treatments-xml/data/9E/3C/FB/9E3CFB429E11FFA51D4AFCC0BED8F936.xml

<document id="0FF735C4D5DED1A0FEB2D9C687B0E2E2" ID-CLB-Dataset="3950" ID-DOI="10.5281/zenodo.1414221" ID-GBIF-Dataset="38a7ff37-34bc-4331-8476-7d539028c4ef" ID-ISSN="0035-418" ID-Zenodo-Dep="1414221" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634784310621" checkinUser="plazi" docAuthor="Luu, Vinh Quang, Bonkowski, Michael, Nguyen, Truong Quang, Le, Minh Duc, Calame, Thomas &amp; Ziegler, Thomas" docDate="2018" docId="9E3CFB429E11FFA51D4AFCC0BED8F936" docLanguage="en" docName="8_Luu_etal.pdf" docOrigin="Revue suisse de Zoologie 125 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Lycodon banksi Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler 2018, sp. nov." docType="treatment" docVersion="6" lastPageNumber="271" masterDocId="6205833A9E12FFAD1C0BFF92BA02FFF0" masterDocTitle="A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos" masterLastPageNumber="276" masterPageNumber="263" pageNumber="266" updateTime="1699464091778" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="43D05933C75CC4FC32C411795C1AE2F7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F899BAE98668900832983388C4011E8B">
<mods:title id="B9445A096467462AF71473B178DE7EA5">A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos</mods:title>
</mods:titleInfo>
<mods:name id="2821352E4004335F9EFC38E50DCB2842" type="personal">
<mods:role id="89D55981421F95D86330F83C21BA9DD3">
<mods:roleTerm id="A8EC1CBF0DF9B6B7B583DD57A6147355">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A05FF38C4E1DB56AF5657795B314786D">Luu, Vinh Quang</mods:namePart>
<mods:affiliation id="5F53B6406F1E363A2EBA05B47A1FA9CF">Department of Wildlife, Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam. E-mail: qvinhfuv @ yahoo. com. au</mods:affiliation>
<mods:nameIdentifier id="FE968F8AA981D008C189B6EDFA7A5A77" type="email">qvinhfuv@yahoo.com.au</mods:nameIdentifier>
</mods:name>
<mods:name id="CE97216B9452E85943595DB8791DCC83" type="personal">
<mods:role id="5AD6A68A5A4AA82065739DC4608FE0B6">
<mods:roleTerm id="7E24F6D75391D6ED2D4E93C619FB9AEF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09105FE43666B56856E25327A4FBF916">Bonkowski, Michael</mods:namePart>
<mods:affiliation id="6A70329E494F634E4AB45ACD2FB6015B">Institute of Zoology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany.</mods:affiliation>
</mods:name>
<mods:name id="3FDCC11F0192F848ABBE4F6EB55EFBB9" type="personal">
<mods:role id="B28248D241153A5A23A5DF34B52A8222">
<mods:roleTerm id="0AB6EE32F6F5C7BFBDC1AF9D684771C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F31D80C33E5E9D73D3FB9BE47973CF39">Nguyen, Truong Quang</mods:namePart>
<mods:affiliation id="7A57371A37831EBC4A86575525305ECE">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam. E-mail: nqt 2 @ yahoo. com &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name id="008BF41BBEBD257299EBBC60D5C608E0" type="personal">
<mods:role id="E435EFB9D6272DE6F0FAAE53C27F222A">
<mods:roleTerm id="F013CED50E4A648B8A8B78F5D5D55694">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="42274F1B7AAA36DF2D2F068C228644B3">Le, Minh Duc</mods:namePart>
<mods:affiliation id="A20DACF9C7460C26AD3419F57BB3CF25">Faculty of Environmental Sciences, Hanoi University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, Vietnam. E-mail: le. duc. minh @ hus. edu. v / Central Institute for Natural Resources and Environmental Studies, Hanoi National University, 19 Le Thanh Tong, Hanoi, Vietnam / Department of Herpetology, American Museum of Natural History, Central Park West at 79 th Street, New York 10024</mods:affiliation>
</mods:name>
<mods:name id="0330D2FDD460F5F53397F7D6588FC86F" type="personal">
<mods:role id="3D9B333ED6147BACFED38F18083FE50F">
<mods:roleTerm id="DE67E4B155A11F494CA13E2C5EE5CEDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F0A3F6E9A68DD491E15C7DA710DCBF9">Calame, Thomas</mods:namePart>
<mods:affiliation id="790ABC58F0F9B46D91B8E77CA6E2AAE9">WWF Greater Mekong, House No. 39, Unit 05, Ban Saylom, Vientiane, Lao PDR. E-mail: calame @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="79D14DADFEC135A3A17B3A016CE18C7A" type="personal">
<mods:role id="F0AFBF6BD126376F2CACDE130C06B3B5">
<mods:roleTerm id="EF3AEF8D9F280B80584F432EBF99BAFC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9AA0D79944D700607A761C98F30E6CEF">Ziegler, Thomas</mods:namePart>
<mods:affiliation id="1AA871F4582D8C65F782527CAEB106E5">Institute of Zoology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="89557EFA572A3A4AB26A189A9E0DD9F5">text</mods:typeOfResource>
<mods:relatedItem id="348B0EDAD72E5327F36A0744C73916EC" type="host">
<mods:titleInfo id="A9147E99CDFA4C9DF2C330B6FE8D10A6">
<mods:title id="ACEB47DC04DE35738763B1E35E5B67A4">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="84B29C4886E3B609D39E2B4F8EB460E2">
<mods:date id="9EA5469D6312BD422C59E39E755E2136">2018</mods:date>
<mods:detail id="0384B4D86C139DD9E14D12D7FA6AA5C7" type="pubDate">
<mods:number id="BC406004B6D2334AFF33FA99EA30195B">2018-09-28</mods:number>
</mods:detail>
<mods:detail id="B160FF9279DEA0B39F19FCB2A184959A" type="volume">
<mods:number id="676C174497AF5F5ED820D6E01D76767A">125</mods:number>
</mods:detail>
<mods:detail id="61D678168E129DD9F36187A83BF5256B" type="issue">
<mods:number id="E4283DBCBD5863C613CA2586771BA909">2</mods:number>
</mods:detail>
<mods:extent id="3F7EAB81FA1389D51E1D1EEE29025563" unit="page">
<mods:start id="D8810CB3247958844F6FEED47266B017">263</mods:start>
<mods:end id="987465EE10BA8B9D9ACB4EBF3276FAD4">276</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8A0455DA63DC8B9398DB54911A93DE10">journal article</mods:classification>
<mods:identifier id="F8F2FD4628F5E92B8EBAB790CCE35CEF" type="CLB-Dataset">3950</mods:identifier>
<mods:identifier id="89C527A9CACA8B1AD349AD5B20F392B4" type="DOI">10.5281/zenodo.1414221</mods:identifier>
<mods:identifier id="6511366CCE318888A4BA6CD01DDCB939" type="GBIF-Dataset">38a7ff37-34bc-4331-8476-7d539028c4ef</mods:identifier>
<mods:identifier id="C015829621BA5E9152216A16F58C9F40" type="ISSN">0035-418</mods:identifier>
<mods:identifier id="C2AC01B317830A4A7735CC45513D04AA" type="Zenodo-Dep">1414221</mods:identifier>
</mods:mods>
<treatment id="9E3CFB429E11FFA51D4AFCC0BED8F936" ID-DOI="http://doi.org/10.5281/zenodo.5590023" ID-GBIF-Taxon="189149755" ID-Zenodo-Dep="5590023" LSID="urn:lsid:plazi:treatment:9E3CFB429E11FFA51D4AFCC0BED8F936" httpUri="http://treatment.plazi.org/id/9E3CFB429E11FFA51D4AFCC0BED8F936" lastPageId="8" lastPageNumber="271" pageId="3" pageNumber="266">
<subSubSection id="5E8F19DF9E11FFAE1D4AFCC0B848FC9A" box="[321,586,850,874]" pageId="3" pageNumber="266" type="nomenclature">
<paragraph id="162A4A549E11FFAE1D4AFCC0B848FC9A" blockId="3.[321,586,850,907]" box="[321,586,850,874]" pageId="3" pageNumber="266">
<heading id="4D62FD389E11FFAE1D4AFCC0B848FC9A" bold="true" box="[321,586,850,874]" centered="true" level="2" pageId="3" pageNumber="266" reason="2">
<emphasis id="24E196469E11FFAE1D4AFCC0B848FC9A" bold="true" box="[321,586,850,874]" pageId="3" pageNumber="266">
<taxonomicName id="D19531D79E11FFAE1D4AFCC0BBEDFC9A" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[321,495,850,874]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E11FFAE1D4AFCC0BBEDFC9A" bold="true" box="[321,495,850,874]" italics="true" pageId="3" pageNumber="266">Lycodon banksi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FD22B3D9E11FFAE1DFEFCC1B848FC9A" box="[501,586,851,874]" pageId="3" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5E8F19DF9E11FFAE1D67FCE6B81DFC7B" box="[364,543,884,907]" pageId="3" pageNumber="266" type="description">
<paragraph id="162A4A549E11FFAE1D67FCE6B81DFC7B" blockId="3.[321,586,850,907]" box="[364,543,884,907]" pageId="3" pageNumber="266">
<figureCitation id="8EAE56D19E11FFAE1D67FCE6BBC4FC7B" box="[364,454,884,907]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="6.[151,186,1310,1331]" captionStartId-1="7.[150,185,989,1010]" captionStartId-2="7.[151,186,1456,1477]" captionStartId-3="8.[149,184,1135,1156]" captionTargetBox-0="[151,1436,257,1267]" captionTargetBox-1="[151,1436,267,952]" captionTargetBox-2="[154,754,1093,1400]" captionTargetBox-3="[152,1436,256,1094]" captionTargetId-0="figure-360@6.[151,1436,255,1267]" captionTargetId-1="figure-417@7.[151,1436,257,952]" captionTargetId-2="figure-440@7.[148,758,1086,1417]" captionTargetId-3="figure-408@8.[151,1436,256,1094]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Fig. 2. Adult male holotype of Lycodon banksi sp. nov. (VNUF R.2015.20) in life. (A) dorsolateral view. (B) Head in dorsolateral view. (C) Head in dorsal view. Photos: V. Q. Luu." captionText-1="Fig. 3. Different head views of the adult male holotype of Lycodon banksi sp. nov. (VNUF R.2015.20). Photos V. Q. Luu." captionText-2="Fig. 4. Lateral head view of the adult male holotype of Lycodon banksi sp. nov. (VNUF R.2015.20). Drawing by T. Ziegler." captionText-3="Fig. 5. Additional record of Lycodon banksi sp. nov. in life: (A- B) dorsolateral views. (C) Head in dorsal view. Photos A-C A. Sahu. (D) Dorsal view. Photo P. Jaeger." figureDoi-0="http://doi.org/10.5281/zenodo.5590027" figureDoi-1="http://doi.org/10.5281/zenodo.5590029" figureDoi-2="http://doi.org/10.5281/zenodo.5590031" figureDoi-3="http://doi.org/10.5281/zenodo.5590035" httpUri-0="https://zenodo.org/record/5590027/files/figure.png" httpUri-1="https://zenodo.org/record/5590029/files/figure.png" httpUri-2="https://zenodo.org/record/5590031/files/figure.png" httpUri-3="https://zenodo.org/record/5590035/files/figure.png" pageId="3" pageNumber="266">Figs 2-5</figureCitation>
, 
<tableCitation id="5B177FEF9E11FFAE1DDAFCE6B81DFC7B" box="[465,543,884,907]" captionStart="Table 3" captionStartId="5.[151,204,258,279]" captionTargetPageId="5" captionText="Table 3. Measurements (in mm) and morphological characters of the holotype of Lycodon banksi sp. nov. (measurements in mm; for other abbreviations see material and methods; * tail tip lost)." httpUri="http://table.plazi.org/id/42EA1ADC9E17FFA81C9CFE90B93EFEC4" pageId="3" pageNumber="266" tableUuid="42EA1ADC9E17FFA81C9CFE90B93EFEC4">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="5E8F19DF9E11FFAE1C9CFC31BB9AFBAA" pageId="3" pageNumber="266" type="materials_examined">
<paragraph id="162A4A549E11FFAE1C9CFC31BB9AFBAA" blockId="3.[151,756,931,1114]" pageId="3" pageNumber="266">
<materialsCitation id="A6FD40099E11FFAE1C9CFC31BB9AFBAA" ID-GBIF-Occurrence="3391023301" collectingDate="2015-04-04" collectionCode="VNUF, R" country="Laos" county="Hinboun District" elevation="167" location="Vinh Quang Luu" municipality="Phou Hin Poun" pageId="3" pageNumber="266" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
<emphasis id="24E196469E11FFAE1C9CFC31BB08FC4A" bold="true" box="[151,266,931,954]" pageId="3" pageNumber="266">
<typeStatus id="C92EF4F69E11FFAE1C9CFC31BB07FC4A" box="[151,261,931,954]" pageId="3" pageNumber="266" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="7084D2919E11FFAE1D29FC31BB6EFC4A" box="[290,364,931,954]" pageId="3" pageNumber="266">VNUF</collectionCode>
<collectionCode id="7084D2919E11FFAE1D88FC31BB97FC4A" box="[387,405,931,954]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="266">R</collectionCode>
.2015.20 (field number: TK 20.15), adult male, collected on 
<date id="622B6C949E11FFAE1E08FC51B890FC2A" box="[515,658,963,986]" pageId="3" pageNumber="266" value="2015-04-04">
<collectingDate id="726F957C9E11FFAE1E08FC51B890FC2A" box="[515,658,963,986]" pageId="3" pageNumber="266" value="2015-04-04">4 April 2015</collectingDate>
</date>
by 
<location id="134A1C8F9E11FFAE1ECBFC51BB16FC0A" LSID="urn:lsid:plazi:treatment:9E3CFB429E11FFA51D4AFCC0BED8F936:134A1C8F9E11FFAE1ECBFC51BB16FC0A" country="Laos" county="Hinboun District" municipality="Phou Hin Poun" name="Vinh Quang Luu" pageId="3" pageNumber="266" stateProvince="Khammouan">Vinh Quang Luu</location>
and 
<location id="134A1C8F9E11FFAE1D45FC71B803FC0A" LSID="urn:lsid:plazi:treatment:9E3CFB429E11FFA51D4AFCC0BED8F936:134A1C8F9E11FFAE1D45FC71B803FC0A" box="[334,513,995,1018]" country="Laos" county="Hinboun District" municipality="Phou Hin Poun" name="Thomas Calame" pageId="3" pageNumber="266" stateProvince="Khammouan">Thomas Calame</location>
in the karst forest, at the mouth of a cave, 
<collectingMunicipality id="F64ED02E9E11FFAE1D9FFB91B841FBEA" box="[404,579,1027,1050]" pageId="3" pageNumber="266">Phou Hin Poun</collectingMunicipality>
NPA, 
<collectingCounty id="FF4B32D89E11FFAE1E9DFB91BAE9FBCA" pageId="3" pageNumber="266">Hinboun District</collectingCounty>
, 
<collectingRegion id="D45184B69E11FFAE1CF6FBB1B804FBCA" box="[253,518,1059,1082]" country="Laos" name="Khammouan" pageId="3" pageNumber="266">Khammouane Province</collectingRegion>
, central 
<collectingCountry id="6E820AC49E11FFAE1E64FBB1B8A4FBCA" box="[623,678,1059,1082]" name="Laos" pageId="3" pageNumber="266">Laos</collectingCountry>
, at an elevation of 
<elevation id="9DB8AD679E11FFAE1D17FBD1BB9AFBAA" box="[284,408,1091,1114]" metricMagnitude="2" metricUnit="m" metricValue="1.67" pageId="3" pageNumber="266" unit="m" value="167.0">
<quantity id="D16DE7B19E11FFAE1D17FBD1BB62FBAA" box="[284,352,1091,1114]" metricMagnitude="2" metricUnit="m" metricValue="1.67" pageId="3" pageNumber="266" unit="m" value="167.0">167 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5E8F19DF9E11FFAE1C9CFBE1BB14FA5B" pageId="3" pageNumber="266" type="diagnosis">
<paragraph id="162A4A549E11FFAE1C9CFBE1BB14FA5B" blockId="3.[151,756,1139,1451]" pageId="3" pageNumber="266">
<emphasis id="24E196469E11FFAE1C9CFBE1BB0DFB7A" bold="true" box="[151,271,1139,1162]" pageId="3" pageNumber="266">Diagnosis:</emphasis>
<taxonomicName id="D19531D79E11FFAE1D15FBE1BBCCFB7A" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[286,462,1139,1162]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E11FFAE1D15FBE1BBCCFB7A" box="[286,462,1139,1162]" italics="true" pageId="3" pageNumber="266">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E11FFAE1DD6FBE1B837FB7A" bold="true" box="[477,565,1139,1162]" pageId="3" pageNumber="266">
<taxonomicNameLabel id="3FD22B3D9E11FFAE1DD6FBE1B837FB7A" box="[477,565,1139,1162]" pageId="3" pageNumber="266" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
is characterized by the following morphological characters: (1) dorsal scales in 17-17-15 rows, dorsal scales on the anterior 2/3 of the body length smooth, the six central dorsal scale rows of the posterior 1/3 of the body length feebly keeled; (2) supralabials 8; (3) infralabials 10; (4) loreal entering orbit; (5) cloacal single; (6) ventral scales 241; (7) dorsal surface of body with 87 greyish yellow blotches; (8) ventral surface of body and tail uniformly grey cream.
</paragraph>
</subSubSection>
<subSubSection id="5E8F19DF9E11FFA51C9CFA51B8C1F8C7" lastPageId="8" lastPageNumber="271" pageId="3" pageNumber="266" type="description">
<paragraph id="162A4A549E11FFAE1C9CFA51B97AFE0A" blockId="3.[151,756,1475,2077]" lastBlockId="3.[831,1437,258,731]" pageId="3" pageNumber="266">
<emphasis id="24E196469E11FFAE1C9CFA51BBFEFA2A" bold="true" box="[151,508,1475,1498]" pageId="3" pageNumber="266">
Description of the 
<typeStatus id="C92EF4F69E11FFAE1D84FA51BBF5FA2A" box="[399,503,1475,1498]" pageId="3" pageNumber="266" type="holotype">holotype</typeStatus>
:
</emphasis>
Head elongate (HL 
<quantity id="D16DE7B19E11FFAE1C9CFA71BB03FA0A" box="[151,257,1507,1530]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="3" pageNumber="266" unit="mm" value="15.3">15.3 mm</quantity>
), moderately distinct from the neck, longer than wide (HW/HL ratio 0.71), depressed (HH/HL ratio 0.40), narrow anteriorly (IN/IO ratio 0.65); snout elongate (SnL/HL ratio 0.39); nostril lateral, oval shaped, located in the middle of the nasal; eye large (ED/HL ratio 0.17), pupils vertically elliptic; rostral triangular, much broader than high, hardly visible from above; nasal divided into two scales by a vertical ridge along posterior edge of nostril; two square internasals, as wide as long, bordered by two large, subpentagonal prefrontals posteriorly; frontal single, enlarged, pentagonal, narrowed posteriorly; parietals longer than wide, in contact with each other medially, with upper anterior and posterior temporals, paraparietal laterally and four nuchal scales posteriorly; loreal 1/1, elongate, entering orbit; supralabials 8/8, first and second in contact with nasal, third to fifth entering orbit, sixth largest; infralabials 10/10, first pair in broad contact with each other, first to fifth in contact with first pair of chin shields; first and second pairs of chin shields elongate, of the same size and shape, separated by a medial groove, first pair larger than the second; preocular 1/1; postoculars 2/2, of the same size, bordering anterior temporals; anterior temporals 2/2, posterior temporals 3/3, upper ones smaller than lower ones.
</paragraph>
<paragraph id="162A4A549E11FFAE1F34FD91BE75FD2B" blockId="3.[831,1437,258,731]" pageId="3" pageNumber="266">
Body elongate, SVL 
<quantity id="D16DE7B19E11FFAE1839FD91BE96FDEA" box="[1074,1172,515,538]" metricMagnitude="-1" metricUnit="m" metricValue="4.15" pageId="3" pageNumber="266" unit="mm" value="415.0">415 mm</quantity>
; TaL&gt; 
<quantity id="D16DE7B19E11FFAE18E1FD91BF39FDEA" box="[1258,1339,515,538]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="266" unit="mm" value="50.0">50 mm</quantity>
(tail tip lost); preventrals 2, ventrals 241; subcaudals 26 (tail tip lost), divided, weakly notched laterally; cloacal single; DSR 17-17-15; dorsal scales on the anterior 2/3 of the body length smooth, the six central dorsal scale rows of the posterior 1/3 of the body length feebly keeled; the vertebral scales not enlarged.
</paragraph>
<paragraph id="162A4A549E11FFAE1F34FD61BF0CFBFB" blockId="3.[831,1437,755,1035]" pageId="3" pageNumber="266">
<emphasis id="24E196469E11FFAE1F34FD61BE26FCFA" bold="true" box="[831,1060,755,778]" pageId="3" pageNumber="266">Colouration in life:</emphasis>
Head dark grey, without vertical light nuchal band; dorsal surface of body dark greyyellow with 87 greyish yellow irregular dorsal blotches; first body blotch starting at ventral scale 13, a half vertebral scale covered by this blotch; two yellow stripes on each side, from behind the neck to vent, indistinct posteriorly; ventral scales grey cream; dorsal surface of tail with at least eleven greyish yellow tail blotches, ventral surface of tail grey cream.
</paragraph>
<paragraph id="162A4A549E11FFAE1F34FBB1BEBEFBAA" blockId="3.[831,1437,1059,1114]" pageId="3" pageNumber="266">
<emphasis id="24E196469E11FFAE1F34FBB1B9C0FBCA" bold="true" box="[831,962,1059,1082]" pageId="3" pageNumber="266">Hemipenis:</emphasis>
The left hemipenis is only in part everted but shows a spinose ornamentation.
</paragraph>
<paragraph id="162A4A549E11FFAE1F34FBE1BE15FABB" blockId="3.[831,1437,1139,1355]" pageId="3" pageNumber="266">
<emphasis id="24E196469E11FFAE1F34FBE1BE39FB7A" bold="true" box="[831,1083,1139,1162]" pageId="3" pageNumber="266">Additional specimen:</emphasis>
<materialsCitation id="A6FD40099E11FFAE1843FBE1BE11FABB" ID-GBIF-Occurrence="3391023302" collectingDate="2016-07-22" county="Peter Jaeger" location="Jonas Ewert" municipality="Aloke Sahu" pageId="3" pageNumber="266" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
One specimen which was not collected but detected and photographed on 
<date id="622B6C949E11FFAE194CFB01B977FB3A" pageId="3" pageNumber="266" value="2016-07-22">
<collectingDate id="726F957C9E11FFAE194CFB01B977FB3A" pageId="3" pageNumber="266" value="2016-07-22">22 July 2016</collectingDate>
</date>
by an arachnology team consisting of 
<collectingCounty id="FF4B32D89E11FFAE196EFB21B987FB1A" pageId="3" pageNumber="266">Peter Jaeger</collectingCounty>
, 
<collectingMunicipality id="F64ED02E9E11FFAE1F98FB41BE16FB1A" box="[915,1044,1235,1258]" pageId="3" pageNumber="266">Aloke Sahu</collectingMunicipality>
and 
<location id="134A1C8F9E11FFAE185BFB41BEDBFB1A" LSID="urn:lsid:plazi:treatment:9E3CFB429E11FFA51D4AFCC0BED8F936:134A1C8F9E11FFAE185BFB41BEDBFB1A" box="[1104,1241,1235,1258]" county="Peter Jaeger" municipality="Aloke Sahu" name="Jonas Ewert" pageId="3" pageNumber="266" stateProvince="Khammouan">Jonas Ewert</location>
, in 
<collectingRegion id="D45184B69E11FFAE190DFB41B9A3FAFA" country="Laos" name="Khammouan" pageId="3" pageNumber="266">Khammouane Province</collectingRegion>
, in ca. 
<quantity id="D16DE7B19E11FFAE1FE4FB61BE45FAFA" box="[1007,1095,1267,1290]" metricMagnitude="4" metricUnit="m" metricValue="1.24" pageId="3" pageNumber="266" unit="km" value="12.4">12.4 km</quantity>
distance from the type locality. 
<location id="134A1C8F9E11FFAE1F34FA81B96BFADA" LSID="urn:lsid:plazi:treatment:9E3CFB429E11FFA51D4AFCC0BED8F936:134A1C8F9E11FFAE1F34FA81B96BFADA" box="[831,873,1299,1322]" county="Peter Jaeger" municipality="Aloke Sahu" name="The" pageId="3" pageNumber="266" stateProvince="Khammouan">The</location>
color pattern of this specimen resembles closely that of the 
<typeStatus id="C92EF4F69E11FFAE1FB9FAA6BE11FABB" box="[946,1043,1332,1355]" pageId="3" pageNumber="266" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="162A4A549E11FFAB1F34FAF1BBD0F84C" blockId="3.[831,1437,1379,2077]" lastBlockId="6.[151,756,1445,2076]" lastPageId="6" lastPageNumber="269" pageId="3" pageNumber="266">
<emphasis id="24E196469E11FFAE1F34FAF1B9E4FA8A" bold="true" box="[831,998,1379,1402]" pageId="3" pageNumber="266">Comparisons:</emphasis>
In our phylogenetic analysis, 
<taxonomicName id="D19531D79E11FFAE1F34FA11B9F8FA6A" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[831,1018,1411,1434]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E11FFAE1F34FA11B9F8FA6A" box="[831,1018,1411,1434]" italics="true" pageId="3" pageNumber="266">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E11FFAE181AFA11BE7AFA6A" bold="true" box="[1041,1144,1411,1434]" pageId="3" pageNumber="266">
<taxonomicNameLabel id="3FD22B3D9E11FFAE181AFA11BE7AFA6A" box="[1041,1144,1411,1434]" pageId="3" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is nested in a clade containing 
<taxonomicName id="D19531D79E11FFAE1FCAFA31BE65FA4A" box="[961,1127,1443,1466]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="rufozonatus">
<emphasis id="24E196469E11FFAE1FCAFA31BE65FA4A" box="[961,1127,1443,1466]" italics="true" pageId="3" pageNumber="266">L. rufozonatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="D19531D79E11FFAE1876FA31BF9AFA4A" authority="(Cope)" baseAuthorityName="Cope" baseAuthorityYear="1860" box="[1149,1432,1443,1466]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="semicarinatus">
<emphasis id="24E196469E11FFAE1876FA31BF39FA4A" box="[1149,1339,1443,1466]" italics="true" pageId="3" pageNumber="266">L. semicarinatus</emphasis>
(Cope)
</taxonomicName>
, ‘ 
<emphasis id="24E196469E11FFAE1F43FA51B9F8FA2A" box="[840,1018,1475,1499]" italics="true" pageId="3" pageNumber="266">L. flavozonatus’</emphasis>
, 
<taxonomicName id="D19531D79E11FFAE1807FA51BEAFFA2A" box="[1036,1197,1475,1498]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="futsingensis">
<emphasis id="24E196469E11FFAE1807FA51BEAFFA2A" box="[1036,1197,1475,1498]" italics="true" pageId="3" pageNumber="266">L. futsingensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="D19531D79E11FFAE18E7FA51BF97FA2A" box="[1260,1429,1475,1498]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="24E196469E11FFAE18E7FA51BF97FA2A" box="[1260,1429,1475,1498]" italics="true" pageId="3" pageNumber="266">L. meridionalis</emphasis>
</taxonomicName>
. The new species differs from the similar 
<taxonomicName id="D19531D79E11FFAE18F2FA71BF9EFA0A" box="[1273,1436,1507,1530]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="24E196469E11FFAE18F2FA71BF9EFA0A" box="[1273,1436,1507,1530]" italics="true" pageId="3" pageNumber="266">L. meridionalis</emphasis>
</taxonomicName>
by having loreal entering the orbit (
<emphasis id="24E196469E11FFAE18EEF991BF28F9EA" box="[1253,1322,1539,1562]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
separated from the orbit), dorsal scales on the anterior 2/3 of the body length smooth, the six central dorsal scale rows on the posterior body third feebly keeled (
<emphasis id="24E196469E11FFAE18E7F9F6BF32F98B" box="[1260,1328,1636,1659]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
distinctly keeled), dorsal head pattern uniform dark grey (
<emphasis id="24E196469E11FFAE1953F916BF9EF96B" box="[1368,1436,1668,1691]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
with yellow-black marbling in 
<taxonomicName id="D19531D79E11FFAE18A0F936BF55F94B" box="[1195,1367,1700,1723]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="24E196469E11FFAE18A0F936BF55F94B" box="[1195,1367,1700,1723]" italics="true" pageId="3" pageNumber="266">L. meridionalis</emphasis>
</taxonomicName>
), and ventral surface grey cream (
<emphasis id="24E196469E11FFAE188CF956BECEF92B" box="[1159,1228,1732,1755]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
yellow with dark spots posteriorly) (see 
<bibRefCitation id="720437A59E11FFAE1832F976BED0F90B" author="Bourret R." box="[1081,1234,1764,1787]" pageId="3" pageNumber="266" pagination="1934 - 1935" refId="ref7323" refString="Bourret R. 1935. Notes herpetologiques sur l'Indochine francaise. IX. Les Serpents de Chapa. Bulletin general de l'Instruction publique, 14 (1934 - 1935), 7 (Mars), 239 - 251 [Reprint: 5 - 17]." type="journal article" year="1935">Bourret, 1935</bibRefCitation>
; 
<bibRefCitation id="720437A59E11FFAE18EBF976B97BF8EC" author="Orlov N. &amp; Ryabov S. A." pageId="3" pageNumber="266" pagination="181 - 197" refId="ref8364" refString="Orlov N. &amp; Ryabov S. A. 2004. Revalidization and change of taxonomic status of Dinodon rufozonatus meridionale Bourret, 1935 (Serpentes: Colubridae: Colubrinae). Russian Journal Herpetology 11 (3): 181 - 197." type="journal article" year="2004">Orlov &amp; Ryabov, 2004</bibRefCitation>
); from 
<taxonomicName id="D19531D79E11FFAE1FD9F897BE71F8EC" box="[978,1139,1797,1820]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="rufozonatus">
<emphasis id="24E196469E11FFAE1FD9F897BE71F8EC" box="[978,1139,1797,1820]" italics="true" pageId="3" pageNumber="266">L. rufozonatus</emphasis>
</taxonomicName>
by having loreal entering the orbit (
<emphasis id="24E196469E11FFAE1FBBF8B7B9F6F8CC" box="[944,1012,1829,1852]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
usually separated), a distinctly higher ventral scale count (241 
<emphasis id="24E196469E11FFAE188AF8D7BEC4F8AC" box="[1153,1222,1861,1884]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
185-204), dorsal scales feebly keeled in the posterior body part (
<emphasis id="24E196469E11FFAE1953F8F7BF9EF88C" box="[1368,1436,1893,1916]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
all smooth), dorsal head pattern uniform dark grey (
<emphasis id="24E196469E11FFAE1F42F837B98CF84C" box="[841,910,1957,1980]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
dark brown with yellow borders), and body pattern blotched (
<emphasis id="24E196469E11FFAE181AF857BE54F82C" box="[1041,1110,1989,2012]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
banded) (
<bibRefCitation id="720437A59E11FFAE18DFF857BF92F82C" author="Boulenger G. A." box="[1236,1424,1989,2012]" pageId="3" pageNumber="266" refId="ref7283" refString="Boulenger G. A. 1893. Catalogue of the Snakes in the British Museum (Natural History). Volume I. Trustees of the British Museum, London, xiii + 448 pp., 28 plates." type="book" year="1893">Boulenger 1893</bibRefCitation>
); from 
<taxonomicName id="D19531D79E11FFAE1F8AF877BE38F80C" baseAuthorityName="Cope" baseAuthorityYear="1860" box="[897,1082,2021,2044]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="266" phylum="Chordata" rank="species" species="semicarinatus">
<emphasis id="24E196469E11FFAE1F8AF877BE38F80C" box="[897,1082,2021,2044]" italics="true" pageId="3" pageNumber="266">L. semicarinatus</emphasis>
</taxonomicName>
by having loreal touching the orbit (
<emphasis id="24E196469E11FFAE1F83F794B9CFF7ED" box="[904,973,2054,2077]" italics="true" pageId="3" pageNumber="266">versus</emphasis>
separated), a higher ventral scale count (241 
<emphasis id="24E196469E14FFAB1CDFFA37BB1BFA4C" box="[212,281,1445,1468]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
211-234), dorsal scale rows keeled along posterior 1/3 (
<emphasis id="24E196469E14FFAB1D3CFA57BB7EFA2C" box="[311,380,1477,1500]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
keeled along anterior half), belly pattern uniform grey cream (
<emphasis id="24E196469E14FFAB1DD0FA77B822FA0C" box="[475,544,1509,1532]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
yellow), and body pattern blotched (
<emphasis id="24E196469E14FFAB1D63F997BBAFF9EC" box="[360,429,1541,1564]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
banded) (
<bibRefCitation id="720437A59E14FFAB1E20F997B8E5F9EC" author="Boulenger G. A." box="[555,743,1541,1564]" pageId="6" pageNumber="269" refId="ref7283" refString="Boulenger G. A. 1893. Catalogue of the Snakes in the British Museum (Natural History). Volume I. Trustees of the British Museum, London, xiii + 448 pp., 28 plates." type="book" year="1893">Boulenger 1893</bibRefCitation>
); from 
<taxonomicName id="D19531D79E14FFAB1CDCF9B7BB7CF9CC" authorityName="' GP" authorityYear="1939" box="[215,382,1572,1596]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="flavozonatus">
<emphasis id="24E196469E14FFAB1CDCF9B7BB7CF9CC" box="[215,382,1572,1596]" italics="true" pageId="6" pageNumber="269">L. flavozonatus</emphasis>
</taxonomicName>
by having loreal in contact with the orbit (
<emphasis id="24E196469E14FFAB1D1CF9D7BB5EF9AC" box="[279,348,1605,1628]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
separated), cloacal single (
<emphasis id="24E196469E14FFAB1EA4F9D7B8F6F9AC" box="[687,756,1605,1628]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
divided), six dorsal scale rows on the posterior third of the body feebly keeled (
<emphasis id="24E196469E14FFAB1DC1F917B80CF96C" box="[458,526,1669,1692]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
10-12 keeled dorsal scale rows at midbody), dorsal head dark grey (
<emphasis id="24E196469E14FFAB1EBBF937B8F6F94C" box="[688,756,1701,1724]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
black with light markings), and belly pattern uniform grey cream (
<emphasis id="24E196469E14FFAB1D3AF977BB74F90C" box="[305,374,1765,1788]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
yellow with large black spots); from 
<taxonomicName id="D19531D79E14FFAB1CDFF897BB73F8EC" box="[212,369,1797,1820]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="futsingensis">
<emphasis id="24E196469E14FFAB1CDFF897BB73F8EC" box="[212,369,1797,1820]" italics="true" pageId="6" pageNumber="269">L. futsingensis</emphasis>
</taxonomicName>
by having loreal entering the orbit (
<emphasis id="24E196469E14FFAB1CABF8B7BAE7F8CC" box="[160,229,1829,1852]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
separated), a higher ventral scale count (241 
<emphasis id="24E196469E14FFAB1C9CF8D7BADEF8AC" box="[151,220,1861,1884]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
<quantity id="D16DE7B19E14FFAB1CECF8D7BB63F8AC" box="[231,353,1861,1884]" metricMagnitude="0" metricUnit="m" metricValue="5.0292" metricValueMax="5.1562" metricValueMin="4.9022" pageId="6" pageNumber="269" unit="in" value="198.0" valueMax="203.0" valueMin="193.0">193-203 in</quantity>
males), dorsal scales feebly keeled in the posterior body part (
<emphasis id="24E196469E14FFAB1DB0F8F7BBFDF88C" box="[443,511,1893,1916]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
all smooth), and body pattern blotched (
<emphasis id="24E196469E14FFAB1D6BF817BBA7F86C" box="[352,421,1925,1948]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
banded) (
<bibRefCitation id="720437A59E14FFAB1E17F817B8F2F86C" author="Vogel G. &amp; Nguyen T. Q. &amp; Kingsada P. &amp; Ziegler T." box="[540,752,1925,1948]" pageId="6" pageNumber="269" pagination="344 - 352" refId="ref9006" refString="Vogel G., Nguyen T. Q., Kingsada P. &amp; Ziegler T. 2012. A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology 8: 344 - 352." type="journal article" year="2012">
Vogel 
<emphasis id="24E196469E14FFAB1E63F817B89FF86C" box="[616,669,1925,1948]" italics="true" pageId="6" pageNumber="269">et al</emphasis>
., 2012
</bibRefCitation>
; 
<bibRefCitation id="720437A59E14FFAB1C9CF837BB5FF84C" author="Neang T. &amp; Hartmann T. &amp; Hun S. &amp; Souter N. J. &amp; Furey N. M." box="[151,349,1957,1980]" pageId="6" pageNumber="269" pagination="1" refId="ref8289" refString="Neang T., Hartmann T., Hun S., Souter N. J. &amp; Furey N. M. 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068 - 080. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
Neang 
<emphasis id="24E196469E14FFAB1CE8F837BB10F84C" box="[227,274,1957,1980]" italics="true" pageId="6" pageNumber="269">et al</emphasis>
., 2014
</bibRefCitation>
) (
<tableCitation id="5B177FEF9E14FFAB1D79F837BBC7F84C" box="[370,453,1957,1980]" captionStart="Table 4" captionStartId="11.[149,202,657,678]" captionTargetPageId="11" captionText="Table 4. Diagnostic characters separating Lycodon banksi sp. nov. from closely related species and the remaining Lycodon species recorded from Laos (data obtained from Boulenger, 1893; Smith, 1943; Taylor, 1965; Deuve, 1970; Orlov &amp; Ryabov, 2004; Vogel et al., 2009, 2012; Siler et al., 2013; Luu et al., 2013; Teynié et al., 2014; Neang et al., 2014)." httpUri="http://table.plazi.org/id/42EA1ADC9E19FFA61C9EFD03BD29FD33" pageId="6" pageNumber="269" tableUuid="42EA1ADC9E19FFA61C9EFD03BD29FD33">Table 4</tableCitation>
).
</paragraph>
<caption id="42EA1ADC9E16FFA91C92F875BEB4F7E9" ID-DOI="http://doi.org/10.5281/zenodo.5590025" ID-Zenodo-Dep="5590025" httpUri="https://zenodo.org/record/5590025/files/figure.png" pageId="4" pageNumber="267" startId="4.[153,188,2023,2044]" targetBox="[181,1396,258,1978]" targetPageId="4">
<paragraph id="162A4A549E16FFA91C92F875BEB4F7E9" blockId="4.[153,1438,2023,2073]" pageId="4" pageNumber="267">
Fig. 1. Bayesian cladogram based on the partial cytochrome 
<emphasis id="24E196469E16FFA91EECF875B8F1F80C" box="[743,755,2023,2044]" italics="true" pageId="4" pageNumber="267">b</emphasis>
gene. Numbers above and below branches are bootstrap values of MP/ ML analyses (&gt;50%) and Bayesian posterior probabilities, respectively. Asterisk denotes 100% value.
</paragraph>
</caption>
<caption id="42EA1ADC9E17FFA81C9CFE90B93EFEC4" ID-Table-UUID="42EA1ADC9E17FFA81C9CFE90B93EFEC4" httpUri="http://table.plazi.org/id/42EA1ADC9E17FFA81C9CFE90B93EFEC4" pageId="5" pageNumber="268" startId="5.[151,204,258,279]" targetBox="[166,1369,353,1931]" targetIsTable="true" targetPageId="5">
<paragraph id="162A4A549E17FFA81C9CFE90B93EFEC4" blockId="5.[151,1436,258,308]" pageId="5" pageNumber="268">
Table 3. Measurements (in mm) and morphological characters of the holotype of 
<taxonomicName id="D19531D79E17FFA81FA4FE90BE44FEE7" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[943,1094,258,279]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="268" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E17FFA81FA4FE90BE44FEE7" box="[943,1094,258,279]" italics="true" pageId="5" pageNumber="268">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E17FFA81847FE90BE9AFEE7" bold="true" box="[1100,1176,258,279]" pageId="5" pageNumber="268">
<taxonomicNameLabel id="3FD22B3D9E17FFA81847FE90BE9AFEE7" box="[1100,1176,258,279]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(measurements in mm; for other abbreviations see material and methods; * tail tip lost).
</paragraph>
</caption>
<paragraph id="162A4A549E17FFA81CADFEF3BE96F87B" pageId="5" pageNumber="268">
<table id="6495B8F49E1700521CADFEF3BF5BF87B" box="[166,1369,353,1931]" gridcols="2" gridrows="46" pageId="5" pageNumber="268">
<tr id="A8A548169E1700521CADFEF3BF5BFE85" box="[166,1369,353,373]" gridrow="0" pageId="5" pageNumber="268">
<th id="EB74216A9E1700521CADFEF3BBEAFE85" box="[166,488,353,373]" gridcol="0" gridrow="0" pageId="5" pageNumber="268">Character</th>
<th id="EB74216A9E1700521FBFFEF3BF5BFE85" box="[948,1369,353,373]" gridcol="1" gridrow="0" pageId="5" pageNumber="268">
<taxonomicName id="D19531D79E17FFA81FCEFEF3BE51FE85" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[965,1107,353,373]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="268" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E17FFA81FCEFEF3BE51FE85" box="[965,1107,353,373]" italics="true" pageId="5" pageNumber="268">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E17FFA81852FEF3BEA3FE85" bold="true" box="[1113,1185,353,373]" pageId="5" pageNumber="268">
<taxonomicNameLabel id="3FD22B3D9E17FFA81852FEF3BEA3FE85" box="[1113,1185,353,373]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
VNUF R.2015.20
</th>
</tr>
<tr id="A8A548169E1700521CADFE11BF5BFE67" box="[166,1369,387,407]" gridrow="1" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFE11BBEAFE67" box="[166,488,387,407]" gridcol="0" gridrow="1" pageId="5" pageNumber="268">Sex</td>
<td id="EB74216A9E1700521FBFFE11BF5BFE67" box="[948,1369,387,407]" gridcol="1" gridrow="1" pageId="5" pageNumber="268">Male</td>
</tr>
<tr id="A8A548169E1700521CADFE37BF5BFE49" box="[166,1369,421,441]" gridrow="2" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFE37BBEAFE49" box="[166,488,421,441]" gridcol="0" gridrow="2" pageId="5" pageNumber="268">SVL</td>
<td id="EB74216A9E1700521FBFFE37BF5BFE49" box="[948,1369,421,441]" gridcol="1" gridrow="2" pageId="5" pageNumber="268">415.0</td>
</tr>
<tr id="A8A548169E1700521CADFE55BF5BFE2B" box="[166,1369,455,475]" gridrow="3" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFE55BBEAFE2B" box="[166,488,455,475]" gridcol="0" gridrow="3" pageId="5" pageNumber="268">TaL</td>
<td id="EB74216A9E1700521FBFFE55BF5BFE2B" box="[948,1369,455,475]" gridcol="1" gridrow="3" pageId="5" pageNumber="268">50.0*</td>
</tr>
<tr id="A8A548169E1700521CADFE7BBF5BFE0D" box="[166,1369,489,509]" gridrow="4" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFE7BBBEAFE0D" box="[166,488,489,509]" gridcol="0" gridrow="4" pageId="5" pageNumber="268">TL</td>
<td id="EB74216A9E1700521FBFFE7BBF5BFE0D" box="[948,1369,489,509]" gridcol="1" gridrow="4" pageId="5" pageNumber="268">465.0</td>
</tr>
<tr id="A8A548169E1700521CADFD99BF5BFDEF" box="[166,1369,523,543]" gridrow="5" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFD99BBEAFDEF" box="[166,488,523,543]" gridcol="0" gridrow="5" pageId="5" pageNumber="268">HL</td>
<td id="EB74216A9E1700521FBFFD99BF5BFDEF" box="[948,1369,523,543]" gridcol="1" gridrow="5" pageId="5" pageNumber="268">15.3</td>
</tr>
<tr id="A8A548169E1700521CADFDBFBF5BFDB1" box="[166,1369,557,577]" gridrow="6" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFDBFBBEAFDB1" box="[166,488,557,577]" gridcol="0" gridrow="6" pageId="5" pageNumber="268">HW</td>
<td id="EB74216A9E1700521FBFFDBFBF5BFDB1" box="[948,1369,557,577]" gridcol="1" gridrow="6" pageId="5" pageNumber="268">10.8</td>
</tr>
<tr id="A8A548169E1700521CADFDDDBF5BFD93" box="[166,1369,591,611]" gridrow="7" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFDDDBBEAFD93" box="[166,488,591,611]" gridcol="0" gridrow="7" pageId="5" pageNumber="268">HH</td>
<td id="EB74216A9E1700521FBFFDDDBF5BFD93" box="[948,1369,591,611]" gridcol="1" gridrow="7" pageId="5" pageNumber="268">6.1</td>
</tr>
<tr id="A8A548169E1700521CADFDE3BF5BFD75" box="[166,1369,625,645]" gridrow="8" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFDE3BBEAFD75" box="[166,488,625,645]" gridcol="0" gridrow="8" pageId="5" pageNumber="268">IO</td>
<td id="EB74216A9E1700521FBFFDE3BF5BFD75" box="[948,1369,625,645]" gridcol="1" gridrow="8" pageId="5" pageNumber="268">6.3</td>
</tr>
<tr id="A8A548169E1700521CADFD01BF5BFD57" box="[166,1369,659,679]" gridrow="9" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFD01BBEAFD57" box="[166,488,659,679]" gridcol="0" gridrow="9" pageId="5" pageNumber="268">EN</td>
<td id="EB74216A9E1700521FBFFD01BF5BFD57" box="[948,1369,659,679]" gridcol="1" gridrow="9" pageId="5" pageNumber="268">3.3</td>
</tr>
<tr id="A8A548169E1700521CADFD27BF5BFD39" box="[166,1369,693,713]" gridrow="10" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFD27BBEAFD39" box="[166,488,693,713]" gridcol="0" gridrow="10" pageId="5" pageNumber="268">IN</td>
<td id="EB74216A9E1700521FBFFD27BF5BFD39" box="[948,1369,693,713]" gridcol="1" gridrow="10" pageId="5" pageNumber="268">4.1</td>
</tr>
<tr id="A8A548169E1700521CADFD45BF5BFD1B" box="[166,1369,727,747]" gridrow="11" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFD45BBEAFD1B" box="[166,488,727,747]" gridcol="0" gridrow="11" pageId="5" pageNumber="268">ED</td>
<td id="EB74216A9E1700521FBFFD45BF5BFD1B" box="[948,1369,727,747]" gridcol="1" gridrow="11" pageId="5" pageNumber="268">2.6</td>
</tr>
<tr id="A8A548169E1700521CADFD6BBF5BFCFD" box="[166,1369,761,781]" gridrow="12" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFD6BBBEAFCFD" box="[166,488,761,781]" gridcol="0" gridrow="12" pageId="5" pageNumber="268">SnL</td>
<td id="EB74216A9E1700521FBFFD6BBF5BFCFD" box="[948,1369,761,781]" gridcol="1" gridrow="12" pageId="5" pageNumber="268">6.0</td>
</tr>
<tr id="A8A548169E1700521CADFC89BF5BFCDF" box="[166,1369,795,815]" gridrow="13" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFC89BF5BFCDF" box="[166,1369,795,815]" colspan="2" colspanRight="1" gridcol="0" gridrow="13" pageId="5" pageNumber="268">
<emphasis id="24E196469E17FFA81CADFC89BAD1FCDF" bold="true" box="[166,211,795,815]" pageId="5" pageNumber="268">DSR</emphasis>
</td>
</tr>
<tr id="A8A548169E1700521CADFCAFBF5BFCA1" box="[166,1369,829,849]" gridrow="14" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFCAFBBEAFCA1" box="[166,488,829,849]" gridcol="0" gridrow="14" pageId="5" pageNumber="268">ASR</td>
<td id="EB74216A9E1700521FBFFCAFBF5BFCA1" box="[948,1369,829,849]" gridcol="1" gridrow="14" pageId="5" pageNumber="268">17</td>
</tr>
<tr id="A8A548169E1700521CADFCCDBF5BFC83" box="[166,1369,863,883]" gridrow="15" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFCCDBBEAFC83" box="[166,488,863,883]" gridcol="0" gridrow="15" pageId="5" pageNumber="268">MSR</td>
<td id="EB74216A9E1700521FBFFCCDBF5BFC83" box="[948,1369,863,883]" gridcol="1" gridrow="15" pageId="5" pageNumber="268">17</td>
</tr>
<tr id="A8A548169E1700521CADFC13BF5BFC65" box="[166,1369,897,917]" gridrow="16" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFC13BBEAFC65" box="[166,488,897,917]" gridcol="0" gridrow="16" pageId="5" pageNumber="268">PSR</td>
<td id="EB74216A9E1700521FBFFC13BF5BFC65" box="[948,1369,897,917]" gridcol="1" gridrow="16" pageId="5" pageNumber="268">15</td>
</tr>
<tr id="A8A548169E1700521CADFC31BF5BFC22" box="[166,1369,931,978]" gridrow="17" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFC31BBEAFC22" box="[166,488,931,978]" gridcol="0" gridrow="17" pageId="5" pageNumber="268">Keeling</td>
<td id="EB74216A9E1700521FBFFC31BF5BFC22" box="[948,1369,931,978]" gridcol="1" gridrow="17" pageId="5" pageNumber="268">6 dorsal scale rows on the posterior body third feebly keeled</td>
</tr>
<tr id="A8A548169E1700521CADFC73BF5BFC05" box="[166,1369,993,1013]" gridrow="18" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFC73BBEAFC05" box="[166,488,993,1013]" gridcol="0" gridrow="18" pageId="5" pageNumber="268">VEN</td>
<td id="EB74216A9E1700521FBFFC73BF5BFC05" box="[948,1369,993,1013]" gridcol="1" gridrow="18" pageId="5" pageNumber="268">241</td>
</tr>
<tr id="A8A548169E1700521CADFB91BF5BFBE7" box="[166,1369,1027,1047]" gridrow="19" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFB91BBEAFBE7" box="[166,488,1027,1047]" gridcol="0" gridrow="19" pageId="5" pageNumber="268">PrVEN</td>
<td id="EB74216A9E1700521FBFFB91BF5BFBE7" box="[948,1369,1027,1047]" gridcol="1" gridrow="19" pageId="5" pageNumber="268">2</td>
</tr>
<tr id="A8A548169E1700521CADFBB7BF5BFBC9" box="[166,1369,1061,1081]" gridrow="20" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFBB7BBEAFBC9" box="[166,488,1061,1081]" gridcol="0" gridrow="20" pageId="5" pageNumber="268">Ventral notched</td>
<td id="EB74216A9E1700521FBFFBB7BF5BFBC9" box="[948,1369,1061,1081]" gridcol="1" gridrow="20" pageId="5" pageNumber="268">Yes</td>
</tr>
<tr id="A8A548169E1700521CADFBD5BF5BFBAB" box="[166,1369,1095,1115]" gridrow="21" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFBD5BBEAFBAB" box="[166,488,1095,1115]" gridcol="0" gridrow="21" pageId="5" pageNumber="268">Ventral keeled</td>
<td id="EB74216A9E1700521FBFFBD5BF5BFBAB" box="[948,1369,1095,1115]" gridcol="1" gridrow="21" pageId="5" pageNumber="268">No</td>
</tr>
<tr id="A8A548169E1700521CADFBFBBF5BFB8D" box="[166,1369,1129,1149]" gridrow="22" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFBFBBBEAFB8D" box="[166,488,1129,1149]" gridcol="0" gridrow="22" pageId="5" pageNumber="268">SC</td>
<td id="EB74216A9E1700521FBFFBFBBF5BFB8D" box="[948,1369,1129,1149]" gridcol="1" gridrow="22" pageId="5" pageNumber="268">26*</td>
</tr>
<tr id="A8A548169E1700521CADFB19BF5BFB6F" box="[166,1369,1163,1183]" gridrow="23" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFB19BBEAFB6F" box="[166,488,1163,1183]" gridcol="0" gridrow="23" pageId="5" pageNumber="268">divided</td>
<td id="EB74216A9E1700521FBFFB19BF5BFB6F" box="[948,1369,1163,1183]" gridcol="1" gridrow="23" pageId="5" pageNumber="268">Yes</td>
</tr>
<tr id="A8A548169E1700521CADFB3FBF5BFB31" box="[166,1369,1197,1217]" gridrow="24" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFB3FBBEAFB31" box="[166,488,1197,1217]" gridcol="0" gridrow="24" pageId="5" pageNumber="268">Cloacal</td>
<td id="EB74216A9E1700521FBFFB3FBF5BFB31" box="[948,1369,1197,1217]" gridcol="1" gridrow="24" pageId="5" pageNumber="268">Single</td>
</tr>
<tr id="A8A548169E1700521CADFB5DBF5BFB13" box="[166,1369,1231,1251]" gridrow="25" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFB5DBBEAFB13" box="[166,488,1231,1251]" gridcol="0" gridrow="25" pageId="5" pageNumber="268">Loreal</td>
<td id="EB74216A9E1700521FBFFB5DBF5BFB13" box="[948,1369,1231,1251]" gridcol="1" gridrow="25" pageId="5" pageNumber="268">1/1</td>
</tr>
<tr id="A8A548169E1700521CADFB63BF5BFAF5" box="[166,1369,1265,1285]" gridrow="26" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFB63BBEAFAF5" box="[166,488,1265,1285]" gridcol="0" gridrow="26" pageId="5" pageNumber="268">Loreal entering orbit</td>
<td id="EB74216A9E1700521FBFFB63BF5BFAF5" box="[948,1369,1265,1285]" gridcol="1" gridrow="26" pageId="5" pageNumber="268">Yes</td>
</tr>
<tr id="A8A548169E1700521CADFA81BF5BFAD7" box="[166,1369,1299,1319]" gridrow="27" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFA81BBEAFAD7" box="[166,488,1299,1319]" gridcol="0" gridrow="27" pageId="5" pageNumber="268">SL</td>
<td id="EB74216A9E1700521FBFFA81BF5BFAD7" box="[948,1369,1299,1319]" gridcol="1" gridrow="27" pageId="5" pageNumber="268">8/8</td>
</tr>
<tr id="A8A548169E1700521CADFAA7BF5BFAB9" box="[166,1369,1333,1353]" gridrow="28" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFAA7BBEAFAB9" box="[166,488,1333,1353]" gridcol="0" gridrow="28" pageId="5" pageNumber="268">entering orbit</td>
<td id="EB74216A9E1700521FBFFAA7BF5BFAB9" box="[948,1369,1333,1353]" gridcol="1" gridrow="28" pageId="5" pageNumber="268">3,4,5</td>
</tr>
<tr id="A8A548169E1700521CADFAC5BF5BFA9B" box="[166,1369,1367,1387]" gridrow="29" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFAC5BBEAFA9B" box="[166,488,1367,1387]" gridcol="0" gridrow="29" pageId="5" pageNumber="268">largest SL</td>
<td id="EB74216A9E1700521FBFFAC5BF5BFA9B" box="[948,1369,1367,1387]" gridcol="1" gridrow="29" pageId="5" pageNumber="268">6/6</td>
</tr>
<tr id="A8A548169E1700521CADFAEBBF5BFA7D" box="[166,1369,1401,1421]" gridrow="30" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFAEBBBEAFA7D" box="[166,488,1401,1421]" gridcol="0" gridrow="30" pageId="5" pageNumber="268">IL</td>
<td id="EB74216A9E1700521FBFFAEBBF5BFA7D" box="[948,1369,1401,1421]" gridcol="1" gridrow="30" pageId="5" pageNumber="268">10/10</td>
</tr>
<tr id="A8A548169E1700521CADFA09BF5BFA5F" box="[166,1369,1435,1455]" gridrow="31" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFA09BBEAFA5F" box="[166,488,1435,1455]" gridcol="0" gridrow="31" pageId="5" pageNumber="268">IL in contact with 1st chin shield</td>
<td id="EB74216A9E1700521FBFFA09BF5BFA5F" box="[948,1369,1435,1455]" gridcol="1" gridrow="31" pageId="5" pageNumber="268">1−5</td>
</tr>
<tr id="A8A548169E1700521CADFA2FBF5BFA21" box="[166,1369,1469,1489]" gridrow="32" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFA2FBBEAFA21" box="[166,488,1469,1489]" gridcol="0" gridrow="32" pageId="5" pageNumber="268">PreOc</td>
<td id="EB74216A9E1700521FBFFA2FBF5BFA21" box="[948,1369,1469,1489]" gridcol="1" gridrow="32" pageId="5" pageNumber="268">1/1</td>
</tr>
<tr id="A8A548169E1700521CADFA4DBF5BFA03" box="[166,1369,1503,1523]" gridrow="33" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADFA4DBBEAFA03" box="[166,488,1503,1523]" gridcol="0" gridrow="33" pageId="5" pageNumber="268">PostOc</td>
<td id="EB74216A9E1700521FBFFA4DBF5BFA03" box="[948,1369,1503,1523]" gridcol="1" gridrow="33" pageId="5" pageNumber="268">2/2</td>
</tr>
<tr id="A8A548169E1700521CADF993BF5BF9E5" box="[166,1369,1537,1557]" gridrow="34" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF993BF5BF9E5" box="[166,1369,1537,1557]" colspan="2" colspanRight="1" gridcol="0" gridrow="34" pageId="5" pageNumber="268">Temporal scales</td>
</tr>
<tr id="A8A548169E1700521CADF9B1BF5BF9C7" box="[166,1369,1571,1591]" gridrow="35" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF9B1BBEAF9C7" box="[166,488,1571,1591]" gridcol="0" gridrow="35" pageId="5" pageNumber="268">anterior</td>
<td id="EB74216A9E1700521FBFF9B1BF5BF9C7" box="[948,1369,1571,1591]" gridcol="1" gridrow="35" pageId="5" pageNumber="268">2/2</td>
</tr>
<tr id="A8A548169E1700521CADF9D7BF5BF9A9" box="[166,1369,1605,1625]" gridrow="36" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF9D7BBEAF9A9" box="[166,488,1605,1625]" gridcol="0" gridrow="36" pageId="5" pageNumber="268">posterior</td>
<td id="EB74216A9E1700521FBFF9D7BF5BF9A9" box="[948,1369,1605,1625]" gridcol="1" gridrow="36" pageId="5" pageNumber="268">3/3</td>
</tr>
<tr id="A8A548169E1700521CADF9F5BF5BF98B" box="[166,1369,1639,1659]" gridrow="37" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF9F5BBEAF98B" box="[166,488,1639,1659]" gridcol="0" gridrow="37" pageId="5" pageNumber="268">Scales around paraparietal</td>
<td id="EB74216A9E1700521FBFF9F5BF5BF98B" box="[948,1369,1639,1659]" gridcol="1" gridrow="37" pageId="5" pageNumber="268">5/7</td>
</tr>
<tr id="A8A548169E1700521CADF91BBF5BF96D" box="[166,1369,1673,1693]" gridrow="38" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF91BBBEAF96D" box="[166,488,1673,1693]" gridcol="0" gridrow="38" pageId="5" pageNumber="268">Scales between parietals</td>
<td id="EB74216A9E1700521FBFF91BBF5BF96D" box="[948,1369,1673,1693]" gridcol="1" gridrow="38" pageId="5" pageNumber="268">4</td>
</tr>
<tr id="A8A548169E1700521CADF939BF5BF94F" box="[166,1369,1707,1727]" gridrow="39" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF939BBEAF94F" box="[166,488,1707,1727]" gridcol="0" gridrow="39" pageId="5" pageNumber="268">Nuchal band</td>
<td id="EB74216A9E1700521FBFF939BF5BF94F" box="[948,1369,1707,1727]" gridcol="1" gridrow="39" pageId="5" pageNumber="268">Absent</td>
</tr>
<tr id="A8A548169E1700521CADF95FBF5BF911" box="[166,1369,1741,1761]" gridrow="40" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF95FBBEAF911" box="[166,488,1741,1761]" gridcol="0" gridrow="40" pageId="5" pageNumber="268">Body blotches</td>
<td id="EB74216A9E1700521FBFF95FBF5BF911" box="[948,1369,1741,1761]" gridcol="1" gridrow="40" pageId="5" pageNumber="268">87 (yellow blotches)</td>
</tr>
<tr id="A8A548169E1700521CADF97DBF5BF8F3" box="[166,1369,1775,1795]" gridrow="41" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF97DBBEAF8F3" box="[166,488,1775,1795]" gridcol="0" gridrow="41" pageId="5" pageNumber="268">Tail blotches</td>
<td id="EB74216A9E1700521FBFF97DBF5BF8F3" box="[948,1369,1775,1795]" gridcol="1" gridrow="41" pageId="5" pageNumber="268">15*</td>
</tr>
<tr id="A8A548169E1700521CADF883BF5BF8D5" box="[166,1369,1809,1829]" gridrow="42" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF883BBEAF8D5" box="[166,488,1809,1829]" gridcol="0" gridrow="42" pageId="5" pageNumber="268">Belly pattern</td>
<td id="EB74216A9E1700521FBFF883BF5BF8D5" box="[948,1369,1809,1829]" gridcol="1" gridrow="42" pageId="5" pageNumber="268">uniform grey cream</td>
</tr>
<tr id="A8A548169E1700521CADF8A1BF5BF8B7" box="[166,1369,1843,1863]" gridrow="43" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF8A1BBEAF8B7" box="[166,488,1843,1863]" gridcol="0" gridrow="43" pageId="5" pageNumber="268">Ventral tail pattern</td>
<td id="EB74216A9E1700521FBFF8A1BF5BF8B7" box="[948,1369,1843,1863]" gridcol="1" gridrow="43" pageId="5" pageNumber="268">grey cream</td>
</tr>
<tr id="A8A548169E1700521CADF8C7BF5BF899" box="[166,1369,1877,1897]" gridrow="44" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF8C7BBEAF899" box="[166,488,1877,1897]" gridcol="0" gridrow="44" pageId="5" pageNumber="268">First body blotch position (at VEN)</td>
<td id="EB74216A9E1700521FBFF8C7BF5BF899" box="[948,1369,1877,1897]" gridcol="1" gridrow="44" pageId="5" pageNumber="268">13</td>
</tr>
<tr id="A8A548169E1700521CADF8E5BF5BF87B" box="[166,1369,1911,1931]" gridrow="45" pageId="5" pageNumber="268">
<td id="EB74216A9E1700521CADF8E5BBEAF87B" box="[166,488,1911,1931]" gridcol="0" gridrow="45" pageId="5" pageNumber="268">First blotch width (vertebral scales)</td>
<td id="EB74216A9E1700521FBFF8E5BF5BF87B" box="[948,1369,1911,1931]" gridcol="1" gridrow="45" pageId="5" pageNumber="268">0.5</td>
</tr>
</table>
</paragraph>
<caption id="42EA1ADC9E14FFAB1C9CFA8CB8C2FAA0" ID-DOI="http://doi.org/10.5281/zenodo.5590027" ID-Zenodo-Dep="5590027" httpUri="https://zenodo.org/record/5590027/files/figure.png" pageId="6" pageNumber="269" startId="6.[151,186,1310,1331]" targetBox="[151,1436,257,1267]" targetPageId="6">
<paragraph id="162A4A549E14FFAB1C9CFA8CB8C2FAA0" blockId="6.[151,1436,1310,1360]" pageId="6" pageNumber="269">
Fig. 2. Adult male holotype of 
<taxonomicName id="D19531D79E14FFAB1DC4FA8CB86AFAC3" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[463,616,1310,1331]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E14FFAB1DC4FA8CB86AFAC3" box="[463,616,1310,1331]" italics="true" pageId="6" pageNumber="269">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E14FFAB1E64FA8CB8BFFAC3" bold="true" box="[623,701,1310,1331]" pageId="6" pageNumber="269">
<taxonomicNameLabel id="3FD22B3D9E14FFAB1E64FA8CB8BFFAC3" box="[623,701,1310,1331]" pageId="6" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(VNUF R.2015.20) in life. (A) dorsolateral view. (B) Head in dorsolateral view. (C) Head in dorsal view. Photos: V. Q. Luu.
</paragraph>
</caption>
<paragraph id="162A4A549E14FFAA1C9CF857BF39FB49" blockId="6.[151,756,1445,2076]" lastBlockId="7.[831,1437,1089,2077]" lastPageId="7" lastPageNumber="270" pageId="6" pageNumber="269">
The new species has a loreal entering the orbit and thus differs from the following species and subspecies of the 
<taxonomicName id="D19531D79E14FFAB1C9CF797BB54F7EC" box="[151,342,2053,2076]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="24E196469E14FFAB1C9CF797BB54F7EC" box="[151,342,2053,2076]" italics="true" pageId="6" pageNumber="269">Lycodon ruhstrati</emphasis>
</taxonomicName>
group which have the loreal separated from the orbit: 
<taxonomicName id="D19531D79E14FFAB1FD5FA37BF9AFA4C" authority="Daltry &amp; Wuster, 2002" authorityName="Daltry &amp; Wuster" authorityYear="2002" box="[990,1432,1445,1468]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="24E196469E14FFAB1FD5FA37BE9DFA4C" box="[990,1183,1445,1468]" italics="true" pageId="6" pageNumber="269">L. cardamomensis</emphasis>
<bibRefCitation id="720437A59E14FFAB18AEFA37BF9AFA4C" author="Daltry J. C. &amp; Wuster W." box="[1189,1432,1445,1468]" pageId="6" pageNumber="269" pagination="498 - 504" refId="ref7459" refString="Daltry J. C. &amp; Wuster W. 2002. A new species of wolf snake (Serpentes: Colubridae: Lycodon) from the Cardamom Mountains, southwestern Cambodia. Herpetologica, 58 (4): 498 - 504." type="journal article" year="2002">Daltry &amp; Wüster, 2002</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="D19531D79E14FFAB1F34FA57B99CFA2C" authorityName="Vogel, Nguyen, Kingsada &amp; Ziegler" authorityYear="2012" box="[831,926,1477,1500]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="davidi">
<emphasis id="24E196469E14FFAB1F34FA57B99CFA2C" box="[831,926,1477,1500]" italics="true" pageId="6" pageNumber="269">L. davidi</emphasis>
</taxonomicName>
, 
<taxonomicName id="D19531D79E14FFAB1FA2FA57BEEFFA2C" authority="(Maki, 1931)" baseAuthorityName="Maki" baseAuthorityYear="1931" box="[937,1261,1477,1500]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="multifasciatus">
<emphasis id="24E196469E14FFAB1FA2FA57BE58FA2C" box="[937,1114,1477,1500]" italics="true" pageId="6" pageNumber="269">L. multifasciatus</emphasis>
(
<bibRefCitation id="720437A59E14FFAB186CFA57BEE7FA2C" author="Maki M." box="[1127,1253,1477,1500]" pageId="6" pageNumber="269" refId="ref8268" refString="Maki M. 1931. A monograph of the snakes of Japan. Dai-ichi Bhobo, Tokyo: 247 pp." type="book" year="1931">Maki, 1931</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="D19531D79E14FFAB18FCFA57B9F5F9EC" authority="Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu &amp; Ziegler, 2009" authorityName="Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu &amp; Ziegler" authorityYear="2009" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="ophiophagus">
<emphasis id="24E196469E14FFAB18FCFA57BF9EFA2C" box="[1271,1436,1477,1500]" italics="true" pageId="6" pageNumber="269">L. ophiophagus</emphasis>
<bibRefCitation id="720437A59E14FFAB1F34FA77B9F5F9EC" author="Vogel G. &amp; David P. &amp; Pauwels O. S. G. &amp; Sumontha M. &amp; Norval G. &amp; Hendrix R. &amp; Vu N. T. &amp; Ziegler T." pageId="6" pageNumber="269" pagination="131 - 182" refId="ref8926" refString="Vogel G., David P., Pauwels O. S. G., Sumontha M., Norval G., Hendrix R., Vu N. T. &amp; Ziegler T. 2009. A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology 22: 131 - 182." type="journal article" year="2009">Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu &amp; Ziegler, 2009</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="D19531D79E14FFAB1803F997BF2BF9EC" authority="Rendahl" authorityName="Rendahl" box="[1032,1321,1541,1564]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="paucifasciatus">
<emphasis id="24E196469E14FFAB1803F997BEC1F9EC" box="[1032,1219,1541,1564]" italics="true" pageId="6" pageNumber="269">L. paucifasciatus</emphasis>
Rendahl
</taxonomicName>
in 
<bibRefCitation id="720437A59E14FFAB195DF997B975F9CC" author="Smith M. A." pageId="6" pageNumber="269" refId="ref8594" refString="Smith M. A. 1943. The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp." type="book" year="1943">Smith, 1943</bibRefCitation>
, 
<taxonomicName id="D19531D79E14FFAB1F89F9B7BF08F9CC" authority="ruhstrati (Fischer, 1886)" authorityName="ruhstrati (Fischer" authorityYear="1886" baseAuthorityName="Fischer" baseAuthorityYear="1886" box="[898,1290,1573,1596]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="subSpecies" species="ruhstrati" subSpecies="ruhstrati">
<emphasis id="24E196469E14FFAB1F89F9B7BE5DF9CC" box="[898,1119,1573,1596]" italics="true" pageId="6" pageNumber="269">L. ruhstrati ruhstrati</emphasis>
(
<bibRefCitation id="720437A59E14FFAB1866F9B7BF00F9CC" author="Fischer J. G." box="[1133,1282,1573,1596]" pageId="6" pageNumber="269" pagination="51 - 67" refId="ref7577" refString="Fischer J. G. 1886. Herpetologische Notizen. Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg 9: 51 - 67 (1 - 19)." type="journal article" year="1886">Fischer, 1886</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="D19531D79E14FFAB194AF9B7BEC1F9AC" authority="(Vogel et al., 2009)" baseAuthorityName="Vogel" baseAuthorityYear="2009" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="subSpecies" species="ruhstrati" subSpecies="abditus">
<emphasis id="24E196469E14FFAB194AF9B7B9F3F9AC" italics="true" pageId="6" pageNumber="269">Lycodon ruhstrati abditus</emphasis>
(
<bibRefCitation id="720437A59E14FFAB1FF6F9D7BEB9F9AC" author="Vogel G. &amp; David P. &amp; Pauwels O. S. G. &amp; Sumontha M. &amp; Norval G. &amp; Hendrix R. &amp; Vu N. T. &amp; Ziegler T." box="[1021,1211,1605,1628]" pageId="6" pageNumber="269" pagination="131 - 182" refId="ref8926" refString="Vogel G., David P., Pauwels O. S. G., Sumontha M., Norval G., Hendrix R., Vu N. T. &amp; Ziegler T. 2009. A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology 22: 131 - 182." type="journal article" year="2009">
Vogel 
<emphasis id="24E196469E14FFAB1849F9D7BE6DF9AC" box="[1090,1135,1605,1628]" italics="true" pageId="6" pageNumber="269">et al</emphasis>
., 2009
</bibRefCitation>
)
</taxonomicName>
. In addition, the new species differs from 
<taxonomicName id="D19531D79E14FFAB182AF9F7BEE4F98C" authorityName="Daltry &amp; Wuster" authorityYear="2002" box="[1057,1254,1637,1660]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="24E196469E14FFAB182AF9F7BEE4F98C" box="[1057,1254,1637,1660]" italics="true" pageId="6" pageNumber="269">L. cardamomensis</emphasis>
</taxonomicName>
by having more ventral scales (241 
<emphasis id="24E196469E14FFAB1819F917BE54F96C" box="[1042,1110,1669,1692]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
215), and in body pattern (87 blotches 
<emphasis id="24E196469E14FFAB1F95F937B9E0F94C" box="[926,994,1701,1724]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
12 bands); from 
<taxonomicName id="D19531D79E14FFAB189FF937BEF0F94C" authorityName="Vogel, Nguyen, Kingsada &amp; Ziegler" authorityYear="2012" box="[1172,1266,1701,1724]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="davidi">
<emphasis id="24E196469E14FFAB189FF937BEF0F94C" box="[1172,1266,1701,1724]" italics="true" pageId="6" pageNumber="269">L. davidi</emphasis>
</taxonomicName>
by having more ventral scales (241 
<emphasis id="24E196469E14FFAB1816F957BE63F92C" box="[1053,1121,1733,1756]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
224), six dorsal scale rows on the posterior third of the body feebly keeled (
<emphasis id="24E196469E14FFAB1953F977BF9EF90C" box="[1368,1436,1765,1788]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
dorsal scale rows at midbody slightly keeled, outermost rows entirely smooth throughout body), and belly pattern uniform grey cream (
<emphasis id="24E196469E14FFAB184EF8D7BE8BF8AC" box="[1093,1161,1861,1884]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
anterior third whitishcream, posterior part heavily speckled with dark dots); from 
<taxonomicName id="D19531D79E14FFAB1F8BF817BE3BF86C" baseAuthorityName="Maki" baseAuthorityYear="1931" box="[896,1081,1925,1948]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="multifasciatus">
<emphasis id="24E196469E14FFAB1F8BF817BE3BF86C" box="[896,1081,1925,1948]" italics="true" pageId="6" pageNumber="269">L. multifasciatus</emphasis>
</taxonomicName>
by having more ventral scales (241 
<emphasis id="24E196469E14FFAB1F73F837B9BEF84C" box="[888,956,1957,1980]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
maximum 237), and dorsal pattern blotched (
<emphasis id="24E196469E14FFAB1F43F857B98EF82C" box="[840,908,1989,2012]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
banded); from 
<taxonomicName id="D19531D79E14FFAB1830F857BEE7F82C" authorityName="Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu &amp; Ziegler" authorityYear="2009" box="[1083,1253,1989,2012]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="ophiophagus">
<emphasis id="24E196469E14FFAB1830F857BEE7F82C" box="[1083,1253,1989,2012]" italics="true" pageId="6" pageNumber="269">L. ophiophagus</emphasis>
</taxonomicName>
by having more ventral scales (241 
<emphasis id="24E196469E14FFAB1812F877BE5FF80C" box="[1049,1117,2021,2044]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
211), and dorsal pattern (87 blotches 
<emphasis id="24E196469E14FFAB1FADF797B9E8F7EC" box="[934,1002,2053,2076]" italics="true" pageId="6" pageNumber="269">versus</emphasis>
21-22 bands); from 
<taxonomicName id="D19531D79E14FFAB18D4F797BF9EF7EC" box="[1247,1436,2053,2076]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="269" phylum="Chordata" rank="species" species="paucifasciatus">
<emphasis id="24E196469E14FFAB18D4F797BF9EF7EC" box="[1247,1436,2053,2076]" italics="true" pageId="6" pageNumber="269">L. paucifasciatus</emphasis>
</taxonomicName>
by having fewer dorsal scale rows at neck (17 
<emphasis id="24E196469E15FFAA1EBBF9DFB8F6F994" box="[688,756,1613,1636]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
19), more ventral scale rows (241 
<emphasis id="24E196469E15FFAA1E33F9FDB87EF976" box="[568,636,1647,1670]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
221-222), six dorsal scale rows on the posterior third of the body feebly keeled (
<emphasis id="24E196469E15FFAA1D3CF921BB79F93A" box="[311,379,1715,1738]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
two upper rows plus vertebral row distinctly keeled), and dorsal pattern blotched (
<emphasis id="24E196469E15FFAA1EBBF946B8F6F91B" box="[688,756,1748,1771]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
banded) (
<bibRefCitation id="720437A59E15FFAA1CF4F964BBD4F8FD" author="Neang T. &amp; Hartmann T. &amp; Hun S. &amp; Souter N. J. &amp; Furey N. M." box="[255,470,1782,1805]" pageId="7" pageNumber="270" pagination="1" refId="ref8289" refString="Neang T., Hartmann T., Hun S., Souter N. J. &amp; Furey N. M. 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068 - 080. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
Neang 
<emphasis id="24E196469E15FFAA1D5AF964BB90F8FD" box="[337,402,1782,1805]" italics="true" pageId="7" pageNumber="270">et al.,</emphasis>
2014
</bibRefCitation>
); from 
<taxonomicName id="D19531D79E15FFAA1E27F964B8C0F8FD" authorityName="ruhstrati (Fischer" authorityYear="1886" box="[556,706,1782,1805]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="subSpecies" species="ruhstrati" subSpecies="ruhstrati">
<emphasis id="24E196469E15FFAA1E27F964B8C0F8FD" box="[556,706,1782,1805]" italics="true" pageId="7" pageNumber="270">L. r. ruhstrati</emphasis>
</taxonomicName>
and 
<taxonomicName id="D19531D79E15FFAA1C9CF88ABBAFF8DF" box="[151,429,1816,1839]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="subSpecies" species="ruhstrati" subSpecies="abditus">
<emphasis id="24E196469E15FFAA1C9CF88ABBAFF8DF" box="[151,429,1816,1839]" italics="true" pageId="7" pageNumber="270">Lycodon ruhstrati abditus</emphasis>
</taxonomicName>
by having more ventral scales (241 
<emphasis id="24E196469E15FFAA1CC4F8A8BB11F8A1" box="[207,275,1850,1873]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
211-228; 241 
<emphasis id="24E196469E15FFAA1DBAF8A8BBF7F8A1" box="[433,501,1850,1873]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
206-224, respectively), and dorsal pattern blotched (
<emphasis id="24E196469E15FFAA1DC5F8CEB810F883" box="[462,530,1884,1907]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
banded in the latter) (
<bibRefCitation id="720437A59E15FFAA1C95F8ECBB68F865" author="Vogel G. &amp; Nguyen T. Q. &amp; Kingsada P. &amp; Ziegler T." box="[158,362,1918,1941]" pageId="7" pageNumber="270" pagination="344 - 352" refId="ref9006" refString="Vogel G., Nguyen T. Q., Kingsada P. &amp; Ziegler T. 2012. A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology 8: 344 - 352." type="journal article" year="2012">
Vogel 
<emphasis id="24E196469E15FFAA1CECF8ECBB1BF865" box="[231,281,1918,1941]" italics="true" pageId="7" pageNumber="270">et al</emphasis>
., 2012
</bibRefCitation>
); from 
<taxonomicName id="D19531D79E15FFAA1DB7F8ECB83AF865" box="[444,568,1918,1941]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="synaptor">
<emphasis id="24E196469E15FFAA1DB7F8ECB83AF865" box="[444,568,1918,1941]" italics="true" pageId="7" pageNumber="270">L. synaptor</emphasis>
</taxonomicName>
by having much more ventral scale rows (241 
<emphasis id="24E196469E15FFAA1DE5F832B830F847" box="[494,562,1952,1975]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
201-203), dorsal pattern with 87 blotches (
<emphasis id="24E196469E15FFAA1DA0F850BBEDF829" box="[427,495,1986,2009]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
30-31 bands), and belly pattern uniform grey cream (
<emphasis id="24E196469E15FFAA1DF9F876B834F80B" box="[498,566,2020,2043]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
banded) (
<bibRefCitation id="720437A59E15FFAA1EBFF876BB39F7ED" author="Vogel G. &amp; David P." pageId="7" pageNumber="270" pagination="289 - 296" refId="ref8842" refString="Vogel G. &amp; David P. 2010. A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn zoological Bulletin 57 (2): 289 - 296." type="journal article" year="2010">Vogel &amp; David 2010</bibRefCitation>
); from 
<taxonomicName id="D19531D79E15FFAA1D85F794B83EF7ED" authorityName="Neang, Hartmann, Hun, Souter &amp; Furey" authorityYear="2014" box="[398,572,2054,2077]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="zoosvictoriae">
<emphasis id="24E196469E15FFAA1D85F794B83EF7ED" box="[398,572,2054,2077]" italics="true" pageId="7" pageNumber="270">L. zoosvictoriae</emphasis>
</taxonomicName>
by having more ventral scales (241 
<emphasis id="24E196469E15FFAA1805FBD3BE50FBA8" box="[1038,1106,1089,1112]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
213), dorsal pattern consisting of 87 blotches (
<emphasis id="24E196469E15FFAA1FE4FBF0BE31FB89" box="[1007,1075,1122,1145]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
31), and having six dorsal scale rows on the posterior third of the body feebly keeled (
<emphasis id="24E196469E15FFAA1F43FB30B98EFB49" box="[840,908,1186,1209]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
all weakly keeled) (
<bibRefCitation id="720437A59E15FFAA186EFB30BF2DFB49" author="Neang T. &amp; Hartmann T. &amp; Hun S. &amp; Souter N. J. &amp; Furey N. M." box="[1125,1327,1186,1209]" pageId="7" pageNumber="270" pagination="1" refId="ref8289" refString="Neang T., Hartmann T., Hun S., Souter N. J. &amp; Furey N. M. 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068 - 080. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
Neang 
<emphasis id="24E196469E15FFAA18B9FB30BEE3FB49" box="[1202,1249,1186,1209]" italics="true" pageId="7" pageNumber="270">et al</emphasis>
., 2014
</bibRefCitation>
).
</paragraph>
<caption id="42EA1ADC9E15FFAA1C9DFC4FBF32FC02" ID-DOI="http://doi.org/10.5281/zenodo.5590029" ID-Zenodo-Dep="5590029" box="[150,1328,989,1010]" httpUri="https://zenodo.org/record/5590029/files/figure.png" pageId="7" pageNumber="270" startId="7.[150,185,989,1010]" targetBox="[151,1436,267,952]" targetPageId="7">
<paragraph id="162A4A549E15FFAA1C9DFC4FBF32FC02" blockId="7.[150,1328,989,1010]" box="[150,1328,989,1010]" pageId="7" pageNumber="270">
Fig. 3. Different head views of the adult male holotype of 
<taxonomicName id="D19531D79E15FFAA1EC5FC4FB966FC02" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[718,868,989,1010]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E15FFAA1EC5FC4FB966FC02" box="[718,868,989,1010]" italics="true" pageId="7" pageNumber="270">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E15FFAA1F61FC4FB9B5FC02" bold="true" box="[874,951,989,1010]" pageId="7" pageNumber="270">
<taxonomicNameLabel id="3FD22B3D9E15FFAA1F61FC4FB9B5FC02" box="[874,951,989,1010]" pageId="7" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(VNUF R.2015.20). Photos V. Q. Luu.
</paragraph>
</caption>
<paragraph id="162A4A549E15FFAA1F34FB50BE87F86C" blockId="7.[831,1437,1089,2077]" pageId="7" pageNumber="270">
From the remaining species occurring in 
<collectingCountry id="6E820AC49E15FFAA1908FB50BF3BFB29" box="[1283,1337,1218,1241]" name="Laos" pageId="7" pageNumber="270">Laos</collectingCountry>
, the new species can be distinguished as follows: from 
<taxonomicName id="D19531D79E15FFAA191CFB70BF9EFB09" box="[1303,1436,1250,1273]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="capucinus">
<emphasis id="24E196469E15FFAA191CFB70BF9EFB09" box="[1303,1436,1250,1273]" italics="true" pageId="7" pageNumber="270">L. capucinus</emphasis>
</taxonomicName>
by having more ventrals (241 
<emphasis id="24E196469E15FFAA1897FA90BEE2FAE9" box="[1180,1248,1282,1305]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
182-211), fewer supralabials (8/8 
<emphasis id="24E196469E15FFAA180DFAB0BE48FAC9" box="[1030,1098,1314,1337]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
9-10), cloacal single (
<emphasis id="24E196469E15FFAA1953FAB0BF9EFAC9" box="[1368,1436,1314,1337]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
divided), dorsal blotches 87 (
<emphasis id="24E196469E15FFAA189EFAD1BEDBFAAA" box="[1173,1241,1347,1370]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
reticulated), and greyish yellow blotched body pattern (
<emphasis id="24E196469E15FFAA18D9FAF1BF14FA8A" box="[1234,1302,1379,1402]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
reticulated); from 
<taxonomicName id="D19531D79E15FFAA1F72FA11B9F6FA6A" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[889,1012,1411,1434]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="24E196469E15FFAA1F72FA11B9F6FA6A" box="[889,1012,1411,1434]" italics="true" pageId="7" pageNumber="270">L. fasciatus</emphasis>
</taxonomicName>
by having more ventral scale rows (241 
<emphasis id="24E196469E15FFAA1F34FA31B981FA4A" box="[831,899,1443,1466]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
182-225), dorsal pattern consisting of 87 blotches (
<emphasis id="24E196469E15FFAA1F43FA51B98EFA2A" box="[840,908,1475,1498]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
19-49 bands), and belly pattern uniform grey cream (
<emphasis id="24E196469E15FFAA1F98FA71B9D5FA0A" box="[915,983,1507,1530]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
white with dark blotches) (
<bibRefCitation id="720437A59E15FFAA1907FA71B97BF9EA" author="Neang T. &amp; Hartmann T. &amp; Hun S. &amp; Souter N. J. &amp; Furey N. M." pageId="7" pageNumber="270" pagination="1" refId="ref8289" refString="Neang T., Hartmann T., Hun S., Souter N. J. &amp; Furey N. M. 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068 - 080. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
Neang 
<emphasis id="24E196469E15FFAA1956FA71BF8DFA0A" box="[1373,1423,1507,1530]" italics="true" pageId="7" pageNumber="270">et al</emphasis>
., 2014
</bibRefCitation>
); from 
<taxonomicName id="D19531D79E15FFAA1FC3F991BE3FF9EA" box="[968,1085,1539,1562]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="laoensis">
<emphasis id="24E196469E15FFAA1FC3F991BE3FF9EA" box="[968,1085,1539,1562]" italics="true" pageId="7" pageNumber="270">L. laoensis</emphasis>
</taxonomicName>
by having loreal in contact with the orbit (
<emphasis id="24E196469E15FFAA1FBBF9B1B9F6F9CA" box="[944,1012,1571,1594]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
separated), more ventrals (241 
<emphasis id="24E196469E15FFAA1953F9B1BF9EF9CA" box="[1368,1436,1571,1594]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
169-192), and dorsal scales feebly keeled in the posterior body part (
<emphasis id="24E196469E15FFAA1FBDF9F6B9F8F98B" box="[950,1018,1636,1659]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
all smooth) (
<bibRefCitation id="720437A59E15FFAA1882F9F6BF57F98B" author="Neang T. &amp; Hartmann T. &amp; Hun S. &amp; Souter N. J. &amp; Furey N. M." box="[1161,1365,1636,1659]" pageId="7" pageNumber="270" pagination="1" refId="ref8289" refString="Neang T., Hartmann T., Hun S., Souter N. J. &amp; Furey N. M. 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068 - 080. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
Neang 
<emphasis id="24E196469E15FFAA18D3F9F6BF05F98B" box="[1240,1287,1636,1659]" italics="true" pageId="7" pageNumber="270">et al</emphasis>
., 2014
</bibRefCitation>
); from 
<taxonomicName id="D19531D79E15FFAA1F34F916B9F9F96B" authorityName="Gunther" authorityYear="1875" box="[831,1019,1668,1691]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="24E196469E15FFAA1F34F916B9F9F96B" box="[831,1019,1668,1691]" italics="true" pageId="7" pageNumber="270">L. septentrionalis</emphasis>
</taxonomicName>
by having more infralabials (10 
<emphasis id="24E196469E15FFAA1953F916BF9EF96B" box="[1368,1436,1668,1691]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
7-8), more ventral scales (241 
<emphasis id="24E196469E15FFAA18A7F936BEF2F94B" box="[1196,1264,1700,1723]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
202-217), and dorsal pattern blotched (
<emphasis id="24E196469E15FFAA184DF956BE88F92B" box="[1094,1162,1732,1755]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
banded), as well as belly pattern uniform grey cream (
<emphasis id="24E196469E15FFAA187FF976BEBAF90B" box="[1140,1208,1764,1787]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
white) (
<bibRefCitation id="720437A59E15FFAA1919F976B97BF8EB" author="Neang T. &amp; Hartmann T. &amp; Hun S. &amp; Souter N. J. &amp; Furey N. M." pageId="7" pageNumber="270" pagination="1" refId="ref8289" refString="Neang T., Hartmann T., Hun S., Souter N. J. &amp; Furey N. M. 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068 - 080. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
Neang 
<emphasis id="24E196469E15FFAA196BF976BF8DF90B" box="[1376,1423,1764,1787]" italics="true" pageId="7" pageNumber="270">et al</emphasis>
., 2014
</bibRefCitation>
); from 
<taxonomicName id="D19531D79E15FFAA1FC0F896BE59F8EB" box="[971,1115,1796,1819]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="subcinctus">
<emphasis id="24E196469E15FFAA1FC0F896BE59F8EB" box="[971,1115,1796,1819]" italics="true" pageId="7" pageNumber="270">L. subcinctus</emphasis>
</taxonomicName>
by the presence of preocular scale (
<emphasis id="24E196469E15FFAA1F8EF8B7B9CBF8CC" box="[901,969,1829,1852]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
absent), having cloacal scale single (
<emphasis id="24E196469E15FFAA1953F8B7BF9EF8CC" box="[1368,1436,1829,1852]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
divided), dorsal pattern blotched (
<emphasis id="24E196469E15FFAA18DFF8D7BF1AF8AC" box="[1236,1304,1861,1884]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
banded in anterior part), and more ventral scale rows (241 
<emphasis id="24E196469E15FFAA1953F8F7BF9EF88C" box="[1368,1436,1893,1916]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
129-230) (
<bibRefCitation id="720437A59E15FFAA1FA4F817BE78F86C" author="Neang T. &amp; Hartmann T. &amp; Hun S. &amp; Souter N. J. &amp; Furey N. M." box="[943,1146,1925,1948]" pageId="7" pageNumber="270" pagination="1" refId="ref8289" refString="Neang T., Hartmann T., Hun S., Souter N. J. &amp; Furey N. M. 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068 - 080. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
Neang 
<emphasis id="24E196469E15FFAA1FF6F817BE2EF86C" box="[1021,1068,1925,1948]" italics="true" pageId="7" pageNumber="270">et al</emphasis>
., 2014
</bibRefCitation>
).
</paragraph>
<caption id="42EA1ADC9E15FFAA1C9CFA22BB66FA0F" ID-DOI="http://doi.org/10.5281/zenodo.5590031" ID-Zenodo-Dep="5590031" httpUri="https://zenodo.org/record/5590031/files/figure.png" pageId="7" pageNumber="270" startId="7.[151,186,1456,1477]" targetBox="[154,754,1093,1400]" targetPageId="7">
<paragraph id="162A4A549E15FFAA1C9CFA22BB66FA0F" blockId="7.[151,756,1456,1535]" pageId="7" pageNumber="270">
Fig. 4. Lateral head view of the adult male holotype of 
<taxonomicName id="D19531D79E15FFAA1CE8FA5FBB79FA12" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[227,379,1485,1506]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E15FFAA1CE8FA5FBB79FA12" box="[227,379,1485,1506]" italics="true" pageId="7" pageNumber="270">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E15FFAA1D89FA5FBBCDFA12" bold="true" box="[386,463,1485,1506]" pageId="7" pageNumber="270">
<taxonomicNameLabel id="3FD22B3D9E15FFAA1D89FA5FBBCDFA12" box="[386,463,1485,1506]" pageId="7" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(VNUF R.2015.20). Drawing by T. Ziegler.
</paragraph>
</caption>
<paragraph id="162A4A549E15FFA51F34F837B8C1F8C7" blockId="7.[831,1437,1089,2077]" lastBlockId="8.[151,756,1280,1847]" lastPageId="8" lastPageNumber="271" pageId="7" pageNumber="270">
From the remaining species in the 
<taxonomicName id="D19531D79E15FFAA18CFF837BF21F84C" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[1220,1315,1957,1980]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="24E196469E15FFAA18CFF837BF21F84C" box="[1220,1315,1957,1980]" italics="true" pageId="7" pageNumber="270">fasciatus</emphasis>
</taxonomicName>
group, the new species differs as follows: from 
<taxonomicName id="D19531D79E15FFAA18CFF857BF9EF82C" authority="Boulenger" authorityName="Boulenger" authorityYear="1900" box="[1220,1436,1989,2012]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="270" phylum="Chordata" rank="species" species="butleri">
<emphasis id="24E196469E15FFAA18CFF857BF25F82C" box="[1220,1319,1989,2012]" italics="true" pageId="7" pageNumber="270">L. butleri</emphasis>
Boulenger
</taxonomicName>
by having more ventral scale rows (241 
<emphasis id="24E196469E15FFAA18E2F877BF2FF80C" box="[1257,1325,2021,2044]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
220-227), dorsal pattern blotched (
<emphasis id="24E196469E15FFAA184BF794BE86F7ED" box="[1088,1156,2054,2077]" italics="true" pageId="7" pageNumber="270">versus</emphasis>
banded), and belly pattern uniform grey cream (
<emphasis id="24E196469E1AFFA51D77FA92BBC2FAE7" box="[380,448,1280,1303]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
banded &amp; spotted) (
<bibRefCitation id="720437A59E1AFFA51E90FA92BB23FAC7" author="Grismer L. L. &amp; Quah E. S. H. &amp; Anuar S. &amp; Muin M. A. &amp; Wood P. L. Jr. &amp; Nor S. A. M." pageId="8" pageNumber="271" pagination="1" refId="ref7685" refString="Grismer L. L., Quah E. S. H., Anuar S., Muin M. A., Wood P. L. Jr. &amp; Nor S. A. M. 2014. A diminutive new species of cavedwelling Wolf Snake (Colubridae: Lycodon Boie, 1826) from Peninsular Malaysia. Zootaxa 3815 (1): 051 - 067. http: // dx. doi. org / 10.11646 / zootaxa. 3815.1.3" type="journal article" year="2014">
Grismer 
<emphasis id="24E196469E1AFFA51C9CFAB2BACFFAC7" box="[151,205,1312,1335]" italics="true" pageId="8" pageNumber="271">et al</emphasis>
., 2014
</bibRefCitation>
); from 
<taxonomicName id="D19531D79E1AFFA51D77FAB2B825FAC7" authorityName="Grismer, Quah, Anuar, Muin, Wood &amp; Nor" authorityYear="2014" box="[380,551,1312,1335]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="271" phylum="Chordata" rank="species" species="cavernicolus">
<emphasis id="24E196469E1AFFA51D77FAB2B825FAC7" box="[380,551,1312,1335]" italics="true" pageId="8" pageNumber="271">L. cavernicolus</emphasis>
</taxonomicName>
by having dorsal head uniformly dark grey (
<emphasis id="24E196469E1AFFA51DC0FAD2B80DFAA7" box="[459,527,1344,1367]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
light brown), fewer supralabials (8 
<emphasis id="24E196469E1AFFA51D30FAF2BB7DFA87" box="[315,383,1376,1399]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
9 or 10), more dorsal blotches (87 
<emphasis id="24E196469E1AFFA51C9CFA12BAD9FA67" box="[151,219,1408,1431]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
36-45 bands), dorsal scales on the anterior 2/3 of the body length smooth, the six central dorsal scale rows of the posterior 1/3 of the body length feebly keeled (
<emphasis id="24E196469E1AFFA51CABFA72BAE6FA07" box="[160,228,1504,1527]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
all keeled), and greyish yellow blotched pattern on the body (
<emphasis id="24E196469E1AFFA51D22F992BB6FF9E7" box="[297,365,1536,1559]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
white bands); from 
<taxonomicName id="D19531D79E1AFFA51E47F992B8D3F9E7" authorityName="Vogel &amp; Luo" authorityYear="2011" box="[588,721,1536,1559]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="271" phylum="Chordata" rank="species" species="gongshan">
<emphasis id="24E196469E1AFFA51E47F992B8D3F9E7" box="[588,721,1536,1559]" italics="true" pageId="8" pageNumber="271">L. gongshan</emphasis>
</taxonomicName>
by having six dorsal scale rows on the posterior third of the body feebly keeled (
<emphasis id="24E196469E1AFFA51D74F9D2BBC1F9A7" box="[383,451,1600,1623]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
upper and vertebral dorsal rows keeled), more ventral scale rows (241 
<emphasis id="24E196469E1AFFA51E7DF9F2B8B8F987" box="[630,698,1632,1655]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
210- 216), and dorsal pattern with 87 blotches (
<emphasis id="24E196469E1AFFA51E8BF912B8C6F967" box="[640,708,1664,1687]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
32- 40 bands) (
<bibRefCitation id="720437A59E1AFFA51D19F932BBEFF947" author="Vogel G. &amp; Luo J." box="[274,493,1696,1719]" pageId="8" pageNumber="271" pagination="29 - 40" refId="ref8886" refString="Vogel G. &amp; Luo J. 2011. A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae). Zootaxa 2807: 29 - 40." type="journal article" year="2011">Vogel &amp; Luo, 2011</bibRefCitation>
); from 
<taxonomicName id="D19531D79E1AFFA51E35F932B8F6F947" authorityName="Zhang, Jiang, Vogel &amp; Rao" authorityYear="2011" box="[574,756,1696,1719]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="271" phylum="Chordata" rank="species" species="liuchengchaoi">
<emphasis id="24E196469E1AFFA51E35F932B8F6F947" box="[574,756,1696,1719]" italics="true" pageId="8" pageNumber="271">L. liuchengchaoi</emphasis>
</taxonomicName>
by having cloacal scale single (
<emphasis id="24E196469E1AFFA51DF1F952B83CF927" box="[506,574,1728,1751]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
divided), dorsal pattern consisting of 87 irregular greyish yellow dorsal blotches (
<emphasis id="24E196469E1AFFA51CF4F892BB41F8E7" box="[255,323,1792,1815]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
40 well-defined yellow rings), and more ventral scales (241 
<emphasis id="24E196469E1AFFA51D6DF8B2BBA8F8C7" box="[358,426,1824,1847]" italics="true" pageId="8" pageNumber="271">versus</emphasis>
204) (
<bibRefCitation id="720437A59E1AFFA51DE4F8B2B8B5F8C7" author="Zhang J. &amp; Jiang K. &amp; Vogel G. &amp; Rao D." box="[495,695,1824,1847]" pageId="8" pageNumber="271" pagination="59 - 68" refId="ref9053" refString="Zhang J., Jiang K., Vogel G. &amp; Rao D. 2011. A new species of the genus Lycodon (Squamata, Colubridae) from Sichuan Province, China. Zootaxa 2982: 59 - 68." type="journal article" year="2011">
Zhang 
<emphasis id="24E196469E1AFFA51E37F8B2B869F8C7" box="[572,619,1824,1847]" italics="true" pageId="8" pageNumber="271">et al</emphasis>
., 2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="42EA1ADC9E1AFFA51C9EFBFDB823FB50" ID-DOI="http://doi.org/10.5281/zenodo.5590035" ID-Zenodo-Dep="5590035" httpUri="https://zenodo.org/record/5590035/files/figure.png" pageId="8" pageNumber="271" startId="8.[149,184,1135,1156]" targetBox="[152,1436,256,1094]" targetPageId="8">
<paragraph id="162A4A549E1AFFA51C9EFBFDB823FB50" blockId="8.[149,1437,1135,1184]" pageId="8" pageNumber="271">
Fig. 5. Additional record of 
<taxonomicName id="D19531D79E1AFFA51DA6FBFDB846FB74" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[429,580,1135,1156]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="271" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E1AFFA51DA6FBFDB846FB74" box="[429,580,1135,1156]" italics="true" pageId="8" pageNumber="271">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E1AFFA51E41FBFDB894FB74" bold="true" box="[586,662,1135,1156]" pageId="8" pageNumber="271">
<taxonomicNameLabel id="3FD22B3D9E1AFFA51E41FBFDB894FB74" box="[586,662,1135,1156]" pageId="8" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life: (A- B) dorsolateral views. (C) Head in dorsal view. Photos A-C A. Sahu. (D) Dorsal view. Photo P. Jaeger.
</paragraph>
</caption>
<subSubSection id="5E8F19DF9E1AFFA51C9CF8DDB8C2F856" pageId="8" pageNumber="271" type="distribution">
<paragraph id="162A4A549E1AFFA51C9CF8DDB8C2F856" blockId="8.[151,756,1871,1958]" pageId="8" pageNumber="271">
<emphasis id="24E196469E1AFFA51C9CF8DDBB2CF896" bold="true" box="[151,302,1871,1894]" pageId="8" pageNumber="271">Distribution:</emphasis>
<taxonomicName id="D19531D79E1AFFA51D35F8DDBBF3F896" authority="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler, 2018" authorityName="Luu &amp; Bonkowski &amp; Nguyen &amp; Le &amp; Calame &amp; Ziegler" authorityYear="2018" box="[318,497,1871,1894]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="271" phylum="Chordata" rank="species" species="banksi" status="sp. nov.">
<emphasis id="24E196469E1AFFA51D35F8DDBBF3F896" box="[318,497,1871,1894]" italics="true" pageId="8" pageNumber="271">Lycodon banksi</emphasis>
</taxonomicName>
<emphasis id="24E196469E1AFFA51E0BF8DDB862F896" bold="true" box="[512,608,1871,1894]" pageId="8" pageNumber="271">
<taxonomicNameLabel id="3FD22B3D9E1AFFA51E0BF8DDB862F896" box="[512,608,1871,1894]" pageId="8" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently known only from the 
<typeStatus id="C92EF4F69E1AFFA51D8BF8FDBBACF876" box="[384,430,1903,1926]" pageId="8" pageNumber="271">type</typeStatus>
locality in the Phou Hin Poun NPA, 
<collectingRegion id="D45184B69E1AFFA51CDCF81DBBD4F856" box="[215,470,1935,1958]" country="Laos" name="Khammouan" pageId="8" pageNumber="271">Khammouane Province</collectingRegion>
, central 
<collectingCountry id="6E820AC49E1AFFA51E24F81DB861F856" box="[559,611,1935,1958]" name="Laos" pageId="8" pageNumber="271">Laos</collectingCountry>
(
<figureCitation id="8EAE56D19E1AFFA51E7AF81DB8B1F856" box="[625,691,1935,1958]" captionStart="Fig" captionStartId="9.[150,185,1645,1666]" captionTargetBox="[273,1329,259,1600]" captionTargetId="figure-160@9.[234,1347,241,1645]" captionTargetPageId="9" captionText="Fig. 6. Map showing the type locality of Lycodon banksi sp. nov. in Khammouane Province (black dots) and other records of the genus Lycodon occurring in Laos (after Deuve, 1970; Vogel et al., 2012; Siler et al., 2013; Luu et al., 2013; Teynié et al., 2014)." figureDoi="http://doi.org/10.5281/zenodo.5590039" httpUri="https://zenodo.org/record/5590039/files/figure.png" pageId="8" pageNumber="271">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5E8F19DF9E1AFFA51C9CF82DBED8F936" pageId="8" pageNumber="271" type="etymology">
<paragraph id="162A4A549E1AFFA51C9CF82DBF67FA67" blockId="8.[151,756,1983,2070]" lastBlockId="8.[831,1436,1280,1431]" pageId="8" pageNumber="271">
<emphasis id="24E196469E1AFFA51C9CF82DBB1EF826" bold="true" box="[151,284,1983,2006]" pageId="8" pageNumber="271">Etymology:</emphasis>
The name of the species is dedicated to our friend and colleague Chris Banks, International Coordinator, Philippine Crocodile National Recovery Team, Zoos 
<collectingRegion id="D45184B69E1AFFA51FD3FA92BE30FAE7" box="[984,1074,1280,1303]" country="Australia" name="Victoria" pageId="8" pageNumber="271">Victoria</collectingRegion>
, 
<collectingCountry id="6E820AC49E1AFFA5184FFA92BEA8FAE7" box="[1092,1194,1280,1303]" name="Australia" pageId="8" pageNumber="271">Australia</collectingCountry>
, for his outstanding contributions towards amphibian and reptile conservation, in particular of the Philippine Crocodile. We propose the following common names: Banks’ Wolf Snake (English), Banks Wolfszahnnatter (German).
</paragraph>
<paragraph id="162A4A549E1AFFA51F34FA3DBED8F936" blockId="8.[831,1437,1455,1734]" pageId="8" pageNumber="271">
<emphasis id="24E196469E1AFFA51F34FA3DB9FFFA36" bold="true" box="[831,1021,1455,1478]" pageId="8" pageNumber="271">Natural history:</emphasis>
The 
<typeStatus id="C92EF4F69E1AFFA51837FA3DBE98FA36" box="[1084,1178,1455,1478]" pageId="8" pageNumber="271" type="holotype">holotype</typeStatus>
was found at 20:39 h, crawling on a limestone outcrop in the karst forest, approximately 
<quantity id="D16DE7B19E1AFFA51FE6FA7DBE36F9F6" box="[1005,1076,1519,1542]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="271" unit="m" value="0.3">0.3 m</quantity>
above the forest floor, at an elevation of 
<quantity id="D16DE7B19E1AFFA51FDDF99DBE21F9D6" box="[982,1059,1551,1574]" metricMagnitude="2" metricUnit="m" metricValue="1.67" pageId="8" pageNumber="271" unit="m" value="167.0">167 m</quantity>
a.s.l. The humidity at the time of collection was approximately 85% and the air temperature ranged from 23 to 26 
<superScript id="E1E0E71C9E1AFFA518B7F9DCBED4F996" attach="right" box="[1212,1238,1614,1638]" fontSize="6" pageId="8" pageNumber="271">oC</superScript>
(
<figureCitation id="8EAE56D19E1AFFA518E3F9DDBF2CF996" box="[1256,1326,1615,1638]" captionStart="Fig" captionStartId="10.[151,186,1146,1167]" captionTargetBox="[151,1436,257,1114]" captionTargetId="figure-406@10.[151,1436,257,1114]" captionTargetPageId="10" captionText="Fig. 7. Habitat of Lycodon banksi sp. nov. at the type locality. Photo V. Q. Luu." figureDoi="http://doi.org/10.5281/zenodo.5590041" httpUri="https://zenodo.org/record/5590041/files/figure.png" pageId="8" pageNumber="271">Fig. 7</figureCitation>
). Another specimen was observed 
<quantity id="D16DE7B19E1AFFA51866F9FDBED0F976" box="[1133,1234,1647,1670]" metricMagnitude="4" metricUnit="m" metricValue="1.24" pageId="8" pageNumber="271" unit="km" value="12.4">12.4 km</quantity>
away from the type locality, active on the ground at 23:30 h, near a limestone cliff in the secondary forest.
</paragraph>
</subSubSection>
</treatment>
</document>