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<document id="39C42C3CADDDF53EE89618FE849AE39F" ID-CLB-Dataset="32618" ID-DOI="10.11646/zootaxa.4299.1.3" ID-GBIF-Dataset="043cd50d-f1b9-400f-8c6b-8ba62b24a72c" ID-ISSN="1175-5326" ID-Zenodo-Dep="835103" ID-ZooBank="38508C24-1594-43DF-84FA-EB7DB779C371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501087938908" checkinUser="plazi" docAuthor="Kraus, Fred" docDate="2017" docId="9009D40C1278FFF4FF02F8988E7CFB8F" docLanguage="en" docName="zootaxa.4299.1.3.pdf" docOrigin="Zootaxa 4299 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gerrhopilus depressiceps Sternfeld 1913" docType="treatment" docVersion="7" lastPageNumber="80" masterDocId="6C30AC741279FFF1FF95FFAA8F23FFD7" masterDocTitle="New Species of Blindsnakes (Squamata: Gerrhopilidae) from the offshore islands of Papua New Guinea" masterLastPageNumber="94" masterPageNumber="75" pageNumber="76" updateTime="1698465934296" updateUser="ExternalLinkService">
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<mods:title id="1C14152DDE612B6E1283D340CF4CE70E">New Species of Blindsnakes (Squamata: Gerrhopilidae) from the offshore islands of Papua New Guinea</mods:title>
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<mods:namePart id="9266050F3615BAFBF1EC64C60A9A4E88">Kraus, Fred</mods:namePart>
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<mods:date id="35BE19E75F875A3E383E3E9F0087FE0A">2017</mods:date>
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<treatment id="9009D40C1278FFF4FF02F8988E7CFB8F" ID-DOI="http://doi.org/10.5281/zenodo.6023896" ID-GBIF-Taxon="132583253" ID-Zenodo-Dep="6023896" LSID="urn:lsid:plazi:treatment:9009D40C1278FFF4FF02F8988E7CFB8F" httpUri="http://treatment.plazi.org/id/9009D40C1278FFF4FF02F8988E7CFB8F" lastPageId="5" lastPageNumber="80" pageId="1" pageNumber="76">
<subSubSection id="50BA36911278FFF0FF02F8988E3BF8BA" pageId="1" pageNumber="76" type="nomenclature">
<paragraph id="181F651A1278FFF0FF02F8988EE4F89B" blockId="1.[151,455,1842,1902]" box="[151,455,1842,1868]" pageId="1" pageNumber="76">
<heading id="4357D2761278FFF0FF02F8988EE4F89B" bold="true" box="[151,455,1842,1868]" fontSize="11" level="1" pageId="1" pageNumber="76" reason="1">
<taxonomicName id="DFA01E991278FFF0FF02F8988EE4F89B" authorityName="Sternfeld" authorityYear="1913" box="[151,455,1842,1868]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="76" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B9081278FFF0FF02F8988EE4F89B" bold="true" box="[151,455,1842,1868]" italics="true" pageId="1" pageNumber="76">Gerrhopilus depressiceps</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="181F651A1278FFF0FF02F8FF8E3BF8BA" blockId="1.[151,455,1842,1902]" box="[151,280,1877,1902]" pageId="1" pageNumber="76">
<figureCitation id="809B799F1278FFF0FF02F8FF8FFCF8B9" box="[151,223,1877,1902]" captionStart="FIGURE 1" captionStartId="2.[151,249,1942,1964]" captionTargetBox="[327,1255,221,1865]" captionTargetId="figure@2.[289,1286,195,1896]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal (A) and lateral (B) views of the head of the holotype of Gerrhopilus depressiceps (MZB 23986); dorsal (C) and lateral (D) views of the head of the holotype of Gerrhopilus addisoni (USNM 195953); dorsal (E) and lateral (F) views of the head of the holotype of Gerrhopilus eurydice (MCZ 145954). Scale bars = 5 mm." httpUri="https://zenodo.org/record/835105/files/figure.png" pageId="1" pageNumber="76">Figs. 1</figureCitation>
A, B
</paragraph>
</subSubSection>
<subSubSection id="50BA36911278FFF0FF02F8378D7FF827" pageId="1" pageNumber="76" type="reference_group">
<paragraph id="181F651A1278FFF0FF02F8378D3BF805" blockId="1.[151,604,1949,2032]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="38B042341278FFF0FF02F8378D3BF805" pageId="1" pageNumber="76">
<taxonomicName id="DFA01E991278FFF0FF02F8378D72F864" authority="Sternfeld, 1913: 384" authorityName="Sternfeld" authorityPageNumber="384" authorityYear="1913" box="[151,593,1949,1971]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="76" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B9081278FFF0FF02F8378E5AF864" box="[151,377,1949,1971]" italics="true" pageId="1" pageNumber="76">Typhlops depressiceps</emphasis>
<treatmentCitation id="9901430B1278FFF0FEEAF8378D72F864" author="Sternfeld" box="[383,593,1949,1971]" page="384" pageId="1" pageNumber="76" year="1913">
<bibRefCitation id="7C3118EB1278FFF0FEEAF8378D3DF864" author="Sternfeld" box="[383,542,1949,1971]" pageId="1" pageNumber="76" refString="Sternfeld, R. (1913) Beitrage zur Schlangenfauna Neuguineas und der benachbarten Inselgruppen. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1913, 384 - 389." type="journal article" year="1913">Sternfeld, 1913</bibRefCitation>
: 384
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="DFA01E991278FFF0FF02F8168D37F805" authority="Vogt, 1932: 293" authorityName="Vogt" authorityPageNumber="293" authorityYear="1932" box="[151,532,1980,2002]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="76" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B9081278FFF0FF02F8168E4AF805" box="[151,361,1980,2002]" italics="true" pageId="1" pageNumber="76">Typhlops monochrus</emphasis>
<treatmentCitation id="9901430B1278FFF0FEFAF8178D37F805" author="Vogt" box="[367,532,1980,2002]" page="293" pageId="1" pageNumber="76" year="1932">
<bibRefCitation id="7C3118EB1278FFF0FEFAF8178EC2F805" author="Vogt" box="[367,481,1980,2002]" pageId="1" pageNumber="76" refString="Vogt, T. (1932) Beitrag zur Reptilienfauna der ehemaligen Kolonie Deutsch-Neuguinea. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1932, 281 - 294." type="journal article" year="1932">Vogt, 1932</bibRefCitation>
: 293
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="181F651A1278FFF0FF02F8708D7FF827" blockId="1.[151,604,1949,2032]" box="[151,604,2010,2032]" pageId="1" pageNumber="76">
<treatmentCitationGroup id="38B042341278FFF0FF02F8708D7FF827" box="[151,604,2010,2032]" pageId="1" pageNumber="76">
<taxonomicName id="DFA01E991278FFF0FF02F8708D7BF827" authority="Loveridge, 1948: 320" authorityName="Loveridge" authorityPageNumber="320" authorityYear="1948" box="[151,600,2010,2032]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="76" phylum="Chordata" rank="species" species="monochrous">
<emphasis id="2AD4B9081278FFF0FF02F8708E56F827" box="[151,373,2010,2032]" italics="true" pageId="1" pageNumber="76">Typhlops monochrous</emphasis>
<treatmentCitation id="9901430B1278FFF0FEEEF8708D7BF827" author="Loveridge" box="[379,600,2010,2032]" page="320" pageId="1" pageNumber="76" year="1948">
<bibRefCitation id="7C3118EB1278FFF0FEEEF8708D06F827" author="Loveridge" box="[379,549,2010,2032]" pageId="1" pageNumber="76" refString="Loveridge, A. (1948) New Guinean reptiles and amphibians in the Museum of Comparative Zoology and United States National Museum. Bulletin of the Museum of Comparative Zoology, 101, 305 - 430." type="journal article" year="1948">Loveridge, 1948</bibRefCitation>
: 320
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="4CDF3592127BFFF3FF02F83C8CD4F83D" httpUri="https://zenodo.org/record/835105/files/figure.png" pageId="2" pageNumber="77" targetBox="[327,1255,221,1865]" targetPageId="2">
<paragraph id="181F651A127BFFF3FF02F83C8CD4F83D" blockId="2.[151,1436,1942,2026]" pageId="2" pageNumber="77">
<emphasis id="2AD4B908127BFFF3FF02F83C8E31F87B" bold="true" box="[151,274,1942,1964]" pageId="2" pageNumber="77">FIGURE 1.</emphasis>
Dorsal (A) and lateral (B) views of the head of the holotype of
<taxonomicName id="DFA01E99127BFFF3FC01F83C8BB6F87B" authorityName="Sternfeld" authorityYear="1913" box="[916,1173,1942,1964]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="77" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127BFFF3FC01F83C8BB6F87B" box="[916,1173,1942,1964]" italics="true" pageId="2" pageNumber="77">Gerrhopilus depressiceps</emphasis>
</taxonomicName>
(MZB 23986); dorsal (C) and lateral (D) views of the head of the holotype of
<taxonomicName id="DFA01E99127BFFF3FD3DF81F8CA2F81C" authorityName="Kraus" authorityYear="2017" box="[680,897,1973,1995]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="77" phylum="Chordata" rank="species" species="addisoni">
<emphasis id="2AD4B908127BFFF3FD3DF81F8CA2F81C" box="[680,897,1973,1995]" italics="true" pageId="2" pageNumber="77">Gerrhopilus addisoni</emphasis>
</taxonomicName>
(USNM 195953); dorsal (E) and lateral (F) views of the head of the holotype of
<taxonomicName id="DFA01E99127BFFF3FE38F87E8DA5F83D" authorityName="Kraus" authorityYear="2017" box="[429,646,2004,2026]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="77" phylum="Chordata" rank="species" species="eurydice">
<emphasis id="2AD4B908127BFFF3FE38F87E8DA5F83D" box="[429,646,2004,2026]" italics="true" pageId="2" pageNumber="77">Gerrhopilus eurydice</emphasis>
</taxonomicName>
(MCZ 145954). Scale bars = 5 mm.
