1876 lines
267 KiB
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1876 lines
267 KiB
XML
<document id="CDFE7311504C681B0E633F99074D2B60" ID-CLB-Dataset="293865" ID-DOI="10.5852/ejt.2024.930.2501" ID-GBIF-Dataset="1838fbf7-fdce-4670-a56c-bd0f3e480199" ID-ISSN="2118-9773" ID-Zenodo-Dep="10978023" ID-ZooBank="3FA94665-E6B2-4455-B40D-4ACDB71AAA55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713263296736" checkinUser="plazi" docAuthor="García-Raso, J. Enrique, D'Udekem D'Acoz, Cédric, Moukrim, Abdellatif, Schubart, Christoph D. & Cuesta, Jose A." docDate="2024" docId="F730427DFF98FFF06244FD67EF33FE63" docLanguage="en" docName="EJT.2024.930.277-313.pdf" docOrigin="European Journal of Taxonomy 930 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2501/11189" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.39:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="39" docTitle="Polybius dioscurus García-Raso & Moukrim & Schubart & Cuesta 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="300" masterDocId="0B093A05FF92FFFC6032FFE5EB16FFB0" masterDocTitle="A new cryptic species of Polybiidae (Crustacea: Decapoda: Portunoidea) from the East Atlantic, with considerations on the genus Polybius" masterLastPageNumber="313" masterPageNumber="277" pageNumber="287" updateTime="1713790897190" updateUser="felipe" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0AD94EEA1D960489BC1CB73B02EEA481">A new cryptic species of Polybiidae (Crustacea: Decapoda: Portunoidea) from the East Atlantic, with considerations on the genus Polybius</mods:title>
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<mods:namePart id="B8F344FB8BFE6213AF965D5124F598DC">García-Raso, J. Enrique</mods:namePart>
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<mods:affiliation id="45EA931238473DC292C56A823AAE69C0">Department Biología Animal, Faculty Ciencias, University of Málaga, Campus de Teatinos s / n, 29006 Málaga, Spain.</mods:affiliation>
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<mods:nameIdentifier id="CEB97629B16E45DB5C928DB073550D79" type="email">garciaraso@uma.es</mods:nameIdentifier>
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<mods:namePart id="76FAB935B958D025661559B44D835FCD">D'Udekem D'Acoz, Cédric</mods:namePart>
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<mods:affiliation id="E828401E76EB93637209B2FB008E0010">Service Heritage, Royal Belgian Institute of Natural Sciences, Rue Vautier 29, 1000 Brussels, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="E10313DBE3866F315ACAC0944FB9494B" type="email">cdudekem@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="0C4E34A7603886BF25C52DFE5CF7D91B">Moukrim, Abdellatif</mods:namePart>
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<mods:affiliation id="267B67CBD068E81CDC0E2CA7285910FA">Laboratoire Aquamar, Fac. Sc. Agadir, Université Ibn Zohr, Morocco.</mods:affiliation>
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<mods:nameIdentifier id="BF7E929EED303AA0D51F51A3960FC1F5" type="email">moukrim@uhp.ac.ma</mods:nameIdentifier>
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<mods:namePart id="32D6E045B6333B55C559B60D2886F9DA">Schubart, Christoph D.</mods:namePart>
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<mods:affiliation id="6FFAD840C39E73B7767C64E5B52DCF79">Deceased 21 March 2023. Former address: Zoology and Evolutionary Biology, University of Regensburg, 93040 Regensburg, Germany.</mods:affiliation>
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<mods:namePart id="D4080975B7BF153A4E81AB1954430529">Cuesta, Jose A.</mods:namePart>
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<mods:affiliation id="E25FD4AADDFC6DDBDBC8AFD5329E2868">Instituto de Ciencias Marinas de Andalucía (ICMAN-CSIC), Avda. República Saharaui, 2, 11519 Puerto Real, Cádiz, Spain.</mods:affiliation>
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<mods:nameIdentifier id="6B76E4F429C63E8D4825D579FF5BCD2D" type="email">jose.cuesta@icman.csic.es</mods:nameIdentifier>
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<mods:title id="439CECF1B2775A1838F2037521874D0D">European Journal of Taxonomy</mods:title>
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<mods:date id="20B0C6916A03524168DE1C7C8324A95C">2024</mods:date>
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<treatment id="F730427DFF98FFF06244FD67EF33FE63" ID-DOI="http://doi.org/10.5281/zenodo.10979179" ID-GBIF-Taxon="225690869" ID-Zenodo-Dep="10979179" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63" httpUri="http://treatment.plazi.org/id/F730427DFF98FFF06244FD67EF33FE63" lastPageId="12" lastPageNumber="300" pageId="10" pageNumber="287">
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<taxonomicName id="B89988E8FF98FFF66244FD67E84DFD2D" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[630,859,642,669]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
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<emphasis id="4DED2F79FF98FFF66244FD67E84DFD2D" bold="true" box="[630,859,642,669]" italics="true" pageId="10" pageNumber="287">Polybius dioscurus</emphasis>
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<taxonomicNameLabel id="56DE9202FF98FFF66351FD66E8ABFD2D" box="[867,957,643,669]" pageId="10" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="7F26F36BFF98FFF66259FD2CE8DFFD53" blockId="10.[365,1223,642,739]" box="[619,969,712,739]" pageId="10" pageNumber="287">
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<figureCitation id="E7A2EFEEFF98FFF66259FD2CE9C5FD53" box="[619,723,712,739]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="14.[189,232,1892,1918]" captionStartId-1="15.[189,232,1892,1918]" captionStartId-2="16.[189,232,1928,1954]" captionTargetBox-0="[247,1340,265,1845]" captionTargetBox-1="[278,1309,282,1800]" captionTargetBox-2="[190,1398,265,1875]" captionTargetId-0="figure-16@14.[247,1340,265,1845]" captionTargetId-1="figure-11@15.[278,1309,265,1847]" captionTargetId-2="figure-17@16.[190,1398,265,1875]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." captionText-1="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." captionText-2="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi-0="http://doi.org/10.5281/zenodo.10978033" figureDoi-1="http://doi.org/10.5281/zenodo.10978037" figureDoi-2="http://doi.org/10.5281/zenodo.10978041" httpUri-0="https://zenodo.org/record/10978033/files/figure.png" httpUri-1="https://zenodo.org/record/10978037/files/figure.png" httpUri-2="https://zenodo.org/record/10978041/files/figure.png" pageId="10" pageNumber="287">Figs 3–5</figureCitation>
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,
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<figureCitation id="E7A2EFEEFF98FFF662EDFD2DE83EFD53" box="[735,808,712,739]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[341,1247,265,1812]" captionTargetId="figure-17@18.[341,1247,265,1812]" captionTargetPageId="18" captionText="Fig. 6. Dorsal view of orbital region and adjacent lateral rostral teeth. A. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). B. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). C. Polybius dioscurus sp. nov., ♀ from Málaga (CRUST_ICMAN/3765) D. Polybius vernalis, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species." figureDoi="http://doi.org/10.5281/zenodo.10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="10" pageNumber="287">6A, C</figureCitation>
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,
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<figureCitation id="E7A2EFEEFF98FFF66306FD2CE84FFD53" box="[820,857,713,739]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="10" pageNumber="287">7B</figureCitation>
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,
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||
<figureCitation id="E7A2EFEEFF98FFF66357FD2CE8BBFD53" box="[869,941,713,739]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[218,1370,265,1801]" captionTargetId="figure-17@20.[218,1370,265,1801]" captionTargetPageId="20" captionText="Fig. 8. Left gonopod 1 of male. A–B. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). A. Ventral view. B. Dorsal view. C–D. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C. Ventral view. D. Dorsal view. E–F. Polybius vernalis (Risso, 1827) from Tunisia, Bay of Tunis (RBINS, INV. 187235).E. Ventral view. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10978047" httpUri="https://zenodo.org/record/10978047/files/figure.png" pageId="10" pageNumber="287">8C–D</figureCitation>
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,
|
||
<figureCitation id="E7A2EFEEFF98FFF66388FD2CE8DFFD53" box="[954,969,713,739]" captionStart="Fig" captionStartId="22.[188,231,1963,1989]" captionTargetBox="[189,1398,558,1917]" captionTargetId="figure-108@22.[189,1398,558,1917]" captionTargetPageId="22" captionText="Fig. 9. Polybius dioscurus sp. nov., paratypes from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187230), colour variation (in life) (sex not recorded during photo session)." figureDoi="http://doi.org/10.5281/zenodo.10978049" httpUri="https://zenodo.org/record/10978049/files/figure.png" pageId="10" pageNumber="287">9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="3783A0E0FF98FFF6608FFCEEE852FACA" pageId="10" pageNumber="287" type="reference_group">
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<paragraph id="7F26F36BFF98FFF6608FFCEEEAEAFCDB" blockId="10.[189,1399,778,1403]" pageId="10" pageNumber="287">
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<treatmentCitationGroup id="5F89D445FF98FFF6608FFCEEEAEAFCDB" pageId="10" pageNumber="287">
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<taxonomicName id="B89988E8FF98FFF6608FFCEEEAA1FC94" authorityName="Leach" authorityYear="1814" box="[189,439,779,804]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="marmoreus">
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<emphasis id="4DED2F79FF98FFF6608FFCEEEAA1FC94" box="[189,439,779,804]" italics="true" pageId="10" pageNumber="287">Portunus marmoreus</emphasis>
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</taxonomicName>
|
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‒? Bell 1844–1853: 105, in part, not unnumbered fig. —?
|
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<treatmentCitation id="FE38D57AFF98FFF664A6FCEFEE47FC95" author="White A." box="[1172,1361,778,805]" page="50" pageId="10" pageNumber="287" year="1857">
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<bibRefCitation id="1B088E9AFF98FFF664A6FCEFEE47FC95" author="White A." box="[1172,1361,778,805]" pageId="10" pageNumber="287" refId="ref17726" refString="White A. 1857. A Popular History of British Crustacea; Comprising a Familiar Account of their Classification and Habits. Lovell Reeve, London. https: // doi. org / 10.5962 / bhl. title. 14410" type="book" year="1857">White 1857: 50</bibRefCitation>
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</treatmentCitation>
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||
, in part. —?
|
||
<treatmentCitation id="FE38D57AFF98FFF66156FCC8E92CFCF7" author="Palmer R." box="[356,570,813,839]" page="889" pageId="10" pageNumber="287" year="1927">
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<bibRefCitation id="1B088E9AFF98FFF66156FCC8E92CFCF7" author="Palmer R." box="[356,570,813,839]" pageId="10" pageNumber="287" pagination="877 - 908" refId="ref16457" refString="Palmer R. 1927. A revision of the genus " Portunus " (A. Milne Edwards, Bell, etc). Journal of the Marine Biological Association of the United Kingdom 14: 877 - 908. https: // doi. org / 10.1017 / S 0025315400051134" type="journal article" year="1927">Palmer 1927: 889</bibRefCitation>
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</treatmentCitation>
|
||
, in part (second abnormal specimen). —
|
||
<treatmentCitation id="FE38D57AFF98FFF66415FCC8EFF2FCF7" author="Nobre A." box="[1063,1252,813,839]" page="66" pageId="10" pageNumber="287" year="1931">
|
||
<bibRefCitation id="1B088E9AFF98FFF66415FCC8EFF2FCF7" author="Nobre A." box="[1063,1252,813,839]" pageId="10" pageNumber="287" refId="ref16236" refString="Nobre A. 1931. Crustaceos Decapodes e Stomatopodes Marinhos de Portugal. Imprensa Portuguesa, Porto." type="book" year="1931">Nobre 1931: 66</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 31–32;
|
||
<treatmentCitation id="FE38D57AFF98FFF660D8FCB5EA42FCDA" author="Nobre A." box="[234,340,848,874]" page="36" pageId="10" pageNumber="287" year="1936">
|
||
<bibRefCitation id="1B088E9AFF98FFF660D8FCB5EA42FCDA" author="Nobre A." box="[234,340,848,874]" pageId="10" pageNumber="287" refId="ref16254" refString="Nobre A. 1936. Crustaceos Decapodes e Stomatopodes Marinhos de Portugal. 2 a Edicao. Fauna Marinha de Portugal 4. Porto." type="book" year="1936">1936: 36</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 11 fig. 19.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFC91E83EFC3E" blockId="10.[189,1399,778,1403]" box="[189,808,883,910]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFC91E83EFC3E" box="[189,808,883,910]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFC91EA8AFC3E" authorityName="H. Lucas" authorityYear="1846" box="[189,412,884,910]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="barbarus">
|
||
<emphasis id="4DED2F79FF98FFF6608FFC91EA8AFC3E" box="[189,412,884,910]" italics="true" pageId="10" pageNumber="287">Portunus barbarus</emphasis>
|
||
</taxonomicName>
|
||
‒
|
||
<treatmentCitation id="FE38D57AFF98FFF6618BFC91E99BFC3E" author="Monod T." box="[441,653,883,910]" page="173" pageId="10" pageNumber="287" year="1956">
|
||
<bibRefCitation id="1B088E9AFF98FFF6618BFC91E99BFC3E" author="Monod T." box="[441,653,883,910]" pageId="10" pageNumber="287" pagination="1 - 674" refId="ref16073" refString="Monod T. 1956. Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire (Mem. IFAN) 45: 1 - 674." type="journal article" year="1956">Monod 1956: 173</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 179, in part.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFC7DEE0FFC02" blockId="10.[189,1399,778,1403]" box="[189,1305,919,946]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFC7DEE0FFC02" box="[189,1305,919,946]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFC7DEAABFC01" ID-CoL="7349H" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[189,445,919,945]" class="Malacostraca" family="Polybiidae" genus="Macropipus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="barbarus">
|
||
<emphasis id="4DED2F79FF98FFF6608FFC7DEAABFC01" box="[189,445,919,945]" italics="true" pageId="10" pageNumber="287">Macropipus barbarus</emphasis>
|
||
</taxonomicName>
|
||
‒
|
||
<treatmentCitation id="FE38D57AFF98FFF661E8FC72E81AFC01" author="Forest J. & Guinot D." box="[474,780,919,946]" page="37" pageId="10" pageNumber="287" year="1956">
|
||
<bibRefCitation id="1B088E9AFF98FFF661E8FC72E81AFC01" author="Forest J. & Guinot D." box="[474,780,919,946]" pageId="10" pageNumber="287" pagination="24 - 43" refId="ref14543" refString="Forest J. & Guinot D. 1956. Sur une collection de crustaces decapodes et stomatopodes des mers tunisiennes. Bulletin de la Station oceanographique de Salammbo 53: 24 - 43." type="journal article" year="1956">Forest & Guinot 1956: 37</bibRefCitation>
|
||
</treatmentCitation>
|
||
, in part (Atlantic specimens only), not fig. 5.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFC5EE8B5FC64" blockId="10.[189,1399,778,1403]" box="[189,931,954,981]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFC5EE8B5FC64" box="[189,931,954,981]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFC5EEAA5FC64" ID-CoL="7VR4Y" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[189,435,954,980]" class="Malacostraca" family="Polybiidae" genus="Macropipus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF98FFF6608FFC5EEAA5FC64" box="[189,435,954,980]" italics="true" pageId="10" pageNumber="287">Macropipus holsatus</emphasis>
|
||
</taxonomicName>
|
||
‒
|
||
<treatmentCitation id="FE38D57AFF98FFF661E2FC5FE80EFC65" author="Cardona Bendito A." box="[464,792,954,981]" page="152" pageId="10" pageNumber="287" year="1965">
|
||
<bibRefCitation id="1B088E9AFF98FFF661E2FC5FE80EFC65" author="Cardona Bendito A." box="[464,792,954,981]" pageId="10" pageNumber="287" pagination="133 - 159" refId="ref14027" refString="Cardona Bendito A. 1965. Decapodos marinos de la ria de Vigo. Investigacion Pesquera, Barcelona 28: 133 - 159." type="journal article" year="1965">Cardona Bendito 1965: 152</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 2 fig. d.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFC3BE85DFC47" blockId="10.[189,1399,778,1403]" box="[189,843,989,1016]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFC3BE85DFC47" box="[189,843,989,1016]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFC3BEAC0FC47" ID-CoL="3X8B2" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[189,470,990,1015]" class="Malacostraca" family="Polybiidae" genus="Macropipus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF98FFF6608FFC3BEAC0FC47" box="[189,470,990,1015]" italics="true" pageId="10" pageNumber="287">Macropipus marmoreus</emphasis>
|
||
</taxonomicName>
|
||
–?
|
||
<treatmentCitation id="FE38D57AFF98FFF66235FC38E9CAFC47" author="Bourdon R." box="[519,732,989,1016]" page="29" pageId="10" pageNumber="287" year="1965">
|
||
<bibRefCitation id="1B088E9AFF98FFF66235FC38E9CAFC47" author="Bourdon R." box="[519,732,989,1016]" pageId="10" pageNumber="287" refId="ref13909" refString="Bourdon R. 1965. Inventaire de la Faune marine de Roscoff. Editions de la Station Biologique de Roscoff, Roscoff." type="book" year="1965">Bourdon 1965: 29</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFBE4E86AFBAB" blockId="10.[189,1399,778,1403]" box="[189,892,1024,1051]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFBE4E86AFBAB" box="[189,892,1024,1051]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFBE4EAA6FBAA" ID-CoL="932XM" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[189,432,1024,1050]" class="Malacostraca" family="Polybiidae" genus="Macropipus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF98FFF6608FFBE4EAA6FBAA" box="[189,432,1024,1050]" italics="true" pageId="10" pageNumber="287">Macropipus vernalis</emphasis>
|
||
</taxonomicName>
|
||
‒
|
||
<treatmentCitation id="FE38D57AFF98FFF661FFFBE4E99EFBAB" author="Neves A. M." box="[461,648,1024,1051]" page="32" pageId="10" pageNumber="287" year="1975">
|
||
<bibRefCitation id="1B088E9AFF98FFF661FFFBE4E99EFBAB" author="Neves A. M." box="[461,648,1024,1051]" pageId="10" pageNumber="287" pagination="1 - 48" refId="ref16160" refString="Neves A. M. 1975. Sobre una coleccao de crustaceos decapodes da Bahia de Setubal (Portugal). Estudos Fauna Portuguesa 5: 1 - 48." type="journal article" year="1975">Neves 1975: 32</bibRefCitation>
|
||
</treatmentCitation>
|
||
, in part, figs 10, 11c.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFBC1E9BCFAA1" blockId="10.[189,1399,778,1403]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFBC1E9BCFAA1" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFBC1EABBFB8E" ID-CoL="932MP" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[189,429,1060,1086]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF98FFF6608FFBC1EABBFB8E" box="[189,429,1060,1086]" italics="true" pageId="10" pageNumber="287">Liocarcinus vernalis</emphasis>
|
||
</taxonomicName>
|
||
‒
|
||
<treatmentCitation id="FE38D57AFF98FFF661FAFBC1E9C5FB8E" author="Garcia Raso J. E." box="[456,723,1060,1086]" page="107" pageId="10" pageNumber="287" year="1984">
|
||
<bibRefCitation id="1B088E9AFF98FFF661FAFBC1E9C5FB8E" author="Garcia Raso J. E." box="[456,723,1060,1086]" pageId="10" pageNumber="287" pagination="105 - 113" refId="ref14602" refString="Garcia Raso J. E. 1984. Brachyura of the coast of Southern Spain. Spixiana 7 (2): 105 - 113." type="journal article" year="1984">
|
||
García Raso 1984:
|
||
<quantity id="B8615E8EFF98FFF66295FBC1E9C5FB8E" box="[679,723,1060,1086]" metricMagnitude="0" metricUnit="m" metricValue="2.7178" pageId="10" pageNumber="287" unit="in" value="107.0">107</quantity>
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
in part. —
|
||
<treatmentCitation id="FE38D57AFF98FFF66367FBC1EE3DFB8E" author="Gonzalez Gurriaran E. & Mendez M." box="[853,1323,1060,1086]" page="43" pageId="10" pageNumber="287" year="1986">
|
||
<bibRefCitation id="1B088E9AFF98FFF66367FBC1EE3DFB8E" author="Gonzalez Gurriaran E. & Mendez M." box="[853,1323,1060,1086]" pageId="10" pageNumber="287" refId="ref14628" refString="Gonzalez Gurriaran E. & Mendez M. 1986. Crustaceos Decapodos das Costas de Galicia. I. Brachyura: 1 - 242. Cuadernos da Area de Ciencias Bioloxicas, Seminario de Estudos galegos, vol. 2 (2. a. Ed.), O Castro-Sada, A Coruna." type="book" year="1986">González Gurriarán & Méndez 1986: 43</bibRefCitation>
|
||
</treatmentCitation>
|
||
(key), 96, fig. 30, 210, foto 19. —
|
||
<treatmentCitation id="FE38D57AFF98FFF66218FBA2E897FBD1" author="D'Udekem d'Acoz C." box="[554,897,1095,1121]" page="184" pageId="10" pageNumber="287" year="1989">
|
||
<bibRefCitation id="1B088E9AFF98FFF66218FBA2E897FBD1" author="D'Udekem d'Acoz C." box="[554,897,1095,1121]" pageId="10" pageNumber="287" pagination="166 - 205" refId="ref14147" refString="D'Udekem d'Acoz C. 1989. Seconde note sur les crustaces decapodes de la Bretagne. De Strandvlo 8 (4): 166 - 205." type="journal article" year="1989">d’Udekem d’Acoz 1989: 184</bibRefCitation>
|
||
</treatmentCitation>
|
||
, in part, fig. 16, not fig. 17; 1991: 84, fig. 1; 2011: 78, fig. 1. —
|
||
<treatmentCitation id="FE38D57AFF98FFF661FBFB8FE98CFB34" author="Adema J. P. H. M." box="[457,666,1130,1156]" page="111" pageId="10" pageNumber="287" year="1991">
|
||
<bibRefCitation id="1B088E9AFF98FFF661FBFB8FE98CFB34" author="Adema J. P. H. M." box="[457,666,1130,1156]" pageId="10" pageNumber="287" pagination="110 - 115" refId="ref13758" refString="Adema J. P. H. M. 1991. Een aanvulling op: " De krabben van Nederland en Belgie ": De grijze zwemkrab, Liocarcinus vernalis (Risso, 1816) in de Noordzee gevonden. Het Zeepaard 51 (5): 110 - 115." type="journal article" year="1991">Adema 1991: 111</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 1–2. —
|
||
<treatmentCitation id="FE38D57AFF98FFF66306FB8FEFFDFB35" author="D'Udekem d'Acoz C. & Rappe G." box="[820,1259,1130,1157]" page="95" pageId="10" pageNumber="287" year="1991">
|
||
<bibRefCitation id="1B088E9AFF98FFF66306FB8FEFFDFB35" author="D'Udekem d'Acoz C. & Rappe G." box="[820,1259,1130,1157]" pageId="10" pageNumber="287" pagination="95 - 99" refId="ref14323" refString="D'Udekem d'Acoz C. & Rappe G. 1991. Presence et abondance de Liocarcinus vernalis (Risso, 1816) dans la baie sud de la mer du Nord (Decapoda, Brachyura, Portunidae). Crustaceana 61 (1): 95 - 99. Available from http: // www. jstor. org / stable / 20104677 [accessed 22 Mar. 2024]." type="journal article" year="1991">d’Udekem d’Acoz & Rappé 1991: 95</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 3, 6, 9, 12. —
|
||
<treatmentCitation id="FE38D57AFF98FFF6610EFB6BE90DFB17" author="Fransen C. H. J. M." box="[316,539,1165,1192]" page="124" pageId="10" pageNumber="287" year="1991">
|
||
<bibRefCitation id="1B088E9AFF98FFF6610EFB6BE90DFB17" author="Fransen C. H. J. M." box="[316,539,1165,1192]" pageId="10" pageNumber="287" refId="ref14577" refString="Fransen C. H. J. M. 1991. Crustacea of the CANCAP and MAURITANIA Expeditions. Nationaal Natuurhistorisch Museum, Leiden." type="book" year="1991">Fransen 1991: 124</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="FE38D57AFF98FFF6627FFB68E9EAFB18" author="Ingle R. W." box="[589,764,1165,1192]" page="57" pageId="10" pageNumber="287" year="1996">
|
||
<bibRefCitation id="1B088E9AFF98FFF6627FFB68E9EAFB18" author="Ingle R. W." box="[589,764,1165,1192]" pageId="10" pageNumber="287" pagination="1 - 243" refId="ref15165" refString="Ingle R. W. 1996. Shallow water crabs. Second edition. Synopsis of the British Fauna (new series) 25: 1 - 243." type="journal article" year="1996">Ingle 1996: 57</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 138, fig. 37. —
|
||
<treatmentCitation id="FE38D57AFF98FFF663F6FB68EFFCFB17" author="Ingle R. W. & Clark P. P." box="[964,1258,1165,1191]" page="224" pageId="10" pageNumber="287" year="1998">
|
||
<bibRefCitation id="1B088E9AFF98FFF663F6FB68EFFCFB17" author="Ingle R. W. & Clark P. P." box="[964,1258,1165,1191]" pageId="10" pageNumber="287" pagination="223 - 229" refId="ref15194" refString="Ingle R. W. & Clark P. P. 1998. A swimming crab new to the British fauna, Liocarcinus vernalis (Crustacea: Brachyura: Portunidae). Journal of the Marine Biological Association of the United Kingdom 78: 223 - 229. https: // doi. org / 10.1017 / S 0025315400040042 ICZN 1999. InternationalCodeofZoologicalNomenclature. 4 thedition. TheInternationalTrustforZoological Nomenclature, London. Available from https: // www. iczn. org / the-code / the-code-online / [accessed 22 Mar. 2024]." type="journal article" year="1998">Ingle & Clark 1998: 224</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 1c, 2c,
|
||
<specimenCount id="699F38E2FF98FFF660D8FB55EA1CFB7A" box="[234,266,1200,1226]" count="2" pageId="10" pageNumber="287" type="generic">2f.</specimenCount>
|
||
—
|
||
<treatmentCitation id="FE38D57AFF98FFF66101FB55EAE5FB7B" author="Livory A." box="[307,499,1200,1227]" page="49" pageId="10" pageNumber="287" year="1998">
|
||
<bibRefCitation id="1B088E9AFF98FFF66101FB55EAE5FB7B" author="Livory A." box="[307,499,1200,1227]" pageId="10" pageNumber="287" pagination="18 - 64" refId="ref15679" refString="Livory A. 1998. Crabes de la Manche: le point des connaissances. L'Argiope 18 - 19: 18 - 64." type="journal article" year="1998">Livory 1998: 49</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="FE38D57AFF98FFF66212FB54E880FB7B" author="Lopez de la Rosa I. & Garcia Raso J. E. & Rodriguez A." box="[544,918,1200,1227]" page="87" pageId="10" pageNumber="287" year="2002">
|
||
<bibRefCitation id="1B088E9AFF98FFF66212FB54E880FB7B" author="Lopez de la Rosa I. & Garcia Raso J. E. & Rodriguez A." box="[544,918,1200,1227]" pageId="10" pageNumber="287" pagination="85 - 95" refId="ref15703" refString="Lopez de la Rosa I., Garcia Raso J. E. & Rodriguez A. 2002. Evolution of a decapod community (Crustacea) of shallow soft bottoms with seaweeds from southern Europe. Journal of the Marine Biological Association of the United Kingdom 82 (1): 85 - 95. https: // doi. org / 10.1017 / S 0025315402005209." type="journal article" year="2002">
|
||
López de la Rosa
|
||
<emphasis id="4DED2F79FF98FFF662DEFB54E833FB7A" box="[748,805,1200,1226]" italics="true" pageId="10" pageNumber="287">et al.</emphasis>
|
||
2002: 87
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="FE38D57AFF98FFF663F1FB55EFFFFB7B" author="McCarthy A. M. & McGrath D. & Allen B. M." box="[963,1257,1200,1227]" page="20" pageId="10" pageNumber="287" year="2005">
|
||
<bibRefCitation id="1B088E9AFF98FFF663F1FB55EFFFFB7B" author="McCarthy A. M. & McGrath D. & Allen B. M." box="[963,1257,1200,1227]" pageId="10" pageNumber="287" pagination="20 - 26" refId="ref15921" refString="McCarthy A. M., McGrath D. & Allen B. M. 2005. The grey swimming crab Liocarcinus vernalis (Risso, 1827) in western Irish coastal waters (Decapoda: Brachyura, Portunidae). Irish Naturalists' Journal 28 (1): 20 - 26." type="journal article" year="2005">
|
||
McCarthy
|
||
<emphasis id="4DED2F79FF98FFF66472FB54EF6CFB7A" box="[1088,1146,1200,1226]" italics="true" pageId="10" pageNumber="287">et al.</emphasis>
|
||
2005: 20
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<bibRefCitation id="1B088E9AFF98FFF66521FB55EE63FB7A" author="Ashelby C. W." box="[1299,1397,1200,1226]" pageId="10" pageNumber="287" pagination="1333 - 1345" refId="ref13809" refString="Ashelby C. W. 2006. Contribution to the biogeographical knowledge of some decapod crustaceans in the British Isles. Crustaceana 79 (11): 1333 - 1345. https: // doi. org / 10.1163 / 156854006779277259" type="journal article" year="2006">Ashelby</bibRefCitation>
|
||
2006: 1341. —
|
||
<treatmentCitation id="FE38D57AFF98FFF66193FB31E966FB5E" author="Martin J." box="[417,624,1236,1262]" page="221" pageId="10" pageNumber="287" year="2011">
|
||
<bibRefCitation id="1B088E9AFF98FFF66193FB31E966FB5E" author="Martin J." box="[417,624,1236,1262]" pageId="10" pageNumber="287" refId="ref15816" refString="Martin J. 2011. Les Invertebres marins du Golfe de Gascogne a la Manche orientale. Editions Quae, Versailles." type="book" year="2011">Martin 2011: 221</bibRefCitation>
|
||
</treatmentCitation>
|
||
, unnumbered colour photographs. —
|
||
<bibRefCitation id="1B088E9AFF98FFF66418FB31EE61FB5E" author="De Blauwe H. & Decleer M." box="[1066,1399,1236,1262]" pageId="10" pageNumber="287" refId="ref14098" refString="De Blauwe H. & Decleer M. 2017. Strandvondsten. Stichting Kunnstboek, Oostkamp." type="book" year="2017">De Blauwe & Decleer 2017:</bibRefCitation>
|
||
173, unnumbered colour photographs.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFAFEE8C5FA84" blockId="10.[189,1399,778,1403]" box="[189,979,1306,1333]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFAFEE8C5FA84" box="[189,979,1306,1333]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFAFEE91FFA84" box="[189,521,1306,1333]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="4DED2F79FF98FFF6608FFAFEEA5EFA84" box="[189,328,1307,1332]" italics="true" pageId="10" pageNumber="287">Liocarcinus</emphasis>
|
||
sp. aff.
