="550" masterPageNumber="540" pageNumber="547" updateTime="1685429945776" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="603EF286E96EF77C156C76C919CF0AF8">New species and records of Laena Dejean (Coleoptera: Tenebrionidae: Lagriinae) from Vietnam, southeastern China, and Korea</mods:title>
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<mods:name id="9BFF261B6F584A185AEB35FE9E5998C6" type="personal">
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<mods:roleTerm id="1D3DCA46F5D3200510B253411F8DC71B">Author</mods:roleTerm>
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<mods:namePart id="E2ACC9041D1C25DB4C59F946B04CFA1B">Schawaller, Wolfgang</mods:namePart>
<mods:nameIdentifier id="1DAC75DEB1AAE40A61D00E3FE3B52D38" type="ORCID">0000-0003-1482-7386</mods:nameIdentifier>
<mods:affiliation id="9ECD11889AA6ACD0BB23B529EBC7317D">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany &amp; schawaller. ehrenamt @ smns-bw. de; https: // orcid. org / 0000 - 0003 - 1482 - 7386</mods:affiliation>
<mods:nameIdentifier id="846A01A7A8561D7466BA9C482783249E" type="email">schawaller.ehrenamt@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="A800A51137FF8950A3B049E2A420C01F">Bellersheim, Aron</mods:namePart>
<mods:nameIdentifier id="1125A160A1E5940F27A2E9585AA63A9A" type="ORCID">0000-0002-5607-328X</mods:nameIdentifier>
<mods:affiliation id="24EEF7DFEC8C39AF2944E1FE25341B81">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany &amp; bellersheim. aron @ smns-bw. de; https: // orcid. org / 0000 - 0002 - 5607 - 328 X</mods:affiliation>
<mods:nameIdentifier id="5EC463157D005C0AF00798DAAA1CCD54" type="email">bellersheim.aron@smns-bw.de</mods:nameIdentifier>
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<mods:title id="D7200A9EE0E9FE9472BF4B02158F236E">Zootaxa</mods:title>
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<mods:date id="ABFE6A71D06F6824DA16F3A6BB956F28">2023</mods:date>
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<mods:number id="958A39C8D59A5E0CAF0D4E6C37EC1482">2023-05-30</mods:number>
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<mods:number id="DA08B3D934D242595453A6CBF887DFFB">5296</mods:number>
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<mods:number id="9790A8798919D6F2DCA5F84E244F460C">4</mods:number>
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<treatment id="CB5E87F7FA73FFE9FF69FA0C846DC15D" ID-DOI="http://doi.org/10.5281/zenodo.7984291" ID-Zenodo-Dep="7984291" LSID="urn:lsid:plazi:treatment:CB5E87F7FA73FFE9FF69FA0C846DC15D" httpUri="http://treatment.plazi.org/id/CB5E87F7FA73FFE9FF69FA0C846DC15D" lastPageId="8" lastPageNumber="548" pageId="7" pageNumber="547">
<subSubSection id="0BED656AFA73FFE6FF69FA0C8180C59A" box="[151,403,1471,1498]" pageId="7" pageNumber="547" type="nomenclature">
<paragraph id="434836E1FA73FFE6FF69FA0C8180C59A" blockId="7.[151,403,1471,1534]" box="[151,403,1471,1498]" pageId="7" pageNumber="547">
<heading id="1800818DFA73FFE6FF69FA0C8180C59A" bold="true" box="[151,403,1471,1498]" fontSize="11" level="1" pageId="7" pageNumber="547" reason="1">
<emphasis id="7183EAF3FA73FFE6FF69FA0C8180C59A" bold="true" box="[151,403,1471,1498]" pageId="7" pageNumber="547">
<taxonomicName id="84F74D62FA73FFE6FF69FA0C8159C59A" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[151,330,1471,1498]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="547" phylum="Arthropoda" rank="species" species="lianhua" status="sp. n.">
