133 lines
16 KiB
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133 lines
16 KiB
XML
<document id="CC5C1F594E55F4DA38A0216136EE4D50" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="3AA2B0AF2AB924791387FF2F6A658973" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus mutatus Omer-Cooper 1970" docType="treatment" docVersion="5" lastPageNumber="170" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="169" updateTime="1701314941532" updateUser="plazi">
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<mods:titleInfo id="B018E95E638B9041E34058F070C0A095">
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<mods:title id="0623D52989FAAE48153DE396AE4965E5">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="8C53D116C5FD818804929D25E27F8ACF">Bistroem, Olof</mods:namePart>
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<mods:namePart id="43EF0799C280E5526063DEFED92C38A9">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="EF62E2ACE19B6FA9D772444212ADA878">Bergsten, Johannes</mods:namePart>
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<mods:typeOfResource id="93827A8A8ABB4C160B1C4ED134B8D378">text</mods:typeOfResource>
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<mods:title id="ADD0C3E659ABECAC3FA63793FAE4B60F">ZooKeys</mods:title>
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<mods:date id="D93C5FFDB90A6E776A536EB78DDC97F0">2015</mods:date>
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<mods:detail id="1EAF16DCFC767F3A73169148673AA971" type="volume">
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<mods:number id="63CD84461A50AC7297DC657CDC5BF190">542</mods:number>
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<mods:start id="9B68A404F351AF5764B79526C75B0E6D">1</mods:start>
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<mods:end id="9361BFBCC3CEB2F7B5E597C53A5744CA">379</mods:end>
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<mods:url id="F4B4E25F8C8EA5739737E1655F601CAC">http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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<mods:classification id="C6A581582558B745530321FFA700D69B">journal article</mods:classification>
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<mods:identifier id="FD34E3DC0766CDCB87E2FE4C73DA54E6" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="3AA2B0AF2AB924791387FF2F6A658973" ID-GBIF-Taxon="127862773" LSID="urn:lsid:plazi:treatment:3AA2B0AF2AB924791387FF2F6A658973" httpUri="http://treatment.plazi.org/id/3AA2B0AF2AB924791387FF2F6A658973" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169">
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<subSubSection id="611DEF9C126AC6A4806DCC25F8102A20" pageId="168" pageNumber="169" type="multiple">
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<paragraph id="88A1DA4FE893F1A41CF60A2661EB4B71" pageId="168" pageNumber="169">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="543EC753F8ECC4468BCFD955A40A5A65" pageId="168" pageNumber="169" type="nomenclature">
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<paragraph id="38B2DF73F377C0C06F733C31F854D8D2" pageId="168" pageNumber="169">
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<taxonomicName id="F8C651B80DC1C44BBFA35DE0583579B4" ID-CoL="3RM99" authority="Omer-Cooper, 1970" authorityName="Omer-Cooper" authorityYear="1970" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mutatus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="mutatus">Laccophilus mutatus Omer-Cooper, 1970</taxonomicName>
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Figs 137-138, 323-325, 468-469, 564
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</paragraph>
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</subSubSection>
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<subSubSection id="C908558A297D0AB30E8195B36EA356CD" pageId="168" pageNumber="169" type="reference_group">
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<paragraph id="6C44A836185B07BA033512FCEFE6D732" pageId="168" pageNumber="169">
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<taxonomicName id="7F838CFCAC52C7C888059C6F0C14B8B7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mutatus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="mutatus">Laccophilus mutatus</taxonomicName>
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<bibRefCitation id="9690F99746C3D61D549E3189FD5E7587" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="285 - 294" title="New and little known species of African Laccophilus (Coleoptera: Dytiscidae)." volume="33" year="1970">Omer-Cooper 1970</bibRefCitation>
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: 286, 287 (original description, faunistics); Nilsson and Persson 199a: 79 (status discussion);
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<bibRefCitation id="B072901A4742A15CEBD2444D1B895793" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 247 (catalogue, faunistics);
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<bibRefCitation id="FCF65A900C74E223199D1EC3E775E5A9" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 214 (catalogue, faunistics).
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</paragraph>
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</subSubSection>
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<subSubSection id="08A098C0E8D7C74452AE8BA12D72CC87" pageId="168" pageNumber="169" type="type locality">
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<paragraph id="083E5F573E030BC5D7E30351A8FB1E00" pageId="168" pageNumber="169">Type locality.</paragraph>
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<paragraph id="9D7F284EA0E8873F6811268E7ECAF52C" pageId="168" pageNumber="169">Kenya: Athi River.</paragraph>
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</subSubSection>
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<subSubSection id="311C8501D8EFC72225D55D339839CE13" pageId="168" pageNumber="169" type="type material">
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<paragraph id="6F9DC6FC16DED5FDDE7F0E48DCFB55C4" pageId="168" pageNumber="169">Type material studied</paragraph>
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<paragraph id="B5A7B47B00A095B01F1126388822B7E7" pageId="168" pageNumber="169">
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(4 exs.). Holotype: male: "Athi River 19.X. 1957 (CAS, not examined). - Paratypes: "Paratype /
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<taxonomicName id="78E99D9E6ED45199C452E03D1582AEC2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mutatus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="mutatus">Laccophilus mutatus</taxonomicName>
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O-C. / Kenya Athi River 1530 m X-19-1957 / E.S. Ross & R.E. Leech collectors" (2 exs. AMGS); "Paratype /
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<taxonomicName id="1177F97F97871E8637F25D3EA2873A59" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mutatus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="mutatus">Laccophilus mutatus</taxonomicName>
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O-C. / Kenya 17 mi. SW of Nairobi, 1800 m, XI-24-1957 / E.S. Ross & R.E. Leech collectors" (2 exs. AMGS).
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</paragraph>
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</subSubSection>
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<subSubSection id="71483ABB3DBAE38F5AF59143F2AD830C" pageId="168" pageNumber="169" type="materials_examined">
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<paragraph id="83EFBD99BD51CBDACB4CE1C54572D186" pageId="168" pageNumber="169">Additional material studied</paragraph>
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<paragraph id="41CAB1C97920FC872610C889CCE537D4" pageId="168" pageNumber="169">
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(30 exs.). Kenya: "Ol Toroto Athi Riv., 5.7. 1970 E.S. Brown leg." (2 exs. BMNH; habitus in Figs 468-469); "Nairobi 7.10. 1967 / Reichart leg." (3 exs. USNM); same data, except "16.10. 1967" (2 exs. USNM); same data except "3.11. 1967" (7 exs. USNM, 4 exs. MZH); same data except "Oct. 1968 / Conway leg." (1 ex. USNM); "Kiserian 26.10.1967 / Reichart leg." (3 exs. USNM); "Langata Rd. 4.11. 1967 / Reichart leg."(1 ex. USNM); "10 km N Nyeri Kinganjo 3.11. 1995 Wewalka" (4 exs. CGW, 1 ex. MZH). - Tanzania: "Kilimandj.
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<normalizedToken id="E19C83CD5539985B9525199FC641FF7B" originalValue="Sjöstedt">Sjoestedt</normalizedToken>
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/ Kibonto 1-1200 m /
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<taxonomicName id="20EF69AA160202D2FFE77A15A672C40F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
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Shp" (1 ex. NHRS).
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</paragraph>
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</subSubSection>
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<subSubSection id="A7783E938DBF7D57336B4F6AC78CAE82" pageId="168" pageNumber="169" type="diagnosis">
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<paragraph id="E5CF92580C4BFC38AD9F3799FD569732" pageId="168" pageNumber="169">Diagnosis.</paragraph>
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<paragraph id="181D161341891CE0D34094136E5AEFE5" pageId="168" pageNumber="169">
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<taxonomicName id="A54E00ECDBB2C06886FDB03A694997AD" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mutatus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="mutatus">Laccophilus mutatus</taxonomicName>
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externally resembles of
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<taxonomicName id="6C916330714E24AD93A44B1D8C103F92" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
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. Peculiar features of penis apex (extreme apex provided with two distinct processes in
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<taxonomicName id="A5E102434BB63625239E88C2A32B26D2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mutatus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="mutatus">Laccophilus mutatus</taxonomicName>
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) distinguish
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<taxonomicName id="D09A6F71E3741BF4B1FBFF7B5BB2AAE1" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mutatus" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="mutatus">Laccophilus mutatus</taxonomicName>
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from
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<taxonomicName id="AD88EBFCB9E4B6FD2F2BB6D683B85F9C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
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as well as does appearance of elytral colour pattern in general. Beware of
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<taxonomicName id="E3113D42C34E4E573986825530A45D21" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="168" pageNumber="169" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
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specimens with reduced elytral colour pattern.