|
||
<emphasis id="4DED2F79FF98FFF66195FAFFE91FFA84" box="[423,521,1306,1332]" italics="true" pageId="10" pageNumber="287">holsatus</emphasis>
|
||
</taxonomicName>
|
||
‒
|
||
<bibRefCitation id="1B088E9AFF98FFF66214FAFFE850FA84" author="D'Udekem d'Acoz C." box="[550,838,1306,1332]" pageId="10" pageNumber="287" pagination="84 - 100" refId="ref14115" refString="D'Udekem d'Acoz C. 1986. Etude d'une collection de crustaces decapodes de Bretagne. De Strandvlo [dec. 1985]: 11 (4): 84 - 100." type="journal article" year="1986">d’Udekem d’Acoz 1986</bibRefCitation>
|
||
: figs 17, 20.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFAD8EFCAFAE7" blockId="10.[189,1399,778,1403]" box="[189,1244,1341,1368]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFAD8EFCAFAE7" box="[189,1244,1341,1368]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFAD8E91DFAE7" ID-CoL="935LR" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[189,523,1341,1367]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="vernalis" subGenus="Polybius">
|
||
<emphasis id="4DED2F79FF98FFF6608FFAD8EA32FAE7" box="[189,292,1341,1367]" italics="true" pageId="10" pageNumber="287">Polybius</emphasis>
|
||
(
|
||
<emphasis id="4DED2F79FF98FFF66107FAD8EA8AFAE7" box="[309,412,1341,1367]" italics="true" pageId="10" pageNumber="287">Polybius</emphasis>
|
||
)
|
||
<emphasis id="4DED2F79FF98FFF6619EFAD8E91DFAE7" box="[428,523,1341,1367]" italics="true" pageId="10" pageNumber="287">vernalis</emphasis>
|
||
</taxonomicName>
|
||
‒
|
||
<treatmentCitation id="FE38D57AFF98FFF6621AFAD8E894FAE7" author="D'Udekem d'Acoz C." box="[552,898,1341,1367]" page="222" pageId="10" pageNumber="287" year="1999">
|
||
<bibRefCitation id="1B088E9AFF98FFF6621AFAD8E894FAE7" author="D'Udekem d'Acoz C." box="[552,898,1341,1367]" pageId="10" pageNumber="287" pagination="40" refId="ref14220" refString="D'Udekem d'Acoz C. 1999. Inventaire et distribution des crustaces decapodes de l'Atlantique nordoriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Patrimoines naturels (M. N. H. N. / S. P. N.) 40." type="journal article" year="1999">d’Udekem d’Acoz 1999: 222</bibRefCitation>
|
||
</treatmentCitation>
|
||
, in part, unnumbered fig. p.1.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF6608FFA84E852FACA" blockId="10.[189,1399,778,1403]" box="[189,836,1376,1403]" pageId="10" pageNumber="287">
|
||
<treatmentCitationGroup id="5F89D445FF98FFF6608FFA84E856FACA" box="[189,832,1376,1403]" pageId="10" pageNumber="287">
|
||
<taxonomicName id="B89988E8FF98FFF6608FFA84EAC1FACA" ID-CoL="3V59T" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[189,471,1376,1402]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF98FFF6608FFA84EA5EFACA" box="[189,328,1377,1402]" italics="true" pageId="10" pageNumber="287">Liocarcinus</emphasis>
|
||
cf.
|
||
<emphasis id="4DED2F79FF98FFF66147FA85EAC1FACA" box="[373,471,1376,1402]" italics="true" pageId="10" pageNumber="287">holsatus</emphasis>
|
||
</taxonomicName>
|
||
– Moukrim
|
||
<emphasis id="4DED2F79FF98FFF66258FA84E9B3FACA" box="[618,677,1376,1402]" italics="true" pageId="10" pageNumber="287">et al.</emphasis>
|
||
2010: fig.
|
||
<specimenCount id="699F38E2FF98FFF66316FA85E856FACA" box="[804,832,1376,1402]" count="2" pageId="10" pageNumber="287" type="generic">2f</specimenCount>
|
||
</treatmentCitationGroup>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3783A0E0FF98FFF7608FFA47EFC2FEF6" lastPageId="11" lastPageNumber="288" pageId="10" pageNumber="287" type="diagnosis">
|
||
<paragraph id="7F26F36BFF98FFF6608FFA47EA20FA0C" blockId="10.[189,310,1442,1468]" box="[189,310,1442,1468]" pageId="10" pageNumber="287">
|
||
<heading id="246E4407FF98FFF6608FFA47EA20FA0C" bold="true" box="[189,310,1442,1468]" fontSize="11" level="3" pageId="10" pageNumber="287" reason="3">
|
||
<emphasis id="4DED2F79FF98FFF6608FFA47EA20FA0C" bold="true" box="[189,310,1442,1468]" pageId="10" pageNumber="287">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF98FFF7608FFA34EFC2FEF6" blockId="10.[187,1401,1488,1973]" lastBlockId="11.[189,1399,265,327]" lastPageId="11" lastPageNumber="288" pageId="10" pageNumber="287">Carapace glabrous, wider than long, smooth. Frontal region with three slightly protruding blunt triangular teeth, with a tuberculate rim, continued by the inner-orbital angle, middle tooth rounded, subacute and does not reach same proportions as the lateral ones, which are more developed (wider and rounded), the curvature between rostral teeth is rounded. Anterolateral margins of the carapace curved with five teeth (including the external orbital), the fifth and the fourth being somewhat larger, the third and fourth with a rectilinear outer face. Chelipeds (P1) subequal; propodus outer face of the palm with 3 longitudinal keels along entire length; dorsal edge with a smooth rounded keel ending in a blunt distal tooth; internal face without keels; dactyl externally with a central depression, between two rounded ridges; dorsally, three rounded longitudinal ridges with depressions between them; merus smooth; carpus with one strong and protruding tooth on antero inner dorsal margin, on the anterior outer upper part one well-marked and rounded projection and two others below. Walking legs with merus, carpus and propodus smooth; dactylus quite rectilinear dorsally (P2 wider, robust and with a slightly more curved tip), with a more or less quadrangular section, and with angles defined by rounded longitudinal edges, the dactylus of P4 has a dense row of short setae in the basal third (or a little more), in P5 the dactyl is strongly flattened, paddle-like, oval, ending in a point. First sexual male pleopod (gonopod) with curved distal part, which forms an angle of almost 90° (more curved in large specimens) and ends in a small point.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3783A0E0FF99FFF7608FFE8BE920FD90" pageId="11" pageNumber="288" type="etymology">
|
||
<paragraph id="7F26F36BFF99FFF7608FFE8BEA52FE38" blockId="11.[189,324,366,392]" box="[189,324,366,392]" pageId="11" pageNumber="288">
|
||
<heading id="246E4407FF99FFF7608FFE8BEA52FE38" bold="true" box="[189,324,366,392]" fontSize="11" level="3" pageId="11" pageNumber="288" reason="3">
|
||
<emphasis id="4DED2F79FF99FFF7608FFE8BEA52FE38" bold="true" box="[189,324,366,392]" pageId="11" pageNumber="288">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF99FFF7608FFE79E920FD90" blockId="11.[187,1399,412,544]" pageId="11" pageNumber="288">
|
||
The name of the species is derived from the Greek mythology. Castor and Pollux are twin half-brothers in Greek and Roman mythology, known together as the Dioscuri (plural). The singular form is Dioscurus. The name, which is a noun in apposition, refers to the great similarity between the new species and
|
||
<taxonomicName id="B89988E8FF99FFF7608FFDE3E925FD90" authority="(Risso, 1827)" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[189,563,518,544]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF99FFF7608FFDE3EA9CFD90" box="[189,394,518,544]" italics="true" pageId="11" pageNumber="288">Polybius vernalis</emphasis>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF99FFF761ABFDE3E93FFD90" author="Risso A." box="[409,553,518,544]" pageId="11" pageNumber="288" refId="ref16928" refString="Risso A. 1827 [in Risso, 1826 - 1827]. Histoire naturelle des Principales Productions de l'Europe meridionale et particulierement de celles des Environs de Nice et des Alpes maritimes. Vol. 5 Animaux Articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. F. - G. Levrault, Paris. https: // doi. org / 10.5962 / bhl. title. 58984" type="book" year="1827">Risso, 1827</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3783A0E0FF99FFF0608FFDA2EF33FE63" lastPageId="12" lastPageNumber="289" pageId="11" pageNumber="288" type="materials_examined">
|
||
<paragraph id="7F26F36BFF99FFF7608FFDA2EABCFDD1" blockId="11.[189,426,583,609]" box="[189,426,583,609]" pageId="11" pageNumber="288">
|
||
<heading id="246E4407FF99FFF7608FFDA2EABCFDD1" bold="true" box="[189,426,583,609]" fontSize="11" level="3" pageId="11" pageNumber="288" reason="3">
|
||
<emphasis id="4DED2F79FF99FFF7608FFDA2EABCFDD1" bold="true" box="[189,426,583,609]" pageId="11" pageNumber="288">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF99FFF760D8FD93E90FFD4A" blockId="11.[189,1399,630,762]" pageId="11" pageNumber="288">
|
||
<materialsCitation id="CFF1F936FF99FFF760D8FD93E90FFD4A" ID-GBIF-Occurrence="4608353332" collectedFrom="fine sand bottoms" collectingDate="1996-10-31" collectionCode="CRUST" collectorName="I. Lopez de la Rosa" country="Spain" elevation="5" latitude="36.716667" location="Valdelagrana" longLatPrecision="1185" longitude="-6.233333" municipality="Cadiz" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3759" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="4DED2F79FF99FFF760D8FD93EA4AFD20" bold="true" box="[234,348,630,656]" pageId="11" pageNumber="288">
|
||
<typeStatus id="A0224DC9FF99FFF760D8FD93EA4AFD20" box="[234,348,630,656]" pageId="11" pageNumber="288" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="078EB3FBFF99FFF7608FFD7CEA19FD03" box="[189,271,665,691]" name="Spain" pageId="11" pageNumber="288">SPAIN</collectingCountry>
|
||
•
|
||
<specimenCount id="699F38E2FF99FFF76119FD7CEA57FD03" box="[299,321,665,691]" count="1" pageId="11" pageNumber="288" type="male">♂</specimenCount>
|
||
(
|
||
<quantity id="B8615E8EFF99FFF76160FD7CEAA8FD03" box="[338,446,665,691]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="11" pageNumber="288" unit="mm" value="26.4">26.4 mm</quantity>
|
||
CW); outer Bay of Cádiz,
|
||
<collectingMunicipality id="9F426911FF99FFF76336FD7CE85DFD03" box="[772,843,665,691]" pageId="11" pageNumber="288">Cádiz</collectingMunicipality>
|
||
,
|
||
<location id="7A46A5B0FF99FFF7636AFD7CE8E3FD03" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF7636AFD7CE8E3FD03" box="[856,1013,665,691]" country="Spain" latitude="36.716667" longLatPrecision="1185" longitude="-6.233333" municipality="Cadiz" name="Valdelagrana" pageId="11" pageNumber="288">Valdelagrana</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF76431FD72EF66FD03" box="[1027,1136,663,691]" degrees="36" direction="north" minutes="43" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="36.716667">36°43′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF7644CFD72EFE7FD03" box="[1150,1265,663,691]" degrees="06" direction="west" minutes="14" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="-6.233333">06°14′ W</geoCoordinate>
|
||
;
|
||
<elevation id="F4B41458FF99FFF764CDFD7CEA14FD66" metricMagnitude="0" metricUnit="m" metricValue="4.949999999999999" metricValueMax="6.6" metricValueMin="3.3" pageId="11" pageNumber="288" unit="m" value="4.949999999999999" valueMax="6.6" valueMin="3.3">
|
||
<quantity id="B8615E8EFF99FFF764CDFD7CEE61FD03" box="[1279,1399,665,691]" metricMagnitude="0" metricUnit="m" metricValue="4.949999999999999" metricValueMax="6.6" metricValueMin="3.3" pageId="11" pageNumber="288" unit="m" value="4.949999999999999" valueMax="6.6" valueMin="3.3">3.3–6.6 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF99FFF7613EFD59EAB2FD66" box="[268,420,700,726]" pageId="11" pageNumber="288" value="1996-10-31">
|
||
<collectingDate id="1B632C43FF99FFF7613EFD59EAB2FD66" box="[268,420,700,726]" pageId="11" pageNumber="288" value="1996-10-31">31 Oct. 1996</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF7619CFD58E99CFD67" box="[430,650,700,727]" pageId="11" pageNumber="288">I. López de la Rosa</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF99FFF762FAFD59E881FD66" box="[712,919,700,726]" pageId="11" pageNumber="288">fine sand bottoms</collectedFrom>
|
||
; GenBank:
|
||
<accessionNumber id="60CA6E88FF99FFF7642FFD59EFB2FD67" box="[1053,1188,700,727]" pageId="11" pageNumber="288">OR555887</accessionNumber>
|
||
–16S,
|
||
<accessionNumber id="60CA6E88FF99FFF764D5FD59EE71FD67" box="[1255,1383,700,727]" pageId="11" pageNumber="288">OR557369</accessionNumber>
|
||
– COI;
|
||
<specimenCode id="2F3F5B10FF99FFF760CDFD3AE903FD4A" box="[255,533,735,762]" pageId="11" pageNumber="288">CRUST_ICMAN/3759</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF99FFF760D8FCC4E856FBC7" blockId="11.[189,1399,801,1144]" pageId="11" pageNumber="288">
|
||
<materialsCitation id="CFF1F936FF99FFF760D8FCC4EE72FC31" ID-GBIF-Occurrence="4608353339" collectedFrom="sandy beach" collectingDate="1990-11-10" collectingMethod="net refuse of shrimp fishermen" collectionCode="RBINS" collectorName="C. d'Udekem d'Acoz" country="Belgium" elevation="1" latitude="51.14167" location="Groenendijk" longLatPrecision="580" longitude="2.6833334" pageId="11" pageNumber="288" specimenCode="INV. 187227" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Southern North Sea" typeStatus="paratype">
|
||
<emphasis id="4DED2F79FF99FFF760D8FCC4EA71FC8B" bold="true" box="[234,359,801,827]" pageId="11" pageNumber="288">
|
||
<typeStatus id="A0224DC9FF99FFF760D8FCC4EA71FC8B" box="[234,359,801,827]" pageId="11" pageNumber="288" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="078EB3FBFF99FFF7608FFCA1EA55FCEE" box="[189,323,836,862]" name="Belgium" pageId="11" pageNumber="288">BELGIUM</collectingCountry>
|
||
–
|
||
<emphasis id="4DED2F79FF99FFF7616BFCA1E958FCEE" bold="true" box="[345,590,836,862]" pageId="11" pageNumber="288">
|
||
<collectingRegion id="BD5D3D89FF99FFF7616BFCA1E958FCEE" box="[345,590,836,862]" pageId="11" pageNumber="288">Southern North Sea</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="699F38E2FF99FFF76257FCA1E9B1FCEE" box="[613,679,836,862]" count="3" pageId="11" pageNumber="288" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF76283FCA1E9E4FCEE" box="[689,754,836,862]" count="2" pageId="11" pageNumber="288" type="female">2 ♀♀</specimenCount>
|
||
;
|
||
<location id="7A46A5B0FF99FFF762CCFCA1E883FCEE" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF762CCFCA1E883FCEE" box="[766,917,836,862]" country="Belgium" latitude="51.14167" longLatPrecision="580" longitude="2.6833334" name="Groenendijk" pageId="11" pageNumber="288" stateProvince="Southern North Sea">Groenendijk</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF763ACFCA7EF24FCEE" box="[926,1074,834,862]" degrees="51" direction="north" minutes="08" orientation="latitude" pageId="11" pageNumber="288" precision="15" seconds="30" value="51.14167">51°08′30″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF76409FCA7EF82FCEE" box="[1083,1172,834,862]" degrees="2" direction="east" minutes="41" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="2.6833334">2°41′ E</geoCoordinate>
|
||
;
|
||
<elevation id="F4B41458FF99FFF764ACFCA1EE05FCEE" box="[1182,1299,836,862]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">
|
||
<quantity id="B8615E8EFF99FFF764ACFCA1EFDFFCEE" box="[1182,1225,836,862]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">1 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF99FFF7652EFCA1EBEDFC31" pageId="11" pageNumber="288" value="1990-11-10">
|
||
<collectingDate id="1B632C43FF99FFF7652EFCA1EBEDFC31" pageId="11" pageNumber="288" value="1990-11-10">10 Nov. 1990</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF7613AFC82E910FC31" box="[264,518,871,897]" pageId="11" pageNumber="288">C. d’Udekem d’Acoz</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF99FFF76278FC82E9C8FC31" box="[586,734,871,897]" pageId="11" pageNumber="288">sandy beach</collectedFrom>
|
||
,
|
||
<collectingMethod id="A6D88B7CFF99FFF762D9FC8DEF4CFC31" box="[747,1114,871,898]" pageId="11" pageNumber="288">net refuse of shrimp fishermen</collectingMethod>
|
||
;
|
||
<collectionCode id="19886BAEFF99FFF76454FC82EFA9FC31" box="[1126,1215,871,897]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF764FEFC8DEE72FC31" box="[1228,1380,871,898]" pageId="11" pageNumber="288">INV. 187227</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF7608FFC6FEE1AFC78" ID-GBIF-Occurrence="4608353313" collectedFrom="sandy beach" collectingDate="2018-10-13" collectingMethod="net refuse of shrimp fishermen" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" elevation="1" latitude="51.15" location="between Nieuwpoort and Oostduinkerke" longLatPrecision="1092" longitude="2.7" pageId="11" pageNumber="288" specimenCode="INV. 187223" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="699F38E2FF99FFF7608FFC6FEBFFFC15" box="[189,233,906,933]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
(illustrated);
|
||
<location id="7A46A5B0FF99FFF761BAFC6FE87EFC14" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF761BAFC6FE87EFC14" box="[392,872,906,932]" latitude="51.15" longLatPrecision="1092" longitude="2.7" name="between Nieuwpoort and Oostduinkerke" pageId="11" pageNumber="288">between Nieuwpoort and Oostduinkerke</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF76341FC6CE8C9FC15" box="[883,991,905,933]" degrees="51" direction="north" minutes="09" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="51.15">51°09′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF763D9FC6CEF53FC15" box="[1003,1093,905,933]" degrees="2" direction="east" minutes="42" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="2.7">2°42′ E</geoCoordinate>
|
||
;
|
||
<elevation id="F4B41458FF99FFF76463FC6FEFDCFC14" box="[1105,1226,906,933]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">
|
||
<quantity id="B8615E8EFF99FFF76463FC6FEF68FC15" box="[1105,1150,906,933]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">1 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF99FFF764E4FC6FEE64FC14" box="[1238,1394,906,932]" pageId="11" pageNumber="288" value="2018-10-13">
|
||
<collectingDate id="1B632C43FF99FFF764E4FC6FEE64FC14" box="[1238,1394,906,932]" pageId="11" pageNumber="288" value="2018-10-13">13 Oct. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF7608FFC48EA8FFC77" box="[189,409,941,967]" pageId="11" pageNumber="288">d’Udekem d’Acoz</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF99FFF761D3FC48E961FC77" box="[481,631,941,967]" pageId="11" pageNumber="288">sandy beach</collectedFrom>
|
||
,
|
||
<collectingMethod id="A6D88B7CFF99FFF762B4FC4BE8EAFC77" box="[646,1020,941,968]" pageId="11" pageNumber="288">net refuse of shrimp fishermen</collectingMethod>
|
||
;
|
||
<collectionCode id="19886BAEFF99FFF76438FC4BEF75FC78" box="[1034,1123,942,968]" collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF76440FC4BEE1AFC78" box="[1138,1292,942,968]" pageId="11" pageNumber="288">INV. 187223</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF76518FC4BEA03FB81" ID-GBIF-Occurrence="4608353342" collectedFrom="sandy beach" collectingDate="2018-10-13" collectingMethod="net refuse of shrimp fishermen" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" elevation="1" latitude="51.15" location="between Nieuwpoort and Oostduinkerke" longLatPrecision="1092" longitude="2.7" pageId="11" pageNumber="288" specimenCode="INV. 187230" specimenCount="10" specimenCount-female="6" specimenCount-male="4" typeStatus="paratype">
|
||
<specimenCount id="699F38E2FF99FFF76518FC4BEE67FC78" box="[1322,1393,942,968]" count="4" pageId="11" pageNumber="288" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF7608FFC34EA17FC5B" box="[189,257,977,1003]" count="6" pageId="11" pageNumber="288" type="female">6 ♀♀</specimenCount>
|
||
(colour in life photographed);
|
||
<location id="7A46A5B0FF99FFF76241FC34EF40FC5B" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF76241FC34EF40FC5B" box="[627,1110,977,1003]" latitude="51.15" longLatPrecision="1092" longitude="2.7" name="between Nieuwpoort and Oostduinkerke" pageId="11" pageNumber="288">between Nieuwpoort and Oostduinkerke</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF76451FC2AEFC7FC5B" box="[1123,1233,975,1003]" degrees="51" direction="north" minutes="09" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="51.15">51°09′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF764ECFC2AEE2CFC5B" box="[1246,1338,975,1003]" degrees="2" direction="east" minutes="42" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="2.7">2°42′ E</geoCoordinate>
|
||
;
|
||
<elevation id="F4B41458FF99FFF7657AFC34EA14FBBE" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">
|
||
<quantity id="B8615E8EFF99FFF7657AFC34EE61FC5B" box="[1352,1399,977,1003]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">1 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF99FFF7613FFC11EAB1FBBE" box="[269,423,1012,1038]" pageId="11" pageNumber="288" value="2018-10-13">
|
||
<collectingDate id="1B632C43FF99FFF7613FFC11EAB1FBBE" box="[269,423,1012,1038]" pageId="11" pageNumber="288" value="2018-10-13">13 Oct. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF76180FC11E99DFBBE" box="[434,651,1012,1038]" pageId="11" pageNumber="288">d’Udekem d’Acoz</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF99FFF762F9FC11E84BFBBE" box="[715,861,1012,1038]" pageId="11" pageNumber="288">sandy beach</collectedFrom>
|
||
,
|
||
<collectingMethod id="A6D88B7CFF99FFF7635BFC11EFC6FBBE" box="[873,1232,1012,1038]" pageId="11" pageNumber="288">net refuse of shrimp fishermen</collectingMethod>
|
||
;
|
||
<collectionCode id="19886BAEFF99FFF764E8FC11EE25FBBE" box="[1242,1331,1012,1038]" collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF7650DFC11EA03FB81" pageId="11" pageNumber="288">INV. 187230</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF76103FBF2EFF1FBE4" ID-GBIF-Occurrence="4608353326" collectedFrom="sandy beach" collectingDate="2019-09-27" collectingMethod="net refuse of shrimp fishermen" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" elevation="1" latitude="51.133335" location="between Oostduinkerke and Koksijde" longLatPrecision="1092" longitude="2.65" pageId="11" pageNumber="288" specimenCode="INV. 187229" specimenCount="20" specimenCount-male="20" typeStatus="paratype">
|
||
<specimenCount id="699F38E2FF99FFF76103FBF2EA90FB82" box="[305,390,1047,1074]" count="20" pageId="11" pageNumber="288" type="male">20 ♂♂</specimenCount>
|
||
;
|
||
<location id="7A46A5B0FF99FFF761A6FBF2E840FB81" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF761A6FBF2E840FB81" box="[404,854,1047,1073]" latitude="51.133335" longLatPrecision="1092" longitude="2.65" name="between Oostduinkerke and Koksijde" pageId="11" pageNumber="288">between Oostduinkerke and Koksijde</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF76351FBF3E8C6FB82" box="[867,976,1046,1074]" degrees="51" direction="north" minutes="08" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="51.133335">51°08′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF763ECFBF3EF2CFB82" box="[990,1082,1046,1074]" degrees="2" direction="east" minutes="39" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="2.65">2°39′ E</geoCoordinate>
|
||
;
|
||
<elevation id="F4B41458FF99FFF7647AFBF2EFD2FB81" box="[1096,1220,1047,1074]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">
|
||
<quantity id="B8615E8EFF99FFF7647AFBF2EF60FB82" box="[1096,1142,1047,1074]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">1 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF99FFF764E3FBF2EE64FB81" box="[1233,1394,1047,1073]" pageId="11" pageNumber="288" value="2019-09-27">
|
||
<collectingDate id="1B632C43FF99FFF764E3FBF2EE64FB81" box="[1233,1394,1047,1073]" pageId="11" pageNumber="288" value="2019-09-27">27 Sep. 2019</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF7608FFBDFEA80FBE4" box="[189,406,1082,1108]" pageId="11" pageNumber="288">d’Udekem d’Acoz</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF99FFF761EAFBDFE97CFBE4" box="[472,618,1082,1108]" pageId="11" pageNumber="288">sandy beach</collectedFrom>
|
||
,
|
||
<collectingMethod id="A6D88B7CFF99FFF76244FBDEE8F6FBE4" box="[630,992,1082,1109]" pageId="11" pageNumber="288">net refuse of shrimp fishermen</collectingMethod>
|
||
;
|
||
<collectionCode id="19886BAEFF99FFF763D9FBDFEF52FBE4" box="[1003,1092,1082,1108]" collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF76462FBDEEFF1FBE4" box="[1104,1255,1082,1109]" pageId="11" pageNumber="288">INV. 187229</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF764CDFBDFE82BFBC7" ID-GBIF-Occurrence="4608353318" collectedFrom="sandy beach" collectingDate="2019-09-27" collectingMethod="net refuse of shrimp fishermen" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" elevation="1" latitude="51.133335" location="between Oostduinkerke and Koksijde" longLatPrecision="1092" longitude="2.65" pageId="11" pageNumber="288" specimenCode="INV. 187226" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="699F38E2FF99FFF764CDFBDFEE38FBE5" box="[1279,1326,1082,1109]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="19886BAEFF99FFF7620FFBBBE980FBC8" box="[573,662,1118,1144]" collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF76290FBBBE82BFBC7" box="[674,829,1117,1144]" pageId="11" pageNumber="288">INV. 187226</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF99FFF7608FFB7AEE5AFB4F" blockId="11.[189,1399,1181,1280]" pageId="11" pageNumber="288">
|
||
<materialsCitation id="CFF1F936FF99FFF7608FFB7AEAACFB6C" ID-GBIF-Occurrence="4608353325" collectingDate="1988-03-18" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" country="France" latitude="48.633335" location="Saint Lunaire" longLatPrecision="1109" longitude="-2.1" pageId="11" pageNumber="288" specimenCode="INV. 187222" specimenCount="1" specimenCount-male="1" stateProvince="Brittany">
|
||
<collectingCountry id="078EB3FBFF99FFF7608FFB7AEA27FB09" box="[189,305,1183,1209]" name="France" pageId="11" pageNumber="288">FRANCE</collectingCountry>
|
||
<emphasis id="4DED2F79FF99FFF7610BFB7AEAACFB09" bold="true" box="[313,442,1183,1209]" pageId="11" pageNumber="288">
|
||
–
|
||
<collectingRegion id="BD5D3D89FF99FFF76162FB7AEAACFB09" box="[336,442,1183,1209]" country="France" name="Bretagne" pageId="11" pageNumber="288">Brittany</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="699F38E2FF99FFF761E4FB7AE910FB0A" box="[470,518,1183,1210]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
;
|
||
<location id="7A46A5B0FF99FFF76221FB7AE9A1FB09" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF76221FB7AE9A1FB09" box="[531,695,1183,1209]" country="France" latitude="48.633335" longLatPrecision="1109" longitude="-2.1" name="Saint Lunaire" pageId="11" pageNumber="288" stateProvince="Brittany">Saint Lunaire</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF762F6FB78E827FB0A" box="[708,817,1181,1210]" degrees="48" direction="north" minutes="38" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="48.633335">48°38′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF7630CFB78E8B4FB0A" box="[830,930,1181,1210]" degrees="2" direction="west" minutes="06" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="-2.1">2°06′ W</geoCoordinate>
|
||
;
|
||
<date id="0B27D5ABFF99FFF76382FB7AEF45FB09" box="[944,1107,1183,1210]" pageId="11" pageNumber="288" value="1988-03-18">
|
||
<collectingDate id="1B632C43FF99FFF76382FB7AEF45FB09" box="[944,1107,1183,1210]" pageId="11" pageNumber="288" value="1988-03-18">18 Mar. 1988</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF76452FB7AEE2CFB09" box="[1120,1338,1183,1209]" pageId="11" pageNumber="288">d’Udekem d’Acoz</collectorName>
|
||
leg.;
|
||
<collectionCode id="19886BAEFF99FFF7608FFB27EA00FB6C" box="[189,278,1218,1244]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF76111FB26EAACFB6C" box="[291,442,1218,1245]" pageId="11" pageNumber="288">INV. 187222</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF761E6FB27EE5FFB4F" ID-GBIF-Occurrence="4608353309" collectingDate="2010-03-30" collectingMethod="net refuse of Donax fishermen" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" elevation="1" latitude="48.25" location="Anse de Dinant" longLatPrecision="1111" longitude="-4.55" municipality="Crozon Peninsula" pageId="11" pageNumber="288" specimenCode="INV. 187228" specimenCount="16" specimenCount-female="7" specimenCount-male="9">
|
||
<specimenCount id="699F38E2FF99FFF761E6FB27E90EFB6D" box="[468,536,1218,1245]" count="9" pageId="11" pageNumber="288" type="male">9 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF76214FB26E97FFB6D" box="[550,617,1219,1245]" count="7" pageId="11" pageNumber="288" type="female">7 ♀♀</specimenCount>
|
||
; W of
|
||
<collectingMunicipality id="9F426911FF99FFF76289FB27E898FB6C" box="[699,910,1218,1244]" pageId="11" pageNumber="288">Crozon Peninsula</collectingMunicipality>
|
||
,
|
||
<location id="7A46A5B0FF99FFF763ABFB27EF42FB6C" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF763ABFB27EF42FB6C" box="[921,1108,1218,1244]" country="France" latitude="48.25" longLatPrecision="1111" longitude="-4.55" municipality="Crozon Peninsula" name="Anse de Dinant" pageId="11" pageNumber="288" stateProvince="Bretagne">Anse de Dinant</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF76452FB24EFDBFB6D" box="[1120,1229,1217,1245]" degrees="48" direction="north" minutes="15" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="48.25">48°15′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF764EBFB24EE2BFB6D" box="[1241,1341,1217,1245]" degrees="4" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="-4.55">4°33′ W</geoCoordinate>
|
||
;
|
||
<elevation id="F4B41458FF99FFF7657BFB27EA14FB4F" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">
|
||
<quantity id="B8615E8EFF99FFF7657BFB27EE61FB6D" box="[1353,1399,1218,1245]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">1 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF99FFF7613CFB00EAB8FB4F" box="[270,430,1253,1280]" pageId="11" pageNumber="288" value="2010-03-30">
|
||
<collectingDate id="1B632C43FF99FFF7613CFB00EAB8FB4F" box="[270,430,1253,1280]" pageId="11" pageNumber="288" value="2010-03-30">30 Mar. 2010</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF76188FB00E982FB4F" box="[442,660,1253,1279]" pageId="11" pageNumber="288">d’Udekem d’Acoz</collectorName>
|
||
leg.;
|
||
<collectingMethod id="A6D88B7CFF99FFF762E4FB03EF2BFB4F" box="[726,1085,1253,1280]" pageId="11" pageNumber="288">
|
||
net refuse of
|
||
<taxonomicName id="B89988E8FF99FFF76342FB03E8A8FB4F" box="[880,958,1254,1279]" class="Liliopsida" family="Marantaceae" genus="Donax" kingdom="Plantae" order="Zingiberales" pageId="11" pageNumber="288" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4DED2F79FF99FFF76342FB03E8A8FB4F" box="[880,958,1254,1279]" italics="true" pageId="11" pageNumber="288">Donax</emphasis>
|
||
</taxonomicName>
|
||
fishermen
|
||
</collectingMethod>
|
||
;
|
||
<collectionCode id="19886BAEFF99FFF7647AFB00EFB7FB4F" box="[1096,1185,1253,1279]" collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF7649CFB03EE5FFB4F" box="[1198,1353,1253,1280]" pageId="11" pageNumber="288">INV. 187228</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF99FFF7608FFAC2EFF3FA41" blockId="11.[188,1399,1317,1521]" pageId="11" pageNumber="288">
|
||
<materialsCitation id="CFF1F936FF99FFF7608FFAC2E878FA37" ID-GBIF-Occurrence="4608353308" collectedFrom="sandy bottoms" collectingDate="1999-05" collectionCode="CRUST" collectorName="Gofas & Moukrim" country="Morocco" elevation="16" latitude="30.3" location="South of Oued Souss" longLatPrecision="1222" longitude="-9.616667" municipality="Pte. d'Anza" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3762" specimenCount="1" specimenCount-male="1" stateProvince="Agadir Bay">
|
||
<collectingCountry id="078EB3FBFF99FFF7608FFAC2EA47FAF1" box="[189,337,1319,1345]" name="Morocco" pageId="11" pageNumber="288">MOROCCO</collectingCountry>
|
||
–
|
||
<emphasis id="4DED2F79FF99FFF76158FAC2EAEEFAF1" bold="true" box="[362,504,1319,1345]" pageId="11" pageNumber="288">
|
||
<collectingRegion id="BD5D3D89FF99FFF76158FAC2EAEEFAF1" box="[362,504,1319,1345]" pageId="11" pageNumber="288">Agadir Bay</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="699F38E2FF99FFF76222FAC2E92CFAF1" box="[528,570,1319,1345]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
(36.7 CW); from
|
||
<collectingMunicipality id="9F426911FF99FFF7633BFAC2E887FAF1" box="[777,913,1319,1345]" pageId="11" pageNumber="288">Pte. d’Anza</collectingMunicipality>
|
||
to
|
||
<location id="7A46A5B0FF99FFF76386FAC2EFBCFAF1" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF76386FAC2EFBCFAF1" box="[948,1194,1319,1345]" country="Morocco" latitude="30.3" longLatPrecision="1222" longitude="-9.616667" municipality="Pte. d'Anza" name="South of Oued Souss" pageId="11" pageNumber="288" stateProvince="Agadir Bay">South of Oued Souss</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF76487FAC0EE09FAF1" box="[1205,1311,1317,1345]" degrees="30" direction="north" minutes="26" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="30.433332">30°26′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF76518FAC0EBCEFAD5" degrees="09" direction="west" minutes="40" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="-9.666667">09°40′ W</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="1AAD95ACFF99FFF760C6FAADEA49FAD5" box="[244,351,1352,1381]" degrees="30" direction="north" minutes="18" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="30.3">30°18′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF76158FAADEACDFAD5" box="[362,475,1352,1381]" degrees="09" direction="west" minutes="37" orientation="longitude" pageId="11" pageNumber="288" precision="925" value="-9.616667">09°37′ W</geoCoordinate>
|
||
;
|
||
<elevation id="F4B41458FF99FFF761D4FAAFE99CFAD4" box="[486,650,1354,1381]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="2.5" metricValueMin="0.6" pageId="11" pageNumber="288" unit="m" value="15.5" valueMax="25.0" valueMin="6.0">
|
||
<quantity id="B8615E8EFF99FFF761D4FAAFE929FAD5" box="[486,575,1354,1381]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="2.5" metricValueMin="0.6" pageId="11" pageNumber="288" unit="m" value="15.5" valueMax="25.0" valueMin="6.0">6–25 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF99FFF762A7FAAEE819FAD4" box="[661,783,1354,1381]" pageId="11" pageNumber="288" value="1999-05">
|
||
<collectingDate id="1B632C43FF99FFF762A7FAAEE819FAD4" box="[661,783,1354,1381]" pageId="11" pageNumber="288" value="1999-05">May 1999</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF76329FAAFE877FAD4" box="[795,865,1354,1380]" pageId="11" pageNumber="288">Gofas</collectorName>
|
||
and
|
||
<collectorName id="D26C96BDFF99FFF763AAFAAFEF11FAD4" box="[920,1031,1354,1380]" pageId="11" pageNumber="288">Moukrim</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF99FFF76475FAAFEFE2FAD4" box="[1095,1268,1354,1380]" pageId="11" pageNumber="288">sandy bottoms</collectedFrom>
|
||
; GenBank:
|
||
<accessionNumber id="60CA6E88FF99FFF7608EFA88EA52FA38" box="[188,324,1389,1416]" pageId="11" pageNumber="288">OR555890</accessionNumber>
|
||
–16S,
|
||
<accessionNumber id="60CA6E88FF99FFF761BBFA88E91AFA38" box="[393,524,1389,1416]" pageId="11" pageNumber="288">OR557370</accessionNumber>
|
||
–COI;
|
||
<specimenCode id="2F3F5B10FF99FFF7626EFA88E878FA37" box="[604,878,1389,1415]" pageId="11" pageNumber="288">CRUST_ICMAN/3762</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF763BBFA88E8EAFA1A" ID-GBIF-Occurrence="4608353311" collectedFrom="sandy bottoms" collectingDate="1999-05" collectionCode="CRUST" collectorName="Gofas & Moukrim" country="Morocco" elevation="16" latitude="30.3" location="South of Oued Souss" longLatPrecision="1222" longitude="-9.616667" municipality="Pte. d'Anza" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3763" specimenCount="1" specimenCount-male="1" stateProvince="Agadir Bay">
|
||
<specimenCount id="699F38E2FF99FFF763BBFA88E8A0FA38" box="[905,950,1389,1416]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
(
|
||
<quantity id="B8615E8EFF99FFF763F4FA88EF25FA38" box="[966,1075,1389,1416]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="11" pageNumber="288" unit="mm" value="40.6">40.6 mm</quantity>
|
||
CW); same collection data as for preceding; GenBank:
|
||
<accessionNumber id="60CA6E88FF99FFF7622FFA75E9B1FA1B" box="[541,679,1424,1451]" pageId="11" pageNumber="288">OR555889</accessionNumber>
|
||
-16S;
|
||
<specimenCode id="2F3F5B10FF99FFF762D8FA75E8EAFA1A" box="[746,1020,1424,1450]" pageId="11" pageNumber="288">CRUST_ICMAN/3763</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF76412FA75E9B5FA7E" ID-GBIF-Occurrence="4608353336" collectedFrom="sandy bottoms" collectingDate="1999-05" collectionCode="CRUST" collectorName="Gofas & Moukrim" country="Morocco" elevation="16" latitude="30.3" location="South of Oued Souss" longLatPrecision="1222" longitude="-9.616667" municipality="Pte. d'Anza" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3777" specimenCount="6" specimenCount-male="6" stateProvince="Agadir Bay">
|
||
<specimenCount id="699F38E2FF99FFF76412FA75EF7CFA1B" box="[1056,1130,1424,1451]" count="6" pageId="11" pageNumber="288" type="male">6 ♂♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="2F3F5B10FF99FFF761A3FA51E9B5FA7E" box="[401,675,1460,1486]" pageId="11" pageNumber="288">CRUST_ICMAN/3777</specimenCode>
|
||
</materialsCitation>
|
||
<emphasis id="4DED2F79FF99FFF7629FFA51E9A1FA7E" bold="true" box="[685,695,1460,1486]" pageId="11" pageNumber="288">•</emphasis>
|
||
<materialsCitation id="CFF1F936FF99FFF762F3FA51EA73FA41" ID-GBIF-Occurrence="4608353315" collectedFrom="sandy bottoms" collectingDate="1999-05" collectionCode="MNHN" collectorName="Gofas & Moukrim" country="Morocco" elevation="16" latitude="30.3" location="South of Oued Souss" longLatPrecision="1222" longitude="-9.616667" municipality="Pte. d'Anza" pageId="11" pageNumber="288" specimenCode="MNHN- IU-2022-4081" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Agadir Bay">
|
||
<specimenCount id="699F38E2FF99FFF762F3FA51E9E7FA7E" box="[705,753,1460,1486]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF76332FA51E826FA7E" box="[768,816,1460,1486]" count="1" pageId="11" pageNumber="288" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="2F3F5B10FF99FFF76521FA51EA73FA41" pageId="11" pageNumber="288">MNHN-IU-2022-4081</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF7614CFA32EFF4FA41" ID-GBIF-Occurrence="4608353338" box="[382,1250,1495,1521]" collectedFrom="sandy bottoms" collectingDate="1999-05" collectionCode="NHMUK" collectorName="Gofas & Moukrim" country="Morocco" elevation="16" latitude="30.3" location="South of Oued Souss" longLatPrecision="1222" longitude="-9.616667" municipality="Pte. d'Anza" pageId="11" pageNumber="288" specimenCode="NHMUK 2024.3, NHMUK 2024.4" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Agadir Bay">
|
||
<specimenCount id="699F38E2FF99FFF7614CFA32EABAFA41" box="[382,428,1495,1521]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF7618AFA32EAF3FA41" box="[440,485,1495,1521]" count="1" pageId="11" pageNumber="288" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="2F3F5B10FF99FFF76385FA32EF97FA41" box="[951,1153,1495,1521]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="288" type="Museum">NHMUK 2024.3</specimenCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF764BFFA32EFF4FA41" box="[1165,1250,1495,1521]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="288" type="Museum">2024.4</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF99FFF7608FF9FDEFAEF9C9" blockId="11.[189,1399,1559,1657]" pageId="11" pageNumber="288">
|
||
<materialsCitation id="CFF1F936FF99FFF7608FF9FDEE73F9E6" ID-GBIF-Occurrence="4608353340" collectingDate="1988-07-20" collectingMethod="net refuse of Donax fishermen" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" country="Portugal" elevation="1" latitude="37.133335" location="Praia de Cabanas" longLatPrecision="925" longitude="-7.5916667" municipality="Sacculina" pageId="11" pageNumber="288" specimenCode="INV. 187232" specimenCount="107" specimenCount-male="107" stateProvince="Algarve">
|
||
<collectingCountry id="078EB3FBFF99FFF7608FF9FDEA41F982" box="[189,343,1560,1586]" name="Portugal" pageId="11" pageNumber="288">PORTUGAL</collectingCountry>
|
||
–
|
||
<emphasis id="4DED2F79FF99FFF7614EF9FDEAC9F982" bold="true" box="[380,479,1560,1586]" pageId="11" pageNumber="288">
|
||
<collectingRegion id="BD5D3D89FF99FFF7614EF9FDEAC9F982" box="[380,479,1560,1586]" pageId="11" pageNumber="288">Algarve</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="699F38E2FF99FFF761CDF9FDE975F983" box="[511,611,1560,1587]" count="107" pageId="11" pageNumber="288" type="male">107 ♂♂</specimenCount>
|
||
(one with
|
||
<taxonomicName id="B89988E8FF99FFF762DEF9FDE876F982" box="[748,864,1560,1586]" class="Maxillopoda" family="Sacculinidae" genus="Sacculina" kingdom="Animalia" order="Kentrogonida" pageId="11" pageNumber="288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DED2F79FF99FFF762DEF9FDE876F982" box="[748,864,1560,1586]" italics="true" pageId="11" pageNumber="288">
|
||
<collectingMunicipality id="9F426911FF99FFF762DEF9FDE876F982" box="[748,864,1560,1586]" pageId="11" pageNumber="288">Sacculina</collectingMunicipality>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<location id="7A46A5B0FF99FFF7634FF9FDEF44F982" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF7634FF9FDEF44F982" box="[893,1106,1560,1586]" country="Portugal" latitude="37.133335" longLatPrecision="925" longitude="-7.5916667" municipality="Sacculina" name="Praia de Cabanas" pageId="11" pageNumber="288" stateProvince="Algarve">Praia de Cabanas</location>
|
||
;
|
||
<geoCoordinate id="1AAD95ACFF99FFF76450F9F2EFC4F983" box="[1122,1234,1559,1587]" degrees="37" direction="north" minutes="08" orientation="latitude" pageId="11" pageNumber="288" precision="925" value="37.133335">37°08′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1AAD95ACFF99FFF764D3F9F2EE64F983" box="[1249,1394,1559,1587]" degrees="7" direction="west" minutes="35" orientation="longitude" pageId="11" pageNumber="288" precision="15" seconds="30" value="-7.5916667">7°35′30″ W</geoCoordinate>
|
||
;
|
||
<quantity id="B8615E8EFF99FFF7608FF9D9EBFCF9E6" box="[189,234,1596,1622]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">
|
||
<elevation id="F4B41458FF99FFF7608FF9D9EBFCF9E6" box="[189,234,1596,1622]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="288" unit="m" value="1.0">1 m</elevation>
|
||
</quantity>
|
||
depth;
|
||
<date id="0B27D5ABFF99FFF76173F9D9EAC5F9E6" box="[321,467,1595,1622]" pageId="11" pageNumber="288" value="1988-07-20">
|
||
<collectingDate id="1B632C43FF99FFF76173F9D9EAC5F9E6" box="[321,467,1595,1622]" pageId="11" pageNumber="288" value="1988-07-20">20 Jul. 1988</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF761EDF9DEE9AEF9E5" box="[479,696,1595,1621]" pageId="11" pageNumber="288">d’Udekem d’Acoz</collectorName>
|
||
leg.;
|
||
<collectingMethod id="A6D88B7CFF99FFF762CBF9D9EF49F9E5" box="[761,1119,1595,1622]" pageId="11" pageNumber="288">
|
||
net refuse of
|
||
<taxonomicName id="B89988E8FF99FFF763A0F9D9E8F6F9E5" box="[914,992,1596,1621]" class="Liliopsida" family="Marantaceae" genus="Donax" kingdom="Plantae" order="Zingiberales" pageId="11" pageNumber="288" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4DED2F79FF99FFF763A0F9D9E8F6F9E5" box="[914,992,1596,1621]" italics="true" pageId="11" pageNumber="288">Donax</emphasis>
|
||
</taxonomicName>
|
||
fishermen
|
||
</collectingMethod>
|
||
;
|
||
<collectionCode id="19886BAEFF99FFF76458F9D9EFD5F9E6" box="[1130,1219,1596,1622]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF764FDF9D9EE73F9E6" box="[1231,1381,1596,1622]" pageId="11" pageNumber="288">INV. 187232</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF7608FF9BAEFA2F9C9" ID-GBIF-Occurrence="4608353316" box="[189,1204,1631,1657]" collectingDate="1988-07-20" collectingMethod="net refuse of Donax fishermen" collectionCode="RBINS" collectorName="d'Udekem d'Acoz" country="Portugal" elevation="1" latitude="37.133335" location="Praia de Cabanas" longLatPrecision="925" longitude="-7.5916667" municipality="Sacculina" pageId="11" pageNumber="288" specimenCode="INV. 187233" specimenCount="67" specimenCount-female="67" stateProvince="Algarve">
|
||
<specimenCount id="699F38E2FF99FFF7608FF9BAEA18F9C9" box="[189,270,1631,1657]" count="67" pageId="11" pageNumber="288" type="female">67 ♀♀</specimenCount>
|
||
(some ovigerous); same collection data as for preceding;
|
||
<collectionCode id="19886BAEFF99FFF76386F9BAEF1BF9C9" box="[948,1037,1631,1657]" collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="288" type="Museum">RBINS</collectionCode>
|
||
,
|
||
<specimenCode id="2F3F5B10FF99FFF7642BF9BAEFA2F9C9" box="[1049,1204,1631,1657]" pageId="11" pageNumber="288">INV. 187233</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF99FFF0608FF945EF33FE63" blockId="11.[189,1406,1696,2004]" lastBlockId="12.[189,1400,265,467]" lastPageId="12" lastPageNumber="289" pageId="11" pageNumber="288">
|
||
<materialsCitation id="CFF1F936FF99FFF7608FF945EAD9F96D" ID-GBIF-Occurrence="4608353302" collectedFrom="fine sand bottoms" collectingDate="1996-10-31" collectionCode="CRUST" collectorName="I. Lopez de la Rosa" country="Spain" elevation="5" latitude="36.716667" location="SPAIN" longLatPrecision="1185" longitude="-6.233333" municipality="Cadiz" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3760" specimenCount="1" specimenCount-female="1">
|
||
<collectingCountry id="078EB3FBFF99FFF7608FF945EA19F90A" box="[189,271,1696,1722]" name="Spain" pageId="11" pageNumber="288">SPAIN</collectingCountry>
|
||
•
|
||
<specimenCount id="699F38E2FF99FFF76107F945EA71F90B" box="[309,359,1696,1723]" count="1" pageId="11" pageNumber="288" type="female">1 ♀</specimenCount>
|
||
(
|
||
<quantity id="B8615E8EFF99FFF7614CF945EAF8F90B" box="[382,494,1696,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="11" pageNumber="288" unit="mm" value="19.3">19.3 mm</quantity>
|
||
CW); same collection data as for holotype; GenBank:
|
||
<accessionNumber id="60CA6E88FF99FFF7649EF945EE26F90B" box="[1196,1328,1696,1723]" pageId="11" pageNumber="288">OR557368</accessionNumber>
|
||
–COI;
|
||
<specimenCode id="2F3F5B10FF99FFF7608FF926EAD9F96D" box="[189,463,1731,1757]" pageId="11" pageNumber="288">CRUST_ICMAN/3760</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF761D5F926E9EFF8B1" ID-GBIF-Occurrence="4608353310" collectedFrom="fine muddy sand bottoms with Caulerpa prolifera" collectingDate="1994" collectingDateMax="1995" collectingDateMin="1994" collectionCode="CRUST" location="Cadiz Bay" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3778" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
|
||
<specimenCount id="699F38E2FF99FFF761D5F926E93FF96E" box="[487,553,1731,1758]" count="2" pageId="11" pageNumber="288" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF76205F926E973F96E" box="[567,613,1731,1758]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
; inner part of the
|
||
<location id="7A46A5B0FF99FFF76301F926E8A4F96E" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF76301F926E8A4F96E" box="[819,946,1731,1758]" country="Spain" name="Cadiz Bay" pageId="11" pageNumber="288">Cádiz Bay</location>
|
||
;
|
||
<collectingDate id="1B632C43FF99FFF7638FF926E8EFF96E" box="[957,1017,1731,1758]" pageId="11" pageNumber="288">1994</collectingDate>
|
||
–
|
||
<collectingDate id="1B632C43FF99FFF7643BF926EF52F96E" box="[1033,1092,1731,1758]" pageId="11" pageNumber="288">1995</collectingDate>
|
||
;
|
||
<collectedFrom id="CC85B096FF99FFF76462F926EAC1F8B0" pageId="11" pageNumber="288">
|
||
fine muddy sand bottoms with
|
||
<taxonomicName id="B89988E8FF99FFF760CBF903EAC1F8B0" box="[249,471,1766,1792]" class="Ulvophyceae" family="Caulerpaceae" genus="Caulerpa" kingdom="Plantae" order="Bryopsidales" pageId="11" pageNumber="288" phylum="Chlorophyta" rank="species" species="prolifera">
|
||
<emphasis id="4DED2F79FF99FFF760CBF903EAC1F8B0" box="[249,471,1766,1792]" italics="true" pageId="11" pageNumber="288">Caulerpa prolifera</emphasis>
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
;
|
||
<specimenCode id="2F3F5B10FF99FFF761D5F902E9EFF8B1" box="[487,761,1767,1793]" pageId="11" pageNumber="288">CRUST_ICMAN/3778</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF76326F902EAD8F894" ID-GBIF-Occurrence="4608353347" collectionCode="CRUST" location="Chipiona" municipality="Cadiz" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3779" specimenCount="14" specimenCount-female="8" specimenCount-male="6">
|
||
<specimenCount id="699F38E2FF99FFF76326F902E841F8B1" box="[788,855,1767,1793]" count="8" pageId="11" pageNumber="288" type="female">8 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF76354F903E8BEF8B1" box="[870,936,1766,1793]" count="6" pageId="11" pageNumber="288" type="male">6 ♂♂</specimenCount>
|
||
(30.0 mm max CW);
|
||
<collectingMunicipality id="9F426911FF99FFF7649DF903EFE0F8B0" box="[1199,1270,1766,1792]" pageId="11" pageNumber="288">Cádiz</collectingMunicipality>
|
||
,
|
||
<location id="7A46A5B0FF99FFF76531F903EE64F8B0" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF76531F903EE64F8B0" box="[1283,1394,1766,1792]" municipality="Cadiz" name="Chipiona" pageId="11" pageNumber="288">Chipiona</location>
|
||
;
|
||
<specimenCode id="2F3F5B10FF99FFF7608FF8EFEAD8F894" box="[189,462,1802,1828]" pageId="11" pageNumber="288">CRUST_ICMAN/3779</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF761D5F8EFE9E3F8F7" ID-GBIF-Occurrence="4608353341" collectingDate="2021-01-14" collectionCode="CRUST" collectorName="Helena Marco-Herrera" location="Isla Canela" municipality="Huelva" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3772" specimenCount="1" specimenCount-male="1" stateProvince="Gulf of Cadiz">
|
||
<specimenCount id="699F38E2FF99FFF761D5F8EFE900F894" box="[487,534,1802,1828]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="BD5D3D89FF99FFF76211F8EFE9D0F894" box="[547,710,1802,1828]" pageId="11" pageNumber="288">Gulf of Cádiz</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="9F426911FF99FFF762E0F8EFE83CF894" box="[722,810,1802,1828]" pageId="11" pageNumber="288">Huelva</collectingMunicipality>
|
||
,
|
||
<location id="7A46A5B0FF99FFF76304F8EFE8AAF894" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF76304F8EFE8AAF894" box="[822,956,1802,1828]" municipality="Huelva" name="Isla Canela" pageId="11" pageNumber="288" stateProvince="Gulf of Cadiz">Isla Canela</location>
|
||
;
|
||
<date id="0B27D5ABFF99FFF763FAF8EFEF76F894" box="[968,1120,1802,1828]" pageId="11" pageNumber="288" value="2021-01-14">
|
||
<collectingDate id="1B632C43FF99FFF763FAF8EFEF76F894" box="[968,1120,1802,1828]" pageId="11" pageNumber="288" value="2021-01-14">14 Jan. 2021</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF99FFF7645EF8EFEE61F894" box="[1132,1399,1802,1828]" pageId="11" pageNumber="288">Helena Marco-Herrera</collectorName>
