328 lines
39 KiB
XML
328 lines
39 KiB
XML
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<mods:title id="0599DBDDA4B29C12D60A17FBC9150002">Revision of the enigmatic South African Cryptolaryngini (Coleoptera, Curculionidae), with description of a new genus and twenty-two new species</mods:title>
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<mods:namePart id="F48575D0BC396DEECA6D0478D8302A8B">Haran, Julien M.</mods:namePart>
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<mods:affiliation id="9AAEFF92FB28A2BC52FBEBF450962862">Montpellier SupAgro, INRAe, IRD, Montpellier University, Montpellier, France.</mods:affiliation>
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<mods:namePart id="556B9AABC247FA2FF3C7A98A2F4DB86A">Marvaldi, Adriana E.</mods:namePart>
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<mods:affiliation id="CB1D6BA233B0E29A381FA562FD34B22A">División Entomología, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, Buenos Aires, Argentina.</mods:affiliation>
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<mods:nameIdentifier id="9E1CFB9940226CF478922AA65F4CA7DC" type="email">Marvaldi@fcnym.unlp.edu.ar</mods:nameIdentifier>
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<mods:namePart id="D407F9BF93D7372CE8A9DCD53D823B90">Benoit, Laure</mods:namePart>
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<mods:affiliation id="F4CB3ED28A3AABB7E5B2424876B3F63D">Montpellier SupAgro, INRAe, IRD, Montpellier University, Montpellier, France. & Laure.</mods:affiliation>
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<mods:nameIdentifier id="6F9468A7D367E02CED1D4187D7F5ADEE" type="email">Laure.Benoit@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="EDE7DE18160500579C1517F995036133">Oberlander, Kenneth</mods:namePart>
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<mods:affiliation id="1D6008E708F30710BFDDEB145963A6B6">Schweickerdt Herbarium, Department of Plant and Soil Sciences, Plant Sciences Complex, University of Pretoria, Private Bag X 20, Hatfield 0024, South Africa. & Kenneth.</mods:affiliation>
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<mods:nameIdentifier id="A001134D0A67F443474EE3CC5F6162C9" type="email">Kenneth.Oberlander@up.ac.za</mods:nameIdentifier>
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<mods:namePart id="697B834D75CB169CED9637D987F0ECD1">Stals, Riaan</mods:namePart>
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<mods:affiliation id="DD4F720EBA4ABB67FCCC714E47DD9B77">South African National Collection of Insects, Agricultural Research Council, Private Bag X 134, Queenswood 0121, South Africa.</mods:affiliation>
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<mods:nameIdentifier id="549559B00238B70E8329D54E5853560A" type="email">StalsR@arc.agric.za</mods:nameIdentifier>
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<mods:namePart id="A9D2B4192A894508CCD57C92DCBF8B97">Oberprieler, Rolf G.</mods:namePart>
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<mods:affiliation id="111472B3421FD1082C47E07DCBB513CB">Australian National Insect Collection, G. P. O. Box 1700, Canberra, A. C. T. 2601, Australia.</mods:affiliation>
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<mods:nameIdentifier id="F6C1B922A3B603DFC55CD289C4439BC1" type="email">curculio@homemail.com.au</mods:nameIdentifier>
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<mods:title id="CC212B299BD923314FFFDA81549A2CE1">European Journal of Taxonomy</mods:title>
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<mods:part id="990324914D13F90091ADC0F593CA5C45">
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<mods:date id="99CA7EC6C568E9B04BE2F07EC19E44C1">2023</mods:date>
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17.
