210 lines
21 KiB
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210 lines
21 KiB
XML
<document id="0FFDAD9C96FE16BFD2D2A21AD20A4EC9" ID-CLB-Dataset="30929" ID-DOI="http://dx.doi.org/10.3897/zookeys.732.21677" ID-GBIF-Dataset="05db50e8-b4e5-422a-b78f-9676d74ac319" ID-PMC="PMC5799741" ID-Pensoft-Pub="1313-2970-732-1" ID-PubMed="29416404" ID-ZooBank="9995702B61464BA1BB5323DC9BA9650F" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-732-1" ModsDocOrigin="ZooKeys 732" ModsDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" checkinTime="1516831538731" checkinUser="pensoft" docAuthor="Mattia, Willy De, Neiber, Marco T. & Groh, Klaus" docDate="2018" docId="BBF5E34F86A6C1A3C7942CDFEB251710" docLanguage="en" docName="ZooKeys 732: 1-125" docOrigin="ZooKeys 732" docSource="http://dx.doi.org/10.3897/zookeys.732.21677" docTitle="Hystricella echinoderma Wollaston 1878" docType="treatment" docVersion="7" lastPageNumber="23" masterDocId="FFA3FF812B32FFA8FFBAFFCDFF9AFF92" masterDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" masterLastPageNumber="125" masterPageNumber="1" pageNumber="22" updateTime="1701378359498" updateUser="ExternalLinkService">
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<mods:title id="8B9ED2E31367C727AB9AA29F9D75EFF2">Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)</mods:title>
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<mods:namePart id="0C085889E8E8E5AF477C054E6473D58A">Mattia, Willy De</mods:namePart>
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<mods:namePart id="753BEC6C4EF8ACED2CE867464FB1F5B3">Neiber, Marco T.</mods:namePart>
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<mods:namePart id="DE91782B6CD8A00514BD00B3C51F7E24">Groh, Klaus</mods:namePart>
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<mods:date id="16FE86C171B17F397C3E0A5FECA5A0D9">2018</mods:date>
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<mods:detail id="C7D98C4990C3078A7B43852FD80F8DB4" type="volume">
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<mods:number id="BDF4423BF1ADC064EF131C27D0F383C1">732</mods:number>
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<mods:classification id="2F01BAC4034565CA15E9EDA4D990F32F">journal article</mods:classification>
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<mods:identifier id="3E7D5DCDC795B070C33EABC14A07FD17" type="DOI">http://dx.doi.org/10.3897/zookeys.732.21677</mods:identifier>
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<treatment id="BBF5E34F86A6C1A3C7942CDFEB251710" ID-GBIF-Taxon="140417750" LSID="urn:lsid:plazi:treatment:BBF5E34F86A6C1A3C7942CDFEB251710" httpUri="http://treatment.plazi.org/id/BBF5E34F86A6C1A3C7942CDFEB251710" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection id="0B62B17A3C2A3C5379E60AD5C13E6999" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="4D82E4CD2C82BB938A99A8A3EE3C8DA9" pageId="21" pageNumber="22">
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†
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<taxonomicName id="7CE39F50DFE9CBA5A7A8E481A8D21B37" ID-CoL="7RQGS" authority="Wollaston, 1878" authorityName="Wollaston" authorityYear="1878" class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Hystricella echinoderma (Wollaston, 1878)</taxonomicName>
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Figs 95, 96
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</paragraph>
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</subSubSection>
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<subSubSection id="774D43D22634A1E94CE6BD9D7AB76E8D" pageId="21" pageNumber="22" type="list of synonyms">
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<paragraph id="1981AF1DF4A7DE30CE0DB7B0AEEE0ACF" pageId="21" pageNumber="22">List of synonyms.</paragraph>
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<paragraph id="4560CF8B06FEDB27F717CF061FB0271B" pageId="21" pageNumber="22">
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1878
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<taxonomicName id="EA90671546D327D1B2613BC7F14DC90C" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Hystricella) echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma" subGenus="Hystricella">Helix (Hystricella) echinoderma</taxonomicName>
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Wollaston: 159-160.