|
||
leg.; FEMP-04: stn 10C;
|
||
<specimenCode id="2F3F5B10FF99FFF761D1F8C8E9E3F8F7" box="[483,757,1837,1863]" pageId="11" pageNumber="288">CRUST_ICMAN/3772</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF7633FF8C8EA74F8DA" ID-GBIF-Occurrence="4608353320" collectingDate="2021-01-14" collectionCode="CRUST" collectorName="Helena Marco-Herrera" location="Isla Canela" municipality="Huelva" pageId="11" pageNumber="288" specimenCode="CRUST_ ICMAN/3773" specimenCount="1" specimenCount-female="1" stateProvince="Gulf of Cadiz">
|
||
<specimenCount id="699F38E2FF99FFF7633FF8C8E82CF8F7" box="[781,826,1837,1863]" count="1" pageId="11" pageNumber="288" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="2F3F5B10FF99FFF7653BF8C8EA74F8DA" pageId="11" pageNumber="288">CRUST_ ICMAN/3773</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF761B4F8B5EA74F83D" ID-GBIF-Occurrence="4608353321" collectingDate="2021-01-14" collectionCode="CRUST" collectorName="Helena Marco-Herrera" location="Isla Canela" municipality="Huelva" pageId="11" pageNumber="288" specimenCode="CRUST_ ICMAN/3764" specimenCount="1" specimenCount-female="1" stateProvince="Gulf of Cadiz">
|
||
<specimenCount id="699F38E2FF99FFF761B4F8B5EAAFF8DB" box="[390,441,1872,1899]" count="1" pageId="11" pageNumber="288" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding; GenBank:
|
||
<accessionNumber id="60CA6E88FF99FFF76405F8B5EFA9F8DB" box="[1079,1215,1872,1899]" pageId="11" pageNumber="288">OR555886</accessionNumber>
|
||
–16S;
|
||
<specimenCode id="2F3F5B10FF99FFF7653BF8B5EA74F83D" pageId="11" pageNumber="288">CRUST_ ICMAN/3764</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF7614FF896E81FF800" ID-GBIF-Occurrence="4608353329" collectingDate="2021-04-12" collectionCode="CRUST" location="Donana National Park" municipality="Huelva" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3774, CRUST_ICMAN/3775, CRUST_ICMAN/3776" specimenCount="3" specimenCount-female="3" stateProvince="Gulf of Cadiz">
|
||
<specimenCount id="699F38E2FF99FFF7614FF896EAD4F83E" box="[381,450,1907,1934]" count="3" pageId="11" pageNumber="288" type="female">3 ♀♀</specimenCount>
|
||
;
|
||
<collectingRegion id="BD5D3D89FF99FFF761E0F896E96FF83D" box="[466,633,1907,1933]" pageId="11" pageNumber="288">Gulf of Cádiz</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="9F426911FF99FFF762B4F896E9C8F83D" box="[646,734,1907,1933]" pageId="11" pageNumber="288">Huelva</collectingMunicipality>
|
||
,
|
||
<location id="7A46A5B0FF99FFF762D9F891E8E1F83D" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF99FFF762D9F891E8E1F83D" box="[747,1015,1907,1934]" municipality="Huelva" name="Donana National Park" pageId="11" pageNumber="288" stateProvince="Gulf of Cadiz">Doñana National Park</location>
|
||
;
|
||
<date id="0B27D5ABFF99FFF76436F896EFB4F83D" box="[1028,1186,1907,1933]" pageId="11" pageNumber="288" value="2021-04-12">
|
||
<collectingDate id="1B632C43FF99FFF76436F896EFB4F83D" box="[1028,1186,1907,1933]" pageId="11" pageNumber="288" value="2021-04-12">12 Apr. 2021</collectingDate>
|
||
</date>
|
||
; FEMP-04: stn 6;
|
||
<specimenCode id="2F3F5B10FF99FFF7608FF872E81FF800" box="[189,777,1942,1969]" pageId="11" pageNumber="288">CRUST_ICMAN/3774 to CRUST_ICMAN/3776</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF76316F872EAD9F864" ID-GBIF-Occurrence="4608353317" collectingDate="2021-04-12" collectionCode="CRUST" location="Donana National Park" municipality="Huelva" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3780" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Gulf of Cadiz">
|
||
<specimenCount id="699F38E2FF99FFF76316F872E87EF801" box="[804,872,1943,1969]" count="4" pageId="11" pageNumber="288" type="female">4 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="699F38E2FF99FFF76345F872E8B1F801" box="[887,935,1943,1969]" count="1" pageId="11" pageNumber="288" type="male">1 ♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="2F3F5B10FF99FFF7608FF85FEAD9F864" box="[189,463,1978,2004]" pageId="11" pageNumber="288">CRUST_ICMAN/3780</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF99FFF061DFF85FEF2DFE93" ID-GBIF-Occurrence="4608353334" collectedFrom="fine sandy bottoms" collectingDate="2021-01-12" collectionCode="CRUST" elevation="8" lastPageId="12" lastPageNumber="289" location="Donana National Park" municipality="Huelva" pageId="11" pageNumber="288" specimenCode="CRUST_ICMAN/3781" specimenCount="6" specimenCount-female="6" stateProvince="Gulf of Cadiz">
|
||
<specimenCount id="699F38E2FF99FFF761DFF85FE924F864" box="[493,562,1978,2004]" count="6" pageId="11" pageNumber="288" type="female">6 ♀♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<date id="0B27D5ABFF99FFF76427F85FEFA4F864" box="[1045,1202,1978,2004]" pageId="11" pageNumber="288" value="2021-01-12">
|
||
<collectingDate id="1B632C43FF99FFF76427F85FEFA4F864" box="[1045,1202,1978,2004]" pageId="11" pageNumber="288" value="2021-01-12">12 Jan. 2021</collectingDate>
|
||
</date>
|
||
;
|
||
<elevation id="F4B41458FF99FFF764F3F85CEE64F864" box="[1217,1394,1977,2004]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="11" pageNumber="288" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">
|
||
<quantity id="B8615E8EFF99FFF764F3F85CEE32F864" box="[1217,1316,1977,2004]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="11" pageNumber="288" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 m</quantity>
|
||
depth
|
||
</elevation>
|
||
; FEMP-04: stns 10C, 6 and 2;
|
||
<collectedFrom id="CC85B096FF9EFFF06203FEECE80CFE93" box="[561,794,265,291]" pageId="12" pageNumber="289">fine sandy bottoms</collectedFrom>
|
||
;
|
||
<specimenCode id="2F3F5B10FF9EFFF0631BFEECEF2DFE93" box="[809,1083,265,291]" pageId="12" pageNumber="289">CRUST_ICMAN/3781</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF9EFFF06469FEECEE4CFEDA" ID-GBIF-Occurrence="4608353301" collectedFrom="from Donax trunculus Linnaeus, 1758 fisheries" collectingDate="2021-03-24" collectionCode="CRUST" collectorName="Garcia Raso" elevation="3" location="Marbella" municipality="Malaga" pageId="12" pageNumber="289" specimenCode="CRUST_ICMAN/3765" specimenCount="1" specimenCount-female="1" stateProvince="Alboran Sea">
|
||
<specimenCount id="699F38E2FF9EFFF06469FEECEF9AFE93" box="[1115,1164,265,291]" count="1" pageId="12" pageNumber="289" type="female">1 ♀</specimenCount>
|
||
; W Mediterranean,
|
||
<collectingRegion id="BD5D3D89FF9EFFF0608FFEC9EA45FEF6" box="[189,339,300,326]" pageId="12" pageNumber="289">Alboran Sea</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="9F426911FF9EFFF06152FEC9EAADFEF6" box="[352,443,300,326]" pageId="12" pageNumber="289">Málaga</collectingMunicipality>
|
||
,
|
||
<location id="7A46A5B0FF9EFFF061FAFEC9E920FEF6" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF9EFFF061FAFEC9E920FEF6" box="[456,566,300,326]" municipality="Malaga" name="Marbella" pageId="12" pageNumber="289" stateProvince="Alboran Sea">Marbella</location>
|
||
;
|
||
<elevation id="F4B41458FF9EFFF06271FEC9E9CAFEF6" box="[579,732,300,327]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="12" pageNumber="289" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">
|
||
<quantity id="B8615E8EFF9EFFF06271FEC9E998FEF7" box="[579,654,300,327]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="12" pageNumber="289" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2–4 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF9EFFF062DBFEC9E89AFEF6" box="[745,908,300,327]" pageId="12" pageNumber="289" value="2021-03-24">
|
||
<collectingDate id="1B632C43FF9EFFF062DBFEC9E89AFEF6" box="[745,908,300,327]" pageId="12" pageNumber="289" value="2021-03-24">24 Mar. 2021</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF9EFFF063ABFEC9EF3DFEF7" box="[921,1067,300,327]" pageId="12" pageNumber="289">García Raso</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF9EFFF0645DFEC9EAE3FED9" pageId="12" pageNumber="289">
|
||
from
|
||
<taxonomicName id="B89988E8FF9EFFF06480FEC8EA6BFEDA" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Liliopsida" family="Marantaceae" genus="Donax" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="289" phylum="Tracheophyta" rank="species" species="trunculus">
|
||
<emphasis id="4DED2F79FF9EFFF06480FEC8EE6EFEF6" box="[1202,1400,300,326]" italics="true" pageId="12" pageNumber="289">Donax trunculus</emphasis>
|
||
<bibRefCitation id="1B088E9AFF9EFFF0608FFEAAEA6BFEDA" author="Linnaeus C." box="[189,381,335,362]" pageId="12" pageNumber="289" refId="ref15623" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Laurentii Salvii, Stockholm [Holmiae]. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
</taxonomicName>
|
||
fisheries
|
||
</collectedFrom>
|
||
; GenBank:
|
||
<accessionNumber id="60CA6E88FF9EFFF062A7FEAAE80DFEDA" box="[661,795,335,362]" pageId="12" pageNumber="289">OR555891</accessionNumber>
|
||
–16S,
|
||
<accessionNumber id="60CA6E88FF9EFFF0635EFEAAE8F8FEDA" box="[876,1006,335,362]" pageId="12" pageNumber="289">OR557371</accessionNumber>
|
||
–COI;
|
||
<specimenCode id="2F3F5B10FF9EFFF0647AFEAAEE4CFEDA" box="[1096,1370,335,362]" pageId="12" pageNumber="289">CRUST_ICMAN/3765</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF9EFFF0608FFE96EAD9FE00" ID-GBIF-Occurrence="4608353304" collectedFrom="from Donax trunculus fisheries" collectingDate="1983-08" collectionCode="CRUST" collectorName="Garcia Raso" elevation="3" location="Fuengirola" municipality="Malaga" pageId="12" pageNumber="289" specimenCode="CRUST_ICMAN/3782" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="699F38E2FF9EFFF0608FFE96EBFBFE3D" box="[189,237,371,397]" count="1" pageId="12" pageNumber="289" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingMunicipality id="9F426911FF9EFFF060C9FE97EA40FE3C" box="[251,342,370,396]" pageId="12" pageNumber="289">Málaga</collectingMunicipality>
|
||
,
|
||
<location id="7A46A5B0FF9EFFF06151FE97EAF0FE3C" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF9EFFF06151FE97EAF0FE3C" box="[355,486,370,396]" municipality="Malaga" name="Fuengirola" pageId="12" pageNumber="289">Fuengirola</location>
|
||
;
|
||
<elevation id="F4B41458FF9EFFF061C1FE96E99AFE3C" box="[499,652,370,397]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="12" pageNumber="289" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">
|
||
<quantity id="B8615E8EFF9EFFF061C1FE96E929FE3D" box="[499,575,371,397]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="12" pageNumber="289" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2–4 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="0B27D5ABFF9EFFF062AAFE96E80EFE3D" box="[664,792,371,397]" pageId="12" pageNumber="289" value="1983-08">
|
||
<collectingDate id="1B632C43FF9EFFF062AAFE96E80EFE3D" box="[664,792,371,397]" pageId="12" pageNumber="289" value="1983-08">Aug. 1983</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="D26C96BDFF9EFFF06314FE96E8AEFE3D" box="[806,952,371,397]" pageId="12" pageNumber="289">García Raso</collectorName>
|
||
leg.;
|
||
<collectedFrom id="CC85B096FF9EFFF063CEFE97EE65FE3C" box="[1020,1395,370,396]" pageId="12" pageNumber="289">
|
||
from
|
||
<taxonomicName id="B89988E8FF9EFFF0640CFE96EE12FE3C" authorityName="Linnaeus" authorityYear="1758" box="[1086,1284,370,396]" class="Liliopsida" family="Marantaceae" genus="Donax" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="289" phylum="Tracheophyta" rank="species" species="trunculus">
|
||
<emphasis id="4DED2F79FF9EFFF0640CFE96EE12FE3C" box="[1086,1284,370,396]" italics="true" pageId="12" pageNumber="289">Donax trunculus</emphasis>
|
||
</taxonomicName>
|
||
fisheries
|
||
</collectedFrom>
|
||
;
|
||
<specimenCode id="2F3F5B10FF9EFFF0608FFE73EAD9FE00" box="[189,463,406,432]" pageId="12" pageNumber="289">CRUST_ICMAN/3782</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="CFF1F936FF9EFFF061DBFE73EF34FE63" ID-GBIF-Occurrence="4608353312" collectedFrom="from Donax trunculus fisheries" collectingDate="1979" collectingDateMax="1981" collectingDateMin="1979" collectionCode="CRUST" collectorName="Garcia Raso" location="Malaga" pageId="12" pageNumber="289" specimenCode="CRUST_ICMAN/3783" specimenCount="33" specimenCount-female="19" specimenCount-male="14">
|
||
<specimenCount id="699F38E2FF9EFFF061DBFE73E92CFE00" box="[489,570,406,432]" count="18" pageId="12" pageNumber="289" type="female">18 ♀♀</specimenCount>
|
||
(smallest ovigerous
|
||
<specimenCount id="699F38E2FF9EFFF0631DFE73E851FE00" box="[815,839,406,432]" count="1" pageId="12" pageNumber="289" type="female">♀</specimenCount>
|
||
:
|
||
<quantity id="B8615E8EFF9EFFF06367FE73E8A9FE00" box="[853,959,406,432]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="12" pageNumber="289" unit="mm" value="16.4">16.4 mm</quantity>
|
||
width),
|
||
<specimenCount id="699F38E2FF9EFFF06410FE73EF60FE00" box="[1058,1142,406,432]" count="14" pageId="12" pageNumber="289" type="male">14 ♂♂</specimenCount>
|
||
;
|
||
<location id="7A46A5B0FF9EFFF064B1FE73EFC9FE00" LSID="urn:lsid:plazi:treatment:F730427DFF98FFF06244FD67EF33FE63:7A46A5B0FF9EFFF064B1FE73EFC9FE00" box="[1155,1247,406,432]" name="Malaga" pageId="12" pageNumber="289">Málaga</location>
|
||
;
|
||
<collectingDate id="1B632C43FF9EFFF064D9FE73EE31FE00" box="[1259,1319,406,432]" pageId="12" pageNumber="289">1979</collectingDate>
|
||
–
|
||
<collectingDate id="1B632C43FF9EFFF0650AFE73EE64FE00" box="[1336,1394,406,432]" pageId="12" pageNumber="289">1981</collectingDate>
|
||
,
|
||
<collectorName id="D26C96BDFF9EFFF0608FFE5CEA5BFE63" box="[189,333,441,467]" pageId="12" pageNumber="289">García Raso</collectorName>
|
||
leg.,
|
||
<collectedFrom id="CC85B096FF9EFFF061BCFE5CE816FE63" box="[398,768,441,467]" pageId="12" pageNumber="289">
|
||
from
|
||
<taxonomicName id="B89988E8FF9EFFF061FDFE5FE985FE63" authorityName="Linnaeus" authorityYear="1758" box="[463,659,441,467]" class="Liliopsida" family="Marantaceae" genus="Donax" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="289" phylum="Tracheophyta" rank="species" species="trunculus">
|
||
<emphasis id="4DED2F79FF9EFFF061FDFE5FE985FE63" box="[463,659,441,467]" italics="true" pageId="12" pageNumber="289">Donax trunculus</emphasis>
|
||
</taxonomicName>
|
||
fisheries
|
||
</collectedFrom>
|
||
;
|
||
<specimenCode id="2F3F5B10FF9EFFF0633EFE5CEF34FE63" box="[780,1058,441,467]" pageId="12" pageNumber="289">CRUST_ICMAN/3783</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3783A0E0FF9FFFED608FFE37EFE7F80E" lastPageId="17" lastPageNumber="294" pageId="13" pageNumber="290" type="description">
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<paragraph id="7F26F36BFF9FFFF1608FFE37EA58FE5C" blockId="13.[189,334,466,492]" box="[189,334,466,492]" pageId="13" pageNumber="290">
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<heading id="246E4407FF9FFFF1608FFE37EA58FE5C" bold="true" box="[189,334,466,492]" fontSize="11" level="3" pageId="13" pageNumber="290" reason="3">
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||
<emphasis id="4DED2F79FF9FFFF1608FFE37EA58FE5C" bold="true" box="[189,334,466,492]" pageId="13" pageNumber="290">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF9FFFF1608FFDE4E99FFDAB" blockId="13.[189,649,513,539]" box="[189,649,513,539]" pageId="13" pageNumber="290">
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<smallCapsWord id="79C065B7FF9FFFF1608FFDE4EA26FDA9" baselines="532,532" box="[189,304,513,539]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Maximum" pageId="13" pageNumber="290">MAXIMUM</smallCapsWord>
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<smallCapsWord id="79C065B7FF9FFFF16105FDE3EA75FDA9" baselines="532" box="[311,355,518,537]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="size" pageId="13" pageNumber="290">SIZE</smallCapsWord>
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. CL/CW: 30.7/
|
||
<quantity id="B8615E8EFF9FFFF1622BFDE4E993FDAB" box="[537,645,513,539]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="13" pageNumber="290" unit="mm" value="40.6">40.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF9FFFF1608FFDA6EF5AFB66" blockId="13.[189,1401,578,1238]" pageId="13" pageNumber="290">
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<smallCapsWord id="79C065B7FF9FFFF1608FFDA6EA0BFDEB" baselines="598,598" box="[189,285,579,605]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="13" pageNumber="290">HABITUS</smallCapsWord>
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. Carapace glabrous, wider than long (length/width relationships, CL/CW = 0.75 to 0.84 (
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||
<typeStatus id="A0224DC9FF9FFFF160F7FD80EA24FDCF" box="[197,306,613,639]" pageId="13" pageNumber="290" type="holotype">holotype</typeStatus>
|
||
: 0.78) (
|
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<figureCitation id="E7A2EFEEFF9FFFF161BCFD80EAE4FD30" box="[398,498,613,640]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="13" pageNumber="290">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16233FD83E931FD30" box="[513,551,614,640]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="13" pageNumber="290">4A</figureCitation>
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,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16204FD80E94BFD30" box="[566,605,613,640]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="13" pageNumber="290">5A</figureCitation>
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||
), somewhat domed (convex), nearly smooth, but with some small tubercles especially located in the protogastric areas (in small aggregations) and in the anterior regions of the branchial areas (forming striae). Frontal region flat, with three slightly protruding blunt triangular teeth (
|
||
<figureCitation id="E7A2EFEEFF9FFFF1613BFD2AEA99FD5A" box="[265,399,719,746]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="13" pageNumber="290">Figs 3A, F</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF161ADFD2AEAD3FD59" box="[415,453,719,745]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="13" pageNumber="290">4A</figureCitation>
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,
|
||
<figureCitation id="E7A2EFEEFF9FFFF161E7FD2AEAEDFD5A" box="[469,507,719,746]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="13" pageNumber="290">5A</figureCitation>
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,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16239FD2AE941FD59" box="[523,599,719,745]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[341,1247,265,1812]" captionTargetId="figure-17@18.[341,1247,265,1812]" captionTargetPageId="18" captionText="Fig. 6. Dorsal view of orbital region and adjacent lateral rostral teeth. A. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). B. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). C. Polybius dioscurus sp. nov., ♀ from Málaga (CRUST_ICMAN/3765) D. Polybius vernalis, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species." figureDoi="http://doi.org/10.5281/zenodo.10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="13" pageNumber="290">6A, C</figureCitation>
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||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16255FD2AE99DFD59" box="[615,651,719,745]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="13" pageNumber="290">7B</figureCitation>
|
||
), with a tuberculate rim, continued by the inner-orbital angle (
|
||
<figureCitation id="E7A2EFEEFF9FFFF160F4FD17EA5BFCBC" box="[198,333,754,780]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[341,1247,265,1812]" captionTargetId="figure-17@18.[341,1247,265,1812]" captionTargetPageId="18" captionText="Fig. 6. Dorsal view of orbital region and adjacent lateral rostral teeth. A. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). B. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). C. Polybius dioscurus sp. nov., ♀ from Málaga (CRUST_ICMAN/3765) D. Polybius vernalis, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species." figureDoi="http://doi.org/10.5281/zenodo.10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="13" pageNumber="290">Fig. 6A, C</figureCitation>
|
||
). This region is antero-dorsally delimited by a clear band; the middle tooth is rounded, subacute and does not reach the same proportions as the lateral ones, which are more developed (wider and rounded). Relative distance between external orbital teeth (apex to apex) and the distance between lateral rostral teeth (apex to apex) =
|
||
<quantity id="B8615E8EFF9FFFF162B1FCB9E82FFCC6" box="[643,825,860,887]" metricMagnitude="-2" metricUnit="m" metricValue="8.5979" metricValueMax="9.0424" metricValueMin="8.1534" pageId="13" pageNumber="290" unit="in" value="3.385" valueMax="3.56" valueMin="3.21">3.21 to 3.56 in</quantity>
|
||
males (
|
||
<typeStatus id="A0224DC9FF9FFFF163AEFCB9EF1FFCC6" box="[924,1033,860,886]" pageId="13" pageNumber="290" type="holotype">holotype</typeStatus>
|
||
: 3.23; mean: 3.38),
|
||
<quantity id="B8615E8EFF9FFFF16530FCB9EBC2FC29" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="9.0424" metricValueMin="8.7376" pageId="13" pageNumber="290" unit="in" value="3.5" valueMax="3.56" valueMin="3.44">3.44–3.56 in</quantity>
|
||
females (mean 3.50). Orbital area circular, concave, dorsally with two short longitudinal incisions, the internal longer (
|
||
<figureCitation id="E7A2EFEEFF9FFFF1619EFC47E926FC0D" box="[428,560,930,957]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="13" pageNumber="290">Figs 3A, F</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF1620CFC47E972FC0C" box="[574,612,930,956]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="13" pageNumber="290">4A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16241FC47E98FFC0D" box="[627,665,930,957]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="13" pageNumber="290">5A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16295FC47E9D8FC0C" box="[679,718,930,956]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[341,1247,265,1812]" captionTargetId="figure-17@18.[341,1247,265,1812]" captionTargetPageId="18" captionText="Fig. 6. Dorsal view of orbital region and adjacent lateral rostral teeth. A. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). B. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). C. Polybius dioscurus sp. nov., ♀ from Málaga (CRUST_ICMAN/3765) D. Polybius vernalis, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species." figureDoi="http://doi.org/10.5281/zenodo.10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="13" pageNumber="290">6A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF162EEFC47E817FC0C" box="[732,769,930,956]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="13" pageNumber="290">7B</figureCitation>
|
||
); ventral face with a very pronounced outer incision and a well protruding antero-internal lobe. Anterolateral borders of the carapace (
|
||
<figureCitation id="E7A2EFEEFF9FFFF16492FC20EE13FC50" box="[1184,1285,965,992]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="13" pageNumber="290">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16527FC23EE2DFC50" box="[1301,1339,966,992]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="13" pageNumber="290">4A</figureCitation>
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||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16579FC20EE66FC50" box="[1355,1392,965,992]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="13" pageNumber="290">5A</figureCitation>
|
||
) well curved, with a length more or less similar than the postero-lateral ones (0.95 to 1.17), which are converging. Anterolateral edges with five teeth (including the external orbital), the fifth and the fourth being somewhat larger, the third and fourth with a rectilinear outer face. Ventrally, the anterior margins (anterolateral teeth region) are delimited by a dense row of short setae; with a fringe of setae behind the pterygostomial sulcus. Branchial regions slightly domed, like the protogastric and centrogastric ones; concave urogastric region (depressed); cardiac region, with two tuberculate protruding areas anteriorly (and sometimes with two smaller ones behind) and flattened intestinal region.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF9FFFF1608FFB1BE8ACFAA7" blockId="13.[189,954,1277,1304]" box="[189,954,1277,1304]" pageId="13" pageNumber="290">
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. With two inclined longitudinal keels, one on each side.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF9FFFF1608FFADAEFB3FACC" blockId="13.[189,1398,1343,1404]" pageId="13" pageNumber="290">
|
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|
||
(A1). With a broad transversely folded basal segment in a fossa below the frontal area, with the ventral edge showing a projecting lobe located towards the third of the outer half.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF9FFFF1608FFA41EF43FA50" blockId="13.[189,1399,1443,1505]" pageId="13" pageNumber="290">
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|
||
(A2). Basal segments narrow, with a flagellum of about 26 segments, longer than the orbital concavity, reaching the basal half of the first anterolateral tooth (outer orbital).