<emphasis id="7183EAF3FA73FFE6FF69FA0C8159C59A" bold="true" box="[151,330,1471,1498]" italics="true" pageId="7" pageNumber="547">Laena lianhua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AB05788FA73FFE6FEAFFA0C8180C59A" box="[337,403,1472,1498]" pageId="7" pageNumber="547" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA73FFE6FF69FA288109C5BE" box="[151,282,1508,1534]" pageId="7" pageNumber="547" type="description">
<paragraph id="434836E1FA73FFE6FF69FA288109C5BE" blockId="7.[151,403,1471,1534]" box="[151,282,1508,1534]" pageId="7" pageNumber="547">
(
<figureCitation id="DBCC2A64FA73FFE6FF61FA2880F4C5BE" box="[159,231,1508,1534]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1801,1825]" captionTargetBox="[192,1388,207,1748]" captionTargetId="figure-17@2.[151,1436,179,1775]" captionTargetPageId="2" captionText="FIGURES 1–7. Dorsal view of Laena species from Vietnam and southeastern China. 1. L. faillei sp. n., ♀ paratype, Vietnam, SMNS. 2. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 3. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 4. L. hajeki sp. n., ♁ paratype, Guangdong, SMNS. 5. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 6. L. logshengica sp. n., ♁ paratype, Guangxi, NMPC. 7. L. honba sp. n., ♁ holotype, Vietnam, SMNS. Scale line: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984384" httpUri="https://zenodo.org/record/7984384/files/figure.png" pageId="7" pageNumber="547">Figs 3</figureCitation>
,
<figureCitation id="DBCC2A64FA73FFE6FF0AFA288102C5BE" box="[244,273,1508,1534]" captionStart="FIGURES 11–20" captionStartId="5.[152,270,1254,1278]" captionTargetBox="[152,1435,185,1226]" captionTargetId="figure-20@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURES 11–20. Aedeagi of Laena species from Vietnam and southeastern China. 11. L. tramtonica sp. n., ♁ holotype, Vietnam, NHMW. 12. L. tamdaoensis Masumoto, 1996, non-type, Vietnam, NHMW. 13. L. fanjingshanana Ren &amp; Hua, 2006, ♁ non-type, Guangxi, SMNS. 14. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 15. L. faillei sp. n., ♁ holotype, Vietnam, SMNS. 16. L. logshengica sp. n., ♁ holotype, Guangxi, NMPC. 17. L. honba sp. n., ♁ holotype, Vietnam, SMNS. 18. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 19. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 20. L. hajeki sp. n., ♁ holotype, Guangdong, NMPC. Scale line: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984388" httpUri="https://zenodo.org/record/7984388/files/figure.png" pageId="7" pageNumber="547">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA73FFE6FF69F9E082AAC62A" pageId="7" pageNumber="547" type="materials_examined">
<paragraph id="434836E1FA73FFE6FF69F9E082AAC62A" blockId="7.[151,1437,1580,2039]" pageId="7" pageNumber="547">
<emphasis id="7183EAF3FA73FFE6FF69F9E08155C606" bold="true" box="[151,326,1580,1606]" pageId="7" pageNumber="547">Type material.</emphasis>
<materialsCitation id="F39F3CBCFA73FFE6FEB0F9E082AAC62A" collectingDate="2004-05-30" collectingDateMax="2004-06-06" collectingDateMin="2004-05-30" collectionCode="SMNS" collectorName="J. Turna" country="China" location="Lianhua Forest Park" pageId="7" pageNumber="547" specimenCount="1" stateProvince="Jiangxi" typeStatus="holotype">
<emphasis id="7183EAF3FA73FFE6FEB0F9E081D7C606" bold="true" box="[334,452,1580,1606]" pageId="7" pageNumber="547">
<typeStatus id="9C4C8843FA73FFE6FEB0F9E081ACC606" box="[334,447,1580,1606]" pageId="7" pageNumber="547">Holotype</typeStatus>
:
</emphasis>
♁,
<collectingCountry id="3BE07671FA73FFE6FE0EF9E08225C606" box="[496,566,1580,1606]" name="China" pageId="7" pageNumber="547">China</collectingCountry>
, N
<collectingRegion id="8133F803FA73FFE6FDA3F9E082A2C606" box="[605,689,1580,1606]" country="China" name="Jiangxi" pageId="7" pageNumber="547">Jiangxi</collectingRegion>
,