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</paragraph>
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<paragraph id="4D915F0D9BACBF95A0AEDB6DE2A53F4F" pageId="21" pageNumber="22">
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1894
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<taxonomicName id="4EABA15E4868356758926C3194F264C6" class="Gastropoda" family="Geomitridae" genus="Geomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geomitra echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Geomitra echinoderma</taxonomicName>
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- Pilsbry in
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<bibRefCitation id="B04211DB9416CD89CE5A9034A377169E" pageId="21" pageNumber="22">Pilsbry 1893-1895</bibRefCitation>
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: 242.
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</paragraph>
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<paragraph id="D0B206209AAD235B7B1A5CCE88AAA7C8" pageId="21" pageNumber="22">
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1931
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<taxonomicName id="AA19AD919ED71FECCE836A8EE185FA0F" class="Gastropoda" family="Geomitridae" genus="Geomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geomitra (Actinella) echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma" subGenus="Actinella">Geomitra (Actinella) echinoderma</taxonomicName>
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- Nobre: 88.
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</paragraph>
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<paragraph id="81A43DFCA9D734F29FCD11E8E9ED02C0" pageId="21" pageNumber="22">
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1950
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<taxonomicName id="34DD71FEF4219706C71D0418B3B3F541" class="Gastropoda" family="Geomitridae" genus="Discula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discula (Hystricella) echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma" subGenus="Hystricella">Discula (Hystricella) echinoderma</taxonomicName>
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-
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<bibRefCitation id="0830DDF0A7FACAC2881BB5EB9FC201CC" author="Mandahl-Barth, G" journalOrPublisher="Abhandlungen der senckenbergischen naturforschenden Gesellschaft" pageId="75" pageNumber="76" pagination="1 - 93" title="Systematische Untersuchungen ueber die Heliciden-Fauna von Madeira." volume="469" year="1950">Mandahl-Barth 1950</bibRefCitation>
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: 31, 55.
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</paragraph>
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<paragraph id="4095709091A6E1C64B07AE3789799964" pageId="21" pageNumber="22">
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1983
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<taxonomicName id="8E4F87EDBF5D52E9163CC4F2936A1E40" class="Gastropoda" family="Geomitridae" genus="Discula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discula (Hystricella) echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma" subGenus="Hystricella">Discula (Hystricella) echinoderma</taxonomicName>
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-
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<normalizedToken id="4B095D8C76584209C351D3DFE263FB6D" originalValue="Waldén">Walden</normalizedToken>
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: 267.
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</paragraph>
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<paragraph id="AFC14FEFEE8B2E490F3EDC4A52FB049A" pageId="21" pageNumber="22">
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2002
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<taxonomicName id="3C684273ADA5FEB387B6FCDCAB94EACF" class="Gastropoda" family="Geomitridae" genus="Geomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geomitra echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Geomitra echinoderma</taxonomicName>
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- Bank et al.: 124.
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</paragraph>
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<paragraph id="7D5FB81AB98A758B0B3F934652763886" pageId="21" pageNumber="22">
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2008
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<taxonomicName id="E977F8388DD29A41E29E477A5591B3E4" class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Hystricella echinoderma</taxonomicName>
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- Seddon: 80.