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<taxonomicName id="E093BBB0FFCDFFB6FDB36086FC7B04DF" authority="Haran" authorityName="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler" authorityYear="2023" box="[554,980,1432,1459]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="cederbergensis" status="sp. nov.">
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<emphasis id="15E71C21FFCDFFB6FDB36086FC2B04DF" bold="true" box="[554,900,1432,1459]" italics="true" pageId="50" pageNumber="51">Cryptolarynx cederbergensis</emphasis>
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Haran
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<taxonomicNameLabel id="0ED4A15AFFCDFFB6FC426087FB9904DF" box="[987,1078,1433,1459]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="272CC033FFCDFFB6FD1F60C1FC020495" blockId="50.[375,1212,1432,1529]" box="[646,941,1502,1529]" pageId="50" pageNumber="51">
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<figureCitation id="BFA8DCB6FFCDFFB6FD1F60C1FD4A0495" box="[646,741,1503,1529]" captionStart="Fig" captionStartId="9.[190,233,1845,1871]" captionTargetId="figure-13@10.[308,1280,265,2019]" captionTargetPageId="10" captionText="Fig. 1 (Part I) (next page). Habitus of males of species of Cryptolarynx Van Schalkwyk, 1966 (dorsal view). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113555" httpUri="https://zenodo.org/record/8113555/files/figure.png" pageId="50" pageNumber="51">Figs 1Q</figureCitation>
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,
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<figureCitation id="BFA8DCB6FFCDFFB6FD6B60C1FCB60495" box="[754,793,1503,1529]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetId="figure-13@16.[201,1387,265,2019]" captionTargetPageId="16" captionText="Fig. 2 (Part I) (next page). Male genitalia of species of Cryptolarynx Van Schalkwyk, 1966. Penis in dorsal and lateral view (left), tegmen in dorsal view (middle) and details of apex of a parameroid lobe and spiculum gastrale in dorsal view (right). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113561" httpUri="https://zenodo.org/record/8113561/files/figure.png" pageId="50" pageNumber="51">2Q</figureCitation>
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,
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<figureCitation id="BFA8DCB6FFCDFFB6FCBC60C1FCE40495" box="[805,843,1503,1529]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetId="figure-13@20.[189,1398,300,1983]" captionTargetPageId="20" captionText="Fig. 3 (Part I) (next page). Habitus of males of species of Cryptolarynx Van Schalkwyk, 1966 (lateral view). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113565" httpUri="https://zenodo.org/record/8113565/files/figure.png" pageId="50" pageNumber="51">3Q</figureCitation>
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,
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<figureCitation id="BFA8DCB6FFCDFFB6FCCE60C1FCD10495" box="[855,894,1503,1529]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetId="figure-12@26.[226,1361,265,2019]" captionTargetPageId="26" captionText="Fig. 4 (Part I) (next page). Head of males of species of Cryptolarynx Van Schalkwyk, 1966. A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113569" httpUri="https://zenodo.org/record/8113569/files/figure.png" pageId="50" pageNumber="51">4Q</figureCitation>
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,
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<figureCitation id="BFA8DCB6FFCDFFB6FC1360C0FC020495" box="[906,941,1502,1529]" captionStart="Fig" captionStartId="31.[189,232,1822,1848]" captionTargetBox="[191,1387,271,1758]" captionTargetId="figure-12@31.[188,1397,262,1759]" captionTargetPageId="31" captionText="Fig. 5 (Part I). Ventrites of males of species of Cryptolarynx Van Schalkwyk, 1966. A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957), with details of a cuticular carina (white arrows). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113573" httpUri="https://zenodo.org/record/8113573/files/figure.png" pageId="50" pageNumber="51">5Q</figureCitation>
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</paragraph>
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<subSubSection id="6F8993B8FFCDFFB6FF24633EFD970799" pageId="50" pageNumber="51" type="diagnosis">
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<paragraph id="272CC033FFCDFFB6FF24633EFE660756" blockId="50.[189,457,1568,1594]" box="[189,457,1568,1594]" pageId="50" pageNumber="51">
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<heading id="7C64775FFFCDFFB6FF24633EFE660756" bold="true" box="[189,457,1568,1594]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
