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<document id="0F60B3289C882C78A8C17E5EF1C8CF0D" ID-DOI="10.11646/zootaxa.4000.5.4" ID-GBIF-Dataset="b4b5f1f3-28f6-4440-b9ea-d1113bcdd647" ID-ISSN="1175-5326" ID-Zenodo-Dep="236743" ID-ZooBank="F13F1205-EBDB-41D0-83B7-228D9C710357" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461192076420" checkinUser="plazi" docAuthor="Chan, Benny K. K. &amp; Cheang, Chi Chiu" docDate="2015" docId="3D156A32FFE50A15FF2BFC3B25F9FCA6" docLanguage="en" docName="zt04000p558.pdf" docOrigin="Zootaxa 4000 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chthamalus williamsi Chan &amp; Cheang, 2015, sp. nov." docType="treatment" docUuid="C06B7AAB-4455-4FB2-99CE-6718853453EF" docUuidSource="ZooBank" docVersion="8" lastPageNumber="550" masterDocId="C12C124AFFE70A16FFBCFFF72608FFFA" masterDocTitle="A new Chthamalus (Crustacea: Cirripedia) from the challengeri subgroup on Taiwan rocky intertidal shores" masterLastPageNumber="558" masterPageNumber="547" pageNumber="549" updateTime="1698600706428" updateUser="plazi">
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<mods:title id="410C495B0A2BD18E9CBE3CCB64B16812">A new Chthamalus (Crustacea: Cirripedia) from the challengeri subgroup on Taiwan rocky intertidal shores</mods:title>
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<mods:namePart id="EE49C99EADFFF6E6640BB917527B5738">Chan, Benny K. K.</mods:namePart>
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<mods:namePart id="212DD51B322B39077159F0F41974714B">Cheang, Chi Chiu</mods:namePart>
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<mods:title id="06A988B6B5A350764186035EB73665CB">Zootaxa</mods:title>
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<mods:date id="0635C13105F7FB2089728D42C1526CDE">2015</mods:date>
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<treatment id="3D156A32FFE50A15FF2BFC3B25F9FCA6" ID-DOI="http://doi.org/10.5281/zenodo.5671803" ID-GBIF-Taxon="119629203" ID-Zenodo-Dep="5671803" LSID="urn:lsid:zoobank.org:act:C06B7AAB-4455-4FB2-99CE-6718853453EF" httpUri="http://treatment.plazi.org/id/3D156A32FFE50A15FF2BFC3B25F9FCA6" lastPageId="3" lastPageNumber="550" pageId="2" pageNumber="549">
<subSubSection id="FDA688AFFFE50A14FF2BFC3B2714FBF3" pageId="2" pageNumber="549" type="nomenclature">
<paragraph id="B503DB24FFE50A14FF2BFC3B240DFC1D" blockId="2.[151,517,972,1033]" box="[151,517,972,999]" pageId="2" pageNumber="549">
<heading id="EE4B6C48FFE50A14FF2BFC3B240DFC1D" bold="true" box="[151,517,972,999]" fontSize="11" level="1" pageId="2" pageNumber="549" reason="1">
<emphasis id="87C80736FFE50A14FF2BFC3B240DFC1D" bold="true" box="[151,517,972,999]" pageId="2" pageNumber="549">
<taxonomicName id="72BCA0A7FFE50A14FF2BFC3B2797FC1C" box="[151,415,972,998]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="2" pageNumber="549" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
<emphasis id="87C80736FFE50A14FF2BFC3B2797FC1C" bold="true" box="[151,415,972,998]" italics="true" pageId="2" pageNumber="549">Chthamalus williamsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CFBBA4DFFE50A14FE1BFC3A240DFC1D" box="[423,517,973,999]" pageId="2" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B503DB24FFE50A14FF2BFC072714FBF3" blockId="2.[151,517,972,1033]" box="[151,284,1008,1033]" pageId="2" pageNumber="549">
<figureCitation id="2D87C7A1FFE50A14FF2BFC072714FBF3" box="[151,284,1008,1033]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[151,250,1940,1962]" captionStartId-1="5.[151,250,1979,2001]" captionStartId-2="6.[151,250,1890,1912]" captionStartId-3="7.[151,250,1905,1927]" captionTargetBox-0="[167,1421,190,1902]" captionTargetBox-1="[159,1396,236,1938]" captionTargetBox-2="[164,1425,213,1851]" captionTargetBox-3="[181,1404,222,1905]" captionTargetId-0="figure@4.[151,1435,179,1918]" captionTargetId-1="figure@5.[152,1435,193,1957]" captionTargetId-2="figure@6.[151,1435,193,1869]" captionTargetId-3="figure@7.[175,1420,193,1905]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 2. Chthamalus williamsi sp. nov., shell morphology A. Group of individuals on shell of Megabalanus volcano, top view. B. Uneroded specimen, top view. C. Holotype, external morphology, top view. D. Shell plates, inner view. E. Scutum and tergum, inner surface. F. scutum and tergum, exterior surface. Scale bars in mm." captionText-1="FIGURE 3. Chthamalus williamsi sp. nov. Line drawing of cirri under stereomicroscope. A. Cirrus I, B. Cirrus II, C. Cirrus III, D. Cirrus IV, E. Cirrus V, F. Cirrus VI. Scale bars in µm." captionText-2="FIGURE 4. Chthamalus williamsi sp. nov. Morphology of cirri (SEM—A I; light microscopy—J, K). A. Cirrus I, B. Tip of cirrus I, showing the serrulate setae, C. Serrulate setae on posterior ramus. D. Cirrus II, E. Tip of cirrus II, showing the saw-like serrulate setae, F. Serrulate setae on anterior ramus of cirrus II, G. Cirrus III, H. Intermediate segments of anterior ramus of cirrus III, I. Serrulate setae of cirrus III, J. Intermediate segments of anterior ramus of cirrus V. H. Intermediate segments of anterior ramus of cirrus VI. Scale bars in µm." captionText-3="FIGURE 5. Chthamalus williamsi sp. nov. Morphology of mouth parts, penis and cirral setae (SEM) A. Maxilla, B. Serrulate setae on maxilla, C. Maxillule, D. Cutting edge of maxillule, E. Mandible, F. Lower margin of mandible, G. Labrum, H. Anterior margin of labrum, inner view, I. Penis, terminal part, J. Tip of penis, K. Serrulate setae on cirrus I., L. Serrulate setae on posterior side of cirrus I. Scale bars in µm." httpUri-0="https://zenodo.org/record/236745/files/figure.png" httpUri-1="https://zenodo.org/record/236746/files/figure.png" httpUri-2="https://zenodo.org/record/236747/files/figure.png" httpUri-3="https://zenodo.org/record/236748/files/figure.png" pageId="2" pageNumber="549">Figures 25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FDA688AFFFE50A14FF2BFBCF2292FB46" pageId="2" pageNumber="549" type="materials_examined">
<paragraph id="B503DB24FFE50A14FF2BFBCF2292FB46" blockId="2.[151,1437,1080,2005]" pageId="2" pageNumber="549">
<emphasis id="87C80736FFE50A14FF2BFBCF278DFBAB" bold="true" box="[151,389,1080,1105]" pageId="2" pageNumber="549">Material examined.</emphasis>
<typeStatus id="6A076586FFE50A14FE2DFBCF27F5FBAB" box="[401,509,1080,1105]" pageId="2" pageNumber="549" type="holotype">Holotype</typeStatus>
. NMNS-006534-00001. Intertidal rocks (low shores, at the same tidal zone as
<taxonomicName id="72BCA0A7FFE50A14FF2BFBAB279DFB8F" box="[151,405,1116,1141]" class="Maxillopoda" family="Balanidae" genus="Megabalanus" kingdom="Animalia" order="Sessilia" pageId="2" pageNumber="549" phylum="Arthropoda" rank="species" species="volcano">
<emphasis id="87C80736FFE50A14FF2BFBAB279DFB8F" box="[151,405,1116,1141]" italics="true" pageId="2" pageNumber="549">Megabalanus volcano</emphasis>
</taxonomicName>
) at Shi-Ti-Ping, Hualien,
<collectingCountry id="CDAB9BB4FFE50A14FD6DFBAB252BFB8F" box="[721,803,1116,1141]" name="Taiwan" pageId="2" pageNumber="549">Taiwan</collectingCountry>
(
<geoCoordinate id="D088BDE3FFE50A14FC84FBAB25B7FB8E" box="[824,959,1116,1140]" direction="north" orientation="latitude" pageId="2" pageNumber="549" precision="9" value="23.488333">23°29.30N</geoCoordinate>
,
<geoCoordinate id="D088BDE3FFE50A14FC6DFBAB226AFB8E" box="[977,1122,1116,1140]" direction="east" orientation="longitude" pageId="2" pageNumber="549" precision="9" value="121.505">121°30.30E</geoCoordinate>
,
<date id="C102FDE4FFE50A14FBC8FBAB2313FB8E" box="[1140,1307,1116,1141]" pageId="2" pageNumber="549" value="2009-05-12">12 May 2009</date>
).
<typeStatus id="6A076586FFE50A14FA8FFBAA2391FB8F" box="[1331,1433,1117,1141]" pageId="2" pageNumber="549" type="paratype">Paratype</typeStatus>
. NMNS-006534-00002. Cheng Gong, Taitung,
<collectingCountry id="CDAB9BB4FFE50A14FD78FB77251EFB63" box="[708,790,1152,1177]" name="Taiwan" pageId="2" pageNumber="549">Taiwan</collectingCountry>
(
<geoCoordinate id="D088BDE3FFE50A14FC92FB7725BDFB63" box="[814,949,1152,1177]" direction="north" orientation="latitude" pageId="2" pageNumber="549" precision="9" value="23.091">23°05.46N</geoCoordinate>
,
<geoCoordinate id="D088BDE3FFE50A14FC76FB772253FB62" box="[970,1115,1152,1176]" direction="east" orientation="longitude" pageId="2" pageNumber="549" precision="9" value="121.37217">121°22.33E</geoCoordinate>
, 0
<date id="C102FDE4FFE50A14FBC1FB772310FB62" box="[1149,1304,1152,1176]" pageId="2" pageNumber="549" value="2008-09-02">2 Sep. 2008</date>
).
<typeStatus id="6A076586FFE50A14FA8FFB762391FB63" box="[1331,1433,1153,1177]" pageId="2" pageNumber="549" type="paratype">Paratype</typeStatus>
. ASIZCR-000328. Cheng Gong, Taitung,
<collectingCountry id="CDAB9BB4FFE50A14FDD9FB5324BFFB47" box="[613,695,1188,1213]" name="Taiwan" pageId="2" pageNumber="549">Taiwan</collectingCountry>
(
<geoCoordinate id="D088BDE3FFE50A14FD7AFB532546FB47" box="[710,846,1188,1213]" direction="north" orientation="latitude" pageId="2" pageNumber="549" precision="9" value="23.091">23°05.46N</geoCoordinate>
,
<geoCoordinate id="D088BDE3FFE50A14FCE5FB5325E2FB46" box="[857,1002,1188,1212]" direction="east" orientation="longitude" pageId="2" pageNumber="549" precision="9" value="121.37217">121°22.33E</geoCoordinate>
, 0
<date id="C102FDE4FFE50A14FBB8FB532286FB46" box="[1028,1166,1188,1212]" pageId="2" pageNumber="549" value="2008-09-02">2 Sep. 2008</date>
).
</paragraph>
</subSubSection>
<subSubSection id="FDA688AFFFE50A14FF7BFB3F247CFAD3" pageId="2" pageNumber="549" type="diagnosis">
<paragraph id="B503DB24FFE50A14FF7BFB3F247CFAD3" blockId="2.[151,1437,1080,2005]" pageId="2" pageNumber="549">
<emphasis id="87C80736FFE50A14FF7BFB3F2749FB1B" bold="true" box="[199,321,1224,1249]" pageId="2" pageNumber="549">Diagnosis.</emphasis>
Shell conical; 6 plates, external surfaces white, smooth or with faint ribs; scutum equilateral triangular, tergal articular margin straight, external surface of scutum with horizontal striations; tergum triangular, scutal articular margin straight, spur blunt.
</paragraph>
</subSubSection>
<subSubSection id="FDA688AFFFE50A15FF7BFAC3253DFE9E" lastPageId="3" lastPageNumber="550" pageId="2" pageNumber="549" type="description">
<paragraph id="B503DB24FFE50A14FF7BFAC325BFF96B" blockId="2.[151,1437,1080,2005]" pageId="2" pageNumber="549">
<emphasis id="87C80736FFE50A14FF7BFAC32751FAB7" bold="true" box="[199,345,1332,1357]" pageId="2" pageNumber="549">Description.</emphasis>
Shell small, about 3.0 mm basal diameter, conical; 6 plates (rostrum, carina, paired lateral and paired rostral-lateral;
<figureCitation id="2D87C7A1FFE50A14FE35FAAF27C2FA8A" box="[393,458,1368,1393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1940,1962]" captionTargetBox="[167,1421,190,1902]" captionTargetId="figure@4.[151,1435,179,1918]" captionTargetPageId="4" captionText="FIGURE 2. Chthamalus williamsi sp. nov., shell morphology A. Group of individuals on shell of Megabalanus volcano, top view. B. Uneroded specimen, top view. C. Holotype, external morphology, top view. D. Shell plates, inner view. E. Scutum and tergum, inner surface. F. scutum and tergum, exterior surface. Scale bars in mm." httpUri="https://zenodo.org/record/236745/files/figure.png" pageId="2" pageNumber="549">Fig. 2</figureCitation>
AD), externally white, surface smooth or with faint ribs, inner operculum pale orangecoloured; carina and rostrum with a pair of alae, carina short and wide, alae large; rostrum long and narrow, alae large; rostral-lateral triangular, interior surface with central mid rib, radii wide; lateral base wide, alae and radii present (
<figureCitation id="2D87C7A1FFE50A14FF4BFA332732FA26" box="[247,314,1476,1501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1940,1962]" captionTargetBox="[167,1421,190,1902]" captionTargetId="figure@4.[151,1435,179,1918]" captionTargetPageId="4" captionText="FIGURE 2. Chthamalus williamsi sp. nov., shell morphology A. Group of individuals on shell of Megabalanus volcano, top view. B. Uneroded specimen, top view. C. Holotype, external morphology, top view. D. Shell plates, inner view. E. Scutum and tergum, inner surface. F. scutum and tergum, exterior surface. Scale bars in mm." httpUri="https://zenodo.org/record/236745/files/figure.png" pageId="2" pageNumber="549">Fig. 2</figureCitation>
D); scutum white, triangular, tergal articular margin straight, perpendicular to basal margin, articular ridge not prominent, long with length extending 2/3 length of tergal articular margin, articular furrow narrow, deep, occludent margin inclined with fine teeth, scutal adductor scar oval, obvious, external surface of scutum with horizontal striations (
<figureCitation id="2D87C7A1FFE50A14FE39F9C727C1F9B2" box="[389,457,1584,1609]" captionStart="FIGURE 2" captionStartId="4.[151,250,1940,1962]" captionTargetBox="[167,1421,190,1902]" captionTargetId="figure@4.[151,1435,179,1918]" captionTargetPageId="4" captionText="FIGURE 2. Chthamalus williamsi sp. nov., shell morphology A. Group of individuals on shell of Megabalanus volcano, top view. B. Uneroded specimen, top view. C. Holotype, external morphology, top view. D. Shell plates, inner view. E. Scutum and tergum, inner surface. F. scutum and tergum, exterior surface. Scale bars in mm." httpUri="https://zenodo.org/record/236745/files/figure.png" pageId="2" pageNumber="549">Fig. 2</figureCitation>
E, F); tergum white, triangular, scutal articular margin straight, extending down to spur, spur blunt, wide, about ½ width of basal margin, articular furrow wide, shallow, basal margin slightly concave (
<figureCitation id="2D87C7A1FFE50A14FAD3F9A326AEF96A" captionStart="FIGURE 2" captionStartId="4.[151,250,1940,1962]" captionTargetBox="[167,1421,190,1902]" captionTargetId="figure@4.[151,1435,179,1918]" captionTargetPageId="4" captionText="FIGURE 2. Chthamalus williamsi sp. nov., shell morphology A. Group of individuals on shell of Megabalanus volcano, top view. B. Uneroded specimen, top view. C. Holotype, external morphology, top view. D. Shell plates, inner view. E. Scutum and tergum, inner surface. F. scutum and tergum, exterior surface. Scale bars in mm." httpUri="https://zenodo.org/record/236745/files/figure.png" pageId="2" pageNumber="549">Fig. 2</figureCitation>
E, F), 5 fine depressor crests at basal margin, external surface striated.
</paragraph>
<paragraph id="B503DB24FFE50A15FF7BF96B2586FF2E" blockId="2.[151,1437,1080,2005]" lastBlockId="3.[151,1436,151,861]" lastPageId="3" lastPageNumber="550" pageId="2" pageNumber="549">
Cirrus I without conical spines on dorsal surface, posterior ramus 5-segmented, anterior ramus 6-segmented (
<figureCitation id="2D87C7A1FFE50A14FF23F93726EDF922" box="[159,229,1728,1753]" captionStart="FIGURE 3" captionStartId="5.[151,250,1979,2001]" captionTargetBox="[159,1396,236,1938]" captionTargetId="figure@5.[152,1435,193,1957]" captionTargetPageId="5" captionText="FIGURE 3. Chthamalus williamsi sp. nov. Line drawing of cirri under stereomicroscope. A. Cirrus I, B. Cirrus II, C. Cirrus III, D. Cirrus IV, E. Cirrus V, F. Cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236746/files/figure.png" pageId="2" pageNumber="549">Fig. 3</figureCitation>
A), both rami with serrulate setae, setae without basal guard (
<figureCitation id="2D87C7A1FFE50A14FC7BF9372207F922" box="[967,1039,1728,1753]" captionStart="FIGURE 4" captionStartId="6.[151,250,1890,1912]" captionTargetBox="[164,1425,213,1851]" captionTargetId="figure@6.[151,1435,193,1869]" captionTargetPageId="6" captionText="FIGURE 4. Chthamalus williamsi sp. nov. Morphology of cirri (SEM—A I; light microscopy—J, K). A. Cirrus I, B. Tip of cirrus I, showing the serrulate setae, C. Serrulate setae on posterior ramus. D. Cirrus II, E. Tip of cirrus II, showing the saw-like serrulate setae, F. Serrulate setae on anterior ramus of cirrus II, G. Cirrus III, H. Intermediate segments of anterior ramus of cirrus III, I. Serrulate setae of cirrus III, J. Intermediate segments of anterior ramus of cirrus V. H. Intermediate segments of anterior ramus of cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236747/files/figure.png" pageId="2" pageNumber="549">Fig. 4</figureCitation>
AC); cirrus II, anterior ramus 5- segmented, posterior ramus 7-segmented (
<figureCitation id="2D87C7A1FFE50A14FD35F91324D9F906" box="[649,721,1764,1789]" captionStart="FIGURE 3" captionStartId="5.[151,250,1979,2001]" captionTargetBox="[159,1396,236,1938]" captionTargetId="figure@5.[152,1435,193,1957]" captionTargetPageId="5" captionText="FIGURE 3. Chthamalus williamsi sp. nov. Line drawing of cirri under stereomicroscope. A. Cirrus I, B. Cirrus II, C. Cirrus III, D. Cirrus IV, E. Cirrus V, F. Cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236746/files/figure.png" pageId="2" pageNumber="549">Fig. 3</figureCitation>
B), both rami with 2
<typeStatus id="6A076586FFE50A14FC66F912221DF907" box="[986,1045,1765,1789]" pageId="2" pageNumber="549">types</typeStatus>
of serrulate setae, bidentate and plumose setae, all setae without basal guard (
<figureCitation id="2D87C7A1FFE50A14FD25F8FF24E9F8DA" box="[665,737,1800,1825]" captionStart="FIGURE 4" captionStartId="6.[151,250,1890,1912]" captionTargetBox="[164,1425,213,1851]" captionTargetId="figure@6.[151,1435,193,1869]" captionTargetPageId="6" captionText="FIGURE 4. Chthamalus williamsi sp. nov. Morphology of cirri (SEM—A I; light microscopy—J, K). A. Cirrus I, B. Tip of cirrus I, showing the serrulate setae, C. Serrulate setae on posterior ramus. D. Cirrus II, E. Tip of cirrus II, showing the saw-like serrulate setae, F. Serrulate setae on anterior ramus of cirrus II, G. Cirrus III, H. Intermediate segments of anterior ramus of cirrus III, I. Serrulate setae of cirrus III, J. Intermediate segments of anterior ramus of cirrus V. H. Intermediate segments of anterior ramus of cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236747/files/figure.png" pageId="2" pageNumber="549">Figs 4</figureCitation>
DF, 5K, L); cirrus III with rami equal length, anterior ramus 14-segmented, posterior ramus 12-segmented (
<figureCitation id="2D87C7A1FFE50A14FD13F8DB24F1F8BE" box="[687,761,1836,1861]" captionStart="FIGURE 3" captionStartId="5.[151,250,1979,2001]" captionTargetBox="[159,1396,236,1938]" captionTargetId="figure@5.[152,1435,193,1957]" captionTargetPageId="5" captionText="FIGURE 3. Chthamalus williamsi sp. nov. Line drawing of cirri under stereomicroscope. A. Cirrus I, B. Cirrus II, C. Cirrus III, D. Cirrus IV, E. Cirrus V, F. Cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236746/files/figure.png" pageId="2" pageNumber="549">Figs 3</figureCitation>
C, 4G), intermediate segments of cirrus III composed of 5 pairs of long serrulate setae (
<figureCitation id="2D87C7A1FFE50A14FE47F8A7244CF892" box="[507,580,1872,1897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1890,1912]" captionTargetBox="[164,1425,213,1851]" captionTargetId="figure@6.[151,1435,193,1869]" captionTargetPageId="6" captionText="FIGURE 4. Chthamalus williamsi sp. nov. Morphology of cirri (SEM—A I; light microscopy—J, K). A. Cirrus I, B. Tip of cirrus I, showing the serrulate setae, C. Serrulate setae on posterior ramus. D. Cirrus II, E. Tip of cirrus II, showing the saw-like serrulate setae, F. Serrulate setae on anterior ramus of cirrus II, G. Cirrus III, H. Intermediate segments of anterior ramus of cirrus III, I. Serrulate setae of cirrus III, J. Intermediate segments of anterior ramus of cirrus V. H. Intermediate segments of anterior ramus of cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236747/files/figure.png" pageId="2" pageNumber="549">Fig. 4</figureCitation>
H, I); cirrus IV with anterior ramus 14-segmented, posterior ramus 15- segmented (
<figureCitation id="2D87C7A1FFE50A14FE99F8832766F876" box="[293,366,1908,1933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1979,2001]" captionTargetBox="[159,1396,236,1938]" captionTargetId="figure@5.[152,1435,193,1957]" captionTargetPageId="5" captionText="FIGURE 3. Chthamalus williamsi sp. nov. Line drawing of cirri under stereomicroscope. A. Cirrus I, B. Cirrus II, C. Cirrus III, D. Cirrus IV, E. Cirrus V, F. Cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236746/files/figure.png" pageId="2" pageNumber="549">Fig. 3</figureCitation>
D); cirrus V with anterior ramus 18-segmented, posterior ramus 15-segmented (
<figureCitation id="2D87C7A1FFE50A14FA8DF8832372F876" box="[1329,1402,1908,1933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1979,2001]" captionTargetBox="[159,1396,236,1938]" captionTargetId="figure@5.[152,1435,193,1957]" captionTargetPageId="5" captionText="FIGURE 3. Chthamalus williamsi sp. nov. Line drawing of cirri under stereomicroscope. A. Cirrus I, B. Cirrus II, C. Cirrus III, D. Cirrus IV, E. Cirrus V, F. Cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236746/files/figure.png" pageId="2" pageNumber="549">Fig. 3</figureCitation>
E), intermediate segments of both rami with 3 pairs of long serrulate setae and 2 pairs of shorter setae (
<figureCitation id="2D87C7A1FFE50A14FB4AF86F2333F84A" box="[1270,1339,1944,1969]" captionStart="FIGURE 4" captionStartId="6.[151,250,1890,1912]" captionTargetBox="[164,1425,213,1851]" captionTargetId="figure@6.[151,1435,193,1869]" captionTargetPageId="6" captionText="FIGURE 4. Chthamalus williamsi sp. nov. Morphology of cirri (SEM—A I; light microscopy—J, K). A. Cirrus I, B. Tip of cirrus I, showing the serrulate setae, C. Serrulate setae on posterior ramus. D. Cirrus II, E. Tip of cirrus II, showing the saw-like serrulate setae, F. Serrulate setae on anterior ramus of cirrus II, G. Cirrus III, H. Intermediate segments of anterior ramus of cirrus III, I. Serrulate setae of cirrus III, J. Intermediate segments of anterior ramus of cirrus V. H. Intermediate segments of anterior ramus of cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236747/files/figure.png" pageId="2" pageNumber="549">Fig. 4</figureCitation>
J); cirrus VI with anterior ramus 19-segmented, posterior ramus 17-segmented, intermediate segments of anterior and posterior rami composed of 3 pairs of long serrulate setae and 2 pairs of shorter setae (
<figureCitation id="2D87C7A1FFE40A15FBEBFF602292FF4A" box="[1111,1178,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1890,1912]" captionTargetBox="[164,1425,213,1851]" captionTargetId="figure@6.[151,1435,193,1869]" captionTargetPageId="6" captionText="FIGURE 4. Chthamalus williamsi sp. nov. Morphology of cirri (SEM—A I; light microscopy—J, K). A. Cirrus I, B. Tip of cirrus I, showing the serrulate setae, C. Serrulate setae on posterior ramus. D. Cirrus II, E. Tip of cirrus II, showing the saw-like serrulate setae, F. Serrulate setae on anterior ramus of cirrus II, G. Cirrus III, H. Intermediate segments of anterior ramus of cirrus III, I. Serrulate setae of cirrus III, J. Intermediate segments of anterior ramus of cirrus V. H. Intermediate segments of anterior ramus of cirrus VI. Scale bars in µm." httpUri="https://zenodo.org/record/236747/files/figure.png" pageId="3" pageNumber="550">Fig. 4</figureCitation>
K); penis long, slender, sparse setae along length, tip with dense bundle of setae (
<figureCitation id="2D87C7A1FFE40A15FCA5FF4B2554FF2F" box="[793,860,188,213]" captionStart="FIGURE 5" captionStartId="7.[151,250,1905,1927]" captionTargetBox="[181,1404,222,1905]" captionTargetId="figure@7.[175,1420,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Chthamalus williamsi sp. nov. Morphology of mouth parts, penis and cirral setae (SEM) A. Maxilla, B. Serrulate setae on maxilla, C. Maxillule, D. Cutting edge of maxillule, E. Mandible, F. Lower margin of mandible, G. Labrum, H. Anterior margin of labrum, inner view, I. Penis, terminal part, J. Tip of penis, K. Serrulate setae on cirrus I., L. Serrulate setae on posterior side of cirrus I. Scale bars in µm." httpUri="https://zenodo.org/record/236748/files/figure.png" pageId="3" pageNumber="550">
Fig.
<date id="C102FDE4FFE40A15FCF1FF4B2554FF2F" box="[845,860,188,213]" pageId="3" pageNumber="550">5</date>
</figureCitation>
I, J).
</paragraph>
<paragraph id="B503DB24FFE40A15FF7BFF28253DFE9E" blockId="3.[151,1436,151,861]" pageId="3" pageNumber="550">
Maxilla bilobed, serrulate setae on both lobes (
<figureCitation id="2D87C7A1FFE40A15FD5CFF28252CFF02" box="[736,804,223,248]" captionStart="FIGURE 5" captionStartId="7.[151,250,1905,1927]" captionTargetBox="[181,1404,222,1905]" captionTargetId="figure@7.[175,1420,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Chthamalus williamsi sp. nov. Morphology of mouth parts, penis and cirral setae (SEM) A. Maxilla, B. Serrulate setae on maxilla, C. Maxillule, D. Cutting edge of maxillule, E. Mandible, F. Lower margin of mandible, G. Labrum, H. Anterior margin of labrum, inner view, I. Penis, terminal part, J. Tip of penis, K. Serrulate setae on cirrus I., L. Serrulate setae on posterior side of cirrus I. Scale bars in µm." httpUri="https://zenodo.org/record/236748/files/figure.png" pageId="3" pageNumber="550">Fig. 5</figureCitation>
A, B); maxillule notched, 3 large setae above notch, 16 setae below notch (
<figureCitation id="2D87C7A1FFE40A15FEC7FEF327C9FEE7" box="[379,449,260,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1905,1927]" captionTargetBox="[181,1404,222,1905]" captionTargetId="figure@7.[175,1420,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Chthamalus williamsi sp. nov. Morphology of mouth parts, penis and cirral setae (SEM) A. Maxilla, B. Serrulate setae on maxilla, C. Maxillule, D. Cutting edge of maxillule, E. Mandible, F. Lower margin of mandible, G. Labrum, H. Anterior margin of labrum, inner view, I. Penis, terminal part, J. Tip of penis, K. Serrulate setae on cirrus I., L. Serrulate setae on posterior side of cirrus I. Scale bars in µm." httpUri="https://zenodo.org/record/236748/files/figure.png" pageId="3" pageNumber="550">Fig. 5</figureCitation>
C, D); mandible with 4 teeth, fourth bidentate, lower margin straight, with 14 setae, inferior angle with pair of large setae (
<figureCitation id="2D87C7A1FFE40A15FDE6FED024A8FEBA" box="[602,672,295,320]" captionStart="FIGURE 5" captionStartId="7.[151,250,1905,1927]" captionTargetBox="[181,1404,222,1905]" captionTargetId="figure@7.[175,1420,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Chthamalus williamsi sp. nov. Morphology of mouth parts, penis and cirral setae (SEM) A. Maxilla, B. Serrulate setae on maxilla, C. Maxillule, D. Cutting edge of maxillule, E. Mandible, F. Lower margin of mandible, G. Labrum, H. Anterior margin of labrum, inner view, I. Penis, terminal part, J. Tip of penis, K. Serrulate setae on cirrus I., L. Serrulate setae on posterior side of cirrus I. Scale bars in µm." httpUri="https://zenodo.org/record/236748/files/figure.png" pageId="3" pageNumber="550">Fig. 5</figureCitation>
E, F); labrum concave, notch absent, dense setae at mid region of cutting margin, latter with numerous fine teeth (
<figureCitation id="2D87C7A1FFE40A15FD0CFEBB24F9FE9F" box="[688,753,332,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1905,1927]" captionTargetBox="[181,1404,222,1905]" captionTargetId="figure@7.[175,1420,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Chthamalus williamsi sp. nov. Morphology of mouth parts, penis and cirral setae (SEM) A. Maxilla, B. Serrulate setae on maxilla, C. Maxillule, D. Cutting edge of maxillule, E. Mandible, F. Lower margin of mandible, G. Labrum, H. Anterior margin of labrum, inner view, I. Penis, terminal part, J. Tip of penis, K. Serrulate setae on cirrus I., L. Serrulate setae on posterior side of cirrus I. Scale bars in µm." httpUri="https://zenodo.org/record/236748/files/figure.png" pageId="3" pageNumber="550">Fig. 5</figureCitation>
G, H).
</paragraph>
</subSubSection>
<subSubSection id="FDA688AFFFE40A15FF7BFE98236BFE2A" pageId="3" pageNumber="550" type="etymology">
<paragraph id="B503DB24FFE40A15FF7BFE98236BFE2A" blockId="3.[151,1436,151,861]" pageId="3" pageNumber="550">
<emphasis id="87C80736FFE40A15FF7BFE982746FE72" bold="true" box="[199,334,367,392]" pageId="3" pageNumber="550">Etymology.</emphasis>
<taxonomicName id="72BCA0A7FFE40A15FEE6FE98245CFE72" box="[346,596,367,392]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="87C80736FFE40A15FEE6FE98245CFE72" box="[346,596,367,392]" italics="true" pageId="3" pageNumber="550">Chthamalus williamsi</emphasis>
</taxonomicName>
is named in honour of Prof. Gray A. Williams, Director of the Swire Institute of Marine Sciences, School of Biological Science, the University of
<collectingCountry id="CDAB9BB4FFE40A15FBB9FE622285FE57" box="[1029,1165,405,429]" name="Hong Kong" pageId="3" pageNumber="550">Hong Kong</collectingCountry>
, for life-long friendship and mentorship with the first author and for his contributions to the intertidal ecology in the South
<collectingCountry id="CDAB9BB4FFE40A15FB55FE402324FE2A" box="[1257,1324,439,464]" name="China" pageId="3" pageNumber="550">China</collectingCountry>
Sea.
</paragraph>
</subSubSection>
<subSubSection id="FDA688AFFFE40A15FF7BFE2B25F9FCA6" pageId="3" pageNumber="550" type="distribution">
<paragraph id="B503DB24FFE40A15FF7BFE2B2448FD52" blockId="3.[151,1436,151,861]" pageId="3" pageNumber="550">
<emphasis id="87C80736FFE40A15FF7BFE2B2768FE0F" bold="true" box="[199,352,476,501]" pageId="3" pageNumber="550">Distribution.</emphasis>
<taxonomicName id="72BCA0A7FFE40A15FEDBFE2B2454FE0F" box="[359,604,476,501]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
<emphasis id="87C80736FFE40A15FEDBFE2B2454FE0F" box="[359,604,476,501]" italics="true" pageId="3" pageNumber="550">Chthamalus williamsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CFBBA4DFFE40A15FDDEFE2B24B4FE0F" box="[610,700,476,501]" pageId="3" pageNumber="550" rank="species">
<emphasis id="87C80736FFE40A15FDDEFE2B24B4FE0F" bold="true" box="[610,700,476,501]" pageId="3" pageNumber="550">sp. nov.</emphasis>
</taxonomicNameLabel>
is common inside crevices of the mid shore of the intertidal zone, and on the shell surfaces of the barnacle
<taxonomicName id="72BCA0A7FFE40A15FDCEFE082217FDE2" authority="Pilsbry, 1916" authorityName="Pilsbry" authorityYear="1916" box="[626,1055,511,536]" class="Maxillopoda" family="Balanidae" genus="Megabalanus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="volcano">
<emphasis id="87C80736FFE40A15FDCEFE082565FDE2" box="[626,877,511,536]" italics="true" pageId="3" pageNumber="550">Megabalanus volcano</emphasis>
(
<bibRefCitation id="D12DA6D5FFE40A15FCC2FE08221EFDE2" author="Pilsbry" box="[894,1046,511,536]" pageId="3" pageNumber="550" refString="Pilsbry, H. A. (1916) The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum, including a monograph of the American species. Bulletin of the United States National Museum, 93, v - xi, 1 - 366, figs. 1 - 99, pls. 1 - 76." type="journal article" year="1916">Pilsbry, 1916</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="72BCA0A7FFE40A15FBE6FDF626DFFDC7" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Maxillopoda" family="Balanidae" genus="Megabalanus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="tintinnabulum">
<emphasis id="87C80736FFE40A15FBE6FDF62317FDE2" box="[1114,1311,511,536]" italics="true" pageId="3" pageNumber="550">M. tintinnabulum</emphasis>
(
<bibRefCitation id="D12DA6D5FFE40A15FA8CFE0826C7FDC7" author="Linnaeus" pageId="3" pageNumber="550" refString="Linnaeus, C. (1758) Systema Naturae. Vol. 10. Impensis direct Laurentii Salvii, Stockholm, Sweden, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
on low shores of the eastern waters of
<collectingCountry id="CDAB9BB4FFE40A15FD35FDD324D7FDC7" box="[649,735,548,573]" name="Taiwan" pageId="3" pageNumber="550">Taiwan</collectingCountry>
.
<taxonomicName id="72BCA0A7FFE40A15FD55FDD325D6FDC7" box="[745,990,548,573]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="87C80736FFE40A15FD55FDD325D6FDC7" box="[745,990,548,573]" italics="true" pageId="3" pageNumber="550">Chthamalus williamsi</emphasis>
</taxonomicName>
has not been identified previously from
<collectingCountry id="CDAB9BB4FFE40A15FF2BFDB026E1FD9A" box="[151,233,583,608]" name="Taiwan" pageId="3" pageNumber="550">Taiwan</collectingCountry>
as it is not easily spotted, due to its small size and low shore location. This species is absent from the N and NE coasts of
<collectingCountry id="CDAB9BB4FFE40A15FE80FD9B2786FD7F" box="[316,398,620,645]" name="Taiwan" pageId="3" pageNumber="550">Taiwan</collectingCountry>
although intensive sampling was conducted (
<figureCitation id="2D87C7A1FFE40A15FC16FD9B25F9FD7E" box="[938,1009,620,645]" captionStart="FIGURE 1" captionStartId="1.[151,250,1246,1268]" captionTargetBox="[457,1129,265,1224]" captionTargetId="figure@1.[457,1129,265,1225]" captionTargetPageId="1" captionText="FIGURE 1. Map of Taiwan, showing collection sites of Chthamalus williamsi sp. nov. Empty circles indicate C. williamsi sp. nov. was absent from samples. Solid circle indicates sites where C. williamsi sp. nov. was collected." httpUri="https://zenodo.org/record/236744/files/figure.png" pageId="3" pageNumber="550">Fig. 1</figureCitation>
), suggesting its distribution may be associated with the Kuroshio Current.
</paragraph>
<paragraph id="B503DB24FFE40A15FF7BFD4325F9FCA6" blockId="3.[151,1436,151,861]" pageId="3" pageNumber="550">
<emphasis id="87C80736FFE40A15FF7BFD4327B0FD37" bold="true" box="[199,440,692,717]" pageId="3" pageNumber="550">Molecular analysis.</emphasis>
From both maximum likelihood and Bayesian inference trees of a COI region,
<taxonomicName id="72BCA0A7FFE40A15FF2BFD202787FD0A" box="[151,399,727,752]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="87C80736FFE40A15FF2BFD202787FD0A" box="[151,399,727,752]" italics="true" pageId="3" pageNumber="550">Chthamalus williamsi</emphasis>
</taxonomicName>
shows a distinct divergence from
<taxonomicName id="72BCA0A7FFE40A15FCA2FD2F25C8FD0A" box="[798,960,727,752]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="challengeri">
<emphasis id="87C80736FFE40A15FCA2FD2F25C8FD0A" box="[798,960,727,752]" italics="true" pageId="3" pageNumber="550">C. challengeri</emphasis>
</taxonomicName>
,
<taxonomicName id="72BCA0A7FFE40A15FC6DFD2F2220FD0A" box="[977,1064,727,752]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="dalli">
<emphasis id="87C80736FFE40A15FC6DFD2F2220FD0A" box="[977,1064,727,752]" italics="true" pageId="3" pageNumber="550">C. dalli</emphasis>
</taxonomicName>
,
<taxonomicName id="72BCA0A7FFE40A15FB85FD2F229EFD0A" box="[1081,1174,728,752]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="moro">
<emphasis id="87C80736FFE40A15FB85FD2F229EFD0A" box="[1081,1174,728,752]" italics="true" pageId="3" pageNumber="550">C. moro</emphasis>
</taxonomicName>
,
<taxonomicName id="72BCA0A7FFE40A15FB1BFD2F234FFD0A" box="[1191,1351,727,752]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="87C80736FFE40A15FB1BFD2F234FFD0A" box="[1191,1351,727,752]" italics="true" pageId="3" pageNumber="550">C. malayensis</emphasis>
</taxonomicName>
and
<taxonomicName id="72BCA0A7FFE40A15FA3EFD2F26F8FCEE" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="87C80736FFE40A15FA3EFD2F26F8FCEE" italics="true" pageId="3" pageNumber="550">C. sinensis</emphasis>
</taxonomicName>
(sequence divergence&gt;0.13 K2P distance from all the
<taxonomicName id="72BCA0A7FFE40A15FC0CFD0B223FFCEF" box="[944,1079,764,789]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="genus">
<emphasis id="87C80736FFE40A15FC0CFD0B223FFCEF" box="[944,1079,764,789]" italics="true" pageId="3" pageNumber="550">Chthamalus</emphasis>
</taxonomicName>
species compared;
<tableCitation id="F83EEE9FFFE40A15FA93FD0B2383FCEE" box="[1327,1419,764,789]" captionStart="TABLE 1" captionStartId="8.[119,195,545,565]" captionTargetBox="[119,1905,614,1617]" captionTargetPageId="8" captionText="TABLE 1. Gene bank accession number, DNA Barcoding reference number and the locality of all samples collected in present study. * refers to the holotype of the species; # refers to sequences newly generated from this study." httpUri="http://table.plazi.org/id/E1C38BACFFEF0A1EFFCBFDD627ADFDB5" pageId="3" pageNumber="550" tableUuid="E1C38BACFFEF0A1EFFCBFDD627ADFDB5">Table 1</tableCitation>
).
<taxonomicName id="72BCA0A7FFE40A15FF2BFCE8278AFCC2" box="[151,386,799,824]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="87C80736FFE40A15FF2BFCE8278AFCC2" box="[151,386,799,824]" italics="true" pageId="3" pageNumber="550">Chthamalus sinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="72BCA0A7FFE40A15FE7CFCD72446FCC2" box="[448,590,799,824]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="87C80736FFE40A15FE7CFCD72446FCC2" box="[448,590,799,824]" italics="true" pageId="3" pageNumber="550">C. neglectus</emphasis>
</taxonomicName>
grouped in the same clade, suggesting
<taxonomicName id="72BCA0A7FFE40A15FB9DFCD722A7FCC2" box="[1057,1199,799,824]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="87C80736FFE40A15FB9DFCD722A7FCC2" box="[1057,1199,799,824]" italics="true" pageId="3" pageNumber="550">C. neglectus</emphasis>
</taxonomicName>
is a synonym of
<taxonomicName id="72BCA0A7FFE40A15FA3EFCD726F8FCA6" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="87C80736FFE40A15FA3EFCD726F8FCA6" italics="true" pageId="3" pageNumber="550">C. sinensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="D12DA6D5FFE40A15FE97FCB327E9FCA6" author="Liu" box="[299,481,836,861]" pageId="3" pageNumber="550" refString="Liu, R. Y. &amp; Ren, X. Q. (2007) Fauna Sinica. Invertebrata. Vol. 42 Crustacea Cirripedia Thoracica. Science Press, Beijing, 633 pp." type="book" year="2007">Liu &amp; Ren 2007</bibRefCitation>
for synonyms of
<taxonomicName id="72BCA0A7FFE40A15FD1BFCB22539FCA7" box="[679,817,836,861]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="87C80736FFE40A15FD1BFCB22539FCA7" box="[679,817,836,861]" italics="true" pageId="3" pageNumber="550">C. neglectus</emphasis>
</taxonomicName>
and
<taxonomicName id="72BCA0A7FFE40A15FCD4FCB225E9FCA6" box="[872,993,837,860]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="87C80736FFE40A15FCD4FCB225E9FCA6" box="[872,993,837,860]" italics="true" pageId="3" pageNumber="550">C. sinensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>