<locationDeviation id="52FA2FC3FA73FFE6FD43F9E08382C606" box="[701,913,1580,1607]" location="Jiujiang" pageId="7" pageNumber="547">
<quantity id="840F9B04FA73FFE6FD43F9E08316C606" box="[701,773,1580,1607]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="547" unit="km" value="15.0">15 km</quantity>
SE Jiujiang
</locationDeviation>
,
<location id="4628603AFA73FFE6FC63F9E08497C606" LSID="urn:lsid:plazi:treatment:CB5E87F7FA73FFE9FF69FA0C846DC15D:4628603AFA73FFE6FC63F9E08497C606" box="[925,1156,1580,1607]" country="China" name="Lianhua Forest Park" pageId="7" pageNumber="547" stateProvince="Jiangxi">Lianhua Forest Park</location>
,
<date id="37491021FA73FFE6FB6EF9E0855EC606" box="[1168,1357,1580,1606]" pageId="7" pageNumber="547" value="2004-05-30" valueMax="2004-06-06" valueMin="2004-05-30">
<collectingDate id="270DE9C9FA73FFE6FB6EF9E0855EC606" box="[1168,1357,1580,1606]" pageId="7" pageNumber="547" value="2004-05-30" valueMax="2004-06-06" valueMin="2004-05-30">30.V.–6.VI.2004</collectingDate>
</date>
, leg.
<collectorName id="EE025337FA73FFE6FA74F9E180CFC62B" pageId="7" pageNumber="547">J. Turna</collectorName>
,
<collectionCode id="25E6AE24FA73FFE6FF19F99C8120C62A" box="[231,307,1616,1642]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="7" pageNumber="547">SMNS</collectionCode>
(antennae and legs partly broken).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA73FFE6FF39F9B881C5C792" pageId="7" pageNumber="547" type="description">
<paragraph id="434836E1FA73FFE6FF39F9B881C5C792" blockId="7.[151,1437,1580,2039]" pageId="7" pageNumber="547">
<emphasis id="7183EAF3FA73FFE6FF39F9B8814AC6CE" bold="true" box="[199,345,1652,1678]" pageId="7" pageNumber="547">Description.</emphasis>
Body length
<quantity id="840F9B04FA73FFE6FE0FF9B88259C6CF" box="[497,586,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="547" unit="mm" value="5.8">5.8 mm</quantity>
, dorsal surface dark brown without metallic reflection. Eyes not prominent. Pronotum cordiform, widest across anterior third, posterior margin only somewhat narrower than the anterior one; with separated large punctures, distance 1–3 puncture diameters, most punctures with long erect setae; surface convex and shining, disc basally with weak impression; lateral margins not beaded; posterior corners rounded; prothoracic hypomera with similar punctures as on pronotum, but without visible setation. Elytra ovate with rows of punctures in weak striae, punctures of rows of similar size as punctures on pronotum, punctures with long erect setae; intervals with a few punctures without setae, interval 9 with 3 indistinct setiferous pores, internal intervals feebly convex, external intervals distinctly convex; surface shining. Apex of elytra not prolonged. All femora with small but distinct teeth. Tibiae in males simple, without peculiarities.Apicale of aedeagus long and narrow triangular with acute apex, see
<figureCitation id="DBCC2A64FA73FFE6FE7EF87481C1C792" box="[384,466,1976,2002]" captionStart="FIGURES 11–20" captionStartId="5.[152,270,1254,1278]" captionTargetBox="[152,1435,185,1226]" captionTargetId="figure-20@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURES 11–20. Aedeagi of Laena species from Vietnam and southeastern China. 11. L. tramtonica sp. n., ♁ holotype, Vietnam, NHMW. 12. L. tamdaoensis Masumoto, 1996, non-type, Vietnam, NHMW. 13. L. fanjingshanana Ren &amp; Hua, 2006, ♁ non-type, Guangxi, SMNS. 14. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 15. L. faillei sp. n., ♁ holotype, Vietnam, SMNS. 16. L. logshengica sp. n., ♁ holotype, Guangxi, NMPC. 17. L. honba sp. n., ♁ holotype, Vietnam, SMNS. 18. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 19. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 20. L. hajeki sp. n., ♁ holotype, Guangdong, NMPC. Scale line: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984388" httpUri="https://zenodo.org/record/7984388/files/figure.png" pageId="7" pageNumber="547">Fig. 14</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA73FFE9FF39F8108121C0B9" lastPageId="8" lastPageNumber="548" pageId="7" pageNumber="547" type="diagnosis">
<paragraph id="434836E1FA73FFE9FF39F8108121C0B9" blockId="7.[151,1437,1580,2039]" lastBlockId="8.[151,1437,151,285]" lastPageId="8" lastPageNumber="548" pageId="7" pageNumber="547">
<emphasis id="7183EAF3FA73FFE6FF39F8108151C7B6" bold="true" box="[199,322,2012,2038]" pageId="7" pageNumber="547">Diagnosis.</emphasis>
<taxonomicName id="84F74D62FA73FFE6FEB5F81181D1C7B6" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[331,450,2012,2038]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="547" phylum="Arthropoda" rank="species" species="lianhua" status="sp. n.">
<emphasis id="7183EAF3FA73FFE6FEB5F81181D1C7B6" box="[331,450,2012,2038]" italics="true" pageId="7" pageNumber="547">L. lianhua</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA73FFE6FE35F810821EC7B6" bold="true" box="[459,525,2012,2038]" pageId="7" pageNumber="547">
<taxonomicNameLabel id="6AB05788FA73FFE6FE35F810821EC7B6" box="[459,525,2012,2038]" pageId="7" pageNumber="547" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
shares with
<taxonomicName id="84F74D62FA73FFE6FD5CF81183F0C7B6" authority="Schawaller, 2008" authorityName="Schawaller" authorityYear="2008" box="[674,995,2012,2038]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="547" phylum="Arthropoda" rank="species" species="hlavaci">
<emphasis id="7183EAF3FA73FFE6FD5CF8118306C7B6" box="[674,789,2012,2038]" italics="true" pageId="7" pageNumber="547">L. hlavaci</emphasis>
Schawaller, 2008
</taxonomicName>
also from
<collectingRegion id="8133F803FA73FFE6FB9BF81084A5C7B6" box="[1125,1206,2012,2038]" country="China" name="Jiangxi" pageId="7" pageNumber="547">Jiangxi</collectingRegion>
the armature of the femora, the dorsal erect setation, and the unbeaded lateral margins of the pronotum, but in
<taxonomicName id="84F74D62FA7CFFE9FB4FFF5B8535C0F1" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[1201,1318,151,177]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="548" phylum="Arthropoda" rank="species" species="lianhua" status="sp. n.">
<emphasis id="7183EAF3FA7CFFE9FB4FFF5B8535C0F1" box="[1201,1318,151,177]" italics="true" pageId="8" pageNumber="548">L. lianhua</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA7CFFE9FAD1FF5B8562C0F1" bold="true" box="[1327,1393,151,177]" pageId="8" pageNumber="548">
<taxonomicNameLabel id="6AB05788FA7CFFE9FAD1FF5B8562C0F1" box="[1327,1393,151,177]" pageId="8" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the elytra are broader, the pronotum is also broader with denser punctation, and the apicale of the aedeagus is longer and narrower.
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA7CFFE9FF39FECF846DC15D" box="[199,1150,259,285]" pageId="8" pageNumber="548" type="etymology">
<paragraph id="434836E1FA7CFFE9FF39FECF846DC15D" blockId="8.[151,1437,151,285]" box="[199,1150,259,285]" pageId="8" pageNumber="548">
<emphasis id="7183EAF3FA7CFFE9FF39FECF815DC15D" bold="true" box="[199,334,259,285]" pageId="8" pageNumber="548">Etymology.</emphasis>
Named after the Lianhua Forest Park, where the
<typeStatus id="9C4C8843FA7CFFE9FC88FECF83CBC15D" box="[886,984,259,285]" pageId="8" pageNumber="548">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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