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF9FFFED608FF9EDEA51FE00" blockId="13.[188,1400,1544,1993]" lastBlockId="17.[188,1400,265,432]" lastPageId="17" lastPageNumber="294" pageId="13" pageNumber="290">
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(P1) (
|
||
<figureCitation id="E7A2EFEEFF9FFFF1614CF9EDE917F992" box="[382,513,1544,1570]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="13" pageNumber="290">Figs 3A–C</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF1623CF9EDE941F992" box="[526,599,1544,1570]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="13" pageNumber="290">4A–C</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF16251F9EDE9BDF992" box="[611,683,1544,1571]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="13" pageNumber="290">5A, D</figureCitation>
|
||
). Subequal. Propodus with a rounded ventral rim on its outer face, extending from the apex of the fixed finger to approximately the middle of the palm. Outer face of the palm with 3 longitudinal keels along entire length (one towards the middle zone and two in the dorsal zone, slightly more separated from each other, the upper one ends in the propo-carpal articular condyle); dorsal edge with a smooth rounded keel ending in a blunt distal tooth; internal face without keels, with the central area bulging (longitudinally) leaving two depressions (upper and lower). The palm (without the fixed finger) is 1.7 times as long as high and 1.48 times as long as the dactyl. Dactyl (mobile finger) with the distal part curved, externally presenting a central longitudinal depression, between two rounded and projecting ridges; dorsally, three rounded longitudinal ridges can be seen: an upper external one (which is the upper rounded edge of the external face), a central dorsal (somewhat more protruding, especially in its basal part) and an upper internal one, with depressions between them; inner face with a central longitudinal depression delimited by two rounded edges. Carpus (
|
||
<figureCitation id="E7A2EFEEFF9FFFF1640BF86EEF8CF815" box="[1081,1178,1931,1957]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="13" pageNumber="290">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF1649AF86EEFDBF815" box="[1192,1229,1931,1957]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="13" pageNumber="290">4C</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF9FFFF164E8F86EEFE9F816" box="[1242,1279,1931,1958]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="13" pageNumber="290">5D</figureCitation>
|
||
) with one strong and protruding tooth on antero inner dorsal margin; on the anterior outer upper part (coinciding with the superior articulation with the propodus) there is one well-marked and rounded projection and two others below: a not very prominent middle one and a bit more prominent posterior one. The external face ornamented with slightly raised areas and soft depressions. Merus smooth, with short setae along the entire upper edge (less in the most distal area) and inner edge of lower face (more developed in the distal area).
|
||
</paragraph>
|
||
<caption id="2BE6A3E3FF9CFFF2608FF881E91BF858" ID-DOI="http://doi.org/10.5281/zenodo.10978033" ID-Zenodo-Dep="10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="14" pageNumber="291" startId="14.[189,232,1892,1918]" targetBox="[247,1340,265,1845]" targetPageId="14">
|
||
<paragraph id="7F26F36BFF9CFFF2608FF881E91BF858" blockId="14.[187,1399,1892,2024]" pageId="14" pageNumber="291">
|
||
<emphasis id="4DED2F79FF9CFFF2608FF881EA07F8CF" bold="true" box="[189,273,1892,1919]" pageId="14" pageNumber="291">Fig. 3.</emphasis>
|
||
<taxonomicName id="B89988E8FF9CFFF2612FF880E915F8CF" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[285,515,1893,1919]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="291" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF9CFFF2612FF880E915F8CF" box="[285,515,1893,1919]" italics="true" pageId="14" pageNumber="291">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF9CFFF2623DF880E964F8CF" box="[527,626,1893,1919]" pageId="14" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759).
|
||
<emphasis id="4DED2F79FF9CFFF2640FF86DEF44F812" bold="true" box="[1085,1106,1928,1954]" pageId="14" pageNumber="291">A</emphasis>
|
||
. Dorsal habitus.
|
||
<emphasis id="4DED2F79FF9CFFF26521F86DEE31F812" bold="true" box="[1299,1319,1928,1954]" pageId="14" pageNumber="291">B</emphasis>
|
||
. Right cheliped, facial view.
|
||
<emphasis id="4DED2F79FF9CFFF2618FF84EEAC4F875" bold="true" box="[445,466,1963,1989]" pageId="14" pageNumber="291">C</emphasis>
|
||
. Carpus of left cheliped.
|
||
<emphasis id="4DED2F79FF9CFFF262C8F84EE819F875" bold="true" box="[762,783,1963,1989]" pageId="14" pageNumber="291">D</emphasis>
|
||
. Carpus to dactylus of right pereiopod 5.
|
||
<emphasis id="4DED2F79FF9CFFF264C5F84EEE1DF875" bold="true" box="[1271,1291,1963,1989]" pageId="14" pageNumber="291">E</emphasis>
|
||
. Body in ventral view, abdomen view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2BE6A3E3FF9DFFF3608FF881EACAF858" ID-DOI="http://doi.org/10.5281/zenodo.10978037" ID-Zenodo-Dep="10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="15" pageNumber="292" startId="15.[189,232,1892,1918]" targetBox="[278,1309,282,1800]" targetPageId="15">
|
||
<paragraph id="7F26F36BFF9DFFF3608FF881EACAF858" blockId="15.[188,1399,1891,2025]" pageId="15" pageNumber="292">
|
||
<emphasis id="4DED2F79FF9DFFF3608FF881EA1AF8CF" bold="true" box="[189,268,1892,1919]" pageId="15" pageNumber="292">Fig. 4.</emphasis>
|
||
<taxonomicName id="B89988E8FF9DFFF36127F880EAE0F8CF" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[277,502,1893,1919]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="292" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF9DFFF36127F880EAE0F8CF" box="[277,502,1893,1919]" italics="true" pageId="15" pageNumber="292">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF9DFFF361CDF880E94BF8CF" box="[511,605,1893,1919]" pageId="15" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762).
|
||
<emphasis id="4DED2F79FF9DFFF36491F86DEFAEF812" bold="true" box="[1187,1208,1928,1954]" pageId="15" pageNumber="292">A</emphasis>
|
||
. Dorsal habitus.
|
||
<emphasis id="4DED2F79FF9DFFF3608EF84EEBC6F875" bold="true" box="[188,208,1963,1989]" pageId="15" pageNumber="292">B</emphasis>
|
||
. Right cheliped, facial view.
|
||
<emphasis id="4DED2F79FF9DFFF3620BF84EE958F875" bold="true" box="[569,590,1963,1989]" pageId="15" pageNumber="292">C</emphasis>
|
||
. Carpus of left cheliped.
|
||
<emphasis id="4DED2F79FF9DFFF363BAF84EE88BF875" bold="true" box="[904,925,1963,1989]" pageId="15" pageNumber="292">D</emphasis>
|
||
. Dactylus of right pereiopods 5 and 4.
|
||
<emphasis id="4DED2F79FF9DFFF3608FF82BEBC7F858" bold="true" box="[189,209,1998,2024]" pageId="15" pageNumber="292">E</emphasis>
|
||
. Body in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2BE6A3E3FF82FFEC608FF86DEA3EF858" ID-DOI="http://doi.org/10.5281/zenodo.10978041" ID-Zenodo-Dep="10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="16" pageNumber="293" startId="16.[189,232,1928,1954]" targetBox="[190,1398,265,1875]" targetPageId="16">
|
||
<paragraph id="7F26F36BFF82FFEC608FF86DEA3EF858" blockId="16.[189,1399,1928,2024]" pageId="16" pageNumber="293">
|
||
<emphasis id="4DED2F79FF82FFEC608FF86DEA1CF812" bold="true" box="[189,266,1928,1954]" pageId="16" pageNumber="293">Fig. 5.</emphasis>
|
||
<taxonomicName id="B89988E8FF82FFEC6123F86DEAE6F812" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[273,496,1928,1954]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="293" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF82FFEC6123F86DEAE6F812" box="[273,496,1928,1954]" italics="true" pageId="16" pageNumber="293">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF82FFEC61C4F86DE944F812" box="[502,594,1928,1954]" pageId="16" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223).
|
||
<emphasis id="4DED2F79FF82FFEC61D7F84EEAECF875" bold="true" box="[485,506,1963,1989]" pageId="16" pageNumber="293">A</emphasis>
|
||
. Dorsal habitus.
|
||
<emphasis id="4DED2F79FF82FFEC62FEF84EE9F6F875" bold="true" box="[716,736,1963,1989]" pageId="16" pageNumber="293">B</emphasis>
|
||
. Facial habitus.
|
||
<emphasis id="4DED2F79FF82FFEC6398F84EE8A9F875" bold="true" box="[938,959,1963,1989]" pageId="16" pageNumber="293">C</emphasis>
|
||
. Ventral habitus.
|
||
<emphasis id="4DED2F79FF82FFEC64A5F84EEFBAF875" bold="true" box="[1175,1196,1963,1989]" pageId="16" pageNumber="293">D</emphasis>
|
||
. Carpus of right cheliped.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7F26F36BFF83FFED608FFE3DEAA6FDCE" blockId="17.[188,1399,471,638]" pageId="17" pageNumber="294">
|
||
<smallCapsWord id="79C065B7FF83FFED608FFE3DEA3EFE40" baselines="491,491" box="[189,296,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Walking" pageId="17" pageNumber="294">WALKING</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED6102FE38EA72FE40" baselines="491" box="[304,356,477,496]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="legs" pageId="17" pageNumber="294">LEGS</smallCapsWord>
|
||
. Morphometric values: merus length (ML): P3 ≈ 2 ≈ 4> 5. The ML/MH (merus length/ height) ratio of P2–P3–P4 is between 2.9 to 3.1 and about
|
||
<quantity id="B8615E8EFF83FFED635BFE1EE8BAFDA4" box="[873,940,506,533]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="17" pageNumber="294" unit="in" value="1.8">1.8 in</quantity>
|
||
P5. The ratio MLP2/P5, MLP3/P5 and MLP4/P5 is 1.82–1.88–1.8, respectively. Propodus length (PL): P3 slightly> P4 slightly> P2. Dactylus length (DL): P2 somewhat> P3 ≈ P4. The dactylus of all walking legs (P2 to P4) are longer than their respective propodus.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF83FFED608FFD43E8AEFBD6" blockId="17.[187,1400,677,1126]" pageId="17" pageNumber="294">
|
||
<smallCapsWord id="79C065B7FF83FFED608FFD43EA02FD0E" baselines="697,697" box="[189,276,678,704]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Second" pageId="17" pageNumber="294">SECOND</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED612DFD4EEA8BFD0E" baselines="697" box="[287,413,683,702]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pereiopods" pageId="17" pageNumber="294">PEREIOPODS</smallCapsWord>
|
||
(P2). Merus smooth, with setae along the basal half of the dorsal margin, ventral margin without setae. Carpus and propodus smooth. Both segments with two dorsal ridges. On the propodus, external (anterior) face delimited ventrally by a row of very short setae, not extending onto carpus, and another row on internal (posterior) face, both separated by a dorsal longitudinal depression (wider in carpus). On the ventral edge of both segments, and along their entire length, there is a dense row of short setae. Inner face of carpus smooth and that of the propodus with a slight central longitudinal depression at anterior part. Dactylus of P2 quite rectilinear dorsally, and morphologically similar to those of P3 and P4, but wider, robust and with a slightly more curved tip; with a more or less quadrangular section, and with angles defined by rounded longitudinal edges, which leave a central longitudinal depression on dorsal, ventral and lateral sides (external and internal). Two dense rows of short setae are present on the anterior (external) face: one ventral (on the ventral edge) that exceeds half the length of the dactyl, up to approximately ⅔ basal, and another dorsal, somewhat shorter (on the dorsal edge) row, which runs from the basal part of the dactyl to half or little more.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF83FFED608FFB6BE801FACB" blockId="17.[188,1399,1165,1403]" pageId="17" pageNumber="294">
|
||
<smallCapsWord id="79C065B7FF83FFED608FFB6BEA14FB16" baselines="1185,1185" box="[189,258,1166,1192]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Third" pageId="17" pageNumber="294">THIRD</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED613AFB76EA23FB16" baselines="1185" box="[264,309,1171,1190]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="17" pageNumber="294">AND</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED6109FB76EA9BFB16" baselines="1185" box="[315,397,1171,1190]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="fourth" pageId="17" pageNumber="294">FOURTH</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED61A1FB76E907FB16" baselines="1185" box="[403,529,1171,1190]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pereiopods" pageId="17" pageNumber="294">PEREIOPODS</smallCapsWord>
|
||
(P3 and P4). Similar to the second one (P2) and to each other. Both present setae on the basal half (P3), or on the entire (P4) dorsal (or superior) edge of the merus, and are glabrous on the ventral edge. Carpus and propodus smooth (without setae on the dorsal and ventral faces). The dactyli are morphologically similar to those of P2, but somewhat less robust and narrower (2
|
||
<superScript id="88EC5E23FF83FFED653CFB13EE09FAB4" attach="left" box="[1294,1311,1270,1284]" fontSize="6" pageId="17" pageNumber="294">nd</superScript>
|
||
> 3
|
||
<superScript id="88EC5E23FF83FFED657DFB13EE4BFAB4" attach="left" box="[1359,1373,1270,1284]" fontSize="6" pageId="17" pageNumber="294">rd</superScript>
|
||
> 4
|
||
<superScript id="88EC5E23FF83FFED60FEFAFCEBCFFA97" attach="right" box="[204,217,1305,1319]" fontSize="6" pageId="17" pageNumber="294">th</superScript>
|
||
). In P3 the row of short ventral setae goes from the basal part to half the length of the dactyl, or little more; while in P4 they are only found in the basal third (or a little more) (
|
||
<figureCitation id="E7A2EFEEFF83FFED642FFAD8EF6DFAE7" box="[1053,1147,1341,1367]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="17" pageNumber="294">Fig. 4D</figureCitation>
|
||
), not reaching half of the dactyl, the row of dorsal setae is like that of P2.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF83FFED608FFA47E924F9F9" blockId="17.[189,1399,1442,1609]" pageId="17" pageNumber="294">
|
||
<smallCapsWord id="79C065B7FF83FFED608FFA47EBEDFA0B" baselines="1461,1462" box="[189,251,1442,1468]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Fifth" pageId="17" pageNumber="294">FIFTH</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED6131FA42EA97FA0A" baselines="1461" box="[259,385,1447,1466]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pereiopods" pageId="17" pageNumber="294">PEREIOPODS</smallCapsWord>
|
||
(P5). Merus smooth, with long setae all over the dorsal border, and in the distal fourth of the ventral outer part; ML/MH ratio: 1.63–1.85 (males), 1.76–2.0 (females). Smooth carpus, with long setae all over the dorsal and ventral edge. Propodus and dactylus flattened, with long setae on all edges (dorsal and ventral). Dactyl strongly flattened, paddle-like, oval (
|
||
<figureCitation id="E7A2EFEEFF83FFED63CCF9EEEF95F996" box="[1022,1155,1547,1574]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="17" pageNumber="294">Figs 3A, D</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF83FFED64BCF9E9EFCEF996" box="[1166,1240,1548,1574]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="17" pageNumber="294">4A, D</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF83FFED64D6F9EEEE3DF996" box="[1252,1323,1547,1574]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="17" pageNumber="294">5A, C</figureCitation>
|
||
) (2.11 to 2.37 L/W), ending in a point.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF83FFED608FF994E8E9F91D" blockId="17.[189,1399,1648,1710]" pageId="17" pageNumber="294">
|
||
<smallCapsWord id="79C065B7FF83FFED608FF994EBECF939" baselines="1668,1668" box="[189,250,1649,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="First" pageId="17" pageNumber="294">FIRST</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED6137F993EA41F939" baselines="1668" box="[261,343,1654,1673]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="sexual" pageId="17" pageNumber="294">SEXUAL</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED6150F993EA8BF939" baselines="1668" box="[354,413,1654,1673]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="male" pageId="17" pageNumber="294">MALE</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED6195F993E912F939" baselines="1668" box="[423,516,1654,1673]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pleopod" pageId="17" pageNumber="294">PLEOPOD</smallCapsWord>
|
||
(
|
||
<smallCapsWord id="79C065B7FF83FFED622BF993E968F939" baselines="1668" box="[537,638,1654,1673]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="gonopod" pageId="17" pageNumber="294">GONOPOD</smallCapsWord>
|
||
). With curved distal part, which forms an angle of almost 90° (
|
||
<figureCitation id="E7A2EFEEFF83FFED60F4F976EA50F91D" box="[198,326,1683,1709]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[218,1370,265,1801]" captionTargetId="figure-17@20.[218,1370,265,1801]" captionTargetPageId="20" captionText="Fig. 8. Left gonopod 1 of male. A–B. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). A. Ventral view. B. Dorsal view. C–D. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C. Ventral view. D. Dorsal view. E–F. Polybius vernalis (Risso, 1827) from Tunisia, Bay of Tunis (RBINS, INV. 187235).E. Ventral view. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10978047" httpUri="https://zenodo.org/record/10978047/files/figure.png" pageId="17" pageNumber="294">Fig. 8C–D</figureCitation>
|
||
), more curved in large specimen, and ends in a small point.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF83FFED608FF930EA26F885" blockId="17.[189,1399,1748,1845]" pageId="17" pageNumber="294">
|
||
<smallCapsWord id="79C065B7FF83FFED608FF930EA12F95D" baselines="1768,1768" box="[189,260,1749,1775]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pleon" pageId="17" pageNumber="294">PLEON</smallCapsWord>
|
||
. Male pleon embedded in the sternum (
|
||
<figureCitation id="E7A2EFEEFF83FFED62C1F930E843F95F" box="[755,853,1749,1775]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="17" pageNumber="294">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF83FFED6357F930E89FF95F" box="[869,905,1749,1775]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="17" pageNumber="294">4E</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF83FFED63AAF930E8ABF940" box="[920,957,1749,1776]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="17" pageNumber="294">5C</figureCitation>
|
||
), with the 3
|
||
<superScript id="88EC5E23FF83FFED6466F931EF74F952" attach="left" box="[1108,1122,1748,1762]" fontSize="6" pageId="17" pageNumber="294">rd</superScript>
|
||
to 5
|
||
<superScript id="88EC5E23FF83FFED64ADF931EFBAF952" attach="left" box="[1183,1196,1748,1762]" fontSize="6" pageId="17" pageNumber="294">th</superScript>
|
||
segments fused, narrowing strongly from the beginning of the 6
|
||
<superScript id="88EC5E23FF83FFED62CFF912E81CF8B5" attach="left" box="[765,778,1783,1797]" fontSize="6" pageId="17" pageNumber="294">th</superScript>
|
||
and ending in a triangular telson. Rounded female abdomen.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF83FFED608FF8B8EFE7F80E" blockId="17.[189,1398,1885,1982]" pageId="17" pageNumber="294">
|
||
<smallCapsWord id="79C065B7FF83FFED608FF8B8EA0EF8C5" baselines="1904,1904" box="[189,280,1885,1911]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="17" pageNumber="294">COLOUR</smallCapsWord>
|
||
<smallCapsWord id="79C065B7FF83FFED6110F887EA6DF8C5" baselines="1904" box="[290,379,1890,1909]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pattern" pageId="17" pageNumber="294">PATTERN</smallCapsWord>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF83FFED61BCF8B8EAC1F8C7" box="[398,471,1885,1911]" captionStart="Fig" captionStartId="22.[188,231,1963,1989]" captionTargetBox="[189,1398,558,1917]" captionTargetId="figure-108@22.[189,1398,558,1917]" captionTargetPageId="22" captionText="Fig. 9. Polybius dioscurus sp. nov., paratypes from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187230), colour variation (in life) (sex not recorded during photo session)." figureDoi="http://doi.org/10.5281/zenodo.10978049" httpUri="https://zenodo.org/record/10978049/files/figure.png" pageId="17" pageNumber="294">Fig. 9</figureCitation>
|
||
). Body and legs usually greyish with a finely speckled motive; a large white, brown or black mark is occasionally present on the anterior part of carapace. Some specimens have a marbled colour pattern. The legs never exhibit the orange hue, which is usual in
|
||
<taxonomicName id="B89988E8FF83FFED645EF841EFFDF80D" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1132,1259,1955,1981]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="294" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF83FFED645EF841EFFDF80D" box="[1132,1259,1955,1981]" italics="true" pageId="17" pageNumber="294">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2BE6A3E3FF80FFEE608FF8A4EF81F858" ID-DOI="http://doi.org/10.5281/zenodo.10978043" ID-Zenodo-Dep="10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="18" pageNumber="295" startId="18.[189,232,1857,1883]" targetBox="[341,1247,265,1812]" targetPageId="18">
|
||
<paragraph id="7F26F36BFF80FFEE608FF8A4EF81F858" blockId="18.[189,1399,1857,2025]" pageId="18" pageNumber="295">
|
||
<emphasis id="4DED2F79FF80FFEE608FF8A4EA1BF8EB" bold="true" box="[189,269,1857,1883]" pageId="18" pageNumber="295">Fig. 6.</emphasis>
|
||
Dorsal view of orbital region and adjacent lateral rostral teeth.
|
||
<emphasis id="4DED2F79FF80FFEE6434F8A4EF0CF8EB" bold="true" box="[1030,1050,1857,1883]" pageId="18" pageNumber="295">A</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF80FFEE641BF8A4EE1DF8EB" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[1065,1291,1857,1883]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="295" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF80FFEE641BF8A4EE1DF8EB" box="[1065,1291,1857,1883]" italics="true" pageId="18" pageNumber="295">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF80FFEE6526F8A7EE64F8EC" box="[1300,1394,1858,1884]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223).
|
||
<emphasis id="4DED2F79FF80FFEE64DDF880EE15F8CF" bold="true" box="[1263,1283,1893,1919]" pageId="18" pageNumber="295">B</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF80FFEE6522F880EADFF812" authority="(Risso, 1827)" baseAuthorityName="Risso" baseAuthorityYear="1827" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="295" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF80FFEE6522F880EA0AF812" italics="true" pageId="18" pageNumber="295">Polybius vernalis</emphasis>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF80FFEE611CF86DEAA9F812" author="Risso A." box="[302,447,1928,1954]" pageId="18" pageNumber="295" refId="ref16928" refString="Risso A. 1827 [in Risso, 1826 - 1827]. Histoire naturelle des Principales Productions de l'Europe meridionale et particulierement de celles des Environs de Nice et des Alpes maritimes. Vol. 5 Animaux Articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. F. - G. Levrault, Paris. https: // doi. org / 10.5962 / bhl. title. 58984" type="book" year="1827">Risso, 1827</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235).
|
||
<emphasis id="4DED2F79FF80FFEE645CF86DEF95F812" bold="true" box="[1134,1155,1928,1954]" pageId="18" pageNumber="295">C</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF80FFEE64A6F86DEE60F812" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[1172,1398,1928,1954]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="295" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF80FFEE64A6F86DEE60F812" box="[1172,1398,1928,1954]" italics="true" pageId="18" pageNumber="295">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF80FFEE608FF849EA08F876" box="[189,286,1964,1990]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀ from Málaga (CRUST_ICMAN/3765)
|
||
<emphasis id="4DED2F79FF80FFEE631BF84EE828F875" bold="true" box="[809,830,1963,1989]" pageId="18" pageNumber="295">D</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF80FFEE6360F84EEF34F875" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[850,1058,1963,1989]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="295" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF80FFEE6360F84EEF34F875" box="[850,1058,1963,1989]" italics="true" pageId="18" pageNumber="295">Polybius vernalis</emphasis>
|
||
</taxonomicName>
|
||
, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2BE6A3E3FF81FFEF608FF8FBEF06F858" ID-DOI="http://doi.org/10.5281/zenodo.10978045" ID-Zenodo-Dep="10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="19" pageNumber="296" startId="19.[189,232,1822,1848]" targetBox="[228,1359,265,1773]" targetPageId="19">
|
||
<paragraph id="7F26F36BFF81FFEF608FF8FBEF06F858" blockId="19.[189,1405,1822,2025]" pageId="19" pageNumber="296">
|
||
<emphasis id="4DED2F79FF81FFEF608FF8FBEA1DF888" bold="true" box="[189,267,1822,1848]" pageId="19" pageNumber="296">Fig. 7.</emphasis>
|
||
Frontal region in dorsal view, male.
|
||
<emphasis id="4DED2F79FF81FFEF6289F8FBE9C6F888" bold="true" box="[699,720,1822,1848]" pageId="19" pageNumber="296">A</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF81FFEF62EDF8FBEF6BF888" authority="(Leach, 1814)" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[735,1149,1822,1848]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="296" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF81FFEF62EDF8FBE8C4F888" box="[735,978,1822,1848]" italics="true" pageId="19" pageNumber="296">Polybius marmoreus</emphasis>
|
||
(Leach, 1814)
|
||
</taxonomicName>
|
||
from Belgium, Heist (RBINS, INV. 187218).
|
||
<emphasis id="4DED2F79FF81FFEF61E4F8A7EAFCF8EC" bold="true" box="[470,490,1858,1884]" pageId="19" pageNumber="296">B</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF81FFEF61CAF8A4E9C1F8EB" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[504,727,1857,1883]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="296" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF81FFEF61CAF8A4E9C1F8EB" box="[504,727,1857,1883]" italics="true" pageId="19" pageNumber="296">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF81FFEF62EFF8A7E82FF8EC" box="[733,825,1858,1884]" pageId="19" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223).
|
||
<emphasis id="4DED2F79FF81FFEF62FEF881E814F8CE" bold="true" box="[716,770,1892,1918]" pageId="19" pageNumber="296">C–E</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF81FFEF6321F880EF98F8CF" authority="(Risso, 1827)" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[787,1166,1893,1919]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="296" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF81FFEF6321F880E8F7F8CF" box="[787,993,1893,1919]" italics="true" pageId="19" pageNumber="296">Polybius vernalis</emphasis>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF81FFEF63C0F880EF93F8CF" author="Risso A." box="[1010,1157,1893,1919]" pageId="19" pageNumber="296" refId="ref16928" refString="Risso A. 1827 [in Risso, 1826 - 1827]. Histoire naturelle des Principales Productions de l'Europe meridionale et particulierement de celles des Environs de Nice et des Alpes maritimes. Vol. 5 Animaux Articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. F. - G. Levrault, Paris. https: // doi. org / 10.5962 / bhl. title. 58984" type="book" year="1827">Risso, 1827</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="4DED2F79FF81FFEF64AFF881EFA6F8CE" bold="true" box="[1181,1200,1892,1918]" pageId="19" pageNumber="296">C</emphasis>
|
||
. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235).
|
||
<emphasis id="4DED2F79FF81FFEF62D9F86DE9E9F812" bold="true" box="[747,767,1928,1954]" pageId="19" pageNumber="296">D</emphasis>
|
||
. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238).
|
||
<emphasis id="4DED2F79FF81FFEF6146F84EEA9EF875" bold="true" box="[372,392,1963,1989]" pageId="19" pageNumber="296">E</emphasis>
|
||
. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236).
|
||
<emphasis id="4DED2F79FF81FFEF644DF84EEF87F875" bold="true" box="[1151,1169,1963,1989]" pageId="19" pageNumber="296">F</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF81FFEF6491F84EEA9CF858" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="296" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF81FFEF6491F84EEE61F875" box="[1187,1399,1963,1989]" italics="true" pageId="19" pageNumber="296">Polybius holsatus</emphasis>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF81FFEF60F7F82BEA97F858" author="Fabricius J. C." box="[197,385,1998,2024]" pageId="19" pageNumber="296" refId="ref14488" refString="Fabricius J. C. 1798. Entomologia Systematica emendata et aucta, secundum classes, ordines, genera, species adjectis synonimis locis observationibus descriptionibus. Supplementum. Impensis Christ. Gottl. Proft, Copenhagen [Hafniae]. https: // doi. org / 10.5962 / bhl. title. 125869" type="book" year="1798">Fabricius, 1798</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂ from Belgium, Duinbergen (RBINS, INV. 187221).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2BE6A3E3FF86FFE8608FF8A4EAAEF858" ID-DOI="http://doi.org/10.5281/zenodo.10978047" ID-Zenodo-Dep="10978047" httpUri="https://zenodo.org/record/10978047/files/figure.png" pageId="20" pageNumber="297" startId="20.[189,232,1857,1883]" targetBox="[218,1370,265,1801]" targetPageId="20">
|
||
<paragraph id="7F26F36BFF86FFE8608FF8A4EAAEF858" blockId="20.[186,1399,1857,2025]" pageId="20" pageNumber="297">
|
||
<emphasis id="4DED2F79FF86FFE8608FF8A4EA1CF8EB" bold="true" box="[189,266,1857,1883]" pageId="20" pageNumber="297">Fig. 8.</emphasis>
|
||
Left gonopod 1 of male.
|
||
<emphasis id="4DED2F79FF86FFE86202F8A4E971F8EB" bold="true" box="[560,615,1857,1883]" pageId="20" pageNumber="297">A–B</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF86FFE86244F8A4EF07F8EC" authority="(Leach, 1814)" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[630,1041,1857,1884]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="297" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF86FFE86244F8A4E87EF8EB" box="[630,872,1857,1883]" italics="true" pageId="20" pageNumber="297">Polybius marmoreus</emphasis>
|
||
(Leach, 1814)
|
||
</taxonomicName>
|
||
from Belgium, Heist (RBINS, INV. 187218).
|
||
<emphasis id="4DED2F79FF86FFE86143F881EA90F8CE" bold="true" box="[369,390,1892,1918]" pageId="20" pageNumber="297">A</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="4DED2F79FF86FFE86272F880E944F8CF" bold="true" box="[576,594,1893,1919]" pageId="20" pageNumber="297">B</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="4DED2F79FF86FFE86331F881E82AF8CE" bold="true" box="[771,828,1892,1918]" pageId="20" pageNumber="297">C–D</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF86FFE8637FF880EF26F8CF" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[845,1072,1893,1919]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="297" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF86FFE8637FF880EF26F8CF" box="[845,1072,1893,1919]" italics="true" pageId="20" pageNumber="297">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF86FFE86408F880EF8FF8CF" box="[1082,1177,1893,1919]" pageId="20" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223).
|
||
<emphasis id="4DED2F79FF86FFE86409F86DEF46F812" bold="true" box="[1083,1104,1928,1954]" pageId="20" pageNumber="297">C</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="4DED2F79FF86FFE86536F86DEE0FF812" bold="true" box="[1284,1305,1928,1954]" pageId="20" pageNumber="297">D</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="4DED2F79FF86FFE86133F84EEA22F875" bold="true" box="[257,308,1963,1989]" pageId="20" pageNumber="297">E–F</emphasis>
|
||
.
|
||
<taxonomicName id="B89988E8FF86FFE86173F84EE9BAF875" authority="(Risso, 1827)" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[321,684,1963,1989]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="297" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF86FFE86173F84EE91AF875" box="[321,524,1963,1989]" italics="true" pageId="20" pageNumber="297">Polybius vernalis</emphasis>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF86FFE8622BF84EE9B2F875" author="Risso A." box="[537,676,1963,1989]" pageId="20" pageNumber="297" refId="ref16928" refString="Risso A. 1827 [in Risso, 1826 - 1827]. Histoire naturelle des Principales Productions de l'Europe meridionale et particulierement de celles des Environs de Nice et des Alpes maritimes. Vol. 5 Animaux Articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. F. - G. Levrault, Paris. https: // doi. org / 10.5962 / bhl. title. 58984" type="book" year="1827">Risso, 1827</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from Tunisia, Bay of Tunis (RBINS, INV. 187235).
|
||
<emphasis id="4DED2F79FF86FFE86532F84EEE02F875" bold="true" box="[1280,1300,1963,1989]" pageId="20" pageNumber="297">E</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="4DED2F79FF86FFE86131F82BEA03F858" bold="true" box="[259,277,1998,2024]" pageId="20" pageNumber="297">F</emphasis>
|
||
. Dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3783A0E0FF87FFE9608FFEEDEFB2FD01" pageId="21" pageNumber="298" type="distribution">
|
||
<paragraph id="7F26F36BFF87FFE9608FFEEDEA40FE92" blockId="21.[189,342,264,290]" box="[189,342,264,290]" pageId="21" pageNumber="298">
|
||
<heading id="246E4407FF87FFE9608FFEEDEA40FE92" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="21" pageNumber="298" reason="3">
|
||
<emphasis id="4DED2F79FF87FFE9608FFEEDEA40FE92" bold="true" box="[189,342,264,290]" pageId="21" pageNumber="298">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF87FFE9608FFEDDEFB2FD01" blockId="21.[189,1399,311,690]" pageId="21" pageNumber="298">
|
||
Its exact geographical distribution is not well known, since in the papers (taxonomic, genetic and ecological) the identification of the species of
|
||
<taxonomicName id="B89988E8FF87FFE962DFFEBFE842FEC4" authorityName="Leach" authorityYear="1820" box="[749,852,346,372]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DED2F79FF87FFE962DFFEBFE842FEC4" box="[749,852,346,372]" italics="true" pageId="21" pageNumber="298">Polybius</emphasis>
|
||
</taxonomicName>
|
||
is not at all clear (there is a lot of confusion between
|
||
<taxonomicName id="B89988E8FF87FFE96118FE9BEABAFE27" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[298,428,381,407]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE96118FE9BEABAFE27" box="[298,428,381,407]" italics="true" pageId="21" pageNumber="298">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B89988E8FF87FFE9618CFE9BE975FE27" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[446,611,382,407]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF87FFE9618CFE9BE975FE27" box="[446,611,382,407]" italics="true" pageId="21" pageNumber="298">P. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B89988E8FF87FFE96247FE9BE9E2FE27" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[629,756,381,407]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF87FFE96247FE9BE9E2FE27" box="[629,756,381,407]" italics="true" pageId="21" pageNumber="298">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B89988E8FF87FFE96306FE9BE8D0FE27" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[820,966,381,407]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF87FFE96306FE9BE8D0FE27" box="[820,966,381,407]" italics="true" pageId="21" pageNumber="298">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF87FFE963E3FE9BEF25FE28" box="[977,1075,382,408]" pageId="21" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). According to our data, it is present in the Atlantic Ocean: British Isles (
|
||
<bibRefCitation id="1B088E9AFF87FFE962C5FE44E8FAFE0B" author="Ingle R. W. & Clark P. P." box="[759,1004,417,443]" pageId="21" pageNumber="298" pagination="223 - 229" refId="ref15194" refString="Ingle R. W. & Clark P. P. 1998. A swimming crab new to the British fauna, Liocarcinus vernalis (Crustacea: Brachyura: Portunidae). Journal of the Marine Biological Association of the United Kingdom 78: 223 - 229. https: // doi. org / 10.1017 / S 0025315400040042 ICZN 1999. InternationalCodeofZoologicalNomenclature. 4 thedition. TheInternationalTrustforZoological Nomenclature, London. Available from https: // www. iczn. org / the-code / the-code-online / [accessed 22 Mar. 2024]." type="journal article" year="1998">Ingle & Clark 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1B088E9AFF87FFE963CEFE44EE10FE0B" author="McCarthy A. M. & McGrath D. & Allen B. M." box="[1020,1286,416,443]" pageId="21" pageNumber="298" pagination="20 - 26" refId="ref15921" refString="McCarthy A. M., McGrath D. & Allen B. M. 2005. The grey swimming crab Liocarcinus vernalis (Risso, 1827) in western Irish coastal waters (Decapoda: Brachyura, Portunidae). Irish Naturalists' Journal 28 (1): 20 - 26." type="journal article" year="2005">
|
||
McCarthy
|
||
<emphasis id="4DED2F79FF87FFE9644DFE44EFABFE0B" box="[1151,1213,417,443]" italics="true" pageId="21" pageNumber="298">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1B088E9AFF87FFE96527FE44EBEBFE6E" author="Ashelby C. W." pageId="21" pageNumber="298" pagination="1333 - 1345" refId="ref13809" refString="Ashelby C. W. 2006. Contribution to the biogeographical knowledge of some decapod crustaceans in the British Isles. Crustaceana 79 (11): 1333 - 1345. https: // doi. org / 10.1163 / 156854006779277259" type="journal article" year="2006">Ashelby 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1B088E9AFF87FFE96138FE21E93FFE6E" author="D'Udekem d'Acoz C." box="[266,553,452,478]" pageId="21" pageNumber="298" pagination="78 - 79" refId="ref14272" refString="D'Udekem d'Acoz C. 2011. Occurrence of the grey swimming crab Polybius vernalis (Risso, 1827) on Rathlin Island, Northern Ireland: northernmost record so far (Crustacea, Decapoda, Brachyura). De Strandvlo 31 (3 - 4): 78 - 79." type="journal article" year="2011">d’Udekem d’Acoz 2011</bibRefCitation>
|
||
),
|
||
<collectingCountry id="078EB3FBFF87FFE9620FFE21E9E1FE6E" box="[573,759,452,478]" name="Netherlands" pageId="21" pageNumber="298">the Netherlands</collectingCountry>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF87FFE96334FE21E8B4FE6E" author="Adema J. P. H. M." box="[774,930,452,478]" pageId="21" pageNumber="298" pagination="110 - 115" refId="ref13758" refString="Adema J. P. H. M. 1991. Een aanvulling op: " De krabben van Nederland en Belgie ": De grijze zwemkrab, Liocarcinus vernalis (Risso, 1816) in de Noordzee gevonden. Het Zeepaard 51 (5): 110 - 115." type="journal article" year="1991">Adema 1991</bibRefCitation>
|
||
),
|
||
<collectingCountry id="078EB3FBFF87FFE9638AFE21EF0BFE6E" box="[952,1053,452,478]" name="Belgium" pageId="21" pageNumber="298">Belgium</collectingCountry>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF87FFE9641EFE21EBEDFDB1" author="D'Udekem d'Acoz C. & Rappe G." pageId="21" pageNumber="298" pagination="95 - 99" refId="ref14323" refString="D'Udekem d'Acoz C. & Rappe G. 1991. Presence et abondance de Liocarcinus vernalis (Risso, 1816) dans la baie sud de la mer du Nord (Decapoda, Brachyura, Portunidae). Crustaceana 61 (1): 95 - 99. Available from http: // www. jstor. org / stable / 20104677 [accessed 22 Mar. 2024]." type="journal article" year="1991">d’Udekem d’Acoz & Rappé 1991</bibRefCitation>
|
||
), Normandy (
|
||
<bibRefCitation id="1B088E9AFF87FFE961AFFE02E922FDB1" author="Livory A." box="[413,564,487,513]" pageId="21" pageNumber="298" pagination="18 - 64" refId="ref15679" refString="Livory A. 1998. Crabes de la Manche: le point des connaissances. L'Argiope 18 - 19: 18 - 64." type="journal article" year="1998">Livory 1998</bibRefCitation>
|
||
), Bretagne (
|
||
<bibRefCitation id="1B088E9AFF87FFE962F0FE02E8F5FDB1" author="D'Udekem d'Acoz C." box="[706,995,487,513]" pageId="21" pageNumber="298" pagination="84 - 100" refId="ref14115" refString="D'Udekem d'Acoz C. 1986. Etude d'une collection de crustaces decapodes de Bretagne. De Strandvlo [dec. 1985]: 11 (4): 84 - 100." type="journal article" year="1986">d’Udekem d’Acoz 1986</bibRefCitation>
|
||
: fig. 10), Bay of Biscay (González Gurriarán & Ménez 1986),
|
||
<collectingCountry id="078EB3FBFF87FFE961CCFDEFE977FD94" box="[510,609,522,548]" name="Portugal" pageId="21" pageNumber="298">Portugal</collectingCountry>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF87FFE96243FDEFE816FD94" author="Nobre A." box="[625,768,522,548]" pageId="21" pageNumber="298" refId="ref16236" refString="Nobre A. 1931. Crustaceos Decapodes e Stomatopodes Marinhos de Portugal. Imprensa Portuguesa, Porto." type="book" year="1931">Nobre 1931</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1B088E9AFF87FFE9633EFDEFE85CFD94" author="Nobre A." box="[780,842,522,548]" pageId="21" pageNumber="298" refId="ref16254" refString="Nobre A. 1936. Crustaceos Decapodes e Stomatopodes Marinhos de Portugal. 2 a Edicao. Fauna Marinha de Portugal 4. Porto." type="book" year="1936">1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1B088E9AFF87FFE96364FDEEE8F3FD95" author="Neves A. M." box="[854,997,522,549]" pageId="21" pageNumber="298" pagination="1 - 48" refId="ref16160" refString="Neves A. M. 1975. Sobre una coleccao de crustaceos decapodes da Bahia de Setubal (Portugal). Estudos Fauna Portuguesa 5: 1 - 48." type="journal article" year="1975">Neves 1975</bibRefCitation>
|
||
), Gulf of Cádiz (
|
||
<bibRefCitation id="1B088E9AFF87FFE96498FDEFEA56FDF8" author="Lopez de la Rosa I. & Garcia Raso J. E. & Rodriguez A." pageId="21" pageNumber="298" pagination="85 - 95" refId="ref15703" refString="Lopez de la Rosa I., Garcia Raso J. E. & Rodriguez A. 2002. Evolution of a decapod community (Crustacea) of shallow soft bottoms with seaweeds from southern Europe. Journal of the Marine Biological Association of the United Kingdom 82 (1): 85 - 95. https: // doi. org / 10.1017 / S 0025315402005209." type="journal article" year="2002">
|
||
López de la Rosa
|
||
<emphasis id="4DED2F79FF87FFE9608FFDCBEBEFFDF7" box="[189,249,557,583]" italics="true" pageId="21" pageNumber="298">et al.</emphasis>
|
||
2002
|
||
</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="B89988E8FF87FFE96142FDCBE977FDF7" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[368,609,557,583]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF87FFE96142FDCBE977FDF7" box="[368,609,557,583]" italics="true" pageId="21" pageNumber="298">Liocarcinus vernalis</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<collectingCountry id="078EB3FBFF87FFE96249FDCBE9FCFDF8" box="[635,746,558,584]" name="Morocco" pageId="21" pageNumber="298">Morocco</collectingCountry>
|
||
: Bay of
|
||
<collectingCountry id="078EB3FBFF87FFE9637CFDC8E889FDF7" box="[846,927,557,583]" name="Morocco" pageId="21" pageNumber="298">Agadir</collectingCountry>
|
||
(Moukrim
|
||
<emphasis id="4DED2F79FF87FFE9641EFDCBEF7EFDF7" box="[1068,1128,557,583]" italics="true" pageId="21" pageNumber="298">et al.</emphasis>
|
||
2010; fig. 2f), “
|
||
<collectingCountry id="078EB3FBFF87FFE96515FDCBEE60FDF8" box="[1319,1398,558,584]" name="Morocco" pageId="21" pageNumber="298">Maroc</collectingCountry>
|
||
atlantique” (materiel examine by d’Udekem d’Acoz),
|
||
<collectingCountry id="078EB3FBFF87FFE96350FDB4EF36FDDB" box="[866,1056,593,619]" name="Western Sahara" pageId="21" pageNumber="298">Western Sahara</collectingCountry>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF87FFE96404FDB4EFC0FDDB" author="Monod T." box="[1078,1238,592,619]" pageId="21" pageNumber="298" pagination="1 - 674" refId="ref16073" refString="Monod T. 1956. Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire (Mem. IFAN) 45: 1 - 674." type="journal article" year="1956">Monod 1956</bibRefCitation>
|
||
as
|
||
<taxonomicName id="B89988E8FF87FFE96539FDB4EA3FFD3E" authorityName="H. Lucas" authorityYear="1846" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="barbarus">
|
||
<emphasis id="4DED2F79FF87FFE96539FDB4EA3FFD3E" italics="true" pageId="21" pageNumber="298">Portunus barbarus</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<collectingCountry id="078EB3FBFF87FFE96176FD91EAD3FD3E" box="[324,453,628,654]" name="Mauritania" pageId="21" pageNumber="298">Mauritania</collectingCountry>
|
||
and Canary Islands (
|
||
<bibRefCitation id="1B088E9AFF87FFE962F4FD91E87AFD3E" author="Fransen C. H. J. M." box="[710,876,628,654]" pageId="21" pageNumber="298" refId="ref14577" refString="Fransen C. H. J. M. 1991. Crustacea of the CANCAP and MAURITANIA Expeditions. Nationaal Natuurhistorisch Museum, Leiden." type="book" year="1991">Fransen 1991</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="B89988E8FF87FFE963ACFD90EF08FD3E" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[926,1054,628,654]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF87FFE963ACFD90EF08FD3E" box="[926,1054,628,654]" italics="true" pageId="21" pageNumber="298">L. vernalis</emphasis>
|
||
</taxonomicName>
|
||
?) and in the Mediterranean Sea: Alboran Sea, and Málaga (Marbella and Fuengirola) (
|
||
<bibRefCitation id="1B088E9AFF87FFE96359FD72EF56FD01" author="Garcia Raso J. E." box="[875,1088,663,690]" pageId="21" pageNumber="298" pagination="105 - 113" refId="ref14602" refString="Garcia Raso J. E. 1984. Brachyura of the coast of Southern Spain. Spixiana 7 (2): 105 - 113." type="journal article" year="1984">
|
||
García Raso
|
||
<quantity id="B8615E8EFF87FFE96437FD72EF56FD01" box="[1029,1088,663,689]" metricMagnitude="1" metricUnit="m" metricValue="5.03936" pageId="21" pageNumber="298" unit="in" value="1984.0">1984</quantity>
|
||
</bibRefCitation>
|
||
in part).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3783A0E0FF87FFE9608FFD3DE8BBFBA7" pageId="21" pageNumber="298" type="biology_ecology">
|
||
<paragraph id="7F26F36BFF87FFE9608FFD3DEA0BFD42" blockId="21.[189,285,728,754]" box="[189,285,728,754]" pageId="21" pageNumber="298">
|
||
<heading id="246E4407FF87FFE9608FFD3DEA0BFD42" bold="true" box="[189,285,728,754]" fontSize="11" level="3" pageId="21" pageNumber="298" reason="3">
|
||
<emphasis id="4DED2F79FF87FFE9608FFD3DEA0BFD42" bold="true" box="[189,285,728,754]" pageId="21" pageNumber="298">Habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF87FFE9608FFCEDE8BBFBA7" blockId="21.[189,1399,775,1047]" pageId="21" pageNumber="298">
|
||
It lives between
|
||
<quantity id="B8615E8EFF87FFE961BAFCE2E915FC92" box="[392,515,775,802]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="21" pageNumber="298" unit="m" value="15.0" valueMax="25.0" valueMin="5.0">5 to 25 m</quantity>
|
||
, on bottoms of fine sand (range of median 0.17–0.32), with a low pelite content near the shore which increases to ca 10% at
|
||
<quantity id="B8615E8EFF87FFE9630EFCCFE86CFCF5" box="[828,890,810,837]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="298" unit="m" value="20.0">20 m</quantity>
|
||
off
|
||
<collectingCountry id="078EB3FBFF87FFE96382FCCEEF0BFCF5" box="[944,1053,811,837]" name="Morocco" pageId="21" pageNumber="298">Morocco</collectingCountry>
|
||
, although at Oued Souss the pelite content was higher at a shallow depth (30% at
|
||
<quantity id="B8615E8EFF87FFE96305FCABE86EFCD8" box="[823,888,846,872]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="298" unit="m" value="10.0">10 m</quantity>
|
||
). In Spain, it inhabits the Gulf of Cádiz on sandy bottoms with
|
||
<taxonomicName id="B89988E8FF87FFE96198FC94E83AFC3B" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[426,812,880,907]" class="Magnoliopsida" family="Ericaceae" genus="Callista" kingdom="Plantae" order="Ericales" pageId="21" pageNumber="298" phylum="Tracheophyta" rank="species" species="chione">
|
||
<emphasis id="4DED2F79FF87FFE96198FC94E94AFC3B" box="[426,604,881,907]" italics="true" pageId="21" pageNumber="298">Callista chione</emphasis>
|
||
(
|
||
<bibRefCitation id="1B088E9AFF87FFE9625EFC94E832FC3B" author="Linnaeus C." box="[620,804,880,907]" pageId="21" pageNumber="298" refId="ref15623" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Laurentii Salvii, Stockholm [Holmiae]. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="B8615E8EFF87FFE9630EFC95E8BBFC3B" box="[828,941,880,907]" metricMagnitude="0" metricUnit="m" metricValue="5.25" metricValueMax="10.0" metricValueMin="0.5" pageId="21" pageNumber="298" unit="m" value="5.25" valueMax="10.0" valueMin="0.5">0.5–10 m</quantity>
|
||
) and on fine muddy sand bottoms with
|
||
<taxonomicName id="B89988E8FF87FFE9608FFC71E9C3FC1E" authority="(Forsskal) J. V. Lamouroux" authorityName="J. V. Lamouroux" baseAuthorityName="Forsskal" box="[189,725,915,942]" class="Ulvophyceae" family="Caulerpaceae" genus="Caulerpa" kingdom="Plantae" order="Bryopsidales" pageId="21" pageNumber="298" phylum="Chlorophyta" rank="species" species="prolifera">
|
||
<emphasis id="4DED2F79FF87FFE9608FFC71EA8DFC1E" box="[189,411,916,942]" italics="true" pageId="21" pageNumber="298">Caulerpa prolifera</emphasis>
|
||
(Forsskål) J.V.Lamouroux
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="B8615E8EFF87FFE962D7FC71E849FC1E" box="[741,863,916,942]" metricMagnitude="0" metricUnit="m" metricValue="4.949999999999999" metricValueMax="6.6" metricValueMin="3.3" pageId="21" pageNumber="298" unit="m" value="4.949999999999999" valueMax="6.6" valueMin="3.3">3.3–6.6 m</quantity>
|
||
) and at Málaga it lives on sandy bottoms and has been caught during the harvesting of “coquina” (
|
||
<taxonomicName id="B89988E8FF87FFE96373FC5DEFDAFC62" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[833,1228,951,978]" class="Liliopsida" family="Marantaceae" genus="Donax" kingdom="Plantae" order="Zingiberales" pageId="21" pageNumber="298" phylum="Tracheophyta" rank="species" species="trunculus">
|
||
<emphasis id="4DED2F79FF87FFE96373FC5DEF1FFC61" box="[833,1033,951,977]" italics="true" pageId="21" pageNumber="298">Donax trunculus</emphasis>
|
||
<bibRefCitation id="1B088E9AFF87FFE96421FC52EFDAFC62" author="Linnaeus C." box="[1043,1228,951,978]" pageId="21" pageNumber="298" refId="ref15623" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Laurentii Salvii, Stockholm [Holmiae]. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
</taxonomicName>
|
||
) at
|
||
<quantity id="B8615E8EFF87FFE964CDFC52EE59FC62" box="[1279,1359,951,978]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="21" pageNumber="298" unit="m" value="4.0" valueMax="6.0" valueMin="2.0">
|
||
2–
|
||
<specimenCount id="699F38E2FF87FFE9652FFC52EE59FC62" box="[1309,1359,951,978]" count="6" pageId="21" pageNumber="298" type="generic">6 m</specimenCount>
|
||
</quantity>
|
||
. In
|
||
<collectingCountry id="078EB3FBFF87FFE9608FFC3FEA33FC44" box="[189,293,986,1012]" name="Belgium" pageId="21" pageNumber="298">Belgium</collectingCountry>
|
||
, it lives on fine sand just below the tide marks and in shallow waters. So far, it has not been recorded in enclosed bays and environments of reduced salinity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3783A0E0FF87FFEB608FFBDAE920F8B3" lastPageId="23" lastPageNumber="300" pageId="21" pageNumber="298" type="discussion">
|
||
<paragraph id="7F26F36BFF87FFE9608FFBDAEA39FBE9" blockId="21.[189,303,1087,1113]" box="[189,303,1087,1113]" pageId="21" pageNumber="298">
|
||
<heading id="246E4407FF87FFE9608FFBDAEA39FBE9" bold="true" box="[189,303,1087,1113]" fontSize="11" level="3" pageId="21" pageNumber="298" reason="3">
|
||
<emphasis id="4DED2F79FF87FFE9608FFBDAEA39FBE9" bold="true" box="[189,303,1087,1113]" pageId="21" pageNumber="298">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF87FFE9608FFB88E901F8B1" blockId="21.[188,1399,1133,1794]" pageId="21" pageNumber="298">
|
||
The new species,
|
||
<taxonomicName id="B89988E8FF87FFE961A1FB88E963FB37" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[403,629,1133,1159]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF87FFE961A1FB88E963FB37" box="[403,629,1133,1159]" italics="true" pageId="21" pageNumber="298">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF87FFE9624CFB8BE9CAFB38" box="[638,732,1134,1160]" pageId="21" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, belongs to the group formed by
|
||
<taxonomicName id="B89988E8FF87FFE96459FB8BEFFFFB37" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1131,1257,1133,1159]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE96459FB8BEFFFFB37" box="[1131,1257,1133,1159]" italics="true" pageId="21" pageNumber="298">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B89988E8FF87FFE964C8FB8BEE61FB37" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1274,1399,1133,1159]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF87FFE964C8FB8BEE61FB37" box="[1274,1399,1133,1159]" italics="true" pageId="21" pageNumber="298">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B89988E8FF87FFE960C3FB74EA82FB1A" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[241,404,1169,1194]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF87FFE960C3FB74EA82FB1A" box="[241,404,1169,1194]" italics="true" pageId="21" pageNumber="298">P. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
, which are morphologically and genetically related (
|
||
<figureCitation id="E7A2EFEEFF87FFE9643EFB75EF6EFB1B" box="[1036,1144,1168,1195]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1857,1883]" captionStartId-1="8.[189,232,1821,1847]" captionTargetBox-0="[348,1048,260,1786]" captionTargetBox-1="[320,1087,255,1734]" captionTargetId-0="graphics-40@7.[394,654,294,1724]" captionTargetId-1="graphics-32@8.[410,709,297,1664]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 1. Neighbor-Joning (NJ) phylogenetic tree based on mitochondrial 16S rRNA sequences of species of Polybius Leach, 1820 and Liocarcinus Stimpson, 1871, using 2000 Bootstrap replicates. Number on nodes represent NJ Bootstrap values, only values>70 are included. Carcinus aestuarii Nardo, 1847 and C. maenas (Linnaeus, 1758) are used as outgroup. GenBank accession codes are indicated for each species, except those with codes XX## which sequences are listed in Supp. file 1." captionText-1="Fig. 2. Maximum likelihood (ML) phylogenetic tree based on mitochondrial cytochrome c oxidase subunit I sequences of species of Polybius Leach, 1820 and Liocarcinus Stimpson, 1871, using 2000 Bootstrap replicates. Number on nodes represent ML Bootstrap values, only values>70 are included. Carcinus aestuarii Nardo, 1847 and C. maenas (Linnaeus, 1758) are used as outgroup. GenBank accession codes are indicated for each species, except those with codes XX## which sequences are listed in Supp. file 1." figureDoi-0="http://doi.org/10.5281/zenodo.10978025" figureDoi-1="http://doi.org/10.5281/zenodo.10978029" httpUri-0="https://zenodo.org/record/10978025/files/figure.png" httpUri-1="https://zenodo.org/record/10978029/files/figure.png" pageId="21" pageNumber="298">Figs 1–2</figureCitation>
|
||
), but with significant differences. They have often been misidentified, because they share a combination of morphological characters, including some variability.
|
||
<bibRefCitation id="1B088E9AFF87FFE962A2FB32E82AFB41" author="Palmer R." box="[656,828,1239,1265]" pageId="21" pageNumber="298" pagination="877 - 908" refId="ref16457" refString="Palmer R. 1927. A revision of the genus " Portunus " (A. Milne Edwards, Bell, etc). Journal of the Marine Biological Association of the United Kingdom 14: 877 - 908. https: // doi. org / 10.1017 / S 0025315400051134" type="journal article" year="1927">Palmer (1927)</bibRefCitation>
|
||
analyzed the morphology of
|
||
<taxonomicName id="B89988E8FF87FFE96496FB32EE61FB41" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1188,1399,1239,1265]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE96496FB32EE61FB41" box="[1188,1399,1239,1265]" italics="true" pageId="21" pageNumber="298">Polybius holsatus</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="B89988E8FF87FFE960D9FB1EEAD2FAA4" authorityName="Fabricius" authorityYear="1798" box="[235,452,1274,1300]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE960D9FB1EEAD2FAA4" box="[235,452,1274,1300]" italics="true" pageId="21" pageNumber="298">Portunus holsatus</emphasis>
|
||
</taxonomicName>
|
||
) and compared it with that of
|
||
<taxonomicName id="B89988E8FF87FFE96376FB1EE8FCFAA4" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[836,1002,1275,1300]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF87FFE96376FB1EE8FCFAA4" box="[836,1002,1275,1300]" italics="true" pageId="21" pageNumber="298">P. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
, showing a variability, which in some cases may correspond to characters of the new species. In 1986, d’Udekem d’Acoz raised the possibility that it was a new species, naming it
|
||
<taxonomicName id="B89988E8FF87FFE962C1FAA4EF01FAEA" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[755,1047,1344,1370]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE962C1FAA4E868FAEA" box="[755,894,1345,1370]" italics="true" pageId="21" pageNumber="298">Liocarcinus</emphasis>
|
||
aff.
|
||
<emphasis id="4DED2F79FF87FFE96387FAA5EF01FAEA" box="[949,1047,1344,1370]" italics="true" pageId="21" pageNumber="298">holsatus</emphasis>
|
||
</taxonomicName>
|
||
, although he later considered it to be an Atlantic form of
|
||
<taxonomicName id="B89988E8FF87FFE96220FA81E9F4FACE" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[530,738,1380,1406]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF87FFE96220FA81E9F4FACE" box="[530,738,1380,1406]" italics="true" pageId="21" pageNumber="298">Polybius vernalis</emphasis>
|
||
</taxonomicName>
|
||
, with small population differences (
|
||
<bibRefCitation id="1B088E9AFF87FFE964AAFA81EBEAFA11" author="D'Udekem d'Acoz C." pageId="21" pageNumber="298" pagination="166 - 205" refId="ref14147" refString="D'Udekem d'Acoz C. 1989. Seconde note sur les crustaces decapodes de la Bretagne. De Strandvlo 8 (4): 166 - 205." type="journal article" year="1989">d’Udekem d’Acoz 1989</bibRefCitation>
|
||
). These considerations were not mentioned in later reports (
|
||
<bibRefCitation id="1B088E9AFF87FFE963E7FA62EFEAFA11" author="D'Udekem d'Acoz C." box="[981,1276,1415,1441]" pageId="21" pageNumber="298" pagination="40" refId="ref14220" refString="D'Udekem d'Acoz C. 1999. Inventaire et distribution des crustaces decapodes de l'Atlantique nordoriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Patrimoines naturels (M. N. H. N. / S. P. N.) 40." type="journal article" year="1999">d’Udekem d’Acoz 1999</bibRefCitation>
|
||
), with the only exception of the existence of a glabrous versus a pubescent carapace. In the same broad period,
|
||
<bibRefCitation id="1B088E9AFF87FFE9608DFA28E971FA57" author="D'Udekem d'Acoz C. & Rappe G." box="[191,615,1485,1511]" pageId="21" pageNumber="298" pagination="95 - 99" refId="ref14323" refString="D'Udekem d'Acoz C. & Rappe G. 1991. Presence et abondance de Liocarcinus vernalis (Risso, 1816) dans la baie sud de la mer du Nord (Decapoda, Brachyura, Portunidae). Crustaceana 61 (1): 95 - 99. Available from http: // www. jstor. org / stable / 20104677 [accessed 22 Mar. 2024]." type="journal article" year="1991">d’Udekem d’Acoz & Rappé (1991)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="1B088E9AFF87FFE96296FA28E8B3FA57" author="Ingle R. W. & Clark P. P." box="[676,933,1485,1511]" pageId="21" pageNumber="298" pagination="223 - 229" refId="ref15194" refString="Ingle R. W. & Clark P. P. 1998. A swimming crab new to the British fauna, Liocarcinus vernalis (Crustacea: Brachyura: Portunidae). Journal of the Marine Biological Association of the United Kingdom 78: 223 - 229. https: // doi. org / 10.1017 / S 0025315400040042 ICZN 1999. InternationalCodeofZoologicalNomenclature. 4 thedition. TheInternationalTrustforZoological Nomenclature, London. Available from https: // www. iczn. org / the-code / the-code-online / [accessed 22 Mar. 2024]." type="journal article" year="1998">Ingle & Clark (1998)</bibRefCitation>
|
||
carried out two very useful studies on the morphological differences between the three species considered valid in this group:
|
||
<taxonomicName id="B89988E8FF87FFE964DFFA14EE79F9BA" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1261,1391,1520,1546]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE964DFFA14EE79F9BA" box="[1261,1391,1520,1546]" italics="true" pageId="21" pageNumber="298">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B89988E8FF87FFE9608FF9F0EA2EF99E" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[189,312,1556,1582]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF87FFE9608FF9F0EA2EF99E" box="[189,312,1556,1582]" italics="true" pageId="21" pageNumber="298">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
[in using material of
|
||
<taxonomicName id="B89988E8FF87FFE96203F9F0E9A9F99E" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[561,703,1556,1582]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF87FFE96203F9F0E9A9F99E" box="[561,703,1556,1582]" italics="true" pageId="21" pageNumber="298">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF87FFE962F4F9F1E835F99E" box="[710,803,1556,1582]" pageId="21" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
|
||
] and
|
||
<taxonomicName id="B89988E8FF87FFE96352F9F0EF17F99E" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[864,1025,1557,1582]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF87FFE96352F9F0EF17F99E" box="[864,1025,1557,1582]" italics="true" pageId="21" pageNumber="298">P. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
(all as
|
||
<taxonomicName id="B89988E8FF87FFE96466F9F0EFC9F99E" authorityName="Stimpson" authorityYear="1871" box="[1108,1247,1557,1582]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DED2F79FF87FFE96466F9F0EFC9F99E" box="[1108,1247,1557,1582]" italics="true" pageId="21" pageNumber="298">Liocarcinus</emphasis>
|
||
</taxonomicName>
|
||
). This group was genetically validated by
|
||
<bibRefCitation id="1B088E9AFF87FFE9622EF9D2E851F9E1" author="Passamonti M. & Mantovani, B. & Scali V. & Froglia C." box="[540,839,1591,1617]" pageId="21" pageNumber="298" pagination="157 - 164" refId="ref16508" refString="Passamonti M., Mantovani, B., Scali V. & Froglia C. 1997. Genetic differentiation of European species of Liocarcinus (Crustacea, Portunidae): a gene-enzyme study. Zoologischer Anzeiger 235: 157 - 164." type="journal article" year="1997">
|
||
Passamonti
|
||
<emphasis id="4DED2F79FF87FFE9629CF9DDE9FDF9E1" box="[686,747,1591,1617]" italics="true" pageId="21" pageNumber="298">et al.</emphasis>
|
||
(1997)
|
||
</bibRefCitation>
|
||
, who also showed that there are some genetic differences among the populations of
|
||
<taxonomicName id="B89988E8FF87FFE962B4F9BEE80CF9C4" box="[646,794,1626,1652]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="depurator">
|
||
<emphasis id="4DED2F79FF87FFE962B4F9BEE80CF9C4" box="[646,794,1626,1652]" italics="true" pageId="21" pageNumber="298">P. depurator</emphasis>
|
||
</taxonomicName>
|
||
and proposed the possible creation of subspecies.
|
||
<bibRefCitation id="1B088E9AFF87FFE9608FF998EAB4F927" author="Ojeda Martin V. & Serra Elias B. & Lagares C. & Selles Altes M. & Marco-Herrero E. & Garcia-Rodriguez E. & Farre M. & Arenas C. & Abello P. & Mestres F." box="[189,418,1661,1687]" pageId="21" pageNumber="298" pagination="9797" refId="ref16315" refString="Ojeda Martin V., Serra Elias B., Lagares C., Selles Altes M., Marco-Herrero E., Garcia-Rodriguez E., Farre M., Arenas C., Abello P. & Mestres F. 2022. Interannual fluctuations in connectivity among crab populations (Liocarcinus depurator) along the Atlantic-Mediterranean transition. Scientific Reports 12: 9797. https: // doi. org / 10.1038 / s 41598 - 022 - 13941 - 4" type="journal article" year="2022">
|
||
Ojeda
|
||
<emphasis id="4DED2F79FF87FFE9613FF99BEA5CF927" box="[269,330,1661,1687]" italics="true" pageId="21" pageNumber="298">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
also cited differences between Atlantic and Mediterranean populations. Moukrim
|
||
<emphasis id="4DED2F79FF87FFE9608DF944EBEAF90A" box="[191,252,1696,1722]" italics="true" pageId="21" pageNumber="298">et al.</emphasis>
|
||
(2010) captured specimens of the new species on bottoms of fine sand between 5 and
|
||
<quantity id="B8615E8EFF87FFE96535F945EE51F90B" box="[1287,1351,1696,1723]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="298" unit="m" value="25.0">25 m</quantity>
|
||
, off
|
||
<collectingCountry id="078EB3FBFF87FFE9608FF921EA18F96E" box="[189,270,1732,1758]" name="Morocco" pageId="21" pageNumber="298">Agadir</collectingCountry>
|
||
(
|
||
<collectingCountry id="078EB3FBFF87FFE96112F921EA86F96E" box="[288,400,1732,1758]" name="Morocco" pageId="21" pageNumber="298">Morocco</collectingCountry>
|
||
), citing it as
|
||
<taxonomicName id="B89988E8FF87FFE9621FF920E85FF96E" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[557,841,1732,1758]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE9621FF920E9AEF96E" box="[557,696,1733,1758]" italics="true" pageId="21" pageNumber="298">Liocarcinus</emphasis>
|
||
cf.
|
||
<emphasis id="4DED2F79FF87FFE962D5F921E85FF96E" box="[743,841,1732,1758]" italics="true" pageId="21" pageNumber="298">holsatus</emphasis>
|
||
</taxonomicName>
|
||
, since its morphology was considered close to
|
||
<taxonomicName id="B89988E8FF87FFE9608DF90DEA2DF8B1" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[191,315,1767,1793]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE9608DF90DEA2DF8B1" box="[191,315,1767,1793]" italics="true" pageId="21" pageNumber="298">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
, but still different.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF87FFEA608FF8CDEAB8FEF7" blockId="21.[189,1400,1832,2000]" lastBlockId="22.[189,1399,265,327]" lastPageId="22" lastPageNumber="299" pageId="21" pageNumber="298">
|
||
This variability and the combination of morphological features is the reason for misidentifications, including records in GenBank and BOLD, where molecular sequences have been assigned to
|
||
<taxonomicName id="B89988E8FF87FFE9608FF895EA77F839" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[189,353,1904,1929]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF87FFE9608FF895EA77F839" box="[189,353,1904,1929]" italics="true" pageId="21" pageNumber="298">P. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
, for example a specimen from Roscoff (
|
||
<collectingCountry id="078EB3FBFF87FFE9637BF88AE88AF839" box="[841,924,1903,1929]" name="France" pageId="21" pageNumber="298">France</collectingCountry>
|
||
) obtained by
|
||
<bibRefCitation id="1B088E9AFF87FFE96471F88AEE27F839" author="Plagge C. & Thanh Son N. & Ng P. L. K. & Turkay M. & Streit B. & Klaus S." box="[1091,1329,1903,1929]" pageId="21" pageNumber="298" pagination="374 - 388" refId="ref16589" refString="Plagge C., Thanh Son N., Ng P. L. K., Turkay M., Streit B. & Klaus S. 2016. Liocarcinus corrugatus (Pennant, 1777) (Crustacea: Brachyura: Portunidae): a cosmopolitan brachyuran species? Raffles Bulletin of Zoology 64: 374 - 388." type="journal article" year="2016">
|
||
Plagge
|
||
<emphasis id="4DED2F79FF87FFE964ACF895EFCDF839" box="[1182,1243,1903,1929]" italics="true" pageId="21" pageNumber="298">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
(COI sequence KP795929) (see
|
||
<figureCitation id="E7A2EFEEFF87FFE96237F877E946F81C" box="[517,592,1938,1964]" captionStart="Fig" captionStartId="8.[189,232,1821,1847]" captionTargetBox="[320,1087,255,1734]" captionTargetId="graphics-32@8.[410,709,297,1664]" captionTargetPageId="8" captionText="Fig. 2. Maximum likelihood (ML) phylogenetic tree based on mitochondrial cytochrome c oxidase subunit I sequences of species of Polybius Leach, 1820 and Liocarcinus Stimpson, 1871, using 2000 Bootstrap replicates. Number on nodes represent ML Bootstrap values, only values>70 are included. Carcinus aestuarii Nardo, 1847 and C. maenas (Linnaeus, 1758) are used as outgroup. GenBank accession codes are indicated for each species, except those with codes XX## which sequences are listed in Supp. file 1." figureDoi="http://doi.org/10.5281/zenodo.10978029" httpUri="https://zenodo.org/record/10978029/files/figure.png" pageId="21" pageNumber="298">Fig. 2</figureCitation>
|
||
) and another from
|
||
<collectingCountry id="078EB3FBFF87FFE96376F877E8B1F81C" box="[836,935,1938,1964]" name="Portugal" pageId="21" pageNumber="298">Portugal</collectingCountry>
|
||
(sequence not available from BOLD, only early-release). Also, it has been assigned to
|
||
<taxonomicName id="B89988E8FF87FFE9633DF853E898F87F" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[783,910,1973,1999]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="298" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF87FFE9633DF853E898F87F" box="[783,910,1973,1999]" italics="true" pageId="21" pageNumber="298">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
, just like
|
||
<specimenCount id="699F38E2FF87FFE9643AF850EFD1F87F" box="[1032,1223,1973,1999]" count="3" pageId="21" pageNumber="298" type="generic">three specimens</specimenCount>
|
||
from
|
||
<collectingCountry id="078EB3FBFF87FFE96521F850EE60F87F" box="[1299,1398,1973,1999]" name="Portugal" pageId="21" pageNumber="298">Portugal</collectingCountry>
|
||
(sequences not available from BOLD, only early-release); but all of them actually correspond to
|
||
<taxonomicName id="B89988E8FF84FFEA608FFEC8EA5AFEF6" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[189,332,300,326]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="299" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF84FFEA608FFEC8EA5AFEF6" box="[189,332,300,326]" italics="true" pageId="22" pageNumber="299">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF84FFEA6161FEC8EAB8FEF7" box="[339,430,301,327]" pageId="22" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF84FFEB608FFE8BEAF2FB99" blockId="22.[189,1399,365,498]" lastBlockId="23.[188,1400,265,1065]" lastPageId="23" lastPageNumber="300" pageId="22" pageNumber="299">
|
||
The
|
||
<emphasis id="4DED2F79FF84FFEA60C7FE8BEA4DFE38" bold="true" box="[245,347,366,392]" pageId="22" pageNumber="299">rostrum</emphasis>
|
||
of the new species (
|
||
<figureCitation id="E7A2EFEEFF84FFEA6265FE8BE9CDFE38" box="[599,731,366,392]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="22" pageNumber="299">Figs 3A, F</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF84FFEA62D8FE8BE806FE38" box="[746,784,366,392]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="22" pageNumber="299">4A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF84FFEA632DFE88E853FE38" box="[799,837,365,392]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="22" pageNumber="299">5A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF84FFEA6366FE8BE861FE38" box="[852,887,366,392]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="22" pageNumber="299">7B</figureCitation>
|
||
) is similar to that of
|
||
<taxonomicName id="B89988E8FF84FFEA644FFE8BEFECFE38" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1149,1274,366,392]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="299" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF84FFEA644FFE8BEFECFE38" box="[1149,1274,366,392]" italics="true" pageId="22" pageNumber="299">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
, since the middle tooth is somewhat narrower either not, or only barely reaching the lateral ones, which are wider, rounded and more protruding; the central one is usually somewhat more rounded wide and somewhat longer in
|
||
<taxonomicName id="B89988E8FF84FFEA6118FE3DEAAEFE41" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[298,440,471,497]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="299" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF84FFEA6118FE3DEAAEFE41" box="[298,440,471,497]" italics="true" pageId="22" pageNumber="299">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF84FFEA618DFE3DE90EFE42" box="[447,536,472,498]" pageId="22" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF84FFEA6214FE32E992FE41" box="[550,644,471,497]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[341,1247,265,1812]" captionTargetId="figure-17@18.[341,1247,265,1812]" captionTargetPageId="18" captionText="Fig. 6. Dorsal view of orbital region and adjacent lateral rostral teeth. A. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). B. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). C. Polybius dioscurus sp. nov., ♀ from Málaga (CRUST_ICMAN/3765) D. Polybius vernalis, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species." figureDoi="http://doi.org/10.5281/zenodo.10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="22" pageNumber="299">Figs 6C</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF84FFEA62A2FE3DE9A4FE42" box="[656,690,472,498]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="22" pageNumber="299">7B</figureCitation>
|
||
vs 6D, 7C–E). In
|
||
<taxonomicName id="B89988E8FF84FFEA63B3FE3DE8EDFE41" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[897,1019,471,497]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="299" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF84FFEA63B3FE3DE8EDFE41" box="[897,1019,471,497]" italics="true" pageId="22" pageNumber="299">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
a greater variability is observed, mainly in specimens from the Black Sea in which the central rostral tooth is longer (
|
||
<figureCitation id="E7A2EFEEFF85FFEB648FFEECEE08FE93" box="[1213,1310,265,291]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="23" pageNumber="300">Figs 7E</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB651EFEECEE5BFE93" box="[1324,1357,265,291]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[189,1398,270,1883]" captionTargetId="figure-16@26.[189,1398,270,1883]" captionTargetPageId="26" captionText="Fig. 12. Polybius vernalis (Risso, 1827), ♂ from Bulgaria, Varna Bay (RBINS INV. 187236). A. Dorsal habitus. B. Carapace." figureDoi="http://doi.org/10.5281/zenodo.10978057" httpUri="https://zenodo.org/record/10978057/files/figure.png" pageId="23" pageNumber="300">12</figureCitation>
|
||
). It differs from that of
|
||
<taxonomicName id="B89988E8FF85FFEB619BFEC8E95AFEF6" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[425,588,301,326]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF85FFEB619BFEC8E95AFEF6" box="[425,588,301,326]" italics="true" pageId="23" pageNumber="300">P. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
because the middle tooth is similar to the lateral ones, rounded, and with equal length and width in this species (
|
||
<figureCitation id="E7A2EFEEFF85FFEB62F3FEAAE835FEDA" box="[705,803,335,362]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="23" pageNumber="300">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB631DFEAAE873FED9" box="[815,869,335,361]" captionStart="Fig" captionStartId="27.[189,232,1931,1957]" captionTargetBox="[189,1398,271,1884]" captionTargetId="figure-12@27.[189,1398,271,1884]" captionTargetPageId="27" captionText="Fig. 13. Polybius marmoreus (Leach, 1814), ♂ from Belgium, Heist (RBINS, INV. 187218). A. Dorsal habitus. B. Carapace facial view. C. Carpus of right pereiopod. D. Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978059" httpUri="https://zenodo.org/record/10978059/files/figure.png" pageId="23" pageNumber="300">13A</figureCitation>
|
||
). In
|
||
<taxonomicName id="B89988E8FF85FFEB63AAFEB5EF01FED9" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[920,1047,335,361]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB63AAFEB5EF01FED9" box="[920,1047,335,361]" italics="true" pageId="23" pageNumber="300">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
the median tooth overreaches the lateral ones (
|
||
<figureCitation id="E7A2EFEEFF85FFEB61B4FE97EAF5FE3D" box="[390,483,370,397]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="23" pageNumber="300">Figs 7F</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB61C3FE96E931FE3D" box="[497,551,371,397]" captionStart="Fig" captionStartId="28.[189,232,1931,1957]" captionTargetBox="[189,1398,269,1882]" captionTargetId="figure-16@28.[189,1398,269,1882]" captionTargetPageId="28" captionText="Fig. 14. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right pereiopod." figureDoi="http://doi.org/10.5281/zenodo.10978064" httpUri="https://zenodo.org/record/10978064/files/figure.png" pageId="23" pageNumber="300">14A</figureCitation>
|
||
), as in
|
||
<taxonomicName id="B89988E8FF85FFEB62B0FE96E807FE3C" authorityName="Leach" authorityYear="1820" box="[642,785,370,396]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="henslowii">
|
||
<emphasis id="4DED2F79FF85FFEB62B0FE96E807FE3C" box="[642,785,370,396]" italics="true" pageId="23" pageNumber="300">P. henslowii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF85FFEB6316FE97E882FE3D" box="[804,916,370,397]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,267,1879]" captionTargetId="figure-12@29.[189,1398,267,1879]" captionTargetPageId="29" captionText="Fig. 15. Polybius henslowii Leach, 1820, ♂ from France, Bay of Biscay, cruise PELGAS 10, stn O.0627, chalut 68 (RBINS, INV. 187237). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right pereiopod." figureDoi="http://doi.org/10.5281/zenodo.10978066" httpUri="https://zenodo.org/record/10978066/files/figure.png" pageId="23" pageNumber="300">Fig. 15A</figureCitation>
|
||
). The
|
||
<emphasis id="4DED2F79FF85FFEB63D6FE96EF6FFE3D" bold="true" box="[996,1145,371,397]" pageId="23" pageNumber="300">rostral area</emphasis>
|
||
of
|
||
<taxonomicName id="B89988E8FF85FFEB6491FE96EE37FE3C" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1187,1313,370,396]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB6491FE96EE37FE3C" box="[1187,1313,370,396]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="B89988E8FF85FFEB6162FE72EAC8FE00" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[336,478,406,432]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB6162FE72EAC8FE00" box="[336,478,406,432]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
. The inner orbital angles are more projected in
|
||
<taxonomicName id="B89988E8FF85FFEB6439FE72EF90FE00" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1035,1158,406,432]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB6439FE72EF90FE00" box="[1035,1158,406,432]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
, while the curvature between the rostral teeth is more rounded in
|
||
<taxonomicName id="B89988E8FF85FFEB62E4FE5FE871FE63" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[726,871,441,467]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB62E4FE5FE871FE63" box="[726,871,441,467]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
(in
|
||
<taxonomicName id="B89988E8FF85FFEB63ABFE5FEF00FE63" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[921,1046,441,467]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB63ABFE5FEF00FE63" box="[921,1046,441,467]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
the deepest zone is displaced towards the median tooth, not centred,
|
||
<figureCitation id="E7A2EFEEFF85FFEB62BAFE39E9FEFE46" box="[648,744,476,502]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[341,1247,265,1812]" captionTargetId="figure-17@18.[341,1247,265,1812]" captionTargetPageId="18" captionText="Fig. 6. Dorsal view of orbital region and adjacent lateral rostral teeth. A. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). B. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). C. Polybius dioscurus sp. nov., ♀ from Málaga (CRUST_ICMAN/3765) D. Polybius vernalis, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species." figureDoi="http://doi.org/10.5281/zenodo.10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="23" pageNumber="300">Figs 6D</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB62C6FE39E821FE46" box="[756,823,476,502]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[228,1359,265,1773]" captionTargetId="figure-11@19.[228,1359,265,1773]" captionTargetPageId="19" captionText="Fig. 7. Frontal region in dorsal view, male. A. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). B. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C–E. Polybius vernalis (Risso, 1827). C. Specimen from Tunisia, Bay of Tunis (RBINS, INV. 187235). D. Specimen from Spain, Málaga, Fuengirola (RBINS, INV. 187238). E. Specimen from Bulgaria, Varna Bay (RBINS INV. 187236). F. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221)." figureDoi="http://doi.org/10.5281/zenodo.10978045" httpUri="https://zenodo.org/record/10978045/files/figure.png" pageId="23" pageNumber="300">7C–E</figureCitation>
|
||
vs 7B); however, there is some variability in this character. In addition, the tuberculate rostral rim is continuous with that of the inner orbital angle in
|
||
<taxonomicName id="B89988E8FF85FFEB608FFDC6EA5BFD8C" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[189,333,546,572]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB608FFDC6EA5BFD8C" box="[189,333,546,572]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
, while in
|
||
<taxonomicName id="B89988E8FF85FFEB61FAFDC6E953FD8C" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[456,581,546,572]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB61FAFDC6E953FD8C" box="[456,581,546,572]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
it is not so clear because it is projected anteriorly and flexes slightly downward (
|
||
<figureCitation id="E7A2EFEEFF85FFEB6179FDA3EA80FDD0" box="[331,406,582,608]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[341,1247,265,1812]" captionTargetId="figure-17@18.[341,1247,265,1812]" captionTargetPageId="18" captionText="Fig. 6. Dorsal view of orbital region and adjacent lateral rostral teeth. A. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). B. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). C. Polybius dioscurus sp. nov., ♀ from Málaga (CRUST_ICMAN/3765) D. Polybius vernalis, ♂ from Málaga (CRUST_ ICMAN/3789). Arrows show differences in the orbital region between both species." figureDoi="http://doi.org/10.5281/zenodo.10978043" httpUri="https://zenodo.org/record/10978043/files/figure.png" pageId="23" pageNumber="300">Fig. 6</figureCitation>
|
||
), and the inner side of the orbital socket shows a steeper slope in
|
||
<taxonomicName id="B89988E8FF85FFEB6488FDA2EE2FFDD0" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1210,1337,582,608]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB6488FDA2EE2FFDD0" box="[1210,1337,582,608]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<emphasis id="4DED2F79FF85FFEB608DFD8CEA39FD33" bold="true" box="[191,303,617,643]" pageId="23" pageNumber="300">carapace</emphasis>
|
||
in
|
||
<taxonomicName id="B89988E8FF85FFEB6161FD8FEAD8FD33" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[339,462,617,643]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB6161FD8FEAD8FD33" box="[339,462,617,643]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
is frequently covered by short setae (e.g.,
|
||
<figureCitation id="E7A2EFEEFF85FFEB638DFD8CEF05FD33" box="[959,1043,617,643]" captionStart="Fig" captionStartId="25.[189,232,1925,1951]" captionTargetBox="[189,1398,268,1881]" captionTargetId="figure-11@25.[189,1398,268,1881]" captionTargetPageId="25" captionText="Fig. 11. Polybius vernalis (Risso, 1827), ♂ from Spain, Málaga, Fuengirola (RBINS, INV. 187238). A. Dorsal habitus. B. Carapace. C. Carpus of right pereiopod." figureDoi="http://doi.org/10.5281/zenodo.10978055" httpUri="https://zenodo.org/record/10978055/files/figure.png" pageId="23" pageNumber="300">Fig. 11</figureCitation>
|
||
), while in the others species it is glabrous, although we have seen specimens that were only almost glabrous. In addition, the carapace of
|
||
<taxonomicName id="B89988E8FF85FFEB60D2FD55EA49FD79" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[224,351,687,713]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB60D2FD55EA49FD79" box="[224,351,687,713]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
from the Black Sea (
|
||
<figureCitation id="E7A2EFEEFF85FFEB625AFD4AE9D6FD79" box="[616,704,687,713]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[189,1398,270,1883]" captionTargetId="figure-16@26.[189,1398,270,1883]" captionTargetPageId="26" captionText="Fig. 12. Polybius vernalis (Risso, 1827), ♂ from Bulgaria, Varna Bay (RBINS INV. 187236). A. Dorsal habitus. B. Carapace." figureDoi="http://doi.org/10.5281/zenodo.10978057" httpUri="https://zenodo.org/record/10978057/files/figure.png" pageId="23" pageNumber="300">Fig. 12</figureCitation>
|
||
) is nearly smooth and hairless and their frontal teeth are a bit longer, different from those of the Mediterranean Sea (
|
||
<figureCitation id="E7A2EFEEFF85FFEB63A8FD37EF33FD5D" box="[922,1061,722,749]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="24.[189,232,1921,1947]" captionStartId-1="25.[189,232,1925,1951]" captionTargetBox-0="[189,1398,268,1881]" captionTargetBox-1="[189,1398,268,1881]" captionTargetId-0="figure-16@24.[189,1398,268,1881]" captionTargetId-1="figure-11@25.[189,1398,268,1881]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="Fig. 10. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus." captionText-1="Fig. 11. Polybius vernalis (Risso, 1827), ♂ from Spain, Málaga, Fuengirola (RBINS, INV. 187238). A. Dorsal habitus. B. Carapace. C. Carpus of right pereiopod." figureDoi-0="http://doi.org/10.5281/zenodo.10978051" figureDoi-1="http://doi.org/10.5281/zenodo.10978055" httpUri-0="https://zenodo.org/record/10978051/files/figure.png" httpUri-1="https://zenodo.org/record/10978055/files/figure.png" pageId="23" pageNumber="300">Figs 10–11</figureCitation>
|
||
); but no genetic differences were detected, and the shape of their orbital region (the most important character separating them from
|
||
<taxonomicName id="B89988E8FF85FFEB608FFCFFEA47FC83" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[189,337,793,819]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB608FFCFFEA47FC83" box="[189,337,793,819]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
) exhibits no difference in relation to the Mediterranean specimens. The
|
||
<emphasis id="4DED2F79FF85FFEB64E6FCFCEA0EFCE6" bold="true" pageId="23" pageNumber="300">anterolateral margin</emphasis>
|
||
of the carapace is clearly curved, but in
|
||
<taxonomicName id="B89988E8FF85FFEB62CEFCD8E86DFCE6" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[764,891,828,854]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB62CEFCD8E86DFCE6" box="[764,891,828,854]" italics="true" pageId="23" pageNumber="300">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
it is slightly shorter than the posterolateral one (
|
||
<figureCitation id="E7A2EFEEFF85FFEB60CBFCBAEA72FCC9" box="[249,356,863,889]" captionStart="Fig" captionStartId="28.[189,232,1931,1957]" captionTargetBox="[189,1398,269,1882]" captionTargetId="figure-16@28.[189,1398,269,1882]" captionTargetPageId="28" captionText="Fig. 14. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right pereiopod." figureDoi="http://doi.org/10.5281/zenodo.10978064" httpUri="https://zenodo.org/record/10978064/files/figure.png" pageId="23" pageNumber="300">Fig. 14A</figureCitation>
|
||
) (AB/PB: 1.17–1.23). In
|
||
<taxonomicName id="B89988E8FF85FFEB62A0FC85E80BFCC9" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[658,797,863,889]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB62A0FC85E80BFCC9" box="[658,797,863,889]" italics="true" pageId="23" pageNumber="300">P dioscurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF85FFEB631FFCBAE899FCC9" box="[813,911,863,889]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="23" pageNumber="300">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB63AFFCBAE8D5FCC9" box="[925,963,863,889]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="23" pageNumber="300">4A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB63E2FCBAE8E3FCCA" box="[976,1013,863,890]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="23" pageNumber="300">5A</figureCitation>
|
||
) both are more or less similar in length (0.95–1.17) and in
|
||
<taxonomicName id="B89988E8FF85FFEB61D8FC66E973FC2C" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[490,613,898,924]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB61D8FC66E973FC2C" box="[490,613,898,924]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
the posterolateral margin is slight shorter (1.07–1.17), but there are no clear differences between these two species. In
|
||
<taxonomicName id="B89988E8FF85FFEB6330FC42E8B4FC70" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[770,930,935,960]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF85FFEB6330FC42E8B4FC70" box="[770,930,935,960]" italics="true" pageId="23" pageNumber="300">P. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
this ratio is 1.04–1.16. The anterolateral fifth tooth in
|
||
<taxonomicName id="B89988E8FF85FFEB6150FC2FEAF5FC53" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[354,483,969,995]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB6150FC2FEAF5FC53" box="[354,483,969,995]" italics="true" pageId="23" pageNumber="300">L. vernalis</emphasis>
|
||
</taxonomicName>
|
||
projects slightly more outward than in
|
||
<taxonomicName id="B89988E8FF85FFEB63F8FC2FEF4BFC53" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[970,1117,969,995]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB63F8FC2FEF4BFC53" box="[970,1117,969,995]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
(with the tips pointing forwards) (
|
||
<figureCitation id="E7A2EFEEFF85FFEB610DFC09EAA7FBB6" box="[319,433,1004,1030]" captionStart="Fig" captionStartId="24.[189,232,1921,1947]" captionTargetBox="[189,1398,268,1881]" captionTargetId="figure-16@24.[189,1398,268,1881]" captionTargetPageId="24" captionText="Fig. 10. Polybius vernalis (Risso, 1827), ♂ from Tunisia, Bay of Tunis (RBINS, INV. 187235). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.10978051" httpUri="https://zenodo.org/record/10978051/files/figure.png" pageId="23" pageNumber="300">Figs 10A</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB618FFC09E913FBB6" box="[445,517,1004,1030]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="25.[189,232,1925,1951]" captionStartId-1="26.[189,232,1929,1955]" captionTargetBox-0="[189,1398,268,1881]" captionTargetBox-1="[189,1398,270,1883]" captionTargetId-0="figure-11@25.[189,1398,268,1881]" captionTargetId-1="figure-16@26.[189,1398,270,1883]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="Fig. 11. Polybius vernalis (Risso, 1827), ♂ from Spain, Málaga, Fuengirola (RBINS, INV. 187238). A. Dorsal habitus. B. Carapace. C. Carpus of right pereiopod." captionText-1="Fig. 12. Polybius vernalis (Risso, 1827), ♂ from Bulgaria, Varna Bay (RBINS INV. 187236). A. Dorsal habitus. B. Carapace." figureDoi-0="http://doi.org/10.5281/zenodo.10978055" figureDoi-1="http://doi.org/10.5281/zenodo.10978057" httpUri-0="https://zenodo.org/record/10978055/files/figure.png" httpUri-1="https://zenodo.org/record/10978057/files/figure.png" pageId="23" pageNumber="300">11–12</figureCitation>
|
||
vs 3A, 4A, 5A). The
|
||
<emphasis id="4DED2F79FF85FFEB633AFC09EF3DFBB6" bold="true" box="[776,1067,1004,1030]" pageId="23" pageNumber="300">posterolaterals borders</emphasis>
|
||
are convergent, particularly in
|
||
<taxonomicName id="B89988E8FF85FFEB60E9FBF5EA4CFB99" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[219,346,1039,1065]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB60E9FBF5EA4CFB99" box="[219,346,1039,1065]" italics="true" pageId="23" pageNumber="300">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF85FFEB6158FBEAEAC1FB99" box="[362,471,1039,1065]" captionStart="Fig" captionStartId="28.[189,232,1931,1957]" captionTargetBox="[189,1398,269,1882]" captionTargetId="figure-16@28.[189,1398,269,1882]" captionTargetPageId="28" captionText="Fig. 14. Polybius holsatus (Fabricius, 1798), ♂ from Belgium, Duinbergen (RBINS, INV. 187221). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right pereiopod." figureDoi="http://doi.org/10.5281/zenodo.10978064" httpUri="https://zenodo.org/record/10978064/files/figure.png" pageId="23" pageNumber="300">Fig. 14A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2BE6A3E3FF84FFEA608EF84EEFC0F858" ID-DOI="http://doi.org/10.5281/zenodo.10978049" ID-Zenodo-Dep="10978049" httpUri="https://zenodo.org/record/10978049/files/figure.png" pageId="22" pageNumber="299" startId="22.[188,231,1963,1989]" targetBox="[189,1398,558,1917]" targetPageId="22">
|
||
<paragraph id="7F26F36BFF84FFEA608EF84EEFC0F858" blockId="22.[188,1399,1963,2025]" pageId="22" pageNumber="299">
|
||
<emphasis id="4DED2F79FF84FFEA608EF84EEA1DF875" bold="true" box="[188,267,1963,1989]" pageId="22" pageNumber="299">Fig. 9.</emphasis>
|
||
<taxonomicName id="B89988E8FF84FFEA6127F84EEAE0F875" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[277,502,1963,1989]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="299" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF84FFEA6127F84EEAE0F875" box="[277,502,1963,1989]" italics="true" pageId="22" pageNumber="299">Polybius dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF84FFEA61CCF849E94BF876" box="[510,605,1964,1990]" pageId="22" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratypes from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187230), colour variation (in life) (sex not recorded during photo session).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7F26F36BFF85FFEB608FFBB4E9E9F9CB" blockId="23.[188,1402,1104,1659]" pageId="23" pageNumber="300">
|
||
Other features that could be used are:
|
||
<emphasis id="4DED2F79FF85FFEB62B9FBB4EE0CFBDA" bold="true" box="[651,1306,1104,1131]" pageId="23" pageNumber="300">the anterior outer border of the carpus of chelipeds</emphasis>
|
||
, that in
|
||
<taxonomicName id="B89988E8FF85FFEB608DFB90EA59FB3E" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[191,335,1140,1166]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF85FFEB608DFB90EA59FB3E" box="[191,335,1140,1166]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF85FFEB6168FB90EAAEFB3F" box="[346,440,1141,1167]" pageId="23" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shows protruding lobes that are slightly less marked than those of
|
||
<taxonomicName id="B89988E8FF85FFEB64C0FB90EE61FB3E" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1266,1399,1140,1166]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB64C0FB90EE61FB3E" box="[1266,1399,1140,1166]" italics="true" pageId="23" pageNumber="300">L. holsatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF85FFEB60F7FB72EA3CFB01" box="[197,298,1175,1201]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[247,1340,265,1845]" captionTargetId="figure-16@14.[247,1340,265,1845]" captionTargetPageId="14" captionText="Fig. 3. Polybius dioscurus sp. nov., holotype, ♂, 26.4 mm CW, from Valdelagrana, Cádiz, Spain, 36°43′ N, 06°14′ W, 3.3–6.6 m depth, 31 Oct. 1996 (CRUST_ICMAN/3759). A. Dorsal habitus.B. Right cheliped, facial view. C. Carpus of left cheliped. D. Carpus to dactylus of right pereiopod 5. E. Body in ventral view, abdomen view." figureDoi="http://doi.org/10.5281/zenodo.10978033" httpUri="https://zenodo.org/record/10978033/files/figure.png" pageId="23" pageNumber="300">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB610FFB72EA77FB01" box="[317,353,1175,1201]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="23" pageNumber="300">4C</figureCitation>
|
||
,
|
||
<figureCitation id="E7A2EFEEFF85FFEB6141FB72EA81FB02" box="[371,407,1175,1202]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[190,1398,265,1875]" captionTargetId="figure-17@16.[190,1398,265,1875]" captionTargetPageId="16" captionText="Fig. 5. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). A. Dorsal habitus. B. Facial habitus. C. Ventral habitus. D. Carpus of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.10978041" httpUri="https://zenodo.org/record/10978041/files/figure.png" pageId="23" pageNumber="300">5D</figureCitation>
|
||
vs 14D), but closer to those of
|
||
<taxonomicName id="B89988E8FF85FFEB6306FB7DE8A1FB01" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[820,951,1175,1201]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB6306FB7DE8A1FB01" box="[820,951,1175,1201]" italics="true" pageId="23" pageNumber="300">L. vernalis</emphasis>
|
||
</taxonomicName>
|
||
. However, in large specimens of
|
||
<taxonomicName id="B89988E8FF85FFEB6553FB7DEA39FB64" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB6553FB7DEA39FB64" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
these lobes are less developed (
|
||
<figureCitation id="E7A2EFEEFF85FFEB6287FB5FE807FB65" box="[693,785,1210,1237]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="23" pageNumber="300">Fig. 4C</figureCitation>
|
||
) (while in medium-small specimens they are more similar to those of
|
||
<taxonomicName id="B89988E8FF85FFEB61A7FB3BE906FB48" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[405,528,1246,1272]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB61A7FB3BE906FB48" box="[405,528,1246,1272]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
). In
|
||
<taxonomicName id="B89988E8FF85FFEB6274FB3BE9FFFB47" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[582,745,1246,1271]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF85FFEB6274FB3BE9FFFB47" box="[582,745,1246,1271]" italics="true" pageId="23" pageNumber="300">L. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
the outer border is rounded and without any protrusions (
|
||
<figureCitation id="E7A2EFEEFF85FFEB60F7FAE4EA39FAAB" box="[197,303,1281,1307]" captionStart="Fig" captionStartId="27.[189,232,1931,1957]" captionTargetBox="[189,1398,271,1884]" captionTargetId="figure-12@27.[189,1398,271,1884]" captionTargetPageId="27" captionText="Fig. 13. Polybius marmoreus (Leach, 1814), ♂ from Belgium, Heist (RBINS, INV. 187218). A. Dorsal habitus. B. Carapace facial view. C. Carpus of right pereiopod. D. Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978059" httpUri="https://zenodo.org/record/10978059/files/figure.png" pageId="23" pageNumber="300">Fig. 13D</figureCitation>
|
||
) (see
|
||
<bibRefCitation id="1B088E9AFF85FFEB6147FAE4E918FAAB" author="Palmer R." box="[373,526,1281,1307]" pageId="23" pageNumber="300" pagination="877 - 908" refId="ref16457" refString="Palmer R. 1927. A revision of the genus " Portunus " (A. Milne Edwards, Bell, etc). Journal of the Marine Biological Association of the United Kingdom 14: 877 - 908. https: // doi. org / 10.1017 / S 0025315400051134" type="journal article" year="1927">Palmer 1927</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1B088E9AFF85FFEB6228FAE4E82EFAAB" author="D'Udekem d'Acoz C." box="[538,824,1281,1307]" pageId="23" pageNumber="300" pagination="84 - 100" refId="ref14115" refString="D'Udekem d'Acoz C. 1986. Etude d'une collection de crustaces decapodes de Bretagne. De Strandvlo [dec. 1985]: 11 (4): 84 - 100." type="journal article" year="1986">d’Udekem d’Acoz 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1B088E9AFF85FFEB6376FAE4EFC2FAAB" author="D'Udekem d'Acoz C. & Rappe G." box="[836,1236,1281,1307]" pageId="23" pageNumber="300" pagination="95 - 99" refId="ref14323" refString="D'Udekem d'Acoz C. & Rappe G. 1991. Presence et abondance de Liocarcinus vernalis (Risso, 1816) dans la baie sud de la mer du Nord (Decapoda, Brachyura, Portunidae). Crustaceana 61 (1): 95 - 99. Available from http: // www. jstor. org / stable / 20104677 [accessed 22 Mar. 2024]." type="journal article" year="1991">d’Udekem d’Acoz & Rappé 1991</bibRefCitation>
|
||
). The
|
||
<emphasis id="4DED2F79FF85FFEB652FFAE4EA8AFA8E" bold="true" pageId="23" pageNumber="300">ventral setae of dactyl P4</emphasis>
|
||
occupy approximately ⅓ basal in
|
||
<taxonomicName id="B89988E8FF85FFEB630EFAC0E8D9FA8E" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[828,975,1316,1342]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB630EFAC0E8D9FA8E" box="[828,975,1316,1342]" italics="true" pageId="23" pageNumber="300">L. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B89988E8FF85FFEB643EFAC0EF9AFA8E" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1036,1164,1316,1342]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB643EFAC0EF9AFA8E" box="[1036,1164,1316,1342]" italics="true" pageId="23" pageNumber="300">L. vernalis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF85FFEB64ACFAC1EFE8FA8E" box="[1182,1278,1316,1342]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[278,1309,282,1800]" captionTargetId="figure-11@15.[278,1309,265,1847]" captionTargetPageId="15" captionText="Fig. 4. Polybius dioscurus sp. nov., paratype, ♂, 36.7 mm CW, from Agadir Bay, Morocco. 30°26′ N, 09°40′ W – 30°18′ N, 09°37′ W, 6 to 25 m depth, May 1999 (CRUST_ICMAN/3762). A. Dorsal habitus. B. Right cheliped, facial view. C. Carpus of left cheliped. D. Dactylus of right pereiopods 5 and 4. E. Body in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10978037" httpUri="https://zenodo.org/record/10978037/files/figure.png" pageId="23" pageNumber="300">Fig. 4D</figureCitation>
|
||
), while in
|
||
<taxonomicName id="B89988E8FF85FFEB608DFAADEA55FAD1" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[191,323,1351,1377]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB608DFAADEA55FAD1" box="[191,323,1351,1377]" italics="true" pageId="23" pageNumber="300">L. holsatus</emphasis>
|
||
</taxonomicName>
|
||
they exceed half their length (
|
||
<bibRefCitation id="1B088E9AFF85FFEB628BFAA2EF4EFAD1" author="D'Udekem d'Acoz C. & Rappe G." box="[697,1112,1351,1377]" pageId="23" pageNumber="300" pagination="95 - 99" refId="ref14323" refString="D'Udekem d'Acoz C. & Rappe G. 1991. Presence et abondance de Liocarcinus vernalis (Risso, 1816) dans la baie sud de la mer du Nord (Decapoda, Brachyura, Portunidae). Crustaceana 61 (1): 95 - 99. Available from http: // www. jstor. org / stable / 20104677 [accessed 22 Mar. 2024]." type="journal article" year="1991">d’Udekem d’Acoz & Rappé 1991</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="B89988E8FF85FFEB64A1FAADEE2CFAD1" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[1171,1338,1352,1377]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF85FFEB64A1FAADEE2CFAD1" box="[1171,1338,1352,1377]" italics="true" pageId="23" pageNumber="300">L. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
they almost reach ½. However, this character remains somewhat variable. The
|
||
<emphasis id="4DED2F79FF85FFEB6471FA8EEFF6FA35" bold="true" box="[1091,1248,1386,1413]" pageId="23" pageNumber="300">merus of P5</emphasis>
|
||
is distinctly shorter in
|
||
<taxonomicName id="B89988E8FF85FFEB6102FA6BEAA6FA18" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[304,432,1422,1448]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB6102FA6BEAA6FA18" box="[304,432,1422,1448]" italics="true" pageId="23" pageNumber="300">L. holsatus</emphasis>
|
||
</taxonomicName>
|
||
(ratio ML/MH: 1.1–1.3; 1.4–1.6 according
|
||
<bibRefCitation id="1B088E9AFF85FFEB639FFA6BEE23FA18" author="D'Udekem d'Acoz C. & Rappe G." box="[941,1333,1422,1448]" pageId="23" pageNumber="300" pagination="95 - 99" refId="ref14323" refString="D'Udekem d'Acoz C. & Rappe G. 1991. Presence et abondance de Liocarcinus vernalis (Risso, 1816) dans la baie sud de la mer du Nord (Decapoda, Brachyura, Portunidae). Crustaceana 61 (1): 95 - 99. Available from http: // www. jstor. org / stable / 20104677 [accessed 22 Mar. 2024]." type="journal article" year="1991">d’Udekem d’Acoz & Rappé 1991</bibRefCitation>
|
||
) than those of
|
||
<taxonomicName id="B89988E8FF85FFEB6115FA57EAAEFA7B" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[295,440,1457,1483]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB6115FA57EAAEFA7B" box="[295,440,1457,1483]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
(1.63–1.89),
|
||
<taxonomicName id="B89988E8FF85FFEB6269FA57E9CEFA7B" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[603,728,1457,1483]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB6269FA57E9CEFA7B" box="[603,728,1457,1483]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
(1.51–1.76) and
|
||
<taxonomicName id="B89988E8FF85FFEB6395FA57EF5AFA7B" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[935,1100,1458,1483]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="marmoreus">
|
||
<emphasis id="4DED2F79FF85FFEB6395FA57EF5AFA7B" box="[935,1100,1458,1483]" italics="true" pageId="23" pageNumber="300">L. marmoreus</emphasis>
|
||
</taxonomicName>
|
||
(1.65–1.87), whose ratio values overlap.
|
||
<bibRefCitation id="1B088E9AFF85FFEB6144FA31E80EFA5E" author="D'Udekem d'Acoz C. & Rappe G." box="[374,792,1492,1518]" pageId="23" pageNumber="300" pagination="95 - 99" refId="ref14323" refString="D'Udekem d'Acoz C. & Rappe G. 1991. Presence et abondance de Liocarcinus vernalis (Risso, 1816) dans la baie sud de la mer du Nord (Decapoda, Brachyura, Portunidae). Crustaceana 61 (1): 95 - 99. Available from http: // www. jstor. org / stable / 20104677 [accessed 22 Mar. 2024]." type="journal article" year="1991">D’Udekem d’Acoz & Rappé (1991)</bibRefCitation>
|
||
and d’Udekem d’Acoz (1991) found different ratios for the last two species (2.0–2.5, and 1.8–2.1, respectively). The datcylus and propodus are similar in
|
||
<taxonomicName id="B89988E8FF85FFEB608DF9FEEA2CF984" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[191,314,1562,1588]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB608DF9FEEA2CF984" box="[191,314,1562,1588]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B89988E8FF85FFEB6140F9FEE917F984" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[370,513,1562,1588]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus">
|
||
<emphasis id="4DED2F79FF85FFEB6140F9FEE917F984" box="[370,513,1562,1588]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
but somewhat different from those of
|
||
<taxonomicName id="B89988E8FF85FFEB63F7F9FEEF52F984" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[965,1092,1562,1588]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB63F7F9FEEF52F984" box="[965,1092,1562,1588]" italics="true" pageId="23" pageNumber="300">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
. The posterodistal lobe of the propodus is more developed and broader in
|
||
<taxonomicName id="B89988E8FF85FFEB62C5F9DBE861F9E8" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[759,887,1598,1624]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB62C5F9DBE861F9E8" box="[759,887,1598,1624]" italics="true" pageId="23" pageNumber="300">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
than in other species (
|
||
<bibRefCitation id="1B088E9AFF85FFEB64B0F9DBEE78F9E8" author="Ingle R. W. & Clark P. P." box="[1154,1390,1598,1624]" pageId="23" pageNumber="300" pagination="223 - 229" refId="ref15194" refString="Ingle R. W. & Clark P. P. 1998. A swimming crab new to the British fauna, Liocarcinus vernalis (Crustacea: Brachyura: Portunidae). Journal of the Marine Biological Association of the United Kingdom 78: 223 - 229. https: // doi. org / 10.1017 / S 0025315400040042 ICZN 1999. InternationalCodeofZoologicalNomenclature. 4 thedition. TheInternationalTrustforZoological Nomenclature, London. Available from https: // www. iczn. org / the-code / the-code-online / [accessed 22 Mar. 2024]." type="journal article" year="1998">Ingle & Clark 1998</bibRefCitation>
|
||
) and the dactylus is usually broader in
|
||
<taxonomicName id="B89988E8FF85FFEB624BF987E9EEF9CB" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[633,760,1633,1659]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="holsatus">
|
||
<emphasis id="4DED2F79FF85FFEB624BF987E9EEF9CB" box="[633,760,1633,1659]" italics="true" pageId="23" pageNumber="300">P. holsatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7F26F36BFF85FFEB608FF947E920F8B3" blockId="23.[189,1399,1698,1795]" pageId="23" pageNumber="300">
|
||
The distal part of the
|
||
<emphasis id="4DED2F79FF85FFEB61F4F947E9BDF90C" bold="true" box="[454,683,1698,1724]" pageId="23" pageNumber="300">first male pleopod</emphasis>
|
||
is slightly more curved in
|
||
<taxonomicName id="B89988E8FF85FFEB63C4F946EF62F90C" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1014,1140,1698,1724]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="4DED2F79FF85FFEB63C4F946EF62F90C" box="[1014,1140,1698,1724]" italics="true" pageId="23" pageNumber="300">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E7A2EFEEFF85FFEB64B5F947EE15F90C" box="[1159,1283,1698,1724]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[218,1370,265,1801]" captionTargetId="figure-17@20.[218,1370,265,1801]" captionTargetPageId="20" captionText="Fig. 8. Left gonopod 1 of male. A–B. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). A. Ventral view. B. Dorsal view. C–D. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C. Ventral view. D. Dorsal view. E–F. Polybius vernalis (Risso, 1827) from Tunisia, Bay of Tunis (RBINS, INV. 187235).E. Ventral view. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10978047" httpUri="https://zenodo.org/record/10978047/files/figure.png" pageId="23" pageNumber="300">Fig. 8E–F</figureCitation>
|
||
; in larger specimens even somewhat hooked) than in
|
||
<taxonomicName id="B89988E8FF85FFEB62F0F923E844F96F" authority="García-Raso & Moukrim & Schubart & Cuesta, 2024" authorityName="García-Raso & Moukrim & Schubart & Cuesta" authorityYear="2024" box="[706,850,1733,1759]" class="Malacostraca" family="Polybiidae" genus="Polybius" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="300" phylum="Arthropoda" rank="species" species="dioscurus" status="sp. nov.">
|
||
<emphasis id="4DED2F79FF85FFEB62F0F923E844F96F" box="[706,850,1733,1759]" italics="true" pageId="23" pageNumber="300">P. dioscurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="56DE9202FF85FFEB6368F923E8AEF950" box="[858,952,1734,1760]" pageId="23" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, in which it tends to form an angle of approximately 90° (
|
||
<figureCitation id="E7A2EFEEFF85FFEB619AF90CE93FF8B3" box="[424,553,1769,1795]" captionStart="Fig" captionStartId="20.[189,232,1857,1883]" captionTargetBox="[218,1370,265,1801]" captionTargetId="figure-17@20.[218,1370,265,1801]" captionTargetPageId="20" captionText="Fig. 8. Left gonopod 1 of male. A–B. Polybius marmoreus (Leach, 1814) from Belgium, Heist (RBINS, INV. 187218). A. Ventral view. B. Dorsal view. C–D. Polybius dioscurus sp. nov., paratype, ♂ from Belgium, between Nieuwpoort and Oostduinkerke (RBINS, INV. 187223). C. Ventral view. D. Dorsal view. E–F. Polybius vernalis (Risso, 1827) from Tunisia, Bay of Tunis (RBINS, INV. 187235).E. Ventral view. F. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10978047" httpUri="https://zenodo.org/record/10978047/files/figure.png" pageId="23" pageNumber="300">Fig. 8C–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |