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<document ID-DOI="10.11646/zootaxa.5271.1.3" ID-GBIF-Dataset="27e19a5d-bfc7-4434-9478-5fd9ed0e0b0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7864266" ID-ZooBank="39D641B7-1800-4918-8E88-4EC5FF4BB56C" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1682330975026" checkinUser="plazi" docAuthor="Zhang, Jing, Dolibaina, Diego R., Cong, Qian, Shen, Jinhui, Song, Leina, Mielke, Carlos G. C., Casagrande, Mirna M., Mielke, Olaf H. H. &amp; Grishin, Nick V." docDate="2023" docId="F84A87F49B27FFD2FF3CA450BE74FDF1" docLanguage="en" docName="zootaxa.5271.1.3.pdf" docOrigin="Zootaxa 5271 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5271.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vehilius spitzi Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin 2023" docType="treatment" docVersion="3" lastPageNumber="108" masterDocId="0473FF8C9B36FFC3FFABA54BBA00FF32" masterDocTitle="Taxonomic notes on Neotropical Hesperiidae (Lepidoptera)" masterLastPageNumber="114" masterPageNumber="91" pageNumber="108" updateTime="1682444975477" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Taxonomic notes on Neotropical Hesperiidae (Lepidoptera)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Jing</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dolibaina, Diego R.</mods:namePart>
<mods:affiliation>Reserva Serra Bonita, Camacã, Bahia, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cong, Qian</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shen, Jinhui</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Song, Leina</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mielke, Carlos G. C.</mods:namePart>
<mods:affiliation>Caixa postal 1206, 84.145 - 000 Carambeí, Paraná, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Casagrande, Mirna M.</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Universidade Federal do Paraná, Caixa postal 19020, 81531 - 980 Curitiba, Paraná, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mielke, Olaf H. H.</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Universidade Federal do Paraná, Caixa postal 19020, 81531 - 980 Curitiba, Paraná, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grishin, Nick V.</mods:namePart>
<mods:affiliation>National Museum of Natural History, Smithsonian Institution, Washington, DC, USA.</mods:affiliation>
</mods:name>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-24</mods:number>
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<mods:detail type="volume">
<mods:number>5271</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:extent unit="page">
<mods:start>91</mods:start>
<mods:end>114</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5271.1.3</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5271.1.3</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7864340" ID-Zenodo-Dep="7864340" LSID="urn:lsid:plazi:treatment:F84A87F49B27FFD2FF3CA450BE74FDF1" httpUri="http://treatment.plazi.org/id/F84A87F49B27FFD2FF3CA450BE74FDF1" lastPageNumber="108" pageId="17" pageNumber="108">
<subSubSection pageId="17" pageNumber="108" type="nomenclature">
<paragraph blockId="17.[151,1436,283,346]" pageId="17" pageNumber="108">
<heading bold="true" centered="true" fontSize="11" level="1" pageId="17" pageNumber="108" reason="1">
<emphasis bold="true" pageId="17" pageNumber="108">
<taxonomicName authority="(O. Mielke, 1967)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1967" box="[151,544,283,310]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="spitzi">
<emphasis bold="true" box="[151,318,283,310]" italics="true" pageId="17" pageNumber="108">Vehilius spitzi</emphasis>
(O. Mielke, 1967)
</taxonomicName>
,
<taxonomicName authority="(Evans, 1955)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[557,902,283,310]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tanta">
<emphasis bold="true" box="[557,724,283,310]" italics="true" pageId="17" pageNumber="108">Vehilius tanta</emphasis>
(
<bibRefCitation author="Evans, W. H." box="[740,894,284,310]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName authority="(O. Mielke, 1980)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1980" box="[916,1375,283,310]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="campestris">
<emphasis bold="true" box="[916,1150,283,310]" italics="true" pageId="17" pageNumber="108">Vehilius campestris</emphasis>
(O. Mielke, 1980)
</taxonomicName>
, and
<taxonomicName authority="(Evans, 1955)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[151,490,319,346]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tinta">
<emphasis bold="true" box="[151,311,319,346]" italics="true" pageId="17" pageNumber="108">Vehilius tinta</emphasis>
(
<bibRefCitation author="Evans, W. H." box="[327,481,320,346]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
)
</taxonomicName>
, new combinations
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="108" type="description">
<paragraph blockId="17.[151,1437,392,707]" pageId="17" pageNumber="108">
<taxonomicName authority="O. Mielke, 1967" authorityName="O. Mielke" authorityYear="1967" box="[151,477,392,419]" class="Insecta" family="Hesperiidae" genus="Vidius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="spitzi">
<emphasis box="[151,287,392,418]" italics="true" pageId="17" pageNumber="108">Vidius spitzi</emphasis>
O. Mielke, 1967
</taxonomicName>
and
<taxonomicName authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[531,804,392,419]" class="Insecta" family="Hesperiidae" genus="Vidius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tinta">
<emphasis box="[531,658,392,418]" italics="true" pageId="17" pageNumber="108">Vidius tinta</emphasis>
<bibRefCitation author="Evans, W. H." box="[665,804,392,419]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
(
<typeStatus box="[818,868,392,418]" pageId="17" pageNumber="108">type</typeStatus>
locality for the two in
<collectingCountry box="[1124,1197,392,418]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
:
<collectingRegion box="[1207,1276,392,418]" country="Brazil" name="Goias" pageId="17" pageNumber="108">Goiás</collectingRegion>
),
<taxonomicName authority="Evans, 1955" authorityName="Evans" authorityYear="1955" class="Insecta" family="Hesperiidae" genus="Nastra" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tanta">
<emphasis box="[1296,1437,393,418]" italics="true" pageId="17" pageNumber="108">Nastra tanta</emphasis>
<bibRefCitation author="Evans, W. H." box="[151,292,428,455]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
(
<typeStatus box="[309,359,428,454]" pageId="17" pageNumber="108">type</typeStatus>
locality in
<collectingCountry box="[493,566,428,454]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
:
<collectingRegion box="[578,731,428,455]" country="Brazil" name="Mato Grosso" pageId="17" pageNumber="108">Mato Grosso</collectingRegion>
), and
<taxonomicName authority="O. Mielke, 1980" authorityName="O. Mielke" authorityYear="1980" box="[802,1245,428,455]" class="Insecta" family="Hesperiidae" genus="Cymaenes" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="campestris">
<emphasis box="[802,1047,428,454]" italics="true" pageId="17" pageNumber="108">Cymaenes campestris</emphasis>
O. Mielke, 1980
</taxonomicName>
(
<typeStatus box="[1262,1312,428,454]" pageId="17" pageNumber="108">type</typeStatus>
locality in
<collectingCountry box="[151,224,464,490]" name="Brazil" pageId="17" pageNumber="108">Brazil</collectingCountry>
:
<collectingRegion box="[235,313,464,490]" country="Brazil" name="Parana" pageId="17" pageNumber="108">Paraná</collectingRegion>
) (
<figureCitation box="[336,403,464,491]" captionStart="FIGURE 7" captionStartId="18.[151,250,1760,1784]" captionTargetBox="[217,1368,185,1736]" captionTargetId="figure-26@18.[214,1374,181,1736]" captionTargetPageId="18" captionText="FIGURE 7. Nuclear genome tree of selected Moncina: Lerema relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Rectava (red, with Rectava cascatona, comb. nov. labeled in magenta), Lurida (lime, with Lurida zama, comb. nov. labeled in orange), Corra (cyan, with Corra xanthus, comb. nov. and former subspecies of Corra coryna labeled in purple and darker cyan, respectively), Vidius (blue, with species transferred from Cobalopsis labeled in dark blue), Cymaenes (green, with species transferred from Vidius or Nastra and Cymaenes macintyrei, stat. rest. labeled in olive and lilac), and Vehilius (gray, with species transferred from Vidius, Nastra, or Cymaenes labeled in pink). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864286" httpUri="https://zenodo.org/record/7864286/files/figure.png" pageId="17" pageNumber="108">Fig. 7</figureCitation>
pink) originate within the species of
<taxonomicName authority="Godman, 1900" authorityName="Godman" authorityYear="1900" box="[823,1086,464,490]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis box="[823,912,464,490]" italics="true" pageId="17" pageNumber="108">Vehilius</emphasis>
Godman, 1900
</taxonomicName>
(
<typeStatus box="[1101,1151,464,490]" pageId="17" pageNumber="108">type</typeStatus>
species
<taxonomicName authority="Mabille, 1891" authorityName="Mabille" authorityYear="1891" class="Insecta" family="Hesperiidae" genus="Cobalus" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="illudens">
<emphasis box="[1247,1437,464,490]" italics="true" pageId="17" pageNumber="108">Cobalus illudens</emphasis>
Mabille, 1891
</taxonomicName>
, currently a subspecies of
<taxonomicName authority="Butler, 1877" authorityName="Butler" authorityYear="1877" box="[627,1030,500,526]" class="Insecta" family="Hesperiidae" genus="Pamphila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="stictomenes">
<emphasis box="[627,877,500,526]" italics="true" pageId="17" pageNumber="108">Pamphila stictomenes</emphasis>
Butler, 1877
</taxonomicName>
) (
<figureCitation box="[1057,1127,500,527]" captionStart="FIGURE 7" captionStartId="18.[151,250,1760,1784]" captionTargetBox="[217,1368,185,1736]" captionTargetId="figure-26@18.[214,1374,181,1736]" captionTargetPageId="18" captionText="FIGURE 7. Nuclear genome tree of selected Moncina: Lerema relatives. See Fig. 4 for notations. Some genera are labeled at their clades. Taxa discussed in the text are shown in different colors: Rectava (red, with Rectava cascatona, comb. nov. labeled in magenta), Lurida (lime, with Lurida zama, comb. nov. labeled in orange), Corra (cyan, with Corra xanthus, comb. nov. and former subspecies of Corra coryna labeled in purple and darker cyan, respectively), Vidius (blue, with species transferred from Cobalopsis labeled in dark blue), Cymaenes (green, with species transferred from Vidius or Nastra and Cymaenes macintyrei, stat. rest. labeled in olive and lilac), and Vehilius (gray, with species transferred from Vidius, Nastra, or Cymaenes labeled in pink). Green arrows point from the clade of the genus where a species was previous placed to the clade of the species that is being transferred between genera." figureDoi="http://doi.org/10.5281/zenodo.7864286" httpUri="https://zenodo.org/record/7864286/files/figure.png" pageId="17" pageNumber="108">Fig. 7</figureCitation>
gray) as evidenced by the nuclear genome tree. Typical COI barcode differences among these species are around 3%5%: e.g.,
<taxonomicName authorityName="O. Mielke" authorityYear="1967" box="[1272,1357,537,562]" class="Insecta" family="Hesperiidae" genus="Vidius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="spitzi">
<emphasis box="[1272,1357,537,562]" italics="true" pageId="17" pageNumber="108">V. spitzi</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Evans" authorityYear="1955" box="[214,307,573,598]" class="Insecta" family="Hesperiidae" genus="Nastra" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tanta">
<emphasis box="[214,307,573,598]" italics="true" pageId="17" pageNumber="108">N. tanta</emphasis>
</taxonomicName>
by 3% (20 bp) and
<taxonomicName authorityName="O. Mielke" authorityYear="1980" box="[539,694,572,598]" class="Insecta" family="Hesperiidae" genus="Cymaenes" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="campestris">
<emphasis box="[539,694,572,598]" italics="true" pageId="17" pageNumber="108">C. campestris</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Evans" authorityYear="1955" box="[847,927,573,598]" class="Insecta" family="Hesperiidae" genus="Vidius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tinta">
<emphasis box="[847,927,573,598]" italics="true" pageId="17" pageNumber="108">V. tinta</emphasis>
</taxonomicName>
by 4.7% (31 bp), with the largest difference being between
<taxonomicName box="[320,574,608,634]" class="Insecta" family="Hesperiidae" genus="Pamphila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="subSpecies" species="stictomenes" subSpecies="illudens">
<emphasis box="[320,574,608,634]" italics="true" pageId="17" pageNumber="108">V. stictomenes illudens</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="O. Mielke" authorityYear="1980" box="[628,782,608,634]" class="Insecta" family="Hesperiidae" genus="Cymaenes" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="campestris">
<emphasis box="[628,782,608,634]" italics="true" pageId="17" pageNumber="108">C. campestris</emphasis>
</taxonomicName>
: 8.1% (53 bp). Therefore, we place all these species in the same genus as
<taxonomicName authority="(O. Mielke, 1967)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1967" box="[325,700,644,671]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="spitzi" status="comb. nov.">
<emphasis box="[325,482,644,670]" italics="true" pageId="17" pageNumber="108">Vehilius spitzi</emphasis>
(O. Mielke, 1967)
</taxonomicName>
,
<emphasis bold="true" box="[714,845,644,670]" pageId="17" pageNumber="108">
<taxonomicNameLabel box="[714,845,644,670]" pageId="17" pageNumber="108" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName authority="(Evans, 1955)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[861,1187,644,671]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tanta" status="comb. nov.">
<emphasis box="[861,1017,644,670]" italics="true" pageId="17" pageNumber="108">Vehilius tanta</emphasis>
(
<bibRefCitation author="Evans, W. H." box="[1033,1178,644,671]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
)
</taxonomicName>
,
<emphasis bold="true" box="[1201,1331,644,670]" pageId="17" pageNumber="108">
<taxonomicNameLabel box="[1201,1331,644,670]" pageId="17" pageNumber="108" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName authority="(O. Mielke, 1980)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="O. Mielke" baseAuthorityYear="1980" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="campestris" status="comb. nov.">
<emphasis italics="true" pageId="17" pageNumber="108">Vehilius campestris</emphasis>
(O. Mielke, 1980)
</taxonomicName>
,
<emphasis bold="true" box="[496,624,680,706]" pageId="17" pageNumber="108">
<taxonomicNameLabel box="[496,624,680,706]" pageId="17" pageNumber="108" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName authority="(Evans, 1955)" authorityName="Zhang &amp; Dolibaina &amp; Cong &amp; Shen &amp; Song &amp; Mielke &amp; Casagrande &amp; Mielke &amp; Grishin" authorityYear="2023" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[685,999,680,707]" class="Insecta" family="Hesperiidae" genus="Vehilius" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="108" phylum="Arthropoda" rank="species" species="tinta" status="comb. nov.">
<emphasis box="[685,832,680,706]" italics="true" pageId="17" pageNumber="108">Vehilius tinta</emphasis>
(
<bibRefCitation author="Evans, W. H." box="[847,990,680,707]" pageId="17" pageNumber="108" pagination="454 - 488" refId="ref17231" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. The Trustees of the British Museum (Natural History), London, v + 499 pp., pls. 454 - 488." type="book chapter" year="1955">Evans, 1955</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel box="[1011,1140,680,707]" pageId="17" pageNumber="108" rank="species">
<emphasis bold="true" box="[1011,1134,680,706]" pageId="17" pageNumber="108">comb. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>