</paragraph>
</caption>
<caption id="4CDF3592127AFFF2FF02FC578A31FB5A" httpUri="https://zenodo.org/record/835107/files/figure.png" pageId="3" pageNumber="78" targetBox="[229,1360,193,999]" targetPageId="3">
<paragraph id="181F651A127AFFF2FF02FC578A31FB5A" blockId="3.[151,1436,1021,1165]" pageId="3" pageNumber="78">
<emphasis id="2AD4B908127AFFF2FF02FC578E36FBC5" bold="true" box="[151,277,1021,1043]" pageId="3" pageNumber="78">FIGURE 2.</emphasis>
Map of Papua New Guinea, showing the two known localities for
<taxonomicName id="DFA01E99127AFFF2FC4CFC578BFDFBC4" authorityName="Sternfeld" authorityYear="1913" box="[985,1246,1021,1043]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127AFFF2FC4CFC578BFDFBC4" box="[985,1246,1021,1043]" italics="true" pageId="3" pageNumber="78">Gerrhopilus depressiceps</emphasis>
</taxonomicName>
(circles) and type localities for
<taxonomicName id="DFA01E99127AFFF2FE8AFBB18ED9FBE6" authorityName="Kraus" authorityYear="2017" box="[287,506,1051,1073]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="addisoni">
<emphasis id="2AD4B908127AFFF2FE8AFBB18ED9FBE6" box="[287,506,1051,1073]" italics="true" pageId="3" pageNumber="78">Gerrhopilus addisoni</emphasis>
</taxonomicName>
(triangle),
<taxonomicName id="DFA01E99127AFFF2FDFBFBB18C6BFBE6" authorityName="Kraus" authorityYear="2017" box="[622,840,1051,1073]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="eurydice">
<emphasis id="2AD4B908127AFFF2FDFBFBB18C6BFBE6" box="[622,840,1051,1073]" italics="true" pageId="3" pageNumber="78">Gerrhopilus eurydice</emphasis>
</taxonomicName>
(square),
<taxonomicName id="DFA01E99127AFFF2FC24FBB18B8BFBE6" authorityName="Kraus" authorityYear="2017" box="[945,1192,1051,1073]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="persephone">
<emphasis id="2AD4B908127AFFF2FC24FBB18B8BFBE6" box="[945,1192,1051,1073]" italics="true" pageId="3" pageNumber="78">Gerrhopilus persephone</emphasis>
</taxonomicName>
(star), and
<taxonomicName id="DFA01E99127AFFF2FAB4FBB18FECFB87" authorityName="Kraus" authorityYear="2017" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="lestes">
<emphasis id="2AD4B908127AFFF2FAB4FBB18FECFB87" italics="true" pageId="3" pageNumber="78">Gerrhopilus lestes</emphasis>
</taxonomicName>
(diamond). The demarcated northern portion of Papua New Guinea is the former German New Guinea from which the holotype of
<taxonomicName id="DFA01E99127AFFF2FE87FBF38ED0FBB8" authorityName="Sternfeld" authorityYear="1913" box="[274,499,1113,1135]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127AFFF2FE87FBF38ED0FBB8" box="[274,499,1113,1135]" italics="true" pageId="3" pageNumber="78">Typhlops depressiceps</emphasis>
</taxonomicName>
presumably came; the irregular solid line running through northwestern Papua New Guinea shows the course of the Sepik River, along which the holotype of
<taxonomicName id="DFA01E99127AFFF2FCA4FBDD8B20FB5A" authorityName="Vogt" authorityYear="1932" box="[817,1027,1143,1165]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127AFFF2FCA4FBDD8B20FB5A" box="[817,1027,1143,1165]" italics="true" pageId="3" pageNumber="78">Typhlops monochrus</emphasis>
</taxonomicName>
was presumably obtained.
</paragraph>
</caption>
<subSubSection id="50BA3691127AFFF2FF02FB1E8B7AFB1A" box="[151,1113,1204,1229]" pageId="3" pageNumber="78" type="materials_examined">
<paragraph id="181F651A127AFFF2FF02FB1E8B7AFB1A" blockId="3.[151,1437,1204,2022]" box="[151,1113,1204,1229]" pageId="3" pageNumber="78">
<materialsCitation id="A8C86F47127AFFF2FF02FB1E8B7AFB1A" ID-GBIF-Occurrence="1580520907" box="[151,1113,1204,1229]" collectionCode="ZMB" country="Papua New Guinea" location="Neuguinea" pageId="3" pageNumber="78" specimenCount="1" typeStatus="holotype">
<emphasis id="2AD4B908127AFFF2FF02FB1E8E28FB1B" bold="true" box="[151,267,1204,1229]" pageId="3" pageNumber="78">
<typeStatus id="C71BDBB8127AFFF2FF02FB1E8E27FB1A" box="[151,260,1204,1229]" pageId="3" pageNumber="78" type="holotype">Holotype</typeStatus>
.
</emphasis>
Sex unknown,
<collectionCode id="7EB1FDDF127AFFF2FE2CFB1C8ED6FB1A" box="[441,501,1206,1229]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="78">ZMB</collectionCode>
23986, “
<location id="1D7F33C1127AFFF2FDCEFB1E8DFAFB1A" LSID="urn:lsid:plazi:treatment:9009D40C1278FFF4FF02F8988E7CFB8F:1D7F33C1127AFFF2FDCEFB1E8DFAFB1A" box="[603,729,1204,1229]" country="Papua New Guinea" name="Neuguinea" pageId="3" pageNumber="78">Neuguinea</location>
” (= northern
<collectingCountry id="60B7258A127AFFF2FCE5FB1C8B6EFB1A" box="[880,1101,1204,1229]" name="Papua New Guinea" pageId="3" pageNumber="78">Papua New Guinea</collectingCountry>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="50BA3691127AFFF2FF52FB738BF1F882" pageId="3" pageNumber="78" type="diagnosis">
<paragraph id="181F651A127AFFF2FF52FB738A10F9E1" blockId="3.[151,1437,1204,2022]" pageId="3" pageNumber="78">
<emphasis id="2AD4B908127AFFF2FF52FB738E62FB25" bold="true" box="[199,321,1241,1266]" pageId="3" pageNumber="78">Diagnosis.</emphasis>
This species belongs to
<taxonomicName id="DFA01E99127AFFF2FDF6FB738DCFFB25" authorityName="Fitzinger" authorityYear="1843" box="[611,748,1241,1266]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="2AD4B908127AFFF2FDF6FB738DCFFB25" box="[611,748,1241,1266]" italics="true" pageId="3" pageNumber="78">Gerrhopilus</emphasis>
</taxonomicName>
based on the presence of head glands in the centers of the anterior head shields in addition to their anterior margins (
<bibRefCitation id="7C3118EB127AFFF2FCA3FB578CD7FAC2" author="McDowell" box="[822,1012,1277,1302]" pageId="3" pageNumber="78" refString="McDowell, S. B. (1974) A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part I. Scolecophidia. Journal of Herpetology, 8, 1 - 57." type="journal article" year="1974">McDowell 1974</bibRefCitation>
;
<bibRefCitation id="7C3118EB127AFFF2FB97FB578B92FAC1" author="Wallach" box="[1026,1201,1277,1302]" pageId="3" pageNumber="78" refString="Wallach, V. (1996 b) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996" yearSuffix="b">Wallach 1996b</bibRefCitation>
). A relatively robust (adult L/M = 70) species of
<taxonomicName id="DFA01E99127AFFF2FE4DFA8B8D43FAED" authorityName="Fitzinger" authorityYear="1843" box="[472,608,1313,1338]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="2AD4B908127AFFF2FE4DFA8B8D43FAED" box="[472,608,1313,1338]" italics="true" pageId="3" pageNumber="78">Gerrhopilus</emphasis>
</taxonomicName>
having the unique combination of a rostrate snout with a transverse keel on the ventral margin of the rostral that extends ventral to the rictus, angle of pre-oral snout in lateral aspect horizontal, distinct pupil in the eye, preocular barely covers the anterior margin of the eye in lateral view, longitudinal scale rows 24/22/20, transverse scale rows posterior to the rostral 649–664, supralabial imbrication pattern T-V, subocular scale single, presubocular scale absent, posterior margin of rostral shallowly concave, prefrontals and supraoculars subequal in size to frontal and parietals and interparietal, subcaudal scales 27–30, L/W ratio 57–67, and tail spine oriented ventrally at an angle of 70˚ to axis of anteroventral surface of that terminal scale (and, hence, to body axis). Refer to
<tableCitation id="552250A1127AFFF2FDBDF9B78D58F9E2" box="[552,635,1565,1590]" captionStart="TABLE 1" captionStartId="8.[119,197,488,507]" captionText="TABLE 1. Variation in qualitative anđ quantitative attributes among specimens of Gerrhopilus treateđ in this paper. Length measurements are given in millimeters. SVA = supraocular when vieweđ from above." pageId="3" pageNumber="78">Table 1</tableCitation>
for additional diagnostic qualitative and quantitative features.
</paragraph>
<paragraph id="181F651A127AFFF2FF52F9EB8BF1F882" blockId="3.[151,1437,1204,2022]" pageId="3" pageNumber="78">
<emphasis id="2AD4B908127AFFF2FF52F9EB8D6FF98D" bold="true" box="[199,588,1601,1626]" pageId="3" pageNumber="78">Comparisons with other species.</emphasis>
<taxonomicName id="DFA01E99127AFFF2FDC1F9EB8C51F98D" authorityName="Sternfeld" authorityYear="1913" box="[596,882,1601,1626]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127AFFF2FDC1F9EB8C51F98D" box="[596,882,1601,1626]" italics="true" pageId="3" pageNumber="78">Gerrhopilus depressiceps</emphasis>
</taxonomicName>
may be distinguished from all other members of this genus except
<taxonomicName id="DFA01E99127AFFF2FEF3F9CC8ED9F9A9" authorityName="Wallach" authorityYear="1996" box="[358,506,1637,1662]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127AFFF2FEF3F9CC8ED9F9A9" box="[358,506,1637,1662]" italics="true" pageId="3" pageNumber="78">G. mcdowelli</emphasis>
</taxonomicName>
in having a transverse keel on the ventral margin of the rostral, which gives the snout a beaked appearance in lateral aspect, and in having a posterior reduction of four longitudinal scale rows from head to vent. From that species
<taxonomicName id="DFA01E99127AFFF2FDDBF9048DDDF911" authorityName="Sternfeld" authorityYear="1913" box="[590,766,1709,1734]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127AFFF2FDDBF9048DDDF911" box="[590,766,1709,1734]" italics="true" pageId="3" pageNumber="78">G. depressiceps</emphasis>
</taxonomicName>
is distinct in its greater number of mid-dorsal scale rows (649–664 vs.
<quantity id="DF58C8FF127AFFF2FEA4F97B8E92F93D" box="[305,433,1745,1770]" metricMagnitude="1" metricUnit="m" metricValue="1.13665" metricValueMax="1.1785599999999998" metricValueMin="1.09474" pageId="3" pageNumber="78" unit="in" value="447.5" valueMax="464.0" valueMin="431.0">431–464 in</quantity>
<taxonomicName id="DFA01E99127AFFF2FE2DF9788D69F93D" authorityName="Wallach" authorityYear="1996" box="[440,586,1745,1770]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127AFFF2FE2DF9788D69F93D" box="[440,586,1745,1770]" italics="true" pageId="3" pageNumber="78">G. mcdowelli</emphasis>
</taxonomicName>
); greater length (
<quantity id="DF58C8FF127AFFF2FC9BF97B8C86F93E" box="[782,933,1745,1769]" metricMagnitude="-1" metricUnit="m" metricValue="2.555" metricValueMax="3.2299999999999995" metricValueMin="1.88" pageId="3" pageNumber="78" unit="mm" value="255.5" valueMax="323.0" valueMin="188.0">188–323 mm</quantity>
vs.
<quantity id="DF58C8FF127AFFF2FC41F97B8B66F93D" box="[980,1093,1745,1770]" metricMagnitude="0" metricUnit="m" metricValue="3.7211" metricValueMax="5.0546" metricValueMin="2.3876" pageId="3" pageNumber="78" unit="in" value="146.5" valueMax="199.0" valueMin="94.0">94–199 in</quantity>
<taxonomicName id="DFA01E99127AFFF2FBD9F9788BFCF93D" authorityName="Wallach" authorityYear="1996" box="[1100,1247,1745,1770]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127AFFF2FBD9F9788BFCF93D" box="[1100,1247,1745,1770]" italics="true" pageId="3" pageNumber="78">G. mcdowelli</emphasis>
</taxonomicName>
); narrower body (L/W =
<quantity id="DF58C8FF127AFFF2FF61F95F8E7BF8D9" box="[244,344,1781,1806]" metricMagnitude="0" metricUnit="m" metricValue="1.5748" metricValueMax="1.7018" metricValueMin="1.4478" pageId="3" pageNumber="78" unit="in" value="62.0" valueMax="67.0" valueMin="57.0">57–67 in</quantity>
<taxonomicName id="DFA01E99127AFFF2FEF5F95C8D2DF8D9" authorityName="Sternfeld" authorityYear="1913" box="[352,526,1781,1806]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127AFFF2FEF5F95C8D2DF8D9" box="[352,526,1781,1806]" italics="true" pageId="3" pageNumber="78">G. depressiceps</emphasis>
</taxonomicName>
vs.
<quantity id="DF58C8FF127AFFF2FDABF95F8D81F8D9" box="[574,674,1781,1806]" metricMagnitude="0" metricUnit="m" metricValue="1.2319" metricValueMax="1.3462" metricValueMin="1.1176" pageId="3" pageNumber="78" unit="in" value="48.5" valueMax="53.0" valueMin="44.0">44–53 in</quantity>
<taxonomicName id="DFA01E99127AFFF2FD3EF95C8C1DF8D9" authorityName="Wallach" authorityYear="1996" box="[683,830,1781,1806]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127AFFF2FD3EF95C8C1DF8D9" box="[683,830,1781,1806]" italics="true" pageId="3" pageNumber="78">G. mcdowelli</emphasis>
</taxonomicName>
); large, transversely expanded apical spine (vs. spine small and needle-like in
<taxonomicName id="DFA01E99127AFFF2FE38F8B08D1CF8E5" authorityName="Wallach" authorityYear="1996" box="[429,575,1817,1842]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127AFFF2FE38F8B08D1CF8E5" box="[429,575,1817,1842]" italics="true" pageId="3" pageNumber="78">G. mcdowelli</emphasis>
</taxonomicName>
); pre-oral snout oriented horizontally (vs. inclined at 30° from horizontal in
<taxonomicName id="DFA01E99127AFFF2FF02F8948E0BF881" authorityName="Wallach" authorityYear="1996" box="[151,296,1853,1878]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127AFFF2FF02F8948E0BF881" box="[151,296,1853,1878]" italics="true" pageId="3" pageNumber="78">G. mcdowelli</emphasis>
</taxonomicName>
); and rostral keel pointing directly downward (vs. anteroventrally in
<taxonomicName id="DFA01E99127AFFF2FBA5F8948BE2F881" authorityName="Wallach" authorityYear="1996" box="[1072,1217,1853,1878]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127AFFF2FBA5F8948BE2F881" box="[1072,1217,1853,1878]" italics="true" pageId="3" pageNumber="78">G. mcdowelli</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="50BA3691127AFFF5FF52F8CB8CADFAE7" lastPageId="4" lastPageNumber="79" pageId="3" pageNumber="78" type="description">
<paragraph id="181F651A127AFFF5FF52F8CB8ABFFCC2" blockId="3.[151,1437,1204,2022]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="79" pageId="3" pageNumber="78">
<emphasis id="2AD4B908127AFFF2FF52F8CB8D08F8AF" bold="true" box="[199,555,1889,1914]" pageId="3" pageNumber="78">
Redescription of the
<typeStatus id="C71BDBB8127AFFF2FE28F8CB8D00F8AD" box="[445,547,1889,1914]" pageId="3" pageNumber="78" type="holotype">holotype</typeStatus>
.
</emphasis>
L =
<quantity id="DF58C8FF127AFFF2FDCAF8CB8D9EF8AE" box="[607,701,1889,1913]" metricMagnitude="-1" metricUnit="m" metricValue="3.2299999999999995" pageId="3" pageNumber="78" unit="mm" value="323.0">323 mm</quantity>
, SVL =
<quantity id="DF58C8FF127AFFF2FC8FF8CB8C5BF8AE" box="[794,888,1889,1913]" metricMagnitude="-1" metricUnit="m" metricValue="3.13" pageId="3" pageNumber="78" unit="mm" value="313.0">313 mm</quantity>
, TL =
<quantity id="DF58C8FF127AFFF2FC57F8CB8B31F8AE" box="[962,1042,1889,1913]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="78" unit="mm" value="10.0">10 mm</quantity>
, HW =
<quantity id="DF58C8FF127AFFF2FBFDF8CB8BE3F8AE" box="[1128,1216,1889,1914]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="78" unit="mm" value="4.5">4.5 mm</quantity>
, NW =
<quantity id="DF58C8FF127AFFF2FA83F8CB8A4EF8AE" box="[1302,1389,1889,1913]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="78" unit="mm" value="3.9">3.9 mm</quantity>
, SN =
<quantity id="DF58C8FF127AFFF2FF38F82F8E26F84A" box="[173,261,1925,1949]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="78" unit="mm" value="2.7">2.7 mm</quantity>
, SW =
<quantity id="DF58C8FF127AFFF2FECDF82F8E93F84A" box="[344,432,1925,1949]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="78" unit="mm" value="3.8">3.8 mm</quantity>
, PSN =
<quantity id="DF58C8FF127AFFF2FD99F82F8D47F84A" box="[524,612,1925,1949]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="78" unit="mm" value="1.4">1.4 mm</quantity>
, RW =
<quantity id="DF58C8FF127AFFF2FD2CF82F8C3CF84A" box="[697,799,1925,1950]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="3" pageNumber="78" unit="mm" value="1.95">1.95 mm</quantity>
, EW =
<quantity id="DF58C8FF127AFFF2FCE1F82F8CF9F84A" box="[884,986,1925,1950]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="78" unit="mm" value="0.35">0.35 mm</quantity>
, W =
<quantity id="DF58C8FF127AFFF2FB88F82F8B56F84A" box="[1053,1141,1925,1949]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="78" unit="mm" value="4.8">4.8 mm</quantity>
, VW =
<quantity id="DF58C8FF127AFFF2FB58F82F8A07F84A" box="[1229,1316,1925,1950]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="78" unit="mm" value="4.5">4.5 mm</quantity>
, TW =
<quantity id="DF58C8FF127AFFF2FAECF82F8FE6F816" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="78" unit="mm" value="4.2">4.2 mm</quantity>
, L/W 67, M =
<quantity id="DF58C8FF127AFFF2FE16F8038E9CF815" box="[387,447,1961,1986]" metricMagnitude="-3" metricUnit="kg" metricValue="4.6" pageId="3" pageNumber="78" unit="g" value="4.6">4.6 g</quantity>
(in preservative). Snout acutely rounded in dorsal view, rounded but with ventrally directed transverse keel in lateral view, giving rostral a beaked appearance; keel deepest medially, keratinized. Preoral snout horizontal, parallel to body axis, with rostral keel extending ventral to rictus; gular region hidden from lateral view. Rostral large (0.43 head width), oval in shape, with parallel sides dorsally, posterior border extending halfway between eye and naris, posterior margin shallowly concave; ventral surface papillose, with sides slightly diverging anteriorly, slightly concave behind keel. Nasals separated dorsally by prefrontal (
<figureCitation id="809B799F127DFFF5FB07FEAE8BF2FECB" box="[1170,1233,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,249,1942,1964]" captionTargetBox="[327,1255,221,1865]" captionTargetId="figure@2.[289,1286,195,1896]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal (A) and lateral (B) views of the head of the holotype of Gerrhopilus depressiceps (MZB 23986); dorsal (C) and lateral (D) views of the head of the holotype of Gerrhopilus addisoni (USNM 195953); dorsal (E) and lateral (F) views of the head of the holotype of Gerrhopilus eurydice (MCZ 145954). Scale bars = 5 mm." httpUri="https://zenodo.org/record/835105/files/figure.png" pageId="4" pageNumber="79">Fig. 1</figureCitation>
A); superior nasal large, with slightly sinuous posterior margin, crescentic dorsally, acutely rounded ventrally (
<figureCitation id="809B799F127DFFF5FB5CFE8D8A2FFE97" box="[1225,1292,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,249,1942,1964]" captionTargetBox="[327,1255,221,1865]" captionTargetId="figure@2.[289,1286,195,1896]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal (A) and lateral (B) views of the head of the holotype of Gerrhopilus depressiceps (MZB 23986); dorsal (C) and lateral (D) views of the head of the holotype of Gerrhopilus addisoni (USNM 195953); dorsal (E) and lateral (F) views of the head of the holotype of Gerrhopilus eurydice (MCZ 145954). Scale bars = 5 mm." httpUri="https://zenodo.org/record/835105/files/figure.png" pageId="4" pageNumber="79">Fig. 1</figureCitation>
B). External naris compressed, oval shaped, oriented obliquely, close to rostral; superior nasal suture complete, extending horizontally from naris to contact rostral; inferior nasal suture complete, contacting second supralabial just posterior to latter’s contact with first supralabial. Prefrontal, frontal, supraoculars, parietals, and interparietal all subequal in size. Preocular large, triangular; larger than ocular but subequal in size to superior nasal. Ocular large, smaller than preocular, extending dorsally slightly medial to preocular, extending ventrally to ~2/3 depth of preocular, bordered posteroventrally by subocular of less than half its size. Eye with distinct round pupil and iris, situated at widest point of ocular and approximately midway along its height, anterior margin barely covered by preocular plate in lateral view. Four postoculars bordering ocular and subocular between parietal and fourth supralabial. Three supralabials, third the largest, all except first with long axis oblique to long axis of body, first approximately square. Supralabial imbrication pattern T-V, posterior border of second supralabial overlaps anteroventral margin of preocular, that of third supralabial overlaps anteroventral margin of subocular. Mental hexagonal, wider than long, projecting slightly beyond curve of lower jaw and fitting into notch on upper lip when mouth is closed. Infralabials two on each side. Head glands no longer obvious due to bleached nature of specimen.
</paragraph>
<paragraph id="181F651A127DFFF5FF52FCB58B37FC57" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">Longitudinal scale rows 24/22/20; mid-dorsal scale rows between rostral and tail tip 649; subcaudals 30; dorsocaudals 28; apical region with transverse, keratinized spine projecting posteroventrally at approximately 70˚ to axis of anteroventral surface of that terminal scale (and, hence, to body axis).</paragraph>
<paragraph id="181F651A127DFFF5FF52FC278C23FC1F" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">In preservative, dorsum very pale brown, bleached, with no difference between dorsal and ventral surfaces. Iris black; pupil pale gray, with black circle interior to that.</paragraph>
<paragraph id="181F651A127DFFF5FF52FC7E8CADFAE7" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">
<emphasis id="2AD4B908127DFFF5FF52FC7E8E1FFC3A" bold="true" box="[199,316,980,1005]" pageId="4" pageNumber="79">Variation.</emphasis>
The sole additional specimen that is complete is YPM 9066 (AMNH 66739 comprises only the head and anterior portion of the body, cf.
<bibRefCitation id="7C3118EB127DFFF5FDF8FC5D8C0BFBC7" author="McDowell" box="[621,808,1015,1040]" pageId="4" pageNumber="79" refString="McDowell, S. B. (1974) A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part I. Scolecophidia. Journal of Herpetology, 8, 1 - 57." type="journal article" year="1974">McDowell 1974</bibRefCitation>
). This specimen is smaller than the
<typeStatus id="C71BDBB8127DFFF5FB29FC5D8A3DFBC7" box="[1212,1310,1015,1040]" pageId="4" pageNumber="79" type="holotype">holotype</typeStatus>
but retains some vestiges of color pattern. Sex male; gular region visible from side; superior nasal suture inclined at approximately 10˚ from horizontal; ocular extends dorsally well medial to the preocular; approximately 25% of eye covered by preocular. Longitudinal scale rows 24/22/20; mid-dorsal scale rows 664; subcaudals 27; dorsocaudals 24. Head glands visible on rostral, nasals, and preoculars, but barely discernible, uncountable. Other features same as
<typeStatus id="C71BDBB8127DFFF5FF28FB068E00FB12" box="[189,291,1196,1221]" pageId="4" pageNumber="79" type="holotype">holotype</typeStatus>
. In preservative, pale brown, dorsum very slightly darker than ventrum, with color gradually decreasing ventrally; each scale slightly darker anteriorly. L =
<quantity id="DF58C8FF127DFFF5FCF7FB7A8CE0FB3F" box="[866,963,1232,1256]" metricMagnitude="-1" metricUnit="m" metricValue="1.88" pageId="4" pageNumber="79" unit="mm" value="188.0">188 mm</quantity>
, SVL =
<quantity id="DF58C8FF127DFFF5FBB3FB7A8BA5FB3F" box="[1062,1158,1232,1256]" metricMagnitude="-1" metricUnit="m" metricValue="1.8199999999999998" pageId="4" pageNumber="79" unit="mm" value="182.0">182 mm</quantity>
, TL =
<quantity id="DF58C8FF127DFFF5FB42FB658A3FFB3F" box="[1239,1308,1231,1256]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="79" unit="mm" value="6.0">6 mm</quantity>
, HW =
<quantity id="DF58C8FF127DFFF5FAECFB7A8FE6FADB" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="79" unit="mm" value="2.7">2.7 mm</quantity>
, NW =
<quantity id="DF58C8FF127DFFF5FE8BFB5E8E55FADB" box="[286,374,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="79" unit="mm" value="2.7">2.7 mm</quantity>
, SN =
<quantity id="DF58C8FF127DFFF5FE50FB5E8D3EFADB" box="[453,541,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="79" unit="mm" value="1.9">1.9 mm</quantity>
, SW =
<quantity id="DF58C8FF127DFFF5FDE7FB5E8DE9FADB" box="[626,714,1268,1293]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="79" unit="mm" value="2.5">2.5 mm</quantity>
, PSN =
<quantity id="DF58C8FF127DFFF5FCBCFB5E8CA2FADB" box="[809,897,1268,1292]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="79" unit="mm" value="0.9">0.9 mm</quantity>
, RW =
<quantity id="DF58C8FF127DFFF5FC42FB5E8B1DFADB" box="[983,1086,1268,1293]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="79" unit="mm" value="1.45">1.45 mm</quantity>
, EW =
<quantity id="DF58C8FF127DFFF5FB01FB5E8BD9FADB" box="[1172,1274,1268,1292]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="79" unit="mm" value="0.4">0.40 mm</quantity>
, W =
<quantity id="DF58C8FF127DFFF5FAAAFB5E8ABBFADB" box="[1343,1432,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="79" unit="mm" value="3.3">3.3 mm</quantity>
, VW =
<quantity id="DF58C8FF127DFFF5FF76FAB28E18FAE7" box="[227,315,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="79" unit="mm" value="2.8">2.8 mm</quantity>
, TW =
<quantity id="DF58C8FF127DFFF5FE05FAB28ECBFAE7" box="[400,488,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="79" unit="mm" value="2.7">2.7 mm</quantity>
, L/W 57, M =
<quantity id="DF58C8FF127DFFF5FD1AFAB28DE4FAE6" box="[655,711,1304,1329]" metricMagnitude="-3" metricUnit="kg" metricValue="1.3" pageId="4" pageNumber="79" unit="g" value="1.3">1.3 g</quantity>
(in preservative).
</paragraph>
</subSubSection>
<subSubSection id="50BA3691127DFFF5FF52FA968DD8F9DF" pageId="4" pageNumber="79" type="distribution">
<paragraph id="181F651A127DFFF5FF52FA968DD8F9DF" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">
<emphasis id="2AD4B908127DFFF5FF52FA968E43FA82" bold="true" box="[199,352,1340,1365]" pageId="4" pageNumber="79">Distribution.</emphasis>
The exact collection locality of the
<typeStatus id="C71BDBB8127DFFF5FC9FFA968C4FFA82" box="[778,876,1340,1365]" pageId="4" pageNumber="79" type="holotype">holotype</typeStatus>
of
<taxonomicName id="DFA01E99127DFFF5FC0DFA968BB6FA82" authorityName="Sternfeld" authorityYear="1913" box="[920,1173,1340,1365]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127DFFF5FC0DFA968BB6FA82" box="[920,1173,1340,1365]" italics="true" pageId="4" pageNumber="79">Typhlops depressiceps</emphasis>
</taxonomicName>
is unknown.
<bibRefCitation id="7C3118EB127DFFF5FAA3FA968FC2FAAF" author="Sternfeld" pageId="4" pageNumber="79" refString="Sternfeld, R. (1913) Beitrage zur Schlangenfauna Neuguineas und der benachbarten Inselgruppen. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1913, 384 - 389." type="journal article" year="1913">Sternfeld (1913)</bibRefCitation>
merely noted that it came from “Neu
<collectingCountry id="60B7258A127DFFF5FD20FAF58C28FAAF" box="[693,779,1375,1400]" name="Guinea-Bissau" pageId="4" pageNumber="79">Guinea</collectingCountry>
,” which
<bibRefCitation id="7C3118EB127DFFF5FCE3FAF58B67FAAF" author="McDowell" box="[886,1092,1375,1400]" pageId="4" pageNumber="79" refString="McDowell, S. B. (1974) A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part I. Scolecophidia. Journal of Herpetology, 8, 1 - 57." type="journal article" year="1974">McDowell (1974)</bibRefCitation>
sensibly interpreted to mean German New
<collectingCountry id="60B7258A127DFFF5FED5FA2E8EB7FA4A" box="[320,404,1412,1437]" name="Guinea-Bissau" pageId="4" pageNumber="79">Guinea</collectingCountry>
, which was the northern portion of present-day
<collectingCountry id="60B7258A127DFFF5FC4EFA2F8B9CFA4A" box="[987,1215,1412,1437]" name="Papua New Guinea" pageId="4" pageNumber="79">
<collectingRegion id="DA64ABF8127DFFF5FC4EFA2F8B3CFA4B" box="[987,1055,1413,1436]" country="Indonesia" name="Papua Barat" pageId="4" pageNumber="79">Papua</collectingRegion>
New Guinea
</collectingCountry>
(
<figureCitation id="809B799F127DFFF5FB41FA2E8A3CFA4B" box="[1236,1311,1412,1437]" captionStart="FIGURE 2" captionStartId="3.[151,249,1021,1043]" captionTargetBox="[229,1360,193,999]" captionTargetId="figure@3.[227,1360,193,999]" captionTargetPageId="3" captionText="FIGURE 2. Map of Papua New Guinea, showing the two known localities for Gerrhopilus depressiceps (circles) and type localities for Gerrhopilus addisoni (triangle), Gerrhopilus eurydice (square), Gerrhopilus persephone (star), and Gerrhopilus lestes (diamond). The demarcated northern portion of Papua New Guinea is the former German New Guinea from which the holotype of Typhlops depressiceps presumably came; the irregular solid line running through northwestern Papua New Guinea shows the course of the Sepik River, along which the holotype of Typhlops monochrus was presumably obtained." httpUri="https://zenodo.org/record/835107/files/figure.png" pageId="4" pageNumber="79">Fig. 2</figureCitation>
). The two specimens with known data come from Omuru (5.31˚S, 145.70˚E, YPM 9066),
<collectingRegion id="DA64ABF8127DFFF5FB9AFA0D8BFBFA17" box="[1039,1240,1447,1472]" country="Papua New Guinea" name="Madang" pageId="4" pageNumber="79">Madang Province</collectingRegion>
, and Lae (6.73˚S, 146.99˚E, AMNH 66739),
<collectingRegion id="DA64ABF8127DFFF5FE5BFA668DB4FA32" box="[462,663,1484,1509]" country="Papua New Guinea" name="Morobe" pageId="4" pageNumber="79">Morobe Province</collectingRegion>
,
<collectingCountry id="60B7258A127DFFF5FD30FA678CA1FA32" box="[677,898,1484,1509]" name="Papua New Guinea" pageId="4" pageNumber="79">
<collectingRegion id="DA64ABF8127DFFF5FD30FA678DC9FA33" box="[677,746,1485,1508]" country="Indonesia" name="Papua Barat" pageId="4" pageNumber="79">Papua</collectingRegion>
New Guinea
</collectingCountry>
(
<figureCitation id="809B799F127DFFF5FC01FA668CFFFA33" box="[916,988,1484,1509]" captionStart="FIGURE 2" captionStartId="3.[151,249,1021,1043]" captionTargetBox="[229,1360,193,999]" captionTargetId="figure@3.[227,1360,193,999]" captionTargetPageId="3" captionText="FIGURE 2. Map of Papua New Guinea, showing the two known localities for Gerrhopilus depressiceps (circles) and type localities for Gerrhopilus addisoni (triangle), Gerrhopilus eurydice (square), Gerrhopilus persephone (star), and Gerrhopilus lestes (diamond). The demarcated northern portion of Papua New Guinea is the former German New Guinea from which the holotype of Typhlops depressiceps presumably came; the irregular solid line running through northwestern Papua New Guinea shows the course of the Sepik River, along which the holotype of Typhlops monochrus was presumably obtained." httpUri="https://zenodo.org/record/835107/files/figure.png" pageId="4" pageNumber="79">Fig. 2</figureCitation>
). The species is likely restricted to the northern and/or northeastern lowlands of New
<collectingCountry id="60B7258A127DFFF5FD36FA458DD4F9DF" box="[675,759,1519,1544]" name="Guinea-Bissau" pageId="4" pageNumber="79">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="50BA3691127DFFF4FF52F9BE8E7CFB8F" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79" type="discussion">
<paragraph id="181F651A127DFFF5FF52F9BE8BEEF8FE" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="79">
<emphasis id="2AD4B908127DFFF5FF52F9BE8E18F9FA" bold="true" box="[199,315,1556,1581]" pageId="4" pageNumber="79">Remarks.</emphasis>
<bibRefCitation id="7C3118EB127DFFF5FED7F9BE8D28F9FB" author="McDowell" box="[322,523,1556,1581]" pageId="4" pageNumber="79" refString="McDowell, S. B. (1974) A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part I. Scolecophidia. Journal of Herpetology, 8, 1 - 57." type="journal article" year="1974">McDowell (1974)</bibRefCitation>
included
<taxonomicName id="DFA01E99127DFFF5FDEFF9BE8CFFF9FB" authority="Vogt 1932" authorityName="Vogt" authorityYear="1932" box="[634,988,1556,1581]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127DFFF5FDEFF9BE8C42F9FA" box="[634,865,1556,1581]" italics="true" pageId="4" pageNumber="79">Typhlops monochrus</emphasis>
<bibRefCitation id="7C3118EB127DFFF5FCFCF9BF8CFFF9FB" author="Vogt" box="[873,988,1556,1581]" pageId="4" pageNumber="79" refString="Vogt, T. (1932) Beitrag zur Reptilienfauna der ehemaligen Kolonie Deutsch-Neuguinea. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1932, 281 - 294." type="journal article" year="1932">
Vogt
<quantity id="DF58C8FF127DFFF5FC31F9BE8CFFF9FB" box="[932,988,1556,1580]" metricMagnitude="1" metricUnit="m" metricValue="4.907279999999999" pageId="4" pageNumber="79" unit="in" value="1932.0">1932</quantity>
</bibRefCitation>
</taxonomicName>
in the synonymy of
<taxonomicName id="DFA01E99127DFFF5FB51F9BF8A4EF9FA" authorityName="Sternfeld" authorityYear="1913" box="[1220,1389,1556,1581]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127DFFF5FB51F9BF8A4EF9FA" box="[1220,1389,1556,1581]" italics="true" pageId="4" pageNumber="79