284 lines
21 KiB
XML
284 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1086.77408" ID-GBIF-Dataset="dd81d919-f6d9-4fda-bbd1-7c45de307fc2" ID-PMC="PMC8863766" ID-Pensoft-Pub="1313-2970-1086-33" ID-Pensoft-UUID="43AFBD7D61105F598ADA74076D751C11" ID-PubMed="35210912" ID-ZooBank="6C32148CA0144AC488EA82A1509E44F8" ModsDocID="1313-2970-1086-33" checkinTime="1644966834654" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Meng, Kaibaryer & Fekete, Judit" docDate="2022" docId="083421BCC53C5D7A9A2A77EA57F4A71F" docLanguage="en" docName="ZooKeys 1086: 33-76" docOrigin="ZooKeys 1086" docPubDate="2022-02-15" docSource="http://dx.doi.org/10.3897/zookeys.1086.77408" docTitle="Laeocathaica minwui Pall-Gergely 2022, sp. nov." docType="treatment" docUuid="C96CE473-3692-4306-9AB2-2AEFDA307824" docUuidSource="ZooBank" docVersion="5" id="43AFBD7D61105F598ADA74076D751C11" lastPageNumber="33" masterDocId="43AFBD7D61105F598ADA74076D751C11" masterDocTitle="A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae)" masterLastPageNumber="76" masterPageNumber="33" pageNumber="33" updateTime="1668151486723" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022, Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
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<mods:affiliation>Adria setany 10 G 2 / 5., Budapest 1148, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Meng, Kaibaryer</mods:namePart>
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<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fekete, Judit</mods:namePart>
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<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, H- 8200 Veszprem, Egyetem u. 10, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1086</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>76</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1086.77408</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1086.77408</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1086-33</mods:identifier>
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<mods:identifier type="ZooBank">6C32148CA0144AC488EA82A1509E44F8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">43AFBD7D61105F598ADA74076D751C11</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="193377852" LSID="urn:lsid:zoobank.org:act:C96CE473-3692-4306-9AB2-2AEFDA307824" httpUri="http://treatment.plazi.org/id/083421BCC53C5D7A9A2A77EA57F4A71F" lastPageNumber="33" pageId="0" pageNumber="33">
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<subSubSection pageId="0" pageNumber="33" type="nomenclature">
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<paragraph pageId="0" pageNumber="33">
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<taxonomicName LSID="http://zoobank.org/C96CE473-3692-4306-9AB2-2AEFDA307824" authority="Páll-Gergely" authorityName="Pall-Gergely" authorityYear="2022" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica minwui" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="minwui" status="sp. nov.">
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Laeocathaica minwui
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="description">
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<paragraph pageId="0" pageNumber="33">
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Laeocathaica minwui sp. nov. (holotype, SMF 95019). Scale bar: 10 mm. Photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure21" httpUri="https://binary.pensoft.net/fig/646396" pageId="0" pageNumber="33">Figure 21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="type material">
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<paragraph pageId="0" pageNumber="33">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Holotype</emphasis>
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China • China, O. Sy-tshuan, coll. C.R. Boettger ex coll.
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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ex coll. L. Fuchs, SMF 95019 (D = 23.1, H = 9.1) (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Laeocathaica minwui sp. nov. (holotype, SMF 95019). Scale bar: 10 mm. Photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure21" httpUri="https://binary.pensoft.net/fig/646396" pageId="0" pageNumber="33">21</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Paratypes</emphasis>
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China • Yangtze-Tal, coll. Jetschin ex coll. Beddome, SMF 95020/1 (mixed sample with
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. carinifera" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="carinifera">
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<emphasis italics="true" pageId="0" pageNumber="33">L. carinifera</emphasis>
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</taxonomicName>
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) • W. China, Sy-tshuan, coll. Kobelt ex coll.
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, SMF 6920/1 (det. Wu 2008, labelled as
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. christinae" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">L. christinae</emphasis>
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</taxonomicName>
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) • China: Hupei: Kao-cha-hien, coll. O. Boettger ex coll. B. Schmacker, 1893, SMF 24263/1 (det. Wu 2008, labelled as
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. christinae" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">L. christinae</emphasis>
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</taxonomicName>
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) • China, O. Sy-tshuan, coll.
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, SMF 24255a/5 • China: Chang-Yang, coll. O. Boettger ex coll. M. Schmacker, SMF 42563/2 • Moupin, leg.
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<normalizedToken originalValue="Abbé">Abbe</normalizedToken>
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David, MNHN-IM-2014-7939/14 (probably erroneous locality).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="diagnosis">
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<paragraph pageId="0" pageNumber="33">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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A rather large
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<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica</emphasis>
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</taxonomicName>
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species with a sharp keel, a domed dorsal side, an oval aperture and a marmorated ventral side.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="description">
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<paragraph pageId="0" pageNumber="33">Description.</paragraph>
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<paragraph pageId="0" pageNumber="33">Shell sinistral, depressed, with domed dorsal side, keel strong, situated in the middle of body whorl, whitish; dorsal side latte-coloured, with darker and paler areas alternating as the shell grows; ventral side with white and pale brownish (latte) spiral bands forming a marmorated colour pattern; inner side of umbilicus with brownish spiral band; protoconch light brownish, ca. 1.5 whorls, finely granulose, slightly protrudes above first whorls of teleoconch; entire shell consists of six whorls; dorsal side finely ribbed, ventral side smoother, only with growth lines; umbilicus ca. one third of shell width; shows all whorls; periumbilical keel absent; aperture oblique to shell axis, oval, without incision at the position of keel; peristome white, expanded and slightly thickened, but not reflexed (only in direction of umbilicus); parietal callus practically absent, only with some additional translucent calcareous layer.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Measurements (in mm)</emphasis>
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: D = 23.1, H = 9.1 (holotype).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="33">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<taxonomicName authorityName="Pall-Gergely" authorityYear="2022" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica minwui" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="minwui">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica minwui</emphasis>
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</taxonomicName>
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sp. nov. has been confused with
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. christinae" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">L. christinae</emphasis>
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</taxonomicName>
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in museum collections, probably due to the lack of examination of the types of
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. christinae" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">L. christinae</emphasis>
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</taxonomicName>
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. However,
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. christinae" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">L. christinae</emphasis>
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</taxonomicName>
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has a flatter dorsal side, a more upper-situated peripheral keel, a darker brown (instead of latte) colour, a more uniformly white ventral side with a brown spiral band inside the umbilicus, and brownish spots. In contrast, in the new species the ventral side is characterised by a marmorated (marbled-like) pattern resulted by the fusing of whitish and pale brown spiral bands.
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<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica filippina" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="filippina">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica filippina</emphasis>
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</taxonomicName>
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has a notched aperture at the position of the peripheral keel, a more brownish colour, and a less marmorated ventral side. See also Table
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<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Differences between L. carinifera, L. christinae, L. filippina, and L. minwui sp. nov." httpUri="http://table.plazi.org/id/25E5100B9318E8C262B91B73285454BE" pageId="0" pageNumber="33" tableUuid="25E5100B9318E8C262B91B73285454BE">3</tableCitation>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.1086.77408.figure21" httpUri="https://binary.pensoft.net/fig/646396" pageId="0" pageNumber="33" start="Figure 21" startId="F21">
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Figure 21.</emphasis>
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<taxonomicName authorityName="Pall-Gergely" authorityYear="2022" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica minwui" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="minwui">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica minwui</emphasis>
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</taxonomicName>
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sp. nov. (holotype, SMF 95019). Scale bar: 10 mm. Photographs: B.
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption ID-Table-UUID="25E5100B9318E8C262B91B73285454BE" httpUri="http://table.plazi.org/id/25E5100B9318E8C262B91B73285454BE" pageId="0" pageNumber="33" start="Table 3" startId="T3">
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Table 3.</emphasis>
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Differences between
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. carinifera" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="carinifera">
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<emphasis italics="true" pageId="0" pageNumber="33">L. carinifera</emphasis>
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</taxonomicName>
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,
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. christinae" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">L. christinae</emphasis>
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</taxonomicName>
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,
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. filippina" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="filippina">
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<emphasis italics="true" pageId="0" pageNumber="33">L. filippina</emphasis>
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</taxonomicName>
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, and
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. minwui" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="minwui">
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<emphasis italics="true" pageId="0" pageNumber="33">L. minwui</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="33">
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<table pageId="0" pageNumber="33">
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<tr pageId="0" pageNumber="33">
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<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Species</th>
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<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Dorsal surface</th>
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<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Keel on body whorl</th>
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<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Aperture (palatal part)</th>
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<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Colour of dorsal side</th>
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<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Ventral side</th>
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</tr>
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<tr pageId="0" pageNumber="33">
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. carinifera" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="carinifera">
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<emphasis italics="true" pageId="0" pageNumber="33">L. carinifera</emphasis>
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</taxonomicName>
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</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">acute, in the middle of body whorl</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">lighter and darker brownish patches alternate rather abruptly resulting in a mosaic-like structure</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">umbilicus narrow, colour pale, or light and darker stripes alternate</td>
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</tr>
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<tr pageId="0" pageNumber="33">
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. christinae" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">L. christinae</emphasis>
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</taxonomicName>
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</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">nearly flat/ slightly domed</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">blunt to acute, upper part of body whorl</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
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same as in
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<taxonomicName baseAuthorityName="H. Adams" baseAuthorityYear="1870" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica carinifera" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="carinifera">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica carinifera</emphasis>
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</taxonomicName>
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, just brown colour darker and there is usually a brown and a white spiral band
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</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">umbilicus wider, mostly white with slender darker radial stripes and dark dots</td>
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</tr>
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<tr pageId="0" pageNumber="33">
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. filippina" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="filippina">
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<emphasis italics="true" pageId="0" pageNumber="33">L. filippina</emphasis>
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</taxonomicName>
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</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">acute, upper part of body whorl</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">notched</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
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paler than
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<taxonomicName baseAuthorityName="H. Adams" baseAuthorityYear="1870" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica carinifera" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="carinifera">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica carinifera</emphasis>
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</taxonomicName>
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and
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<taxonomicName baseAuthorityName="H. Adams" baseAuthorityYear="1870" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica christinae" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="christinae">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica christinae</emphasis>
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</taxonomicName>
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, light and brown patches alternate smoothly
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</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
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umbilicus wider, similar to
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<taxonomicName authorityName="Pall-Gergely" authorityYear="2022" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica minwui" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="minwui">
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<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica minwui</emphasis>
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</taxonomicName>
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sp. nov., but paler and less nicely marmorated
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</td>
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</tr>
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<tr pageId="0" pageNumber="33">
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
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<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. minwui" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="minwui">
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<emphasis italics="true" pageId="0" pageNumber="33">L. minwui</emphasis>
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</taxonomicName>
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sp. nov.
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</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">acute, upper part of body whorl</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">mostly cream/latte-coloured, lighter and darker patches alternate smoothly</td>
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<td colspan="1" pageId="0" pageNumber="33" rowspan="1">umbilicus wider, latte and white spiral bands form a marmorated pattern</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="etymology">
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<paragraph pageId="0" pageNumber="33">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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This new species is dedicated to and named after Dr. Min Wu, the leading expert of Chinese
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<taxonomicName authorityName="Pilsbry" authorityYear="1895" class="Gastropoda" family="Camaenidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="family">Camaenidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="distribution">
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<paragraph pageId="0" pageNumber="33">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="33">This new species is only known from historical samples from the Yangtze valley. Other samples labelled as being collected from Sichuan are not precise enough to understand their geographic origin.</paragraph>
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</subSubSection>
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</treatment>
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</document> |