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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.544.6074" ID-GBIF-Dataset="0e5e47a0-f8f3-4039-b0e1-c159fef0b9f6" ID-PMC="PMC4714384" ID-Pensoft-Pub="1313-2970-544-1" ID-PubMed="26798289" ID-ZBK="C5978BD0145B40F8ACDEB27371B7B9A4" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-544-1" ModsDocOrigin="ZooKeys 544" ModsDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" checkinTime="1451243640953" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2015" docId="29760A70A3810A10D93C631DE936EEA6" docLanguage="en" docName="ZooKeys 544: 1-407" docOrigin="ZooKeys 544" docSource="http://dx.doi.org/10.3897/zookeys.544.6074" docTitle="Mecyclothorax anchisteus Liebherr, 2015, sp. n." docType="treatment" docUuid="FF5D8759-E3F4-45D6-9FE2-67EEFE7B39B6" docUuidSource="ZooBank" docVersion="5" lastPageNumber="58" masterDocId="FF92FFC2FF9FFFC7191EFFB8FF91AF5D" masterDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" masterLastPageNumber="407" masterPageNumber="1" pageNumber="56" updateTime="1668162300918" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liebherr, James K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>544</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>407</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.544.6074</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.544.6074</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-544-1</mods:identifier>
<mods:identifier type="ZBK">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
<mods:identifier type="ZooBank">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127853611" LSID="urn:lsid:zoobank.org:act:FF5D8759-E3F4-45D6-9FE2-67EEFE7B39B6" httpUri="http://treatment.plazi.org/id/29760A70A3810A10D93C631DE936EEA6" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56">
<subSubSection pageId="55" pageNumber="56" type="multiple">
<paragraph pageId="55" pageNumber="56">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="55" pageNumber="56" type="nomenclature">
<paragraph pageId="55" pageNumber="56">
(019)
<taxonomicName LSID="http://zoobank.org/FF5D8759-E3F4-45D6-9FE2-67EEFE7B39B6" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax anchisteus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="anchisteus">Mecyclothorax anchisteus</taxonomicName>
<taxonomicNameLabel pageId="55" pageNumber="56">sp. n.</taxonomicNameLabel>
Figs 32, 33A
</paragraph>
</subSubSection>
<subSubSection pageId="56" pageNumber="57" type="diagnosis">
<paragraph pageId="56" pageNumber="57">
<pageBreakToken pageId="56" pageNumber="57" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="56" pageNumber="57">
This species is easily diagnosed by the distinctly cordate pronotum, MPW/BPW = 1.57, with glabrous hind angles. The pronotal lateral margins are distinctly convergent anterad the acute and acuminately projected hind angles (Fig. 33A). The head, pronotum, and elytra are a uniformly dark rufous. The eyes are moderately convex, ocular ratio = 1.43, but rather small, covering only
<normalizedToken originalValue="¾">3/4</normalizedToken>
of the little protruded ocular lobe; ocular lobe ratio = 0.76. The elytra are broadly subovoid, with the humeri narrowly rounded; MEW/HuW = 2.0. The discal elytral intervals are covered with irregular transverse-line microsculpture, the lines not joined into a mesh. Setal formula 2 1 2 1[sae]. Standardized body length 5.1 mm.
</paragraph>
</subSubSection>
<subSubSection lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="description">
<paragraph pageId="56" pageNumber="57">Description</paragraph>
<paragraph lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
(n = 1). Head capsule with frontal grooves broad near clypeus, a lateral carina to anterior supraorbital seta; dorsal surface of neck slightly concave; labral anterior margin broadly, shallowly emarginate, antennae filiform, sparse pelage of short setae present on antennomeres 2-3; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW/PL = 1.26; median base depressed relative to disc, with small punctures and lateral wrinkles; basal margin straight, slightly indented posterad laterobasal depressions; median longitudinal impression shallow, very finely incised at depth; anterior transverse impression very shallow, narrow, crossed medially by longitudinal wrinkles; anterior callosity nearly flat, crossed by indistinct wrinkles; front angles slightly projected, tightly rounded; anterior width greater than basal width, APW/BPW = 1.07; lateral marginal depression narrow throughout, margin upturned to finely beaded before sinuation; laterobasal depression smooth, laterally elevated to projected lateral margin. Proepisternum with smooth hind marginal groove; prosternal process medially depressed, a broad lateral marginal bead that is narrowed anteriorly. Elytra with convex disc, sides depressed; basal groove slightly recurved to broadly rounded humeral angle; parascutellar seta present; parascutellar striole continuous, with 4-5 punctures; sutural interval more convex than intervals 2-4, sutural juncture upraised; sutural and 2nd striae of subequal depth from base to apex; striae 1-8 complete, stria 7 slightly shallower, associated intervals moderately convex; discal striae with slightly irregular punctulae basally, smooth and deep apically; 7th and 8th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.27
<normalizedToken originalValue="×">x</normalizedToken>
and 0.49
<normalizedToken originalValue="0.52×">-0.52x</normalizedToken>
elytral length; setal impressions extended over 2/3 width of interval 3; subapical seta present, apical seta absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series 6 setae; elytral marginal depression narrow throughout, margin slightly upraised at humerus; subapical sinuation shallow, broad. Mesepisternum with ~5 punctures in 1 row; metepisternal width to length ratio = 0.81; metepisternum/metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1-5; suture between ventrites 2 and 3 complete; apical ventrite of female with 4 equally spaced marginal setae and a median trapezoid of 4 setae, the basal pair longer. Legs-metatarsomere 1/metatibial length ratio = 0.22; metatarsomere 4 length along outer lobe 1.37
<normalizedToken originalValue="×">x</normalizedToken>
medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci very broad, median area rough to carinate. Microsculpture of vertex with isodiametric to slightly stretched isodiametric sculpticells in transverse rows; pronotal disc with transverse mesh, sculpticell breadth 3
<normalizedToken originalValue="4×">-4x</normalizedToken>
length;
<pageBreakToken pageId="57" pageNumber="58" start="start">pronotal</pageBreakToken>
median base with isodiametric to transverse sculpticells; elytral disc with irregular transverse lines, apex with more regular transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of antennomere 1 flavous, antennomeres 2-3 rufoflavous, 4-11 rufobrunneous; proepipleuron and proepisternum rufobrunneous; elytral apex paler than disc, rufoflavous to position of subapical seta; elytral epipleuron rufoflavous laterally, rufobrunneous ventrally, metepisternum rufobrunneous; abdomen medially rufobrunneous, laterally rufoflavous; abdominal apical ventrite 6 with apical 1/3 paler, flavous; metafemur flavous with piceous cloud covering basal half; metatibia rufoflavous with brunneous cast.
</paragraph>
<paragraph pageId="57" pageNumber="58">Female reproductive tract. The lone female holotype was not dissected.</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="58" type="holotype">
<paragraph pageId="57" pageNumber="58">Holotype.</paragraph>
<paragraph pageId="57" pageNumber="58">
Female (NMNH) labeled: HI:Maui Haleakala N.P. / Kipahulu Vy. West Camp / 20-V-1998 lot01 1950 m / el. pyrethrum fog mossy / ohia D.A. Polhemus // HOLOTYPE /
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
/
<taxonomicName lsidName="anchisteus" pageId="57" pageNumber="58" rank="species" species="anchisteus">anchisteus</taxonomicName>
/ Sharp / J.K. Liebherr 2015 (black-margined red label).
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="58" type="etymology">
<paragraph pageId="57" pageNumber="58">Etymology.</paragraph>
<paragraph pageId="57" pageNumber="58">
The species epithet
<taxonomicName lsidName="anchisteus" pageId="57" pageNumber="58" rank="species" species="anchisteus">anchisteus</taxonomicName>
is Greek for next of kin (
<bibRefCitation author="Jaeger, EC" journalOrPublisher="Charles C Thomas, Publisher, Springfield, IL" pageId="308" pageNumber="309" title="A Source-book of Biological Names and Terms" year="1955">Jaeger 1955</bibRefCitation>
), signifying this
<normalizedToken originalValue="species">species'</normalizedToken>
close affinities to
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cognatus" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="cognatus">Mecyclothorax cognatus</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax affinis" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="affinis">Mecyclothorax affinis</taxonomicName>
, and
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax consanguineus" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="consanguineus">Mecyclothorax consanguineus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="58" type="distribution and habitat">
<paragraph pageId="57" pageNumber="58">Distribution and habitat.</paragraph>
<paragraph pageId="57" pageNumber="58">
This species is known from a single specimen collected at the head of
<normalizedToken originalValue="Kīpahulu">Kīpahulu</normalizedToken>
Valley (Fig. 32) after application of pyrethrin fog to a mossy
<normalizedToken originalValue="ōhia">'ōhi'a</normalizedToken>
tree.
</paragraph>
</subSubSection>
</treatment>
</document>