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</paragraph>
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</subSubSection>
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<subSubSection id="2757FC8742179AF5A9DD375D695B4E7C" pageId="21" pageNumber="22" type="type material">
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<paragraph id="9EA774D1E646EB365491886044718F39" pageId="21" pageNumber="22">Type material.</paragraph>
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<paragraph id="E29004425525C540EB5E09AA6A037731" pageId="21" pageNumber="22">NHM 1875.02.02-52, lectotype (herewith designated) from loc. typ.; NHM 1875.02.02-53 to -54, 2 paralectotypes, from loc. typ.</paragraph>
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</subSubSection>
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<subSubSection id="75A5BF0B3B9DDE2EDAE4E73509F56FAE" pageId="21" pageNumber="22" type="locus typicus">
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<paragraph id="00BFC77C8A2D6E6D29856658A1CE1540" pageId="21" pageNumber="22">Locus typicus.</paragraph>
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<paragraph id="AA0A2F1EC56C4911BC3532BF0DA35E93" pageId="21" pageNumber="22">Portum Sanctum, semifossilis; recens haud observata.</paragraph>
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</subSubSection>
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<subSubSection id="76D590C1BA9BC76FAC990ECCEB9A6A00" pageId="21" pageNumber="22" type="description">
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<paragraph id="1546887EC554FDE0AD82AC741EFB06CC" pageId="21" pageNumber="22">Original description.</paragraph>
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<paragraph id="AAD76EA91344EEBABAB6B3947FD94F17" pageId="21" pageNumber="22">
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From
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<bibRefCitation id="84ACFABE92FDB083910D3AAA044DE78E" author="Wollaston, TV" journalOrPublisher="Reeve, London" pageId="77" pageNumber="78" title="Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena." year="1878">Wollaston 1878</bibRefCitation>
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: T. trochiformis, subtus subplanulata perforata, undiquegranulis magnis obtusis sat dense obsita; spira elevata; anfractibus convexis, subgibbosis, ultimo subtectiformi acute carinato (carina simplici, solum antice gradatim obsolete subduplici); umbilico punctiformi, aperto; apertura subovali-rotundata, labris continuis conjunctis, peristomate simplici, expanso, subrecurvo, tenui, relevato. - Long. axis 2
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<normalizedToken id="27959CF47E091093E51B7CAFB2243CA5" originalValue="½">1/2</normalizedToken>
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lin.; diam. 3
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<normalizedToken id="BF7B96AC6860A0586A05ADE80BB416A3" originalValue="½">1/2</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C6BDB33D7AE1BC4EABE73717F048B9BF" pageId="21" pageNumber="22" type="description">
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<paragraph id="76CE4F7C9514D954C4F2A9C3D8F33E2D" pageId="21" pageNumber="22">Description of the lectotype.</paragraph>
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<paragraph id="80477069BD8F5AC013418D3E88ABAB13" pageId="21" pageNumber="22">
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Shell large for the genus, with
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<normalizedToken id="5D671B4FC549509CE2DBC158BEEC2AA2" originalValue="6¾">63/4</normalizedToken>
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regularly increasing whorls, the protoconch with 1.9 whorls. The form of the shell is high conical, the convex to quadrangular teleoconch whorls showing two more and more expressed angulations which form in the last whorl two rounded keels, the upper less developed than the lower one. The last whorl measures 59%, the penultimate whorl 14% of the total shell height. The lower 70% of the body whorl are situated below the peripheral keel which is slightly constricted by a concavity below the periphery in frontal view. The base is convex, slightly straightening towards the umbilicus. The two keels of the body whorl are located in the upper ⅛ and ⅓ of the total height of the body whorl. The suture between the whorls is simple but deeply sunken. The aperture, which is inclined to the vertical axis of the shell in an angle of 48° and descending in the last 5% of the last whorl in an angle of 38° to the horizontal axis, has an oblique-ovate form, its width amounts to 43% of the total shell width, its height to 34% of the total shell height. It has a slightly reflected lip, which is completely detached from the body whorl. The centered umbilicus, which measures 10% of the
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<normalizedToken id="A3D51291F645CEBA35FDD76C7C436D1D" originalValue="shell’s">shell's</normalizedToken>
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total width, is in the last whorls circular, but completely closed deeper inside. The protoconch is smooth, the teleoconch shows a number of oblique radial ribs, 15 in the penultimate quadrant of the body whorl, and is additionally covered by numerous irregularly arranged, elongate, rough tubercles. The number of tubercles in the standard-quadrate of the base is 81. There are no traces of colouration. See Fig. 95.
