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<document id="83F013857F675F872E53490E60C39639" ID-CLB-Dataset="3155" ID-DOI="10.11646/zootaxa.5072.4.1" ID-GBIF-Dataset="b715a8f1-cf4b-4f80-91bc-9019eb8a3409" ID-ISSN="1175-5326" ID-Zenodo-Dep="5748820" ID-ZooBank="1D561060-A3A6-4F21-80B5-C393744349DB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638424170995" checkinUser="plazi" docAuthor="Kelly, Joseph B. &amp; Thacker, Robert W." docDate="2021" docId="BB4A878FC063FFA9A3C4F8FAFD9E9038" docLanguage="en" docName="zootaxa.5072.4.1.pdf" docOrigin="Zootaxa 5072 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ircinia laeviconulosa Kelly &amp; Thacker 2021, sp. nov." docType="treatment" docUuid="2F579603-36E6-4D5A-9009-FBB43A99247C" docUuidSource="ZooBank" docVersion="7" lastPageNumber="311" masterDocId="4773FFF7C06AFFA3A353FFEEFF809259" masterDocTitle="New shallow water species of Caribbean Ircinia Nardo, 1833 (Porifera: Irciniidae)" masterLastPageNumber="323" masterPageNumber="301" pageNumber="310" updateTime="1699471693658" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DF52B7A054B7F5271445BEA524430B92">New shallow water species of Caribbean Ircinia Nardo, 1833 (Porifera: Irciniidae)</mods:title>
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<mods:namePart id="E69FA2CE90CB9E40AD6D1D35857B370C">Kelly, Joseph B.</mods:namePart>
<mods:affiliation id="9C89B2FF4A13F9F7B375B03BD227A2DE">Stony Brook University, Department of Ecology and Evolution, Stony Brook, NY, USA. &amp; Limnological Institute University Konstanz, Aquatic Ecology and Evolution, Konstanz, Germany.</mods:affiliation>
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<mods:namePart id="36B933D5007D48A65DD594A2ADEEBCEE">Thacker, Robert W.</mods:namePart>
<mods:affiliation id="04F8A1336751CA3A025FDDC9455EA74E">Stony Brook University, Department of Ecology and Evolution, Stony Brook, NY, USA. &amp; Smithsonian Tropical Research Institute, PO Box 0843 - 03092, Balboa, Republic of Panama.</mods:affiliation>
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<mods:title id="1A278D25F3545086B55C2291AA555C82">Zootaxa</mods:title>
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<mods:date id="1140760A7DB1307B0937E3FBE6888EF7">2021</mods:date>
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<mods:number id="6C2DD1B1E0E238D34080E115404D7FF6">5072</mods:number>
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<treatment id="BB4A878FC063FFA9A3C4F8FAFD9E9038" ID-DOI="http://doi.org/10.5281/zenodo.5748806" ID-GBIF-Taxon="190875676" ID-Zenodo-Dep="5748806" LSID="urn:lsid:zoobank.org:act:2F579603-36E6-4D5A-9009-FBB43A99247C" httpUri="http://treatment.plazi.org/id/BB4A878FC063FFA9A3C4F8FAFD9E9038" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
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<taxonomicName id="F4E34D1AC063FFAAA3C4F8FAFE189576" ID-CoL="8TFWW" authority="Kelly &amp; Thacker, 2021" authorityName="Kelly &amp; Thacker" authorityYear="2021" box="[151,408,1812,1839]" class="Demospongiae" family="Irciniidae" genus="Ircinia" kingdom="Animalia" order="Dictyoceratida" pageId="9" pageNumber="310" phylum="Porifera" rank="species" species="laeviconulosa" status="sp. nov.">
<emphasis id="0197EA8BC063FFAAA3C4F8FAFE189576" bold="true" box="[151,408,1812,1839]" italics="true" pageId="9" pageNumber="310">Ircinia laeviconulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA457F0C063FFAAA2CCF8FAFE7D9577" box="[415,509,1812,1838]" pageId="9" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="ABD82A1CC063FFAAA3C4F8D6FE81950B" box="[151,257,1848,1874]" captionStart="FIGURE 8" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[156,1432,633,1787]" captionTargetId="figure-201@10.[155,1432,633,1787]" captionTargetPageId="10" captionText="FIGURE 8. Ircinia laeviconulosa sp. nov. A: USNM 1582283 (holoype), B: USNM 1582285 (paratype), C: USNM 1582288 (paratype), D: USNM 1582286 (paratype). Note the variation in oscular position, from a single large osculum (A) to multiple smaller oscula (B-D)." figureDoi="http://doi.org/10.5281/zenodo.5748842" httpUri="https://zenodo.org/record/5748842/files/figure.png" pageId="9" pageNumber="310">Figures 8</figureCitation>
,
<figureCitation id="ABD82A1CC063FFAAA25DF8D6FE9E950B" box="[270,286,1848,1874]" captionStart="FIGURE 9" captionStartId="11.[151,250,1648,1672]" captionTargetBox="[156,1431,167,1624]" captionTargetId="figure-18@11.[155,1431,166,1624]" captionTargetPageId="11" captionText="FIGURE 9. A,B: Histological sections of Ircinia laeviconulosa sp. nov., specimen voucher USNM 1582285 (paratype). Abbreviations follow Figure 3." figureDoi="http://doi.org/10.5281/zenodo.5748844" httpUri="https://zenodo.org/record/5748844/files/figure.png" pageId="9" pageNumber="310">9</figureCitation>
;
<tableCitation id="7E610322C063FFAAA279F8D6FE08950B" box="[298,392,1848,1874]" captionStart="TABLE 1" captionStartId="2.[151,239,673,699]" captionTargetId="graphics-31@2.[151,2035,809,1539]" captionTargetPageId="2" captionText="TABLE 1. Sampling locations of Ircinia spp." httpUri="http://table.plazi.org/id/679C6611C068FFA1A3C4FD4FFDFC90E2" pageId="9" pageNumber="310" tableUuid="679C6611C068FFA1A3C4FD4FFDFC90E2">Tables 1</tableCitation>
,
<tableCitation id="7E610322C063FFAAA2C5F8D6FE27950B" box="[406,423,1848,1874]" captionStart="TABLE 2" captionStartId="20.[151,239,531,557]" captionTargetPageId="20" captionText="TABLE 2. A morphological comparison of shallow water Caribbean Ircinia." httpUri="http://table.plazi.org/id/679C6611C07EFFB7A3C4FDFDFC4A9074" pageId="9" pageNumber="310" tableUuid="679C6611C07EFFB7A3C4FDFDFC4A9074">2</tableCitation>
.
</paragraph>
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urn:lsid:zoobank.org:act:
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<typeStatus id="EC58883BC063FFAAA3C4F84AFE8895E7" box="[151,264,1956,1982]" pageId="9" pageNumber="310" type="holotype">Holotype</typeStatus>
:
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,
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; appx.
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depth
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; coll.
<collectorName id="9E16534FC063FFAAA749F84BFBE195E6" box="[1050,1121,1957,1983]" name="Joseph B. Kelly" pageId="9" pageNumber="310">J.B.K.</collectorName>
and
<collectionCode id="55F2AE5CC063FFAAA7C4F84BFB2C95E6" box="[1175,1196,1957,1983]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="310">R</collectionCode>
.W.
<collectionCode id="55F2AE5CC063FFAAA780F84BFB6695E6" box="[1235,1254,1957,1983]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="310">T</collectionCode>
.;
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<collectingDate id="5719E9B1C063FFAAA7A6F84AFA0A95E6" box="[1269,1418,1956,1983]" pageId="9" pageNumber="310" value="2016-07-26">26 July 2016</collectingDate>
</date>
)
</materialsCitation>
.
<materialsCitation id="838B3CC4C063FFA9A394F826FF5B92EB" ID-GBIF-Occurrence="3413089321" collectingDate="2016-07-26" collectionCode="R, T" collectorName="Joseph B. Kelly" elevation="5" lastPageId="10" lastPageNumber="311" latitude="9.35133" longLatPrecision="5" longitude="-82.2593" pageId="9" pageNumber="310" specimenCode="USNM 1582285" specimenCount="1" typeStatus="paratype">
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<typeStatus id="EC58883BC063FFAAA394F826FEC395BB" box="[199,323,1992,2018]" pageId="9" pageNumber="310" type="paratype">Paratypes</typeStatus>
:
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,
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; appx.
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depth
</elevation>
; coll.
<collectorName id="9E16534FC063FFAAA73AF827FB3095BA" box="[1129,1200,1993,2019]" name="Joseph B. Kelly" pageId="9" pageNumber="310">J.B.K.</collectorName>
and
<collectionCode id="55F2AE5CC063FFAAA7B9F827FB7F95BA" box="[1258,1279,1993,2019]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="310">R</collectionCode>
.W.
<collectionCode id="55F2AE5CC063FFAAA676F827FAB895BA" box="[1317,1336,1993,2019]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="310">T</collectionCode>
.;
<date id="475D1059C063FFA9A61AF826FF5392EB" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310" value="2016-07-26">
<collectingDate id="5719E9B1C063FFA9A61AF826FF5392EB" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310" value="2016-07-26">26 July 2016</collectingDate>
</date>
)
</materialsCitation>
,
<materialsCitation id="838B3CC4C060FFA9A3B9FF79FA1892EB" ID-GBIF-Occurrence="3413089302" box="[234,1432,150,178]" collectingDate="2016-07-26" collectionCode="R, T" collectorName="Joseph B. Kelly" elevation="5" latitude="9.35133" longLatPrecision="5" longitude="-82.2593" pageId="10" pageNumber="311" specimenCode="USNM 1582286" specimenCount="1" typeStatus="paratype">
<specimenCode id="63459EE2C060FFA9A3B9FF79FE2792E8" box="[234,423,150,177]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="311" type="Museum">USNM 1582286</specimenCode>
(P16x60;
<geoCoordinate id="56D7505EC060FFA9A171FF78FD0092E8" box="[546,640,150,177]" degrees="9.35133" direction="north" orientation="latitude" pageId="10" pageNumber="311" precision="1" value="9.35133">9.35133</geoCoordinate>
,
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; appx.
<elevation id="B8CED1AAC060FFA9A01EFF78FC4792E8" box="[845,967,150,177]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="311" unit="m" value="5.0">
<quantity id="F41B9B7CC060FFA9A01EFF78FCFB92E8" box="[845,891,150,177]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="311" unit="m" value="5.0">5 m</quantity>
depth
</elevation>
; coll.
<collectorName id="9E16534FC060FFA9A743FF79FBD792E8" box="[1040,1111,151,177]" name="Joseph B. Kelly" pageId="10" pageNumber="311">J.B.K.</collectorName>
and
<collectionCode id="55F2AE5CC060FFA9A7C7FF79FB2992E8" box="[1172,1193,151,177]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="311">R</collectionCode>
.W.
<collectionCode id="55F2AE5CC060FFA9A79CFF79FB6292E8" box="[1231,1250,151,177]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="311">T</collectionCode>
.;
<date id="475D1059C060FFA9A7A6FF79FA1092EB" box="[1269,1424,151,178]" pageId="10" pageNumber="311" value="2016-07-26">
<collectingDate id="5719E9B1C060FFA9A7A6FF79FA1092EB" box="[1269,1424,151,178]" pageId="10" pageNumber="311" value="2016-07-26">26 July 2016</collectingDate>
</date>
)
</materialsCitation>
,
<materialsCitation id="838B3CC4C060FFA9A3C4FF55FED1928C" ID-GBIF-Occurrence="3413089307" box="[151,337,186,213]" collectionCode="USNM" pageId="10" pageNumber="311" specimenCode="USNM 1582288" specimenCount="1" typeStatus="paratype">
<specimenCode id="63459EE2C060FFA9A3C4FF55FED1928C" box="[151,337,186,213]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="311" type="Museum">USNM 1582288</specimenCode>
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<materialsCitation id="838B3CC4C060FFA9A20BFF55FA95928F" ID-GBIF-Occurrence="3413089305" box="[344,1301,186,214]" collectingDate="2016-07-26" collectionCode="R, T" collectorName="Joseph B. Kelly" elevation="5" latitude="9.35133" longLatPrecision="5" longitude="-82.2593" pageId="10" pageNumber="311" specimenCount="1" typeStatus="paratype">
(P16x62;
<geoCoordinate id="56D7505EC060FFA9A295FF54FDA4928C" box="[454,548,186,213]" degrees="9.35133" direction="north" orientation="latitude" pageId="10" pageNumber="311" precision="1" value="9.35133">9.35133</geoCoordinate>
,
<geoCoordinate id="56D7505EC060FFA9A17CFF54FD16928C" box="[559,662,186,213]" degrees="82.2593" direction="west" orientation="longitude" pageId="10" pageNumber="311" precision="5" value="-82.2593">-82.2593</geoCoordinate>
; appx.
<elevation id="B8CED1AAC060FFA9A1B5FF54FCDA928C" box="[742,858,186,213]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="311" unit="m" value="5.0">
<quantity id="F41B9B7CC060FFA9A1B5FF54FC91928C" box="[742,785,186,213]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="311" unit="m" value="5.0">5 m</quantity>
depth
</elevation>
; coll.
<collectorName id="9E16534FC060FFA9A0CEFF55FC64928C" box="[925,996,187,213]" name="Joseph B. Kelly" pageId="10" pageNumber="311">J.B.K.</collectorName>
and
<collectionCode id="55F2AE5CC060FFA9A749FF55FBAF928C" box="[1050,1071,187,213]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="311">R</collectionCode>
.W.
<collectionCode id="55F2AE5CC060FFA9A706FF55FBE8928C" box="[1109,1128,187,213]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="311">T</collectionCode>
.;
<date id="475D1059C060FFA9A72BFF55FA8D928F" box="[1144,1293,187,214]" pageId="10" pageNumber="311" value="2016-07-26">
<collectingDate id="5719E9B1C060FFA9A72BFF55FA8D928F" box="[1144,1293,187,214]" pageId="10" pageNumber="311" value="2016-07-26">26 July 2016</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="335C3699C060FFA9A394FF31FD0792A0" blockId="10.[151,1437,150,609]" box="[199,647,222,249]" pageId="10" pageNumber="311">
<emphasis id="0197EA8BC060FFA9A394FF31FEE992A1" bold="true" box="[199,361,222,249]" pageId="10" pageNumber="311">Type locality:</emphasis>
<collectingRegion id="F127F87BC060FFA9A223FF31FD9B92A0" box="[368,539,223,249]" country="Panama" name="Bocas del Toro" pageId="10" pageNumber="311">Bocas del Toro</collectingRegion>
,
<collectingCountry id="4BF47609C060FFA9A174FF31FD0392A0" box="[551,643,223,249]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
.
</paragraph>
<paragraph id="335C3699C060FFA9A394FEEDFBCD933C" blockId="10.[151,1437,150,609]" pageId="10" pageNumber="311">
<emphasis id="0197EA8BC060FFA9A394FEEDFE519344" bold="true" box="[199,465,259,285]" pageId="10" pageNumber="311">External morphology.</emphasis>
<taxonomicName id="F4E34D1AC060FFA9A28EFEEDFDA89345" authorityName="Nardo" authorityYear="1833" box="[477,552,259,284]" class="Demospongiae" family="Irciniidae" genus="Ircinia" kingdom="Animalia" order="Dictyoceratida" pageId="10" pageNumber="311" phylum="Porifera" rank="genus">
<emphasis id="0197EA8BC060FFA9A28EFEEDFDA89345" box="[477,552,259,284]" italics="true" pageId="10" pageNumber="311">Ircinia</emphasis>
</taxonomicName>
with a massive growth form and dark green pinacoderm (
<figureCitation id="ABD82A1CC060FFA9A78CFEEDFAC49344" box="[1247,1348,259,285]" captionStart="FIGURE 8" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[156,1432,633,1787]" captionTargetId="figure-201@10.[155,1432,633,1787]" captionTargetPageId="10" captionText="FIGURE 8. Ircinia laeviconulosa sp. nov. A: USNM 1582283 (holoype), B: USNM 1582285 (paratype), C: USNM 1582288 (paratype), D: USNM 1582286 (paratype). Note the variation in oscular position, from a single large osculum (A) to multiple smaller oscula (B-D)." figureDoi="http://doi.org/10.5281/zenodo.5748842" httpUri="https://zenodo.org/record/5748842/files/figure.png" pageId="10" pageNumber="311">Figure 8</figureCitation>
). Body diameter
<quantity id="F41B9B7CC060FFA9A3ACFEC8FEF19318" box="[255,369,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="10" pageNumber="311" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
. Possesses low conules (
<quantity id="F41B9B7CC060FFA9A12CFEC8FC999318" box="[639,793,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="10" pageNumber="311" unit="mm" value="1.625" valueMax="1.75" valueMin="1.5">1.5–1.75 mm</quantity>
). Oscula flush, sometimes slightly darker than the exterior of the sponge,
<quantity id="F41B9B7CC060FFA9A26FFEA5FE39933C" box="[316,441,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="12.0" metricValueMin="4.0" pageId="10" pageNumber="311" unit="cm" value="0.8" valueMax="1.2" valueMin="0.4">0.4–1.2 cm</quantity>
in diameter, and with a thin dark green or gray membrane.
</paragraph>
<paragraph id="335C3699C060FFA9A394FE80FE769388" blockId="10.[151,1437,150,609]" pageId="10" pageNumber="311">
<emphasis id="0197EA8BC060FFA9A394FE80FE4293D0" bold="true" box="[199,450,366,393]" pageId="10" pageNumber="311">Interior morphology.</emphasis>
Massive fascicular fibers 110–160 µm wide, heavily cored. Interconnecting fibers 20–50 um wide, sparsely cored (
<figureCitation id="ABD82A1CC060FFA9A2ECFE7DFDA493F4" box="[447,548,403,429]" captionStart="FIGURE 9" captionStartId="11.[151,250,1648,1672]" captionTargetBox="[156,1431,167,1624]" captionTargetId="figure-18@11.[155,1431,166,1624]" captionTargetPageId="11" captionText="FIGURE 9. A,B: Histological sections of Ircinia laeviconulosa sp. nov., specimen voucher USNM 1582285 (paratype). Abbreviations follow Figure 3." figureDoi="http://doi.org/10.5281/zenodo.5748844" httpUri="https://zenodo.org/record/5748844/files/figure.png" pageId="10" pageNumber="311">Figure 9</figureCitation>
). Irciniid filaments 1–5 µm wide, terminating in spherical to tear-drop knobs measuring 6–9 µm in diameter.
</paragraph>
</subSubSection>
<subSubSection id="7BF96512C060FFA9A394FE35FB3693AC" box="[199,1206,475,501]" pageId="10" pageNumber="311" type="biology_ecology">
<paragraph id="335C3699C060FFA9A394FE35FB3693AC" blockId="10.[151,1437,150,609]" box="[199,1206,475,501]" pageId="10" pageNumber="311">
<emphasis id="0197EA8BC060FFA9A394FE35FEAC93AC" bold="true" box="[199,300,475,501]" pageId="10" pageNumber="311">Ecology.</emphasis>
This species is found among
<taxonomicName id="F4E34D1AC060FFA9A12BFE35FC9A93AD" box="[632,794,475,501]" class="Liliopsida" family="Hydrocharitaceae" genus="Thalassia" kingdom="Plantae" order="Alismatales" pageId="10" pageNumber="311" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="0197EA8BC060FFA9A12BFE35FD6593AC" box="[632,741,475,501]" italics="true" pageId="10" pageNumber="311">Thalassia</emphasis>
spp.
</taxonomicName>
and coral patches in shallow depths.
</paragraph>
</subSubSection>
<subSubSection id="7BF96512C060FFA9A394FE11FBD09040" box="[199,1104,511,537]" pageId="10" pageNumber="311" type="etymology">
<paragraph id="335C3699C060FFA9A394FE11FBD09040" blockId="10.[151,1437,150,609]" box="[199,1104,511,537]" pageId="10" pageNumber="311">
<emphasis id="0197EA8BC060FFA9A394FE11FECE9040" bold="true" box="[199,334,511,537]" pageId="10" pageNumber="311">Etymology.</emphasis>
The name refers to the texture imparted by the species’ low conules.
</paragraph>
</subSubSection>
<subSubSection id="7BF96512C060FFA9A394FDCDFD9E9038" pageId="10" pageNumber="311" type="discussion">
<paragraph id="335C3699C060FFA9A394FDCDFD9E9038" blockId="10.[151,1437,150,609]" pageId="10" pageNumber="311">
<emphasis id="0197EA8BC060FFA9A394FDCDFEBB9064" bold="true" box="[199,315,547,573]" pageId="10" pageNumber="311">Remarks.</emphasis>
All specimens collected had a globose growth morphology. Referred to as the ‘Massive A green’ growth form in
<bibRefCitation id="57724B68C060FFA9A21BFDA9FD9A9038" author="Kelly, J. B. &amp; Carlson, D. E. &amp; Low, J. S. &amp; Rice, T. &amp; Thacker, R. W." box="[328,538,583,609]" pageId="10" pageNumber="311" pagination="489" refId="ref9793" refString="Kelly, J. B., Carlson, D. E., Low, J. S., Rice, T. &amp; Thacker, R. W. (2021) The relationship between microbiomes and selective regimes in the sponge genus Ircinia. Frontiers in Microbiology, 12, 489. https: // doi. org / 10.3389 / fmicb. 2021.607289" type="journal article" year="2021">
Kelly
<emphasis id="0197EA8BC060FFA9A2DEFDA9FE459038" box="[397,453,583,609]" italics="true" pageId="10" pageNumber="311">et al.</emphasis>
(2021)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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