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<emphasis id="15E71C21FFCDFFB6FF24633EFE660756" bold="true" box="[189,457,1568,1594]" pageId="50" pageNumber="51">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="272CC033FFCDFFB6FF246350FD970799" blockId="50.[188,1399,1614,1782]" pageId="50" pageNumber="51">
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<taxonomicName id="E093BBB0FFCDFFB6FF246350FDBE0704" authority="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler, 2023" authorityName="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler" authorityYear="2023" box="[189,529,1614,1640]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="cederbergensis" status="sp. nov.">
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<emphasis id="15E71C21FFCDFFB6FF246350FDBE0704" box="[189,529,1614,1640]" italics="true" pageId="50" pageNumber="51">Cryptolarynx cederbergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0ED4A15AFFCDFFB6FD806351FDDB0705" box="[537,628,1615,1641]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
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is most similar to
|
||
<taxonomicName id="E093BBB0FFCDFFB6FCC96351FC400704" authority="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler, 2023" authorityName="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler" authorityYear="2023" box="[848,1007,1614,1641]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pyrophilus" status="sp. nov.">
|
||
<emphasis id="15E71C21FFCDFFB6FCC96351FC400704" box="[848,1007,1614,1641]" italics="true" pageId="50" pageNumber="51">C. pyrophilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0ED4A15AFFCDFFB6FC6F6351FBFD0705" box="[1014,1106,1615,1641]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
but can be distinguished from it by the width of its forehead, which is less than twice the width of an eye (equal to twice width in
|
||
<taxonomicName id="E093BBB0FFCDFFB6FF25638BFEF407C3" authorityName="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler" authorityYear="2023" box="[188,347,1685,1711]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pyrophilus">
|
||
<emphasis id="15E71C21FFCDFFB6FF25638BFEF407C3" box="[188,347,1685,1711]" italics="true" pageId="50" pageNumber="51">C. pyrophilus</emphasis>
|
||
</taxonomicName>
|
||
), and by the arrangement of setae at the apex of the parameroid lobes (
|
||
<figureCitation id="BFA8DCB6FFCDFFB6FB3B638BFAAE07C3" box="[1186,1281,1685,1711]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetId="figure-13@16.[201,1387,265,2019]" captionTargetPageId="16" captionText="Fig. 2 (Part I) (next page). Male genitalia of species of Cryptolarynx Van Schalkwyk, 1966. Penis in dorsal and lateral view (left), tegmen in dorsal view (middle) and details of apex of a parameroid lobe and spiculum gastrale in dorsal view (right). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113561" httpUri="https://zenodo.org/record/8113561/files/figure.png" pageId="50" pageNumber="51">Fig. 2Q</figureCitation>
|
||
). The two species were found to have interspecific uncorrected p-distances ranging from 20.9% to 23.3% for
|
||
<emphasis id="15E71C21FFCDFFB6FADD63A7FAD807BF" box="[1348,1399,1721,1747]" italics="true" pageId="50" pageNumber="51">COI</emphasis>
|
||
and 5.2% for
|
||
<emphasis id="15E71C21FFCDFFB6FEC463C5FE3F0799" box="[349,400,1755,1781]" italics="true" pageId="50" pageNumber="51">EF1</emphasis>
|
||
(Supp. file 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F8993B8FFCDFFB6FF246203FB2C06E4" pageId="50" pageNumber="51" type="etymology">
|
||
<paragraph id="272CC033FFCDFFB6FF246203FEEB065B" blockId="50.[189,324,1821,1847]" box="[189,324,1821,1847]" pageId="50" pageNumber="51">
|
||
<heading id="7C64775FFFCDFFB6FF246203FEEB065B" bold="true" box="[189,324,1821,1847]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
|
||
<emphasis id="15E71C21FFCDFFB6FF246203FEEB065B" bold="true" box="[189,324,1821,1847]" pageId="50" pageNumber="51">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCDFFB6FF246255FB2C06E4" blockId="50.[189,1399,1867,1929]" pageId="50" pageNumber="51">
|
||
The species name
|
||
<taxonomicName id="E093BBB0FFCDFFB6FE066255FDE00609" authorityName="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler" authorityYear="2023" box="[415,591,1867,1893]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="cederbergensis">
|
||
<emphasis id="15E71C21FFCDFFB6FE066255FDE00609" box="[415,591,1867,1893]" italics="true" pageId="50" pageNumber="51">cederbergensis</emphasis>
|
||
</taxonomicName>
|
||
refers to the origin of this species, the Cederberg mountains in the
|
||
<collectingRegion id="E5570ED1FFCDFFB6FF266271FE7A06E4" box="[191,469,1902,1929]" country="South Africa" name="Western Cape" pageId="50" pageNumber="51">Western Cape province</collectingRegion>
|
||
. The specific epithet is an adjective in the masculine form.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F8993B8FFCCFFB7FF246416FEA1030A" pageId="51" pageNumber="52" type="materials_examined">
|
||
<paragraph id="272CC033FFCCFFB7FF246416FE05004E" blockId="51.[189,426,264,290]" box="[189,426,264,290]" pageId="51" pageNumber="52">
|
||
<heading id="7C64775FFFCCFFB7FF246416FE05004E" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
|
||
<emphasis id="15E71C21FFCCFFB7FF246416FE05004E" bold="true" box="[189,426,264,290]" pageId="51" pageNumber="52">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF736429FDD200B2" blockId="51.[186,1400,311,478]" pageId="51" pageNumber="52">
|
||
<materialsCitation id="97FBCA6EFFCCFFB7FF736429FDD200B2" ID-GBIF-Occurrence="4145879370" collectedFrom="at base of Oxalis obtusa" collectingDate="2018-08-23" collectionCode="SAMC" collectorName="J. Haran" country="South Africa" latitude="-32.469" location="Forest Station" longLatPrecision="72" longitude="19.206" municipality="Cederberg Wilderness Area" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape Province" typeStatus="holotype">
|
||
<emphasis id="15E71C21FFCCFFB7FF736429FEF3003D" bold="true" box="[234,348,311,337]" pageId="51" pageNumber="52">
|
||
<typeStatus id="F8287E91FFCCFFB7FF736429FEF3003D" box="[234,348,311,337]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="5F8480A3FFCCFFB7FF246444FDFC0018" box="[189,595,346,372]" name="South Africa" pageId="51" pageNumber="52">REPUBLIC OF SOUTH AFRICA</collectingCountry>
|
||
•
|
||
<emphasis id="15E71C21FFCCFFB7FDF46445FD2C001B" bold="true" box="[621,643,347,375]" pageId="51" pageNumber="52">
|
||
<specimenCount id="31950BBAFFCCFFB7FDF46445FD2C001B" box="[621,643,347,375]" pageId="51" pageNumber="52" type="male">♂</specimenCount>
|
||
</emphasis>
|
||
; “REPUBLIC OF SOUTH AFRICA.
|
||
<emphasis id="15E71C21FFCCFFB7FBD16444FAC00018" bold="true" box="[1096,1391,346,372]" pageId="51" pageNumber="52">
|
||
<collectingRegion id="E5570ED1FFCCFFB7FBD16444FAC00018" box="[1096,1391,346,372]" country="South Africa" name="Western Cape" pageId="51" pageNumber="52">Western Cape Province</collectingRegion>
|
||
</emphasis>
|
||
, [
|
||
<collectingMunicipality id="C7485A49FFCCFFB7FF5A6463FDA600F4" box="[195,521,381,408]" pageId="51" pageNumber="52">Cederberg Wilderness Area</collectingMunicipality>
|
||
;] Algeria [
|
||
<location id="224C96E8FFCCFFB7FDE56460FC8C00FB" LSID="urn:lsid:plazi:treatment:AF3A7125FFCDFFB0FE676086FCF6031B:224C96E8FFCCFFB7FDE56460FC8C00FB" box="[636,803,381,408]" country="South Africa" latitude="-32.469" longLatPrecision="72" longitude="19.206" municipality="Cederberg Wilderness Area" name="Forest Station" pageId="51" pageNumber="52" stateProvince="Western Cape Province">Forest Station</location>
|
||
]
|
||
<locationDeviation id="369ED911FFCCFFB7FCB66460FC1B00F4" box="[815,948,381,408]" pageId="51" pageNumber="52">
|
||
ca.
|
||
<quantity id="E06B6DD6FFCCFFB7FCCC6460FC3300FB" box="[853,924,381,408]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="51" pageNumber="52" unit="km" value="20.0">20 km</quantity>
|
||
S
|
||
</locationDeviation>
|
||
.
|
||
<date id="532DE6F3FFCCFFB7FC246463FBE000FB" box="[957,1103,381,407]" pageId="51" pageNumber="52" value="2018-08-23">
|
||
<collectingDate id="43691F1BFFCCFFB7FC246463FBE000FB" box="[957,1103,381,407]" pageId="51" pageNumber="52" value="2018-08-23">23.viii.2018</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="8A66A5E5FFCCFFB7FBCE6460FB1800F4" box="[1111,1207,382,408]" pageId="51" pageNumber="52">J. Haran</collectorName>
|
||
leg.” “
|
||
<geoCoordinate id="42A7A6F4FFCCFFB7FA9A6463FAD900F4" box="[1283,1398,381,408]" degrees="32.469" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="55" value="-32.469">32.469° S</geoCoordinate>
|
||
<geoCoordinate id="42A7A6F4FFCCFFB7FF2364BEFE9B00D7" box="[186,308,416,443]" degrees="19.206" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="55" value="19.206">19.206° E</geoCoordinate>
|
||
,
|
||
<collectedFrom id="948F83CEFFCCFFB7FEDA64BFFDC700D7" box="[323,616,417,443]" pageId="51" pageNumber="52">
|
||
at base of
|
||
<taxonomicName id="E093BBB0FFCCFFB7FE5D64BFFDC700D7" box="[452,616,417,443]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="species" species="obtusa">
|
||
<emphasis id="15E71C21FFCCFFB7FE5D64BFFDC700D7" box="[452,616,417,443]" italics="true" pageId="51" pageNumber="52">Oxalis obtusa</emphasis>
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
. JHAR01422_0101. Cirad-CBGP coll.” “Holotype.
|
||
<emphasis id="15E71C21FFCCFFB7FB4564BFFEDA00B2" italics="true" pageId="51" pageNumber="52">
|
||
<taxonomicName id="E093BBB0FFCCFFB7FB4564BFFEDE00B2" authorityName="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="cederbergensis">Cryptolarynx cederbergensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Haran 2023”;
|
||
<collectionCode id="418258F6FFCCFFB7FDBB64DAFDD600B2" box="[546,633,452,478]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="51" pageNumber="52" type="Museum">SAMC</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF73671BFEA1030A" blockId="51.[189,1398,517,614]" pageId="51" pageNumber="52">
|
||
<materialsCitation id="97FBCA6EFFCCFFB7FF73671BFEA5030A" ID-GBIF-Occurrence="4145879374" collectedFrom="at base of Oxalis obtusa" collectingDate="2018-08-23" collectionCode="CBGP" collectorName="J. Haran" country="South Africa" latitude="-32.469" location="Western Cape" longLatPrecision="72" longitude="19.206" municipality="Cederberg Wilderness Area" pageId="51" pageNumber="52" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
|
||
<emphasis id="15E71C21FFCCFFB7FF73671BFEC80373" bold="true" box="[234,359,517,543]" pageId="51" pageNumber="52">
|
||
<typeStatus id="F8287E91FFCCFFB7FF73671BFEC80373" box="[234,359,517,543]" pageId="51" pageNumber="52" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="5F8480A3FFCCFFB7FF246737FDFE032F" box="[189,593,553,579]" name="South Africa" pageId="51" pageNumber="52">REPUBLIC OF SOUTH AFRICA</collectingCountry>
|
||
–
|
||
<emphasis id="15E71C21FFCCFFB7FDF66737FC8F032E" bold="true" box="[623,800,552,579]" pageId="51" pageNumber="52">
|
||
<collectingRegion id="E5570ED1FFCCFFB7FDF66737FC8F032E" box="[623,800,552,579]" country="South Africa" name="Western Cape" pageId="51" pageNumber="52">Western Cape</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="31950BBAFFCCFFB7FCA06737FCC7032F" box="[825,872,553,579]" pageId="51" pageNumber="52" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="31950BBAFFCCFFB7FCED6737FC17032F" box="[884,952,553,579]" pageId="51" pageNumber="52" type="female">2 ♀♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="418258F6FFCCFFB7FF246752FEA5030A" box="[189,266,588,614]" pageId="51" pageNumber="52">CBGP</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F8993B8FFCCFFB0FF246793FDB10018" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
|
||
<paragraph id="272CC033FFCCFFB7FF246793FED103CB" blockId="51.[189,382,653,679]" box="[189,382,653,679]" pageId="51" pageNumber="52">
|
||
<emphasis id="15E71C21FFCCFFB7FF246793FEE103CB" bold="true" box="[189,334,653,679]" pageId="51" pageNumber="52">Description</emphasis>
|
||
(
|
||
<specimenCount id="31950BBAFFCCFFB7FEC66793FED903CB" box="[351,374,653,679]" pageId="51" pageNumber="52" type="male">♂</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF2467A2FD0603BA" blockId="51.[189,681,700,726]" box="[189,681,700,726]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF2467A2FEC103B8" baselines="719,719" box="[189,366,700,726]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="51" pageNumber="52">MEASUREMENTS</smallCapsWord>
|
||
. Body length
|
||
<quantity id="E06B6DD6FFCCFFB7FD8C67A2FD0A03BA" box="[533,677,700,726]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="51" pageNumber="52" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF2467E3FBD90286" blockId="51.[187,1400,765,1003]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF2467E3FEB7027A" baselines="784,785" box="[189,280,765,791]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="51" pageNumber="52">COLOUR</smallCapsWord>
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FE87661DFEE4027A" baselines="785" box="[286,331,771,790]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="51" pageNumber="52">AND</smallCapsWord>
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FEC8661DFE6D027A" baselines="785" box="[337,450,771,790]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="vestiture" pageId="51" pageNumber="52">VESTITURE</smallCapsWord>
|
||
. Body integument black, basal half of scapes reddish in mature specimens. Dorsal vestiture (pronotum + elytra) consisting of overlapping, recumbent, parallel-sided clothing scales, 2× as long as wide, mostly rounded at apex; colour of scales white or pale brown to dark brown; white scales usually concentrated in a pair of longitudinal lateral stripes on pronotum and elytral interstriae 4, creating a broad darker stripe medially on pronotum and basal ⅔ of elytra; white scales also concentrated in a pair of ill-defined pale spots surrounded by black scales at apical ⅔ of interstria 3; scales of striae recumbent, in lateral view not or only very slightly distinct from rest of vestiture.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF24610CFDDD044E" blockId="51.[188,1400,1042,1315]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF24610CFF530546" baselines="1061,1061" box="[189,252,1042,1068]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="51" pageNumber="52">HEAD</smallCapsWord>
|
||
. Forehead very wide, slightly wider than epifrons near antennal insertions, ca 2 × as wide as width of an eye, scales suberect. Eyes convex, in dorsal view slightly exceeding outline of head, surrounded by a ring of short pale scales, on forehead directed towards occiput; distance between eye and scrobe as large as width of antennal club. Epifrons with distance between antennal insertions 0.8× length of scape, scales at least 3 × as long as wide, suberect, contiguous. Frons with a pair of long erect lateral setae. Epistome with one or two elongate median setae. Antennal funicles with segment 1 elongate; 2 shorter, at most 1.5× as long as wide; 3 longer than wide; 4 globular, isodiametric, compressed, slightly angular on inside; 5–7 globular, isodiametric.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF246054FE3A04EB" blockId="51.[189,1398,1353,1415]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF246054FE99040E" baselines="1373,1373" box="[189,310,1354,1380]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pronotum" pageId="51" pageNumber="52">PRONOTUM</smallCapsWord>
|
||
. Transverse (W:L ratio 1.6), widest posteriorly of midlength, sides arcuate; apex and base subequal in width.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF2460B1FB6704A4" blockId="51.[189,1224,1454,1481]" box="[189,1224,1454,1481]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF2460B1FEBD04AB" baselines="1474,1474" box="[189,274,1455,1481]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Elytra" pageId="51" pageNumber="52">ELYTRA</smallCapsWord>
|
||
. Broadly ovate, isodiametric (W:L ratio 1), sides convex, widest near midlength.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF2460EEFE96073C" blockId="51.[189,1400,1520,1616]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF2460EEFF590764" baselines="1539,1539" box="[189,246,1520,1546]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="51" pageNumber="52">LEGS</smallCapsWord>
|
||
. Tibiae with apical mucro; protibiae with both outer and inner margins straight; metatibiae with inner setal fringe, the setae shorter than segment 5 of metatarsus. Tarsi with segment 2 slightly longer than wide.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF246366FB5307D9" blockId="51.[189,1398,1655,1717]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF246366FE8307FC" baselines="1675,1675" box="[189,300,1656,1682]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="51" pageNumber="52">ABDOMEN</smallCapsWord>
|
||
. Ventrites with creamy-white plumose scales not concealing integument; ventrites 1–4 with long suberect scales; ventrite 5 devoid of scales in apical half, there bearing only erect setae.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCCFFB7FF2463C3FD3C06EF" blockId="51.[186,1399,1756,1923]" pageId="51" pageNumber="52">
|
||
<smallCapsWord id="21CA56EFFFCCFFB7FF2463C3FEEC0799" baselines="1776,1776" box="[189,323,1757,1783]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Terminalia" pageId="51" pageNumber="52">TERMINALIA</smallCapsWord>
|
||
. Body of penis elongate (W:L ratio 0.3–0.4), as long as temones, sides subparallel, converging in apical third; curvature in profile weak and regular, dorsoventrally narrowed before apex. Copulatory sclerite forming a reversed V. Parameroid lobes separate, divided by deep median notch, each lobe broad, bearing a series of long setae directed apicad. Spiculum gastrale with basal arms long, regularly curved, right arm angulate near its base.
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCBFFB0FF246416FE1F004E" blockId="52.[189,432,264,290]" box="[189,432,264,290]" pageId="52" pageNumber="53">
|
||
<heading id="7C64775FFFCBFFB0FF246416FE1F004E" bold="true" box="[189,432,264,290]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
|
||
<emphasis id="15E71C21FFCBFFB0FF246416FE1F004E" bold="true" box="[189,432,264,290]" pageId="52" pageNumber="53">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCBFFB0FF246429FDB10018" blockId="52.[189,1399,311,373]" pageId="52" pageNumber="53">The sexes can be distinguished by the width of the forehead (as wide as or narrower than width of an eye in male, wider in female).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F8993B8FFCBFFB0FF246485FE250364" pageId="52" pageNumber="53" type="biology_ecology">
|
||
<paragraph id="272CC033FFCBFFB0FF246485FEE000D9" blockId="52.[189,335,411,437]" box="[189,335,411,437]" pageId="52" pageNumber="53">
|
||
<heading id="7C64775FFFCBFFB0FF246485FEE000D9" bold="true" box="[189,335,411,437]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
|
||
<emphasis id="15E71C21FFCBFFB0FF246485FEE000D9" bold="true" box="[189,335,411,437]" pageId="52" pageNumber="53">Life history</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCBFFB0FF2464D4FE250364" blockId="52.[189,1399,458,520]" pageId="52" pageNumber="53">
|
||
Specimens of
|
||
<taxonomicName id="E093BBB0FFCBFFB0FEFA64D5FD990088" authority="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler, 2023" authorityName="Haran & Marvaldi & Benoit & Oberlander & Stals & Oberprieler" authorityYear="2023" box="[355,566,458,485]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="cederbergensis" status="sp. nov.">
|
||
<emphasis id="15E71C21FFCBFFB0FEFA64D5FD990088" box="[355,566,458,485]" italics="true" pageId="52" pageNumber="53">C. cederbergensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0ED4A15AFFCBFFB0FDA464D5FD370089" box="[573,664,459,485]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were collected in monospecific stands of
|
||
<taxonomicName id="E093BBB0FFCBFFB0FB1A64D4FA8A0088" box="[1155,1317,458,484]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="obtusa">
|
||
<emphasis id="15E71C21FFCBFFB0FB1A64D4FA8A0088" box="[1155,1317,458,484]" italics="true" pageId="52" pageNumber="53">Oxalis obtusa</emphasis>
|
||
</taxonomicName>
|
||
, in the month of August.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F8993B8FFCBFFB0FF246731FCF6031B" box="[189,857,559,632]" pageId="52" pageNumber="53" type="distribution">
|
||
<paragraph id="272CC033FFCBFFB0FF246731FEF90325" blockId="52.[189,342,559,585]" box="[189,342,559,585]" pageId="52" pageNumber="53">
|
||
<heading id="7C64775FFFCBFFB0FF246731FEF90325" bold="true" box="[189,342,559,585]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
|
||
<emphasis id="15E71C21FFCBFFB0FF246731FEF90325" bold="true" box="[189,342,559,585]" pageId="52" pageNumber="53">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="272CC033FFCBFFB0FF246743FCF6031B" blockId="52.[189,857,605,632]" box="[189,857,605,632]" pageId="52" pageNumber="53">
|
||
The species was found only at the
|
||
<typeStatus id="F8287E91FFCBFFB0FDCB6740FD2A0314" box="[594,645,606,632]" pageId="52" pageNumber="53">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="BFA8DCB6FFCBFFB0FD6D6743FCE4031B" box="[756,843,605,631]" captionStart="Fig" captionStartId="77.[189,232,1963,1989]" captionTargetBox="[366,1201,265,1898]" captionTargetId="figure-11@77.[361,1226,265,1922]" captionTargetPageId="77" captionText="Fig. 13. Distribution maps of species of Cryptolarynx Van Schalkwyk, 1966 and Hadrocryptolarynx Haran gen. nov." figureDoi="http://doi.org/10.5281/zenodo.8110596" httpUri="https://zenodo.org/record/8110596/files/figure.png" pageId="52" pageNumber="53">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |