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<document id="DD204FDEDE5AEBF5BD0341E5D27CC8F4" ID-CLB-Dataset="57636" ID-DOI="10.5852/ejt.2023.877.2151" ID-GBIF-Dataset="74111f75-19b1-4944-b6d0-09db245266fd" ID-ISSN="2118-9773" ID-Zenodo-Dep="8110586" ID-ZooBank="65E8C3F7-5EA4-4013-A09E-37C96B4929FB" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1688395635639" checkinUser="felipe" docAuthor="Haran, Julien M., Marvaldi, Adriana E., Benoit, Laure, Oberlander, Kenneth, Stals, Riaan &amp; Oberprieler, Rolf G." docDate="2023" docId="AF3A7125FFCAFFB3FD876018FCC1003D" docLanguage="en" docName="ejt-877_1-89.pdf" docOrigin="European Journal of Taxonomy 877 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.877.2151" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cryptolarynx marshalli Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler 2023, sp. nov." docType="treatment" docUuid="B4281E9A-F0FF-42D4-8099-9ADA5005C6E5" docUuidSource="ZooBank" docVersion="12" lastPageNumber="56" masterDocId="5303095DFFFFFF84FF99651EFFAF016C" masterDocTitle="Revision of the enigmatic South African Cryptolaryngini (Coleoptera, Curculionidae), with description of a new genus and twenty-two new species" masterLastPageNumber="89" masterPageNumber="1" pageNumber="54" updateTime="1699533238137" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FEF650BD674ABC57DEC50A2F13DBA38C">Revision of the enigmatic South African Cryptolaryngini (Coleoptera, Curculionidae), with description of a new genus and twenty-two new species</mods:title>
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<mods:namePart id="33F58EA39AE16392A7BF9C65015C1580">Haran, Julien M.</mods:namePart>
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<mods:affiliation id="C175A4F466B2CE1D00F018753ADE1BD1">División Entomología, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, Buenos Aires, Argentina.</mods:affiliation>
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<mods:affiliation id="59E4F7C96B15442B845C9A9C27F4B864">Montpellier SupAgro, INRAe, IRD, Montpellier University, Montpellier, France. &amp; Laure.</mods:affiliation>
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<mods:affiliation id="F09738775CE3E1B4753A4AEA5FA114D3">Schweickerdt Herbarium, Department of Plant and Soil Sciences, Plant Sciences Complex, University of Pretoria, Private Bag X 20, Hatfield 0024, South Africa. &amp; Kenneth.</mods:affiliation>
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<mods:affiliation id="EDC3127491BABF4A6C5488DD4D89089C">Australian National Insect Collection, G. P. O. Box 1700, Canberra, A. C. T. 2601, Australia.</mods:affiliation>
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19. 
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<emphasis id="15E71C21FFCAFFB1FDD3601BFCCA044C" bold="true" box="[586,869,1285,1312]" italics="true" pageId="53" pageNumber="54">Cryptolarynx marshalli</emphasis>
Haran
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<taxonomicNameLabel id="0ED4A15AFFCAFFB1FC226018FBB9044C" box="[955,1046,1286,1312]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="BFA8DCB6FFCAFFB1FDF46053FD68040B" box="[621,711,1357,1383]" captionStart="Fig" captionStartId="9.[190,233,1845,1871]" captionTargetId="figure-13@10.[308,1280,265,2019]" captionTargetPageId="10" captionText="Fig. 1 (Part I) (next page). Habitus of males of species of Cryptolarynx Van Schalkwyk, 1966 (dorsal view). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113555" httpUri="https://zenodo.org/record/8113555/files/figure.png" pageId="53" pageNumber="54">Figs 1S</figureCitation>
, 
<figureCitation id="BFA8DCB6FFCAFFB1FD4D6053FD59040B" box="[724,758,1357,1383]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetId="figure-13@16.[201,1387,265,2019]" captionTargetPageId="16" captionText="Fig. 2 (Part I) (next page). Male genitalia of species of Cryptolarynx Van Schalkwyk, 1966. Penis in dorsal and lateral view (left), tegmen in dorsal view (middle) and details of apex of a parameroid lobe and spiculum gastrale in dorsal view (right). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113561" httpUri="https://zenodo.org/record/8113561/files/figure.png" pageId="53" pageNumber="54">2S</figureCitation>
, 
<figureCitation id="BFA8DCB6FFCAFFB1FC9B6053FC8C040B" box="[770,803,1357,1383]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetId="figure-13@20.[189,1398,300,1983]" captionTargetPageId="20" captionText="Fig. 3 (Part I) (next page). Habitus of males of species of Cryptolarynx Van Schalkwyk, 1966 (lateral view). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113565" httpUri="https://zenodo.org/record/8113565/files/figure.png" pageId="53" pageNumber="54">3S</figureCitation>
, 
<figureCitation id="BFA8DCB6FFCAFFB1FCB66053FCFF040B" box="[815,848,1357,1383]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetId="figure-12@26.[226,1361,265,2019]" captionTargetPageId="26" captionText="Fig. 4 (Part I) (next page). Head of males of species of Cryptolarynx Van Schalkwyk, 1966. A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113569" httpUri="https://zenodo.org/record/8113569/files/figure.png" pageId="53" pageNumber="54">4S</figureCitation>
, 
<figureCitation id="BFA8DCB6FFCAFFB1FCC46052FCD1040B" box="[861,894,1356,1383]" captionStart="Fig" captionStartId="31.[189,232,1822,1848]" captionTargetBox="[191,1387,271,1758]" captionTargetId="figure-12@31.[188,1397,262,1759]" captionTargetPageId="31" captionText="Fig. 5 (Part I). Ventrites of males of species of Cryptolarynx Van Schalkwyk, 1966. A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957), with details of a cuticular carina (white arrows). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113573" httpUri="https://zenodo.org/record/8113573/files/figure.png" pageId="53" pageNumber="54">5S</figureCitation>
, 
<figureCitation id="BFA8DCB6FFCAFFB1FC126053FC69040B" box="[907,966,1357,1383]" captionStart="Fig" captionStartId="40.[189,232,1431,1457]" captionTargetId="figure-11@39.[189,1398,265,1901]" captionTargetPageId="39" captionText="Fig. 7 (previous page). Habitus in natura, host plants and biotopes of species of Cryptolarynx Van Schalkwyk, 1966 Cryptolarynx and Hadrocryptolarynx Haran gen. nov. (part 2). A. Habitus of C. oberprieleri Haran sp. nov. on and at base of Oxalis glabra Thunb., Stellenbosch (JHAR01201). B. Patch of O. glabra, biotope of C. oberprieleri Haran sp. nov., Stellenbosch (JHAR01485). C. Burnt area near Montagu with patches of Oxalis pes-caprae L. resprouting; biotope of C. pyrophilus Haran sp. nov. (JHAR01528). D–F. Habitus, Oxalis cf. odorata J.C.Manning &amp; Goldblatt, host and biotope of C. spinicornis Haran sp. nov., Sutherland (JHAR02512). G. Habitus of C. marshalli Haran sp. nov., near Worcester (JHAR02355). H. Oxalis imbricata Eckl. &amp; Zeyh., hosting adults of C. marshalli Haran sp. nov., near Worcester. I. Shale Renosterveld vegetation type on hillsides at Worcester, biotope of C. marshalli Haran sp. nov. (JHAR02355). J. Habitus of C. oberlanderi Haran sp. nov., near Worcester (JHAR02353). K. Oxalis depressa Eckl. &amp; Zeyh., hosting adults of C. oberlanderi Haran sp. nov., near Worcester. L. Habitus of C. san Haran sp. nov., Langebaan Dune Strandveld vegetation type near West Coast National Park (JHAR02484). M. Habitus of male of Hadrocryptolarynx major Haran gen. et sp. nov., Klawer (JHAR02464). N. Habitus of female of Hadrocryptolarynx major Haran gen. et sp. nov., Klawer. O. Gallery formed by Hadrocryptolarynx major Haran gen. et sp. nov. under a leaf of Amaryllidaceae J.St.-Hil. P. Vanrhynsdorp Gannabosveld vegetation type near Vanrhynsdorp, biotope of Hadrocryptolarynx major Haran gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8113714" httpUri="https://zenodo.org/record/8113714/files/figure.png" pageId="53" pageNumber="54">7G–I</figureCitation>
</paragraph>
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<paragraph id="272CC033FFCAFFB1FF246090FE6604C4" blockId="53.[189,457,1422,1448]" box="[189,457,1422,1448]" pageId="53" pageNumber="54">
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<emphasis id="15E71C21FFCAFFB1FF246090FE6604C4" bold="true" box="[189,457,1422,1448]" pageId="53" pageNumber="54">Differential diagnosis</emphasis>
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<taxonomicName id="E093BBB0FFCAFFB1FF2460A3FE6004BB" authority="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler, 2023" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[189,463,1469,1495]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="marshalli" status="sp. nov.">
<emphasis id="15E71C21FFCAFFB1FF2460A3FE6004BB" box="[189,463,1469,1495]" italics="true" pageId="53" pageNumber="54">Cryptolarynx marshalli</emphasis>
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<taxonomicNameLabel id="0ED4A15AFFCAFFB1FE4F60A0FD8004B4" box="[470,559,1470,1496]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
is closely related to 
<taxonomicName id="E093BBB0FFCAFFB1FC8E60A0FC6804BB" authority="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler, 2023" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[791,967,1469,1496]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="oberprieleri" status="sp. nov.">
<emphasis id="15E71C21FFCAFFB1FC8E60A0FC6804BB" box="[791,967,1469,1496]" italics="true" pageId="53" pageNumber="54">C. oberprieleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0ED4A15AFFCAFFB1FC5460A0FB8904B4" box="[973,1062,1470,1496]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
but distinctly more elongate, and the apex of its parameroid lobes is also distinct (
<figureCitation id="BFA8DCB6FFCAFFB1FC8F60FEFC3C0497" box="[790,915,1504,1531]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetId="figure-13@16.[201,1387,265,2019]" captionTargetPageId="16" captionText="Fig. 2 (Part I) (next page). Male genitalia of species of Cryptolarynx Van Schalkwyk, 1966. Penis in dorsal and lateral view (left), tegmen in dorsal view (middle) and details of apex of a parameroid lobe and spiculum gastrale in dorsal view (right). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113561" httpUri="https://zenodo.org/record/8113561/files/figure.png" pageId="53" pageNumber="54">Fig. 2O–S</figureCitation>
). See Differential diagnosis section under species 
<taxonomicName id="E093BBB0FFCAFFB1FE80631AFE660772" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[281,457,1540,1566]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="oberprieleri">
<emphasis id="15E71C21FFCAFFB1FE80631AFE660772" box="[281,457,1540,1566]" italics="true" pageId="53" pageNumber="54">C. oberprieleri</emphasis>
</taxonomicName>
for the genetic distances between these species.
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFCAFFB1FF246358FDB707BA" pageId="53" pageNumber="54" type="etymology">
<paragraph id="272CC033FFCAFFB1FF246358FEEB070C" blockId="53.[189,324,1606,1632]" box="[189,324,1606,1632]" pageId="53" pageNumber="54">
<heading id="7C64775FFFCAFFB1FF246358FEEB070C" bold="true" box="[189,324,1606,1632]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="15E71C21FFCAFFB1FF246358FEEB070C" bold="true" box="[189,324,1606,1632]" pageId="53" pageNumber="54">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFCAFFB1FF24636AFDB707BA" blockId="53.[189,1399,1652,1750]" pageId="53" pageNumber="54">This species is dedicated to the late weevil expert Sir Guy A.K. Marshall, who described the genus and its original two species and discussed its unique characters among known weevils. The specific epithet is a noun in the genitive case.</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFCAFFB2FF2463E3FEA10005" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="272CC033FFCAFFB1FF2463E3FE05067B" blockId="53.[189,426,1789,1815]" box="[189,426,1789,1815]" pageId="53" pageNumber="54">
<heading id="7C64775FFFCAFFB1FF2463E3FE05067B" bold="true" box="[189,426,1789,1815]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="15E71C21FFCAFFB1FF2463E3FE05067B" bold="true" box="[189,426,1789,1815]" pageId="53" pageNumber="54">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFCAFFB1FF736232FBE006DD" blockId="53.[185,1399,1836,1969]" pageId="53" pageNumber="54">
<materialsCitation id="97FBCA6EFFCAFFB1FF736232FBE006DD" ID-GBIF-Occurrence="4145879361" collectedFrom="at base of Oxalis imbricata" collectingDate="2019-07-03" collectionCode="SAMC" collectorName="J. Haran" country="South Africa" latitude="-33.613" location="Worcester" longLatPrecision="72" longitude="19.447" municipality="Worcester" pageId="53" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape Province" typeStatus="holotype">
<emphasis id="15E71C21FFCAFFB1FF736232FEF3062A" bold="true" box="[234,348,1836,1862]" pageId="53" pageNumber="54">
<typeStatus id="F8287E91FFCAFFB1FF736232FEF3062A" box="[234,348,1836,1862]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
</emphasis>
REPUBLIC OF SOUTH AFRICA • 
<emphasis id="15E71C21FFCAFFB1FDF4624EFD2C0600" bold="true" box="[621,643,1872,1900]" pageId="53" pageNumber="54">
<specimenCount id="31950BBAFFCAFFB1FDF4624EFD2C0600" box="[621,643,1872,1900]" pageId="53" pageNumber="54" type="male">♂</specimenCount>
</emphasis>
; “ 
<collectingCountry id="5F8480A3FFCAFFB1FD07624EFB930606" box="[670,1084,1872,1898]" name="South Africa" pageId="53" pageNumber="54">REPUBLIC OF SOUTH AFRICA</collectingCountry>
. 
<emphasis id="15E71C21FFCAFFB1FBD1624EFAC00605" bold="true" box="[1096,1391,1871,1898]" pageId="53" pageNumber="54">
<collectingRegion id="E5570ED1FFCAFFB1FBD1624EFAC00605" box="[1096,1391,1871,1898]" country="South Africa" name="Western Cape" pageId="53" pageNumber="54">Western Cape Province</collectingRegion>
</emphasis>
, 
<collectingMunicipality id="C7485A49FFCAFFB1FF24626DFE9906E1" box="[189,310,1907,1933]" pageId="53" pageNumber="54">Worcester</collectingMunicipality>
. 
<date id="532DE6F3FFCAFFB1FEA5626DFE1806E1" box="[316,439,1907,1933]" pageId="53" pageNumber="54" value="2019-07-03">
<collectingDate id="43691F1BFFCAFFB1FEA5626DFE1806E1" box="[316,439,1907,1933]" pageId="53" pageNumber="54" value="2019-07-03">3.vii.2019</collectingDate>
</date>
. 
<collectorName id="8A66A5E5FFCAFFB1FE27626DFDB206E1" box="[446,541,1907,1933]" pageId="53" pageNumber="54">J. Haran</collectorName>
leg.” “ 
<geoCoordinate id="42A7A6F4FFCAFFB1FDFF626DFD7606E1" box="[614,729,1907,1933]" degrees="33.613" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="55" value="-33.613">33.613° S</geoCoordinate>
<geoCoordinate id="42A7A6F4FFCAFFB1FD44626DFCFD06E1" box="[733,850,1907,1933]" degrees="19.447" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="55" value="19.447">19.447° E</geoCoordinate>
. 
<collectedFrom id="948F83CEFFCAFFB1FCC0626DFB2706E1" box="[857,1160,1907,1933]" pageId="53" pageNumber="54">
at base of 
<taxonomicName id="E093BBB0FFCAFFB1FC5F626DFB2706E1" box="[966,1160,1907,1933]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="imbricata">
<emphasis id="15E71C21FFCAFFB1FC5F626DFB2706E1" box="[966,1160,1907,1933]" italics="true" pageId="53" pageNumber="54">Oxalis imbricata</emphasis>
</taxonomicName>
</collectedFrom>
. JHAR02355_0101. Cirad-CBGP coll.” “Holotype. 
<taxonomicName id="E093BBB0FFCAFFB1FDB26288FC9006DC" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[555,831,1942,1968]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="15E71C21FFCAFFB1FDB26288FC9006DC" box="[555,831,1942,1968]" italics="true" pageId="53" pageNumber="54">Cryptolarynx marshalli</emphasis>
</taxonomicName>
. Haran 2023”; 
<collectionCode id="418258F6FFCAFFB1FC6D6289FBE406DD" box="[1012,1099,1943,1969]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="53" pageNumber="54" type="Museum">SAMC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="272CC033FFC9FFB2FF736417FEA10005" blockId="54.[189,1398,265,361]" pageId="54" pageNumber="55">
<materialsCitation id="97FBCA6EFFC9FFB2FF736417FEA50005" ID-GBIF-Occurrence="4145879438" collectedFrom="at base of Oxalis imbricata" collectingDate="2019-07-03" collectionCode="CBGP" collectorName="J. Haran" country="South Africa" latitude="-33.613" location="Western Cape" longLatPrecision="72" longitude="19.447" municipality="Worcester" pageId="54" pageNumber="55" specimenCount="3" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<emphasis id="15E71C21FFC9FFB2FF736417FEC8004F" bold="true" box="[234,359,265,291]" pageId="54" pageNumber="55">
<typeStatus id="F8287E91FFC9FFB2FF736417FEC8004F" box="[234,359,265,291]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="5F8480A3FFC9FFB2FF246432FDE1002A" box="[189,590,300,326]" name="South Africa" pageId="54" pageNumber="55">REPUBLIC OF SOUTH AFRICA</collectingCountry>
– 
<emphasis id="15E71C21FFC9FFB2FDF16432FCB70029" bold="true" box="[616,792,299,326]" pageId="54" pageNumber="55">
<collectingRegion id="E5570ED1FFC9FFB2FDF16432FCB70029" box="[616,792,299,326]" country="South Africa" name="Western Cape" pageId="54" pageNumber="55">Western Cape</collectingRegion>
</emphasis>
• 
<specimenCount id="31950BBAFFC9FFB2FCA96432FCF2002A" box="[816,861,300,326]" pageId="54" pageNumber="55" type="male">1 ♂</specimenCount>
, 
<specimenCount id="31950BBAFFC9FFB2FCF06432FCD7002A" box="[873,888,300,326]" pageId="54" pageNumber="55">2</specimenCount>
specs; same collection data as for holotype; 
<collectionCode id="418258F6FFC9FFB2FF246451FEA50005" box="[189,266,335,361]" pageId="54" pageNumber="55">CBGP</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFC9FFB2FF24648EFE590651" pageId="54" pageNumber="55" type="description">
<paragraph id="272CC033FFC9FFB2FF24648EFED100C6" blockId="54.[189,382,400,426]" box="[189,382,400,426]" pageId="54" pageNumber="55">
<emphasis id="15E71C21FFC9FFB2FF24648EFEE100C6" bold="true" box="[189,334,400,426]" pageId="54" pageNumber="55">Description</emphasis>
(
<specimenCount id="31950BBAFFC9FFB2FEC6648EFED900C6" box="[351,374,400,426]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
)
</paragraph>
<paragraph id="272CC033FFC9FFB2FF2464A1FD0600B5" blockId="54.[189,681,447,474]" box="[189,681,447,474]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF2464A1FEC100BB" baselines="466,466" box="[189,366,447,473]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="54" pageNumber="55">MEASUREMENTS</smallCapsWord>
. Body length 
<quantity id="E06B6DD6FFC9FFB2FD8C64A1FD0A00B5" box="[533,677,447,474]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="54" pageNumber="55" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF24671EFC9303A6" blockId="54.[188,1399,512,715]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF24671EFEB70375" baselines="531,532" box="[189,280,512,538]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="54" pageNumber="55">COLOUR</smallCapsWord>
<smallCapsWord id="21CA56EFFFC9FFB2FEBD6718FEFE0375" baselines="532" box="[292,337,518,537]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="54" pageNumber="55">AND</smallCapsWord>
<smallCapsWord id="21CA56EFFFC9FFB2FEC46718FE610375" baselines="532" box="[349,462,518,537]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="vestiture" pageId="54" pageNumber="55">VESTITURE</smallCapsWord>
. Body integument black, antennae, tibiae and tarsi reddish. Dorsal vestiture (pronotum + elytra) consisting of overlapping, recumbent, parallel-sided clothing scales, 3× as long as wide, truncate at apex; colour of scales mostly brown, white scales interspersed with pale brown scales concentrated in two longitudinal bands on pronotum, at base of elytral interstriae 4, and in a pair of pale spots surrounded by black scales at apical ⅔ of interstriae 3; scales of striae recumbent, in lateral view not or only very slightly distinct from rest of vestiture.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF2467ECFF56056E" blockId="54.[187,1400,754,1026]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF2467ECFF530266" baselines="773,773" box="[189,252,754,780]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="54" pageNumber="55">HEAD</smallCapsWord>
. Forehead wide, slightly wider than epifrons near antennal insertions, scales suberect. Eyes convex, in dorsal view slightly exceeding outline of head, surrounded by a ring of short scales, on forehead directed towards occiput; distance between eye and scrobe smaller than width of antennal club. Epifrons with distance between antennal insertions slightly smaller than length of scape, scales at least 2 × as long as wide, recumbent, mostly non-contiguous. Frons with 3 pairs of erect lateral setae. Epistome without median seta. Antennal funicles with segment 1 moderately elongate, 1.5 longer than wide; 2 subequal in length to 1; 2 
<figureCitation id="BFA8DCB6FFC9FFB2FE1B66DBFE6C02B3" box="[386,451,965,991]" captionStart="and 4" captionStartId="46.[189,231,1777,1803]" captionTargetBox="[195,1393,274,1677]" captionTargetId="figure-13@46.[189,1398,265,1692]" captionText="and 4. I. Protibia of C. armatus Haran sp. nov., arrow pointing at inner preapical expansion proximally of mucro. J. Penis of C. oberprieleri Haran sp. nov., dorsal view, black arrows indicating length of penis body and temones and white arrows indicating width of penis body as measured; bottom arrow indicating copulatory sclerite. K. Ovipositor, Crypolarynx variabilis Haran sp. nov. L. Sternite VIII, same species. M. Spermatheca, same species. N. Ovipositor, Hadrocryptolarynx major Haran gen. et sp. nov. O. Sternite VIII, same species. P. Spermatheca, same species. Q. Right maxilla, same species. Not to scale." pageId="54" pageNumber="55">and 4</figureCitation>
compressed, slightly angular on inside; 5–7 globular, isodiametric or wider than long.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF246134FE39050B" blockId="54.[189,1398,1065,1127]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF246134FE99052E" baselines="1085,1085" box="[189,310,1066,1092]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pronotum" pageId="54" pageNumber="55">PRONOTUM</smallCapsWord>
. Moderately transverse (W:L ratio 1.2), widest near midlength, sides arcuate; apex and base subequal in width.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF246191FAD805C4" blockId="54.[189,1399,1166,1193]" box="[189,1399,1166,1193]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF246191FEBD05CB" baselines="1186,1186" box="[189,274,1167,1193]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Elytra" pageId="54" pageNumber="55">ELYTRA</smallCapsWord>
. Bullet-shaped, longer than wide (W:L ratio 0.85), sides convex, widest anteriorly of midlength.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF2561CEFE53045C" blockId="54.[188,1398,1232,1329]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF2561CEFF590584" baselines="1251,1251" box="[188,246,1232,1258]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="54" pageNumber="55">LEGS</smallCapsWord>
. Slender. Tibiae with apical mucro; protibiae with outer margin straight, inner margin slightly bisinuate; metatibiae with inner setal fringe, setae shorter than segment 5 of metatarsus. Tarsi with segment 2 wider than long.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF246046FD4604B7" blockId="54.[189,1398,1367,1500]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF246046FE83041C" baselines="1387,1387" box="[189,300,1368,1394]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="54" pageNumber="55">ABDOMEN</smallCapsWord>
. Ventrite with creamy-white plumose scales not fully concealing integument, scales on ventrites 2–5 medially intermixed with long suberect setae, apically bifid or not; ventrite 1 slightly concave medially, impression covered with long setae deeply divided from their bases; ventrite 5 with scales concentrated laterally and on basal third.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF24631DFB3A07C5" blockId="54.[188,1400,1538,1706]" pageId="54" pageNumber="55">
<smallCapsWord id="21CA56EFFFC9FFB2FF24631DFEEC0777" baselines="1558,1558" box="[189,323,1539,1565]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Terminalia" pageId="54" pageNumber="55">TERMINALIA</smallCapsWord>
. Body of penis moderately elongate (W:L ratio 0.45), slightly shorter than temones, sides convex; curvature in profile weak, stronger in basal half, not dorsoventrally narrowed at apex. Copulatory sclerite weakly sclerotised or not discerned in examined specimens. Parameroid lobes separate, divided by modest median notch, each lobe broad, bearing a series of setae directed apicad, median setae longer. Spiculum gastrale with basal arms long, right arm slightly angulate at its midlength.
</paragraph>
<paragraph id="272CC033FFC9FFB2FF2463CEFE1F0786" blockId="54.[189,432,1744,1770]" box="[189,432,1744,1770]" pageId="54" pageNumber="55">
<heading id="7C64775FFFC9FFB2FF2463CEFE1F0786" bold="true" box="[189,432,1744,1770]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="15E71C21FFC9FFB2FF2463CEFE1F0786" bold="true" box="[189,432,1744,1770]" pageId="54" pageNumber="55">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFC9FFB2FF2463E1FE590651" blockId="54.[188,1399,1791,1853]" pageId="54" pageNumber="55">
The sexes can be distinguished by the elytra (shorter in male) and by ventrite 
<quantity id="E06B6DD6FFC9FFB2FBDB63E1FBC10675" box="[1090,1134,1791,1817]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="54" pageNumber="55" unit="in" value="1.0">1 in</quantity>
the female lacking the long, deeply divided setae.
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFC9FFB2FF24627AFAD806C0" pageId="54" pageNumber="55" type="biology_ecology">
<paragraph id="272CC033FFC9FFB2FF24627AFEE00612" blockId="54.[189,335,1892,1918]" box="[189,335,1892,1918]" pageId="54" pageNumber="55">
<heading id="7C64775FFFC9FFB2FF24627AFEE00612" bold="true" box="[189,335,1892,1918]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="15E71C21FFC9FFB2FF24627AFEE00612" bold="true" box="[189,335,1892,1918]" pageId="54" pageNumber="55">Life history</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFC9FFB2FF24628CFAD806C0" blockId="54.[189,1399,1938,1965]" box="[189,1399,1938,1965]" pageId="54" pageNumber="55">
All specimens of this species were collected in July at the base of 
<taxonomicName id="E093BBB0FFC9FFB2FC2A628CFA8C06C0" authority="Eckl. &amp; Zeyh." authorityName="Eckl. &amp; Zeyh." box="[947,1315,1938,1965]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="imbricata">
<emphasis id="15E71C21FFC9FFB2FC2A628CFBD706C0" box="[947,1144,1938,1964]" italics="true" pageId="54" pageNumber="55">Oxalis imbricata</emphasis>
Eckl. &amp; Zeyh.
</taxonomicName>
plants.
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFC8FFB3FF246416FCC1003D" box="[189,878,264,338]" pageId="55" pageNumber="56" type="distribution">
<paragraph id="272CC033FFC8FFB3FF246416FEF9004E" blockId="55.[189,342,264,290]" box="[189,342,264,290]" pageId="55" pageNumber="56">
<heading id="7C64775FFFC8FFB3FF246416FEF9004E" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="55" pageNumber="56" reason="3">
<emphasis id="15E71C21FFC8FFB3FF246416FEF9004E" bold="true" box="[189,342,264,290]" pageId="55" pageNumber="56">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFC8FFB3FF246429FCC1003D" blockId="55.[189,878,311,338]" box="[189,878,311,338]" pageId="55" pageNumber="56">
The species is only known from the 
<typeStatus id="F8287E91FFC8FFB3FDFE6426FD35003E" box="[615,666,312,338]" pageId="55" pageNumber="56">type</typeStatus>
locality (
<figureCitation id="BFA8DCB6FFC8FFB3FC906429FCCF003D" box="[777,864,311,337]" captionStart="Fig" captionStartId="77.[189,232,1963,1989]" captionTargetBox="[366,1201,265,1898]" captionTargetId="figure-11@77.[361,1226,265,1922]" captionTargetPageId="77" captionText="Fig. 13. Distribution maps of species of Cryptolarynx Van Schalkwyk, 1966 and Hadrocryptolarynx Haran gen. nov." figureDoi="http://doi.org/10.5281/zenodo.8110596" httpUri="https://zenodo.org/record/8110596/files/figure.png" pageId="55" pageNumber="56">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>