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</paragraph>
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</subSubSection>
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<subSubSection id="69C740712CB9E09638C1FA2E83BE790A" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169" type="description">
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<paragraph id="9861DADCB3FC8DE25202259DBF17ECE9" pageId="168" pageNumber="169">Description.</paragraph>
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<paragraph id="FF51015C6A613A0E22AC8B2A52487B19" pageId="168" pageNumber="169">Body length 4.2-4.7 mm, width 2.4-2.6 mm. Elytra with, in part, reduced dark longitudinal lines; habitus, as in Figs 468-469.</paragraph>
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<paragraph id="908024DA3C7470E30B5FD92AABF8A1F2" pageId="168" pageNumber="169">Head: Pale ferrugineous. Slightly mat, finely microsculptured. Reticulation double, coarser meshes contain 2-5 smaller meshes. At eyes with some very fine, irregularly distributed punctures.</paragraph>
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<paragraph id="B1CCCDFD0C27CDAA01FA847DFFF17A72" pageId="168" pageNumber="169">Pronotum: Pale ferrugineous, anteriorly in middle with a bilobed, slightly darker spot. Submat, distinctly microsculpturd. Large meshes contain 3-6 fine meshes. Scattered, irregular fine punctures present.</paragraph>
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<paragraph id="C4E9161884DC748D27ACB4BCA1F35E72" pageId="168" pageNumber="169">Elytra: Pale ferrugineous, with dark, in part reduced, longitudinal markings (Figs 468-469). Punctation indistinct, posteriorly with sparse and fine punctation. Very finely microsculptured and reticulation almost simple, of one kind.</paragraph>
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<paragraph id="A6D073CA4C49E6EC350CC68F03E2D457" pageId="169" pageNumber="170">
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<pageBreakToken id="33CE30546D2BD7AA694C593ED443053A" pageId="169" pageNumber="170" start="start">Ventral</pageBreakToken>
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aspect: Pale ferrugineous to pale brown, no distinct colour pattern discernible. Rather shiny, although extensively with fine but in part inditinct microsculpture. Metacoxal plates with some shallow, vague and transverse furrows. Abdomen with some inwards curved striae. Almost impunctate. Apex of prosternal process slender and pointed. Apical ventrite asymmetric, with a small, lateral knob (Fig. 137).
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</paragraph>
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<paragraph id="1D191AA118CEE464E97F1C71FA0E4279" pageId="169" pageNumber="170">Legs: Pro- and mesotarsus rather slender, provided with distinct suckers.</paragraph>
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<paragraph id="9D064CFF1C6E58A8BCF7D3266AD75FC4" pageId="169" pageNumber="170">Male genitalia: Penis, apical portion, with two distinct processes (Fig. 324). In lateral aspect penis curved from base to apex; apically, on external outline, provided with a narrow membranous area (Fig. 323).</paragraph>
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<paragraph id="D6C71A94A7616017D3AA40DF1846E6A1" pageId="169" pageNumber="170">Female: Apical ventrite apically not incised (Fig. 138). Pro- and mesotarsus slender.</paragraph>
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</subSubSection>
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<subSubSection id="EEC679803F224C7FC3BEF5FB71EA6857" pageId="169" pageNumber="170" type="distribution">
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<paragraph id="28937EB4FE27D4EB2B15304D3858735C" pageId="169" pageNumber="170">Distribution.</paragraph>
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<paragraph id="975CC0827A8E729FE1FF44E19EA3F12F" pageId="169" pageNumber="170">Kenya, Tanzania (Fig. 564).</paragraph>
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</subSubSection>
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<subSubSection id="2E0CB4D9F19E41126BF347F3F726F1A5" pageId="169" pageNumber="170" type="collecting circumstances">
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<paragraph id="3BE7603B951B05D55362E82368A728AF" pageId="169" pageNumber="170">Collecting circumstances.</paragraph>
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<paragraph id="A2D1EA69177145725D9EB1D268C2A38E" pageId="169" pageNumber="170">Unknown, not documented.</paragraph>
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</subSubSection>
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</treatment>
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</document> |