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</paragraph>
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<caption id="8DD355493F40A4A9C6C2CA57108A9CDE" pageId="21" pageNumber="22">
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<paragraph id="D5DC715FFD422D487CFAB81D94870D58" pageId="21" pageNumber="22">
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Figure 95. Lectotype of
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<taxonomicName id="36D6079395226F5CAC21EC5F9F2BAE88" class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Hystricella echinoderma</taxonomicName>
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, NHM 1875.02.02-52. Scale bar 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="DA134D5B2EA695CAEDB3922218D5065D" pageId="21" pageNumber="22" type="description">
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<paragraph id="49125A2DD72F2F1317005F8E08C4E38C" pageId="21" pageNumber="22">Measurements.</paragraph>
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<paragraph id="FA61F7ECCBCC59AABFBB0441209CBD16" pageId="21" pageNumber="22">D 7.0 mm; H 5.9 mm; FW 3.5 mm; PA 42.2°; DU 0.5 mm; NT ≈ 35; NW 6.6 (n = 1). Ratio D/H 1.2; ratio FW/H 0.5.</paragraph>
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</subSubSection>
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<subSubSection id="4B618EE97B36E73EA3803800BE6D303B" pageId="21" pageNumber="22" type="distribution">
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<paragraph id="C6C8E33180D3947FB967955E88F75A48" pageId="21" pageNumber="22">Distribution.</paragraph>
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<paragraph id="D38D13CFE39CF927E5C1A1D4D3F6451F" pageId="21" pageNumber="22">
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<taxonomicName id="38AA2CFE2319C634C72253590112B462" class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Hystricella echinoderma</taxonomicName>
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is only known from Fonte da Areia in the northern part of Porto Santo (Fig. 96).
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</paragraph>
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<caption id="49EDD15748B4611468181A3D1D4F409B" pageId="21" pageNumber="22">
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<paragraph id="6C0BD5C300D4B443E3501CDF7D8002D7" pageId="21" pageNumber="22">
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Figure 96. Distribution of
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<taxonomicName id="64D347E46A08D9F61DBDAFB0AEC9D394" class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Hystricella echinoderma</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="96FB52AFF3A3661AC56D43E3CD90522A" pageId="21" pageNumber="22" type="comparisons">
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<paragraph id="18EFA5CE9AB0B52A1A8DD5A8797450B7" pageId="21" pageNumber="22">Comparison and comments.</paragraph>
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<paragraph id="093680F6670863712D835607774CCAF9" pageId="21" pageNumber="22">
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<taxonomicName id="B06FACE58F3C1DF6475773AB7FCE8D9B" class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella echinoderma" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="echinoderma">Hystricella echinoderma</taxonomicName>
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can be confused on first glance with comparatively large-sized species of the genus
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<taxonomicName id="F26D99D3F5216A50C4FA170CCB90A30D" class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">Wollastonia</taxonomicName>
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like
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<taxonomicName id="793040E2358E757E0F613F852DB368DB" lsidName="W. vermetiformis" pageId="21" pageNumber="22" rank="species" species="vermetiformis">W. vermetiformis</taxonomicName>
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,
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<taxonomicName id="F53C92779736694BA3EF35E766D28242" lsidName="W. ripkeni" pageId="21" pageNumber="22" rank="species" species="ripkeni">W. ripkeni</taxonomicName>
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sp. n., and
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<taxonomicName id="80AAC21629A4E873C1209B2A4422A00F" lsidName="W. falknerorum" pageId="21" pageNumber="22" rank="species" species="falknerorum">W. falknerorum</taxonomicName>
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sp. n.; from these it is distinguishable by the lack of one or two sharply pointed keels, respectively, a much rougher sculpture, a non-eccentric umbilicus, and the stepped, high conical form. From the similarly sized
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<taxonomicName id="BC666049A3F39295F63F8C7BCB55E5AE" lsidName="W. subcarinulata" pageId="21" pageNumber="22" rank="species" species="subcarinulata">W. subcarinulata</taxonomicName>
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and
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<taxonomicName id="E30FE33AF17E826DC071254AABA7D0CE" lsidName="W. inexpectata" pageId="21" pageNumber="22" rank="species" species="inexpectata">W. inexpectata</taxonomicName>
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sp. n. it is separated by a wider or non-eccentric umbilicus, a relatively higher shell, the quadrangular instead of convex rounded whorls, a coarser granulation and the oblique-ovate instead of straight horizontally elliptical aperture. From the similar shaped
|
||
<taxonomicName id="1B0F7B27D266508D990FF52D607CC660" lsidName="H. echinulata" pageId="21" pageNumber="22" rank="species" species="echinulata">H. echinulata</taxonomicName>
|
||
and
|
||
<taxonomicName id="24708BAA5F0DB418A2366115217D6C31" lsidName="H. microcarinata" pageId="21" pageNumber="22" rank="species" species="microcarinata">H. microcarinata</taxonomicName>
|
||
sp. n. it is easily distinguishable by the much larger size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="216320D21D7F88A7908826D52F1E760B" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="taxonomic remarks">
|
||
<paragraph id="BF19C4640089FFA66BB19B379E4E6AAF" pageId="21" pageNumber="22">Taxonomic remarks.</paragraph>
|
||
<paragraph id="3E32E43B849B5E155E708DB42E62553B" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<bibRefCitation id="7875F71497D2402FB12A79FE4A9DCD74" author="Wollaston, TV" journalOrPublisher="Reeve, London" pageId="77" pageNumber="78" title="Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena." year="1878">Wollaston (1878)</bibRefCitation>
|
||
compares
|
||
<taxonomicName id="CF8DB30C1FC990AF87C26857E00C98A8" lsidName="H. echinoderma" pageId="21" pageNumber="22" rank="species" species="echinoderma">H. echinoderma</taxonomicName>
|
||
with the rather similar
|
||
<taxonomicName id="8927261DCDB8FFA21BCF3062331D0791" lsidName="H. echinulata" pageId="21" pageNumber="22" rank="species" species="echinulata">H. echinulata</taxonomicName>
|
||
and calls it a monstrous sister species of that species, citing as examples of
|
||
<pageBreakToken id="77CD3C6C0707E5123875A757A6372BEA" pageId="22" pageNumber="23" start="start">pairs</pageBreakToken>
|
||
of nominal species with a distinct size difference
|
||
<taxonomicName id="E06BD95EB2A23AE0951A6D05DF9DB51E" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix vermetiformis" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="vermetiformis">Helix vermetiformis</taxonomicName>
|
||
and
|
||
<taxonomicName id="DB73FCFD2D1ACBF18E60772F45B1686A" lsidName="H. bicarinata" pageId="22" pageNumber="23" rank="species" species="bicarinata">H. bicarinata</taxonomicName>
|
||
, H. Lowei and
|
||
<taxonomicName id="25B45276721F3982B34C23638FAB5332" lsidName="H. portosanctana" pageId="22" pageNumber="23" rank="species" species="portosanctana">H. portosanctana</taxonomicName>
|
||
as well as
|
||
<taxonomicName id="2A8EE95BAD2986D3B4AA58FF909B731A" lsidName="H. bowdichiana" pageId="22" pageNumber="23" rank="species" species="bowdichiana">H. bowdichiana</taxonomicName>
|
||
and
|
||
<taxonomicName id="248969EA110BCBEDDBDB47EE217184F0" lsidName="H. punctulata" pageId="22" pageNumber="23" rank="species" species="punctulata">H. punctulata</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46F74CFE47F853D460AC320983259D35" pageId="22" pageNumber="23" type="status and conservation">
|
||
<paragraph id="954C621B3E87AEC75E9216710D41D9EB" pageId="22" pageNumber="23">Status and conservation.</paragraph>
|
||
<paragraph id="93B491911439F22277BA372EF3B263EF" pageId="22" pageNumber="23">
|
||
Extinct before the
|
||
<normalizedToken id="7C9FC306317DAB5201D771D2AD7EE951" originalValue="islands’">islands'</normalizedToken>
|
||
scientific exploration in the 19th century, possibly already before human settlement.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |