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<document id="8534911DE2F88ADB93800017AFA7E513" ID-CLB-Dataset="25069" ID-DOI="10.5852/ejt.2016.225" ID-GBIF-Dataset="7dda3f39-7ee4-40be-a268-75056e3d2a9a" ID-Zenodo-Dep="830814" ID-ZooBank="C1F4C6FC-2EB6-48D9-B628-F1C8BD1FF2F8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500295631582" checkinUser="jeremy" docAuthor="Huber, Bernhard A., Nuñeza, Olga M. &amp; Leh Moi Ung, Charles" docDate="2016" docId="12428788FF8EFFBFFDB92509A4CE782F" docLanguage="en" docName="ejt-225.pdf.imd" docOrigin="European Journal of Taxonomy 225" docSource="http://www.europeanjournaloftaxonomy.eu/index.php/ejt/article/view/349" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pholcus Walckenaer 1805" docType="treatment" docVersion="13" lastPageNumber="5" masterDocId="EE7BFFF0FF8CFFBBFFC52121A5557B4A" masterDocTitle="The Philippine hair wax spiders and their relatives: revision of the Pholcus bicornutus species group (Araneae, Pholcidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="3" updateTime="1698465665123" updateUser="ExternalLinkService">
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<mods:title id="C9062041880E4685BFA0A601F326CC04">The Philippine hair wax spiders and their relatives: revision of the Pholcus bicornutus species group (Araneae, Pholcidae)</mods:title>
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<paragraph id="9A54369EFF8EFFB9FDB92509A6ED7F09" blockId="2.[636,952,1064,1091]" box="[636,952,1064,1091]" pageId="2" pageNumber="3">
<heading id="C11C81F2FF8EFFB9FDB92509A6ED7F09" box="[636,952,1064,1091]" centered="true" fontSize="11" level="4" pageId="2" pageNumber="3" reason="4">
<taxonomicName id="5DEB4D1DFF8EFFB9FDB92509A6ED7F09" ID-CoL="6NWM" ID-ENA="6931" authority="Walckenaer, 1805" authorityName="Walckenaer" authorityYear="1805" box="[636,952,1064,1091]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A89FEA8CFF8EFFB9FDB92509A78B7F09" bold="true" box="[636,734,1064,1091]" italics="true" pageId="2" pageNumber="3">Pholcus</emphasis>
<bibRefCitation id="FE7A4B6FFF8EFFB9FD202509A6ED7F09" author="Walckenaer C. A." box="[741,952,1064,1091]" journalOrPublisher="E. Dentu, Paris" pageId="2" pageNumber="3" refString="Walckenaer C. A. 1805. Tableau des araneides ou caracteres essentiels des tribus, genres, familles et races que renferme le genre Aranea de Linne, avec la designation des especes comprises dans chacune de ces divisions. E. Dentu, Paris." title="Tableau des araneides ou caracteres essentiels des tribus, genres, familles et races que renferme le genre Aranea de Linne, avec la designation des especes comprises dans chacune de ces divisions" type="book" year="1805">Walckenaer, 1805</bibRefCitation>
</taxonomicName>
</heading>
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<paragraph id="9A54369EFF8EFFB9FF78254BA5AA7FE2" blockId="2.[189,1398,1129,1192]" pageId="2" pageNumber="3">
<taxonomicName id="5DEB4D1DFF8EFFB9FF78254BA7727FCE" ID-CoL="6NWM" authority="Walckenaer, 1805: 80" authorityName="Walckenaer" authorityPageNumber="80" authorityYear="1805" box="[189,551,1129,1156]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A89FEA8CFF8EFFB9FF78254BA4497FCE" box="[189,284,1130,1156]" italics="true" pageId="2" pageNumber="3">Pholcus</emphasis>
<bibRefCitation id="FE7A4B6FFF8EFFB9FEE6254BA4AF7FCE" author="Walckenaer C. A." box="[291,506,1129,1156]" journalOrPublisher="E. Dentu, Paris" pageId="2" pageNumber="3" refString="Walckenaer C. A. 1805. Tableau des araneides ou caracteres essentiels des tribus, genres, familles et races que renferme le genre Aranea de Linne, avec la designation des especes comprises dans chacune de ces divisions. E. Dentu, Paris." title="Tableau des araneides ou caracteres essentiels des tribus, genres, familles et races que renferme le genre Aranea de Linne, avec la designation des especes comprises dans chacune de ces divisions" type="book" year="1805">Walckenaer, 1805</bibRefCitation>
: 80
</taxonomicName>
.
<typeStatus id="4550883CFF8EFFB9FDF6254BA7387FCE" box="[563,621,1130,1156]" pageId="2" pageNumber="3">Type</typeStatus>
species by (assumed) monotypy:
<taxonomicName id="5DEB4D1DFF8EFFB9FBC5254BA5AE7FE2" authority="Fuesslin, 1775" authorityName="Fuesslin" authorityYear="1775" class="Gastropoda" family="Muricidae" genus="Aranea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="phalangioides">
<emphasis id="A89FEA8CFF8EFFB9FBC5254BA0507FCE" box="[1024,1285,1130,1156]" italics="true" pageId="2" pageNumber="3">Aranea phalangioides</emphasis>
Fuesslin, 1775
</taxonomicName>
.
</paragraph>
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<taxonomicName id="5DEB4D1DFF8EFFB9FF7825EFA4497FA2" ID-CoL="6NWM" box="[189,284,1230,1256]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A89FEA8CFF8EFFB9FF7825EFA4497FA2" box="[189,284,1230,1256]" italics="true" pageId="2" pageNumber="3">Pholcus</emphasis>
</taxonomicName>
<bibRefCitation id="FE7A4B6FFF8EFFB9FEFC25EFA4817FA2" author="Huber B. A." box="[313,468,1230,1256]" journalOrPublisher="Bonner zoologische Monographien" pageId="2" pageNumber="3" pagination="1 - 509" part="58" refString="Huber B. A. 2011 a. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1 - 509." title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)" type="journal article" year="2011" yearSuffix="a">Huber 2011a</bibRefCitation>
: 124126.
</paragraph>
</subSubSection>
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<paragraph id="9A54369EFF8EFFB9FD89242EA6B27E60" blockId="2.[588,999,1295,1322]" box="[588,999,1295,1322]" pageId="2" pageNumber="3">
<heading id="C11C81F2FF8EFFB9FD89242EA6B27E60" bold="true" box="[588,999,1295,1322]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="A89FEA8CFF8EFFB9FD89242EA6B27E60" bold="true" box="[588,999,1295,1322]" pageId="2" pageNumber="3">
<taxonomicName id="5DEB4D1DFF8EFFB9FD89242EA6627E60" box="[588,823,1295,1322]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="A89FEA8CFF8EFFB9FD89242EA6627E60" bold="true" box="[588,823,1295,1322]" italics="true" pageId="2" pageNumber="3">Pholcus bicornutus</emphasis>
</taxonomicName>
species group
</emphasis>
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</paragraph>
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<heading id="C11C81F2FF8EFFB9FF782470A4637E21" bold="true" box="[189,310,1361,1387]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="2">
<emphasis id="A89FEA8CFF8EFFB9FF782470A4637E21" bold="true" box="[189,310,1361,1387]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="9A54369EFF8EFFB9FF7824A1A7D87D27" blockId="2.[189,1399,1407,1645]" pageId="2" pageNumber="3">
Large, long-legged
<taxonomicName id="5DEB4D1DFF8EFFB9FE6024A1A7517ED0" box="[421,516,1408,1434]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A89FEA8CFF8EFFB9FE6024A1A7517ED0" box="[421,516,1408,1434]" italics="true" pageId="2" pageNumber="3">Pholcus</emphasis>
</taxonomicName>
(body length ~58, male leg 1 length: ~4060) with cylindrical abdomen, eight eyes, and rather dark coloration; distinguished from similar insular Southeast Asian species groups (
<taxonomicName id="5DEB4D1DFF8EFFB9FF0224E6A4387EAA" box="[199,365,1478,1504]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gracillimus">
<emphasis id="A89FEA8CFF8EFFB9FF0224E6A4387EAA" box="[199,365,1478,1504]" italics="true" pageId="2" pageNumber="3">P. gracillimus</emphasis>
</taxonomicName>
group;
<taxonomicName id="5DEB4D1DFF8EFFB9FE0824E6A7097EAA" box="[461,604,1478,1504]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis id="A89FEA8CFF8EFFB9FE0824E6A7097EAA" box="[461,604,1478,1504]" italics="true" pageId="2" pageNumber="3">P. ancoralis</emphasis>
</taxonomicName>
group; cf.
<bibRefCitation id="FE7A4B6FFF8EFFB9FD2324E7A6D47EAA" author="Huber B. A." box="[742,897,1478,1504]" journalOrPublisher="Bonner zoologische Monographien" pageId="2" pageNumber="3" pagination="1 - 509" part="58" refString="Huber B. A. 2011 a. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1 - 509." title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)" type="journal article" year="2011" yearSuffix="a">Huber 2011a</bibRefCitation>
) by male ocular area with unique pair of processes, each carrying brush of hairs (
<figureCitation id="02D02A1BFF8EFFB9FD5124C8A64E7D49" box="[660,795,1513,1539]" captionStart="Figs 12 15" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[189,1399,582,1888]" captionTargetId="figure@7.[189,1399,582,1888]" captionTargetPageId="7" captionText="Figs 12 15. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 12. Male prosoma, frontal view. 13, 15. Male ocular horns with modiFed and glued or waxed hairs, frontal and lateral views. 14. Female prosoma, frontal view. Scale lines: 12, 14 = 400 µm; 13 = 50 µm; 15 = 60 µm." httpUri="https://zenodo.org/record/830822/files/figure.png" pageId="2" pageNumber="3">Figs 1213</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FCE224C8A6267D49" box="[807,883,1513,1539]" captionStart="Figs 63 70" captionStartId="20.[189,241,1821,1847]" captionTargetBox="[189,1399,387,1778]" captionTargetId="figure@20.[189,1399,387,1778]" captionTargetPageId="20" captionText="Figs 63 70. Pholcus baguio Huber, sp. nov., ZFMK Ar 15507. 63. Male prosoma, frontal view. 64. Male ocular area, frontal (slightly dorsal) view. 65 66. Male ocular horns and modiFed hairs. 67. Male ocular area, dorsal view. 68. Male left palpal tarsal organ. 69. Right male palpal trochanter and procursus, retrolatero-dorsal view. 70. Left procursus, distal view. Scale lines: 63 = 500 µm; 64 = 300 µm; 65 = 80 µm; 66, 70 = 100 µm; 67, 69 = 200 µm; 68 = 30 µm." httpUri="https://zenodo.org/record/830840/files/figure.png" pageId="2" pageNumber="3">6367</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FCBA24C8A6CA7D49" box="[895,927,1513,1539]" captionStart="Figs 77 84" captionStartId="24.[189,241,1859,1885]" captionTargetBox="[189,1399,283,1816]" captionTargetId="figure@24.[189,1399,283,1816]" captionTargetPageId="24" captionText="Figs 77 84. Pholcus arayat Huber, 2011, ZFMK Ar 15509. 77. Male prosoma, frontal view. 78. Male ocular horns and modiFed hairs. 79. Female prosoma, frontal view. 80. Male ocular area, dorsal view. 81. Right genital bulb, prolateral view. 82. Left male palpal trochanter and procursus, dorsal view (arrow points at biFd process). 83 84. Tip of right procursus, prolateral and prolatero-distal views (arrows point at biFd process). Scale lines: 77, 79 = 300 µm; 78 = 50 µm; 80 82 = 200 µm; 83 84 = 100 µm." httpUri="https://zenodo.org/record/830844/files/figure.png" pageId="2" pageNumber="3">78</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FC6924C8A69E7D49" box="[940,971,1513,1539]" captionStart="Figs 91 98" captionStartId="27.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1824]" captionTargetId="figure@27.[189,1399,265,1824]" captionTargetPageId="27" captionText="Figs 91 98. Pholcus pagbilao Huber, 2011, ZFMK Ar 15515 16. 91. Male prosoma, frontal view. 92. Male ocular horns and modiFed hairs. 93. Female prosoma, frontal view. 94 95. Male ocular area, dorsal and latero-dorsal views. 96. Male distal cheliceral apophysis (arrows point at modiFed hairs). 97. Left procursus, dorsal view (arrow points at semitransparent process). 98. Left genital bulb, prolateral view. Scale lines: 91, 93 = 400 µm; 92 = 60 µm; 94 95, 98 = 200 µm; 96 = 20 µm; 97 = 100 µm." httpUri="https://zenodo.org/record/830848/files/figure.png" pageId="2" pageNumber="3">92</figureCitation>
) (
<taxonomicName id="5DEB4D1DFF8EFFB9FC2124CBA1257D49" box="[996,1136,1513,1539]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis id="A89FEA8CFF8EFFB9FC2124CBA1257D49" box="[996,1136,1513,1539]" italics="true" pageId="2" pageNumber="3">P. ancoralis</emphasis>
</taxonomicName>
group also with horns but without brushes of hairs), by male palpal tibia very large relative to femur (in contrast to
<taxonomicName id="5DEB4D1DFF8EFFB9FB29272CA0227D6C" box="[1260,1399,1548,1574]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis id="A89FEA8CFF8EFFB9FB29272CA0227D6C" box="[1260,1399,1548,1574]" italics="true" pageId="2" pageNumber="3">P. ancoralis</emphasis>
</taxonomicName>
species group), and by paired anterior marks in dorsal abdominal pattern (
<figureCitation id="02D02A1BFF8EFFB9FBDD2711A1367D00" box="[1048,1123,1584,1610]" captionStart="Figs 3 6" captionStartId="5.[189,241,1964,1990]" captionTargetBox="[189,1399,942,1923]" captionTargetId="figure@5.[189,1399,942,1923]" captionTargetPageId="5" captionText="Figs 3 6. Live specimens, Pholcus olangapo Huber, sp. nov., Pamulaklakin Forest Trail. 3 4. ♁ and ♀ in their natural habitat. 5 6. ♁ and ♀ photographed on a dead leaf." httpUri="https://zenodo.org/record/830818/files/figure.png" pageId="2" pageNumber="3">Figs 3</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FBB52711A1C57D00" box="[1136,1168,1584,1610]" captionStart="Figs 36 39" captionStartId="12.[189,241,1966,1992]" captionTargetBox="[189,1399,620,1926]" captionTargetId="figure@12.[189,1399,620,1926]" captionTargetPageId="12" captionText="Figs 36 39. Live specimens. 36 37. Pholcus mulu Huber, sp. nov., ♁, Gunung Mulu. 38 39. Pholcus kawit Huber, sp. nov., ♁ and ♀, Mt. Matutum." httpUri="https://zenodo.org/record/830828/files/figure.png" pageId="2" pageNumber="3">36</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FB5E270EA1EE7D00" box="[1179,1211,1583,1610]" captionStart="Figs 50 53" captionStartId="17.[189,241,1904,1930]" captionTargetBox="[189,1399,365,1862]" captionTargetId="figure@17.[189,1399,365,1862]" captionTargetPageId="17" captionText="Figs 50 53. Live specimens. 50 52. Pholcus baguio Huber, sp. nov., ♁, ♀, and ♀ with egg-sac and emerging juveniles, Mt. Kabuyao. 53. Pholcus arayat Huber, 2011, ♁, Pamulaklakin Forest Trail." httpUri="https://zenodo.org/record/830834/files/figure.png" pageId="2" pageNumber="3">50</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FB03270EA1B37D00" box="[1222,1254,1583,1610]" captionStart="Figs 54 57" captionStartId="18.[189,241,1869,1895]" captionTargetBox="[189,1399,418,1829]" captionTargetId="figure@18.[189,1399,418,1829]" captionTargetPageId="18" captionText="Figs 54 57. Live specimens. 54. Pholcus arayat Huber, 2011, ♀, Pamulaklakin Forest Trail. 55 57. Pholcus pagbilao Huber, 2011, ♁ from Loboc (55), ♁ from Mt. Banahaw (56), and ♀ with egg-sac from Loboc (57)." httpUri="https://zenodo.org/record/830836/files/figure.png" pageId="2" pageNumber="3">55</figureCitation>
; unpaired in
<taxonomicName id="5DEB4D1DFF8EFFB9FF782775A4357D27" box="[189,352,1619,1645]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gracillimus">
<emphasis id="A89FEA8CFF8EFFB9FF782775A4357D27" box="[189,352,1619,1645]" italics="true" pageId="2" pageNumber="3">P. gracillimus</emphasis>
</taxonomicName>
and
<taxonomicName id="5DEB4D1DFF8EFFB9FE5D2775A7717D27" box="[408,548,1619,1645]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis id="A89FEA8CFF8EFFB9FE5D2775A7717D27" box="[408,548,1619,1645]" italics="true" pageId="2" pageNumber="3">P. ancoralis</emphasis>
</taxonomicName>
groups).
</paragraph>
</subSubSection>
<subSubSection id="D2F16515FF8EFFBFFF7827B5A6287A03" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="9A54369EFF8EFFB9FF7827B5A75E7DE5" blockId="2.[189,523,1684,1711]" box="[189,523,1684,1711]" pageId="2" pageNumber="3">
<heading id="C11C81F2FF8EFFB9FF7827B5A75E7DE5" bold="true" box="[189,523,1684,1711]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="2">
<emphasis id="A89FEA8CFF8EFFB9FF7827B5A75E7DE5" bold="true" box="[189,523,1684,1711]" pageId="2" pageNumber="3">Description amendments</emphasis>
</heading>
</paragraph>
<paragraph id="9A54369EFF8EFFB8FF7827E2A1C67A03" blockId="2.[189,1399,1731,1969]" lastBlockId="3.[189,1398,267,329]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
The original description of the species group (
<bibRefCitation id="FE7A4B6FFF8EFFB9FD0827E2A6337D97" author="Huber B. A." box="[717,870,1731,1757]" journalOrPublisher="Bonner zoologische Monographien" pageId="2" pageNumber="3" pagination="1 - 509" part="58" refString="Huber B. A. 2011 a. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1 - 509." title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)" type="journal article" year="2011" yearSuffix="a">Huber 2011a</bibRefCitation>
) is still largely valid. The following are minor updates and additions: body length ~58, carapace width 1.31.8. Male distal cheliceral apophyses provided with 23 modified hairs each (
<figureCitation id="02D02A1BFF8EFFB9FD622628A6527C69" box="[679,775,1801,1827]" captionStart="Figs 16 23" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1399,397,1826]" captionTargetId="figure@8.[189,1399,397,1826]" captionTargetPageId="8" captionText="Figs 16 23. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 16. Right genital bulb, prolateral view. 17. Left tarsus and procursus, dorsal view (asterisk marks prolatero-dorsal process of procursus). 18. Tip of left procursus, dorsal view. 19. Female spinnerets. 20. Male cheliceral apophysis (arrows point at modiFed hairs). 21. Male ALS. 22. Male gonopore. 23. Epigynum. Scale lines: 16, 23 = 200 µm; 17 = 300 µm; 18 = 100 µm; 19 = 70 µm; 20 21 = 20 µm; 22 = 40 µm." httpUri="https://zenodo.org/record/830824/files/figure.png" pageId="2" pageNumber="3">Figs 20</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FCD32628A6637C69" box="[790,822,1801,1827]" captionStart="Figs 71 76" captionStartId="22.[189,241,1870,1896]" captionTargetBox="[189,1399,437,1834]" captionTargetId="figure@22.[189,1399,437,1834]" captionTargetPageId="22" captionText="Figs 71 76. Pholcus baguio Huber, sp. nov., ZFMK Ar 15507 08. 71 72. Left genital bulb, prolateral and prolatero-distal views. 73. Male distal cheliceral apophysis (arrows point at modiFed hairs). 74. Male ALS. 75. Epigynum. 76. Female ALS and PMS. Scale lines: 71 72, 75 = 200 µm; 73 74, 76 = 20 µm." httpUri="https://zenodo.org/record/830842/files/figure.png" pageId="2" pageNumber="3">73</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FC832628A6337C6E" box="[838,870,1801,1828]" captionStart="Figs 85 90" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[189,1399,626,1887]" captionTargetId="figure@25.[189,1399,626,1887]" captionTargetPageId="25" captionText="Figs 85 90. Pholcus arayat Huber, 2011, ZFMK Ar 15509. 85. Male distal cheliceral apophysis (arrows point at modiFed hairs). 86. Male gonopore. 87. Male tarsus 4 comb-hairs. 88. Male ALS. 89. Epigynum. 90. Female ALS. Scale lines: 85, 87 88, 90 = 20 µm; 86 = 50 µm; 89 = 200 µm." httpUri="https://zenodo.org/record/830846/files/figure.png" pageId="2" pageNumber="3">85</figureCitation>
,
<figureCitation id="02D02A1BFF8EFFB9FCB32628A6C27C6E" box="[886,919,1801,1828]" captionStart="Figs 91 98" captionStartId="27.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1824]" captionTargetId="figure@27.[189,1399,265,1824]" captionTargetPageId="27" captionText="Figs 91 98. Pholcus pagbilao Huber, 2011, ZFMK Ar 15515 16. 91. Male prosoma, frontal view. 92. Male ocular horns and modiFed hairs. 93. Female prosoma, frontal view. 94 95. Male ocular area, dorsal and latero-dorsal views. 96. Male distal cheliceral apophysis (arrows point at modiFed hairs). 97. Left procursus, dorsal view (arrow points at semitransparent process). 98. Left genital bulb, prolateral view. Scale lines: 91, 93 = 400 µm; 92 = 60 µm; 94 95, 98 = 200 µm; 96 = 20 µm; 97 = 100 µm." httpUri="https://zenodo.org/record/830848/files/figure.png" pageId="2" pageNumber="3">96</figureCitation>
). Tibia 1 length in males ~9.516.0, in females ~7.514.0; male tibia 1 L/d ~6580; legs either with curved hairs (tibiae and metatarsi 12;
<taxonomicName id="5DEB4D1DFF8EFFB9FF782670A4027C20" box="[189,343,1872,1898]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="A89FEA8CFF8EFFB9FF782670A4027C20" box="[189,343,1872,1898]" italics="true" pageId="2" pageNumber="3">P. bicornutus</emphasis>
</taxonomicName>
,
<taxonomicName id="5DEB4D1DFF8EFFB9FEA22670A4A67C20" box="[359,499,1872,1898]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="olangapo" status="sp. nov.">
<emphasis id="A89FEA8CFF8EFFB9FEA22670A4A67C20" box="[359,499,1872,1898]" italics="true" pageId="2" pageNumber="3">P. olangapo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF8EFFB9FE3E2671A7027C20" box="[507,599,1872,1898]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
) or without curved hairs (other species). Abdomen cylindrical, with cuticular color pattern dorsally (also in
<taxonomicName id="5DEB4D1DFF8EFFB9FD5D2655A6617CC7" box="[664,820,1907,1933]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="A89FEA8CFF8EFFB9FD5D2655A6617CC7" box="[664,820,1907,1933]" italics="true" pageId="2" pageNumber="3">P. bicornutus</emphasis>
</taxonomicName>
), anterior element always paired. ALS with one widened, one pointed, and variable number of smaller spigots: either six cylindrically-shaped spigots of usual size (i.e., ~38 µm wide and ~58 µm long;
<figureCitation id="02D02A1BFF8FFFB8FC84202AA6927A6F" box="[833,967,267,293]" captionStart="Figs 16 23" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1399,397,1826]" captionTargetId="figure@8.[189,1399,397,1826]" captionTargetPageId="8" captionText="Figs 16 23. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 16. Right genital bulb, prolateral view. 17. Left tarsus and procursus, dorsal view (asterisk marks prolatero-dorsal process of procursus). 18. Tip of left procursus, dorsal view. 19. Female spinnerets. 20. Male cheliceral apophysis (arrows point at modiFed hairs). 21. Male ALS. 22. Male gonopore. 23. Epigynum. Scale lines: 16, 23 = 200 µm; 17 = 300 µm; 18 = 100 µm; 19 = 70 µm; 20 21 = 20 µm; 22 = 40 µm." httpUri="https://zenodo.org/record/830824/files/figure.png" pageId="3" pageNumber="4">Figs 19, 21</figureCitation>
,
<figureCitation id="02D02A1BFF8FFFB8FC16202AA1687A6F" box="[979,1085,267,293]" captionStart="Figs 99 102" captionStartId="28.[189,241,1964,1990]" captionTargetBox="[189,1399,918,1922]" captionTargetId="figure@28.[189,1399,918,1922]" captionTargetPageId="28" captionText="Figs 99 102. Pholcus pagbilao Huber, 2011, ZFMK Ar 15515 16. 99. Male gonopore. 100. Male ALS. 101. Epigynum. 102. Female ALS. Scale lines: 99 = 50 µm; 100, 102 = 20 µm; 101 = 200 µm." httpUri="https://zenodo.org/record/830850/files/figure.png" pageId="3" pageNumber="4">100, 102</figureCitation>
), or only three to five very small conical spigots (i.e., ~1.53 µm wide and ~36 µm long;
<figureCitation id="02D02A1BFF8FFFB8FC62200FA1787A02" box="[935,1069,302,329]" captionStart="Figs 71 76" captionStartId="22.[189,241,1870,1896]" captionTargetBox="[189,1399,437,1834]" captionTargetId="figure@22.[189,1399,437,1834]" captionTargetPageId="22" captionText="Figs 71 76. Pholcus baguio Huber, sp. nov., ZFMK Ar 15507 08. 71 72. Left genital bulb, prolateral and prolatero-distal views. 73. Male distal cheliceral apophysis (arrows point at modiFed hairs). 74. Male ALS. 75. Epigynum. 76. Female ALS and PMS. Scale lines: 71 72, 75 = 200 µm; 73 74, 76 = 20 µm." httpUri="https://zenodo.org/record/830842/files/figure.png" pageId="3" pageNumber="4">Figs 74, 76</figureCitation>
,
<figureCitation id="02D02A1BFF8FFFB8FBFC200EA1D37A03" box="[1081,1158,303,329]" captionStart="Figs 85 90" captionStartId="25.[189,241,1928,1954]" captionTargetBox="[189,1399,626,1887]" captionTargetId="figure@25.[189,1399,626,1887]" captionTargetPageId="25" captionText="Figs 85 90. Pholcus arayat Huber, 2011, ZFMK Ar 15509. 85. Male distal cheliceral apophysis (arrows point at modiFed hairs). 86. Male gonopore. 87. Male tarsus 4 comb-hairs. 88. Male ALS. 89. Epigynum. 90. Female ALS. Scale lines: 85, 87 88, 90 = 20 µm; 86 = 50 µm; 89 = 200 µm." httpUri="https://zenodo.org/record/830846/files/figure.png" pageId="3" pageNumber="4">88, 90</figureCitation>
).
</paragraph>
<paragraph id="9A54369EFF8FFFB8FF782051A4087AC0" blockId="3.[189,1243,368,441]" box="[189,349,368,394]" pageId="3" pageNumber="4">
<heading id="C11C81F2FF8FFFB8FF782051A4087AC0" bold="true" box="[189,349,368,394]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="6">
<emphasis id="A89FEA8CFF8FFFB8FF782051A4087AC0" bold="true" box="[189,349,368,394]" pageId="3" pageNumber="4">Composition</emphasis>
</heading>
</paragraph>
<paragraph id="9A54369EFF8FFFB8FF7820BFA18E7AF2" blockId="3.[189,1243,368,441]" box="[189,1243,414,441]" pageId="3" pageNumber="4">The eight species assigned to this group are here divided into two operational sub-groups:</paragraph>
<paragraph id="9A54369EFF8FFFB8FF7820C1A1E77984" blockId="3.[189,1399,480,718]" pageId="3" pageNumber="4">
(1) A sub-group of two species (
<taxonomicName id="5DEB4D1DFF8FFFB8FDF120C0A6387AB0" authority="Simon, 1892" authorityName="Simon" authorityYear="1892" box="[564,877,480,506]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="A89FEA8CFF8FFFB8FDF120C0A7987AB0" box="[564,717,480,506]" italics="true" pageId="3" pageNumber="4">P. bicornutus</emphasis>
<bibRefCitation id="FE7A4B6FFF8FFFB8FD1020C1A6387AB0" author="Simon E." box="[725,877,480,506]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="3" pageNumber="4" pagination="35 - 52" part="61" refString="Simon E. 1892. Etude sur les arthropodes cavernicoles de l'ile de Luzon. Annales de la Societe entomologique de France 61: 35 - 52." title="Etude sur les arthropodes cavernicoles de l'ile de Luzon" type="journal article" year="1892">Simon, 1892</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="5DEB4D1DFF8FFFB8FCBC20C0A1517AB0" box="[889,1028,480,506]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="olangapo" status="sp. nov.">
<emphasis id="A89FEA8CFF8FFFB8FCBC20C0A1517AB0" box="[889,1028,480,506]" italics="true" pageId="3" pageNumber="4">P. olangapo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF8FFFB8FBCE20C1A13D7AB0" box="[1035,1128,480,506]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
), sharing the following similarities: absence of male bulbal appendix (
<figureCitation id="02D02A1BFF8FFFB8FD2E2322A6607954" box="[747,821,515,542]" captionStart="Figs 7 11" captionStartId="6.[189,241,1929,1955]" captionTargetBox="[194,1399,481,1883]" captionTargetId="figure@6.[189,1399,481,1888]" captionTargetPageId="6" captionText="Figs 7 11. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 7 8. Left male palp, prolateral and retrolateral views (asterisk marks prolatero-dorsal process of procursus). 9. Male chelicerae, frontal view. 10 11. Cleared female genitalia, ventral and dorsal views. Scale lines: 7 8 = 1 mm; 9 1 1 = 0.5 mm." httpUri="https://zenodo.org/record/830820/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
); procursus with hinged distal element and with distinctive prolatero-dorsal process (
<figureCitation id="02D02A1BFF8FFFB8FDAF2307A7E3790A" box="[618,694,550,576]" captionStart="Figs 7 11" captionStartId="6.[189,241,1929,1955]" captionTargetBox="[194,1399,481,1883]" captionTargetId="figure@6.[189,1399,481,1888]" captionTargetPageId="6" captionText="Figs 7 11. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 7 8. Left male palp, prolateral and retrolateral views (asterisk marks prolatero-dorsal process of procursus). 9. Male chelicerae, frontal view. 10 11. Cleared female genitalia, ventral and dorsal views. Scale lines: 7 8 = 1 mm; 9 1 1 = 0.5 mm." httpUri="https://zenodo.org/record/830820/files/figure.png" pageId="3" pageNumber="4">Figs 8</figureCitation>
,
<figureCitation id="02D02A1BFF8FFFB8FD012307A7B1790A" box="[708,740,550,576]" captionStart="Figs 16 23" captionStartId="8.[189,241,1860,1886]" captionTargetBox="[189,1399,397,1826]" captionTargetId="figure@8.[189,1399,397,1826]" captionTargetPageId="8" captionText="Figs 16 23. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 16. Right genital bulb, prolateral view. 17. Left tarsus and procursus, dorsal view (asterisk marks prolatero-dorsal process of procursus). 18. Tip of left procursus, dorsal view. 19. Female spinnerets. 20. Male cheliceral apophysis (arrows point at modiFed hairs). 21. Male ALS. 22. Male gonopore. 23. Epigynum. Scale lines: 16, 23 = 200 µm; 17 = 300 µm; 18 = 100 µm; 19 = 70 µm; 20 21 = 20 µm; 22 = 40 µm." httpUri="https://zenodo.org/record/830824/files/figure.png" pageId="3" pageNumber="4">17</figureCitation>
); male palpal tibia ventrally unmodified; hairs of ocular processes glued or waxed together (
<figureCitation id="02D02A1BFF8FFFB8FD4F2368A615792E" box="[650,832,585,612]" captionStart="Figs 12 15" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[189,1399,582,1888]" captionTargetId="figure@7.[189,1399,582,1888]" captionTargetPageId="7" captionText="Figs 12 15. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 12. Male prosoma, frontal view. 13, 15. Male ocular horns with modiFed and glued or waxed hairs, frontal and lateral views. 14. Female prosoma, frontal view. Scale lines: 12, 14 = 400 µm; 13 = 50 µm; 15 = 60 µm." httpUri="https://zenodo.org/record/830822/files/figure.png" pageId="3" pageNumber="4">Figs 1213, 15</figureCitation>
); epigynum with small epigynal plate and large membranous area in front of plate (
<figureCitation id="02D02A1BFF8FFFB8FD90234CA7F979CD" box="[597,684,621,647]" captionStart="Figs 24 35" captionStartId="9.[189,241,1894,1920]" captionTargetBox="[189,1399,427,1852]" captionTargetId="figure@9.[189,1399,427,1852]" captionTargetPageId="9" captionText="Figs 24 35. Epigyna, ventral views, and cleared female genitalia, ventral and dorsal view. 24 26. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15498. 27 29. Pholcus mulu Huber, sp. nov., ZFMK Ar 15502. 30 32. Pholcus kawit Huber, sp. nov., ZFMK Ar 15505. 33 35. Pholcus baguio Huber, sp. nov., ZFMK Ar 15508." httpUri="https://zenodo.org/record/830826/files/figure.png" pageId="3" pageNumber="4">Fig. 24</figureCitation>
); proximal lateral processes on male chelicerae large (
<figureCitation id="02D02A1BFF8FFFB8FAE5234CA03279CD" box="[1312,1383,621,647]" captionStart="Figs 7 11" captionStartId="6.[189,241,1929,1955]" captionTargetBox="[194,1399,481,1883]" captionTargetId="figure@6.[189,1399,481,1888]" captionTargetPageId="6" captionText="Figs 7 11. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15497 98. 7 8. Left male palp, prolateral and retrolateral views (asterisk marks prolatero-dorsal process of procursus). 9. Male chelicerae, frontal view. 10 11. Cleared female genitalia, ventral and dorsal views. Scale lines: 7 8 = 1 mm; 9 1 1 = 0.5 mm." httpUri="https://zenodo.org/record/830820/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
); and presence of curved hairs on tibiae and metatarsi 12. A third (unnamed) species occurs on Negros Island (“sp. Phi 190” in
<figureCitation id="02D02A1BFF8FFFB8FE1C2392A7487987" box="[473,541,691,717]" captionStart="Figs 1 2" captionStartId="3.[189,241,2000,2026]" captionTargetBox="[189,1398,1092,1972]" captionTargetId="figure@3.[189,1398,1092,1972]" captionTargetPageId="3" captionText="Figs 1 2. Known distributions of the two subgroups of the Pholcus bicornutus species group." httpUri="https://zenodo.org/record/830816/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
; 1 ♀, 1 juv. from Twin Lakes National Park, in ZFMK).
</paragraph>
<paragraph id="9A54369EFF8FFFBFFF7823D4A6287A03" blockId="3.[189,1399,757,1030]" lastBlockId="4.[189,1398,267,329]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
(2) A sub-group of six species (
<taxonomicName id="5DEB4D1DFF8FFFB8FDF423D4A6627845" authority="Huber, 2011" authorityName="Huber" authorityYear="2011" box="[561,823,757,783]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="arayat">
<emphasis id="A89FEA8CFF8FFFB8FDF423D4A7CE7844" box="[561,667,757,782]" italics="true" pageId="3" pageNumber="4">P. arayat</emphasis>
<bibRefCitation id="FE7A4B6FFF8FFFB8FD6623D4A6627845" author="Huber B. A." box="[675,823,757,783]" journalOrPublisher="Bonner zoologische Monographien" pageId="3" pageNumber="4" pagination="1 - 509" part="58" refString="Huber B. A. 2011 a. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1 - 509." title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)" type="journal article" year="2011">Huber, 2011</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="5DEB4D1DFF8FFFB8FC8123D4A6C97845" box="[836,924,757,783]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mulu" status="sp. nov.">
<emphasis id="A89FEA8CFF8FFFB8FC8123D4A6C97845" box="[836,924,757,783]" italics="true" pageId="3" pageNumber="4">P. mulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF8FFFB8FC6123D4A1577845" box="[932,1026,757,783]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="5DEB4D1DFF8FFFB8FBCB23D4A1287845" box="[1038,1149,757,783]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="baguio" status="sp. nov.">
<emphasis id="A89FEA8CFF8FFFB8FBCB23D4A1287845" box="[1038,1149,757,783]" italics="true" pageId="3" pageNumber="4">P. baguio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF8FFFB8FB4023D4A1B17845" box="[1157,1252,757,783]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="5DEB4D1DFF8FFFB8FB3523D4A01B7845" box="[1264,1358,757,783]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kawit" status="sp. nov.">
<emphasis id="A89FEA8CFF8FFFB8FB3523D4A01B7845" box="[1264,1358,757,783]" italics="true" pageId="3" pageNumber="4">P. kawit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF8FFFB8FA9023D4A5A77878" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="5DEB4D1DFF8FFFB8FF3B2238A7757878" authority="Huber, 2011" authorityName="Huber" authorityYear="2011" box="[254,544,792,818]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pagbilao">
<emphasis id="A89FEA8CFF8FFFB8FF3B2238A4D17878" box="[254,388,792,818]" italics="true" pageId="3" pageNumber="4">P. pagbilao</emphasis>
<bibRefCitation id="FE7A4B6FFF8FFFB8FE492239A7757878" author="Huber B. A." box="[396,544,792,818]" journalOrPublisher="Bonner zoologische Monographien" pageId="3" pageNumber="4" pagination="1 - 509" part="58" refString="Huber B. A. 2011 a. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1 - 509." title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)" type="journal article" year="2011">Huber, 2011</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="5DEB4D1DFF8FFFB8FDE92238A6CD7878" authority="Yao et al., 2014" authorityName="Yao et al." authorityYear="2014" box="[556,920,792,818]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="schawalleri">
<emphasis id="A89FEA8CFF8FFFB8FDE92238A7867878" box="[556,723,792,818]" italics="true" pageId="3" pageNumber="4">P. schawalleri</emphasis>
<bibRefCitation id="FE7A4B6FFF8FFFB8FD1C2239A6CD7878" author="Yao Z. &amp; Li S. &amp; Jager P." box="[729,920,792,818]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="4" pagination="331 - 349" part="3793" publicationUrl="http://dx.doi.org/10.11646/zootaxa.3793.3.2" refString="Yao Z., Li S. &amp; Jager P. 2014. Four new species of pholcine spiders (Araneae: Pholcidae) from Southeast Asia. Zootaxa 3793: 331 - 349. http: // dx. doi. org / 10.11646 / zootaxa. 3793.3.2" title="Four new species of pholcine spiders (Araneae: Pholcidae) from Southeast Asia" type="journal article" year="2014">
Yao
<emphasis id="A89FEA8CFF8FFFB8FCCA2238A61F7878" box="[783,842,792,818]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
, 2014
</bibRefCitation>
</taxonomicName>
), sharing the alternative character states: presence of male bulbal appendix (
<figureCitation id="02D02A1BFF8FFFB8FDA5221AA7E8781F" box="[608,701,827,853]" captionStart="Figs 40 44" captionStartId="13.[189,241,1930,1956]" captionTargetBox="[191,1395,557,1894]" captionTargetId="figure@13.[189,1399,557,1902]" captionTargetPageId="13" captionText="Figs 40 44. Pholcus mulu Huber, sp. nov., ZFMK Ar 15501 02. 40 41. Left male palp, prolateral and retrolateral views. 42. Male chelicerae, frontal view. 43 44. Cleared female genitalia, ventral and dorsal views. Scale lines: 40 41 = 1 mm; 42 44 = 0.5 mm." httpUri="https://zenodo.org/record/830830/files/figure.png" pageId="3" pageNumber="4">Figs 40</figureCitation>
,
<figureCitation id="02D02A1BFF8FFFB8FD0E221AA7B9781C" box="[715,748,827,854]" captionStart="Figs 45 49" captionStartId="15.[189,241,1930,1956]" captionTargetBox="[189,1399,581,1887]" captionTargetId="figure@15.[189,1399,580,1888]" captionTargetPageId="15" captionText="Figs 45 49. Pholcus kawit Huber, sp. nov., ZFMK Ar 15505. 45 46. Left male palp, prolateral and retrolateral views (arrow points at ventral cavity of tibia). 47. Male chelicerae, frontal view. 48 49. Cleared female genitalia, ventral and dorsal views. Scale lines: 45 46 = 1 mm; 47 49 = 0.5 mm." httpUri="https://zenodo.org/record/830832/files/figure.png" pageId="3" pageNumber="4">45</figureCitation>
); procursus without hinged distal element and without dorsal process (
<figureCitation id="02D02A1BFF8FFFB8FEB4227FA49E7832" box="[369,459,862,888]" captionStart="Figs 40 44" captionStartId="13.[189,241,1930,1956]" captionTargetBox="[191,1395,557,1894]" captionTargetId="figure@13.[189,1399,557,1902]" captionTargetPageId="13" captionText="Figs 40 44. Pholcus mulu Huber, sp. nov., ZFMK Ar 15501 02. 40 41. Left male palp, prolateral and retrolateral views. 42. Male chelicerae, frontal view. 43 44. Cleared female genitalia, ventral and dorsal views. Scale lines: 40 41 = 1 mm; 42 44 = 0.5 mm." httpUri="https://zenodo.org/record/830830/files/figure.png" pageId="3" pageNumber="4">Figs 41</figureCitation>
,
<figureCitation id="02D02A1BFF8FFFB8FE13227FA4A27833" box="[470,503,862,889]" captionStart="Figs 45 49" captionStartId="15.[189,241,1930,1956]" captionTargetBox="[189,1399,581,1887]" captionTargetId="figure@15.[189,1399,580,1888]" captionTargetPageId="15" captionText="Figs 45 49. Pholcus kawit Huber, sp. nov., ZFMK Ar 15505. 45 46. Left male palp, prolateral and retrolateral views (arrow points at ventral cavity of tibia). 47. Male chelicerae, frontal view. 48 49. Cleared female genitalia, ventral and dorsal views. Scale lines: 45 46 = 1 mm; 47 49 = 0.5 mm." httpUri="https://zenodo.org/record/830832/files/figure.png" pageId="3" pageNumber="4">46</figureCitation>
); male palpal tibia with small ventral cavity to accommodate proximal bulbal sclerite in resting position (arrow in
<figureCitation id="02D02A1BFF8FFFB8FDA822A0A79078D6" box="[621,709,897,924]" captionStart="Figs 45 49" captionStartId="15.[189,241,1930,1956]" captionTargetBox="[189,1399,581,1887]" captionTargetId="figure@15.[189,1399,580,1888]" captionTargetPageId="15" captionText="Figs 45 49. Pholcus kawit Huber, sp. nov., ZFMK Ar 15505. 45 46. Left male palp, prolateral and retrolateral views (arrow points at ventral cavity of tibia). 47. Male chelicerae, frontal view. 48 49. Cleared female genitalia, ventral and dorsal views. Scale lines: 45 46 = 1 mm; 47 49 = 0.5 mm." httpUri="https://zenodo.org/record/830832/files/figure.png" pageId="3" pageNumber="4">Fig. 45</figureCitation>
); hairs of ocular processes not glued or waxed together (
<figureCitation id="02D02A1BFF8FFFB8FF002284A40478F5" box="[197,337,932,959]" captionStart="Figs 63 70" captionStartId="20.[189,241,1821,1847]" captionTargetBox="[189,1399,387,1778]" captionTargetId="figure@20.[189,1399,387,1778]" captionTargetPageId="20" captionText="Figs 63 70. Pholcus baguio Huber, sp. nov., ZFMK Ar 15507. 63. Male prosoma, frontal view. 64. Male ocular area, frontal (slightly dorsal) view. 65 66. Male ocular horns and modiFed hairs. 67. Male ocular area, dorsal view. 68. Male left palpal tarsal organ. 69. Right male palpal trochanter and procursus, retrolatero-dorsal view. 70. Left procursus, distal view. Scale lines: 63 = 500 µm; 64 = 300 µm; 65 = 80 µm; 66, 70 = 100 µm; 67, 69 = 200 µm; 68 = 30 µm." httpUri="https://zenodo.org/record/830840/files/figure.png" pageId="3" pageNumber="4">Figs 6566</figureCitation>
,
<figureCitation id="02D02A1BFF8FFFB8FEA42284A4D778F5" box="[353,386,933,959]" captionStart="Figs 77 84" captionStartId="24.[189,241,1859,1885]" captionTargetBox="[189,1399,283,1816]" captionTargetId="figure@24.[189,1399,283,1816]" captionTargetPageId="24" captionText="Figs 77 84. Pholcus arayat Huber, 2011, ZFMK Ar 15509. 77. Male prosoma, frontal view. 78. Male ocular horns and modiFed hairs. 79. Female prosoma, frontal view. 80. Male ocular area, dorsal view. 81. Right genital bulb, prolateral view. 82. Left male palpal trochanter and procursus, dorsal view (arrow points at biFd process). 83 84. Tip of right procursus, prolateral and prolatero-distal views (arrows point at biFd process). Scale lines: 77, 79 = 300 µm; 78 = 50 µm; 80 82 = 200 µm; 83 84 = 100 µm." httpUri="https://zenodo.org/record/830844/files/figure.png" pageId="3" pageNumber="4">78</figureCitation>
); epigynum with large epigynal plate (
<figureCitation id="02D02A1BFF8FFFB8FCA52284A17778F5" box="[864,1058,933,959]" captionStart="Figs 24 35" captionStartId="9.[189,241,1894,1920]" captionTargetBox="[189,1399,427,1852]" captionTargetId="figure@9.[189,1399,427,1852]" captionTargetPageId="9" captionText="Figs 24 35. Epigyna, ventral views, and cleared female genitalia, ventral and dorsal view. 24 26. Pholcus olangapo Huber, sp. nov., ZFMK Ar 15498. 27 29. Pholcus mulu Huber, sp. nov., ZFMK Ar 15502. 30 32. Pholcus kawit Huber, sp. nov., ZFMK Ar 15505. 33 35. Pholcus baguio Huber, sp. nov., ZFMK Ar 15508." httpUri="https://zenodo.org/record/830826/files/figure.png" pageId="3" pageNumber="4">Figs 27, 30, 33</figureCitation>
); proximal lateral processes on male chelicerae small (
<figureCitation id="02D02A1BFF8FFFB8FE3D22E9A70378A8" box="[504,598,968,994]" captionStart="Figs 40 44" captionStartId="13.[189,241,1930,1956]" captionTargetBox="[191,1395,557,1894]" captionTargetId="figure@13.[189,1399,557,1902]" captionTargetPageId="13" captionText="Figs 40 44. Pholcus mulu Huber, sp. nov., ZFMK Ar 15501 02. 40 41. Left male palp, prolateral and retrolateral views. 42. Male chelicerae, frontal view. 43 44. Cleared female genitalia, ventral and dorsal views. Scale lines: 40 41 = 1 mm; 42 44 = 0.5 mm." httpUri="https://zenodo.org/record/830830/files/figure.png" pageId="3" pageNumber="4">Figs 42</figureCitation>
,
<figureCitation id="02D02A1BFF8FFFB8FDA122E9A7D078A8" box="[612,645,968,994]" captionStart="Figs 45 49" captionStartId="15.[189,241,1930,1956]" captionTargetBox="[189,1399,581,1887]" captionTargetId="figure@15.[189,1399,580,1888]" captionTargetPageId="15" captionText="Figs 45 49. Pholcus kawit Huber, sp. nov., ZFMK Ar 15505. 45 46. Left male palp, prolateral and retrolateral views (arrow points at ventral cavity of tibia). 47. Male chelicerae, frontal view. 48 49. Cleared female genitalia, ventral and dorsal views. Scale lines: 45 46 = 1 mm; 47 49 = 0.5 mm." httpUri="https://zenodo.org/record/830832/files/figure.png" pageId="3" pageNumber="4">47</figureCitation>
); no curved hairs on legs. A possible further species occurs on Mindanao, but it is represented by two juvenile specimens only (“sp. Phi 254” in
<figureCitation id="02D02A1BFF8FFFB8FB4722CAA19C7F4F" box="[1154,1225,1003,1029]" captionStart="Figs 1 2" captionStartId="3.[189,241,2000,2026]" captionTargetBox="[189,1398,1092,1972]" captionTargetId="figure@3.[189,1398,1092,1972]" captionTargetPageId="3" captionText="Figs 1 2. Known distributions of the two subgroups of the Pholcus bicornutus species group." httpUri="https://zenodo.org/record/830816/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
; from Marilog Distr., Baganihan, in ZFMK). The undescribed species from Mt. Apo mentioned in
<bibRefCitation id="FE7A4B6FFF88FFBFFB55202AA0237A6C" author="Huber B. A." box="[1168,1398,267,294]" journalOrPublisher="Bonner zoologische Monographien" pageId="4" pageNumber="5" pagination="1 - 509" part="58" refString="Huber B. A. 2011 a. Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1 - 509." title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)" type="journal article" year="2011" yearSuffix="a">Huber (2011a: 315)</bibRefCitation>
was restudied and found to belong to another species group.
</paragraph>
</subSubSection>
<caption id="CE946616FF8FFFB8FF7826F1A0527CA1" box="[189,1287,2000,2027]" httpUri="https://zenodo.org/record/830816/files/figure.png" pageId="3" pageNumber="4" targetBox="[189,1398,1092,1972]" targetPageId="3">
<paragraph id="9A54369EFF8FFFB8FF7826F1A0527CA1" blockId="3.[189,1287,2000,2027]" box="[189,1287,2000,2027]" pageId="3" pageNumber="4">
<emphasis id="A89FEA8CFF8FFFB8FF7826F1A4797CA0" bold="true" box="[189,300,2000,2026]" pageId="3" pageNumber="4">Figs 12.</emphasis>
Known distributions of the two subgroups of the
<taxonomicName id="5DEB4D1DFF8FFFB8FCB126F1A1027CA0" box="[884,1111,2000,2026]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="A89FEA8CFF8FFFB8FCB126F1A1027CA0" box="[884,1111,2000,2026]" italics="true" pageId="3" pageNumber="4">Pholcus bicornutus</emphasis>
</taxonomicName>
species group.
</paragraph>
</caption>
<subSubSection id="D2F16515FF88FFBFFF782051A17E7998" pageId="4" pageNumber="5" type="biology_ecology">
<paragraph id="9A54369EFF88FFBFFF782051A42B7AC0" blockId="4.[189,1399,368,723]" box="[189,382,368,394]" pageId="4" pageNumber="5">
<heading id="C11C81F2FF88FFBFFF782051A42B7AC0" bold="true" box="[189,382,368,394]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="6">
<emphasis id="A89FEA8CFF88FFBFFF782051A42B7AC0" bold="true" box="[189,382,368,394]" pageId="4" pageNumber="5">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="9A54369EFF88FFBFFF7820BEA17E7998" blockId="4.[189,1399,368,723]" pageId="4" pageNumber="5">
Seven of the eight named species were observed in the field. They occupied a variety of microhabitats, ranging from caves (close to the ground;
<taxonomicName id="5DEB4D1DFF88FFBFFD7420E2A6187A96" box="[689,845,450,476]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="A89FEA8CFF88FFBFFD7420E2A6187A96" box="[689,845,450,476]" italics="true" pageId="4" pageNumber="5">P. bicornutus</emphasis>
</taxonomicName>
) and wet rock walls (
<taxonomicName id="5DEB4D1DFF88FFBFFB9D20E2A1B37A96" box="[1112,1254,450,476]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="olangapo" status="sp. nov.">
<emphasis id="A89FEA8CFF88FFBFFB9D20E2A1B37A96" box="[1112,1254,450,476]" italics="true" pageId="4" pageNumber="5">P. olangapo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF88FFBFFB3520E3A01B7A96" box="[1264,1358,450,476]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
) to dark sheltered spaces among rocks (
<taxonomicName id="5DEB4D1DFF88FFBFFDB120C7A79A7AB5" box="[628,719,485,511]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mulu" status="sp. nov.">
<emphasis id="A89FEA8CFF88FFBFFDB120C7A79A7AB5" box="[628,719,485,511]" italics="true" pageId="4" pageNumber="5">P. mulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF88FFBFFD1C20C4A66C7AB5" box="[729,825,485,511]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="5DEB4D1DFF88FFBFFC8220C7A6ED7AB5" box="[839,952,485,511]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baguio" status="sp. nov.">
<emphasis id="A89FEA8CFF88FFBFFC8220C7A6ED7AB5" box="[839,952,485,511]" italics="true" pageId="4" pageNumber="5">P. baguio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF88FFBFFC0620C4A1767AB5" box="[963,1059,485,511]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="5DEB4D1DFF88FFBFFBF420C7A1EC7AB5" box="[1073,1209,485,511]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pagbilao">
<emphasis id="A89FEA8CFF88FFBFFBF420C7A1EC7AB5" box="[1073,1209,485,511]" italics="true" pageId="4" pageNumber="5">P. pagbilao</emphasis>
</taxonomicName>
), large holes at tree bases (
<taxonomicName id="5DEB4D1DFF88FFBFFE8D2328A4FD7968" box="[328,424,520,546]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kawit" status="sp. nov.">
<emphasis id="A89FEA8CFF88FFBFFE8D2328A4FD7968" box="[328,424,520,546]" italics="true" pageId="4" pageNumber="5">P. kawit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF88FFBFFE762328A7417969" box="[435,532,521,547]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
), abandoned buildings (
<taxonomicName id="5DEB4D1DFF88FFBFFCFD2328A6957968" box="[824,960,520,546]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pagbilao">
<emphasis id="A89FEA8CFF88FFBFFCFD2328A6957968" box="[824,960,520,546]" italics="true" pageId="4" pageNumber="5">P. pagbilao</emphasis>
</taxonomicName>
), and open space among vegetation exposed to direct sunlight (
<taxonomicName id="5DEB4D1DFF88FFBFFDC7230DA73B790F" box="[514,622,556,581]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arayat">
<emphasis id="A89FEA8CFF88FFBFFDC7230DA73B790F" box="[514,622,556,581]" italics="true" pageId="4" pageNumber="5">P. arayat</emphasis>
</taxonomicName>
). Webs mainly consisted of a domed sheet with a diameter of up to ~
<quantity id="5D139B7BFF88FFBFFF2C236FA4677923" box="[233,306,590,617]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="cm" value="50.0">50 cm</quantity>
(
<taxonomicName id="5DEB4D1DFF88FFBFFE87236EA4CC7923" box="[322,409,591,617]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mulu" status="sp. nov.">
<emphasis id="A89FEA8CFF88FFBFFE87236EA4CC7923" box="[322,409,591,617]" italics="true" pageId="4" pageNumber="5">P. mulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF88FFBFFE5A236EA4A97923" box="[415,508,591,617]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
), but were much smaller in the rock-dwelling
<taxonomicName id="5DEB4D1DFF88FFBFFBD6236EA1CB7923" box="[1043,1182,591,617]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="olangapo" status="sp. nov.">
<emphasis id="A89FEA8CFF88FFBFFBD6236EA1CB7923" box="[1043,1182,591,617]" italics="true" pageId="4" pageNumber="5">P. olangapo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF88FFBFFB61236EA1AB7923" box="[1188,1278,591,617]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(~
<quantity id="5D139B7BFF88FFBFFADB236FA03F7923" box="[1310,1386,590,617]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="cm" value="15.0">15 cm</quantity>
). In
<taxonomicName id="5DEB4D1DFF88FFBFFF1B2352A41879C6" box="[222,333,626,652]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baguio" status="sp. nov.">
<emphasis id="A89FEA8CFF88FFBFFF1B2352A41879C6" box="[222,333,626,652]" italics="true" pageId="4" pageNumber="5">P. baguio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3AC57F7FF88FFBFFE932353A4E179C6" box="[342,436,626,652]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, webs were connected to each other and formed large communal structures up to
<quantity id="5D139B7BFF88FFBFFF7823B4A5B879E5" box="[189,237,661,687]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="m" value="2.0">2 m</quantity>
in length. When disturbed, most species were observed to run toward the periphery of the web, seeking shelter at the substrate rather than staying and whirling in the web.
</paragraph>
</subSubSection>
<subSubSection id="D2F16515FF88FFBFFF7823D8A4CE782F" pageId="4" pageNumber="5" type="distribution">
<paragraph id="9A54369EFF88FFBFFF7823D8A4037859" blockId="4.[189,342,761,787]" box="[189,342,761,787]" pageId="4" pageNumber="5">
<heading id="C11C81F2FF88FFBFFF7823D8A4037859" bold="true" box="[189,342,761,787]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="2">
<emphasis id="A89FEA8CFF88FFBFFF7823D8A4037859" bold="true" box="[189,342,761,787]" pageId="4" pageNumber="5">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9A54369EFF88FFBFFF782209A4CE782F" blockId="4.[189,1398,808,870]" pageId="4" pageNumber="5">
The
<taxonomicName id="5DEB4D1DFF88FFBFFF332209A4887808" box="[246,477,808,834]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bicornutus">
<emphasis id="A89FEA8CFF88FFBFFF332209A4887808" box="[246,477,808,834]" italics="true" pageId="4" pageNumber="5">Pholcus bicornutus</emphasis>
</taxonomicName>
group is largely restricted to the Philippines, with only one species in NE Borneo (
<figureCitation id="02D02A1BFF88FFBFFEE6226AA4D8782F" box="[291,397,843,869]" captionStart="Figs 1 2" captionStartId="3.[189,241,2000,2026]" captionTargetBox="[189,1398,1092,1972]" captionTargetId="figure@3.[189,1398,1092,1972]" captionTargetPageId="3" captionText="Figs 1 2. Known distributions of the two subgroups of the Pholcus bicornutus species group." httpUri="https://zenodo.org/record/830816/files/figure.png" pageId="4" pageNumber="5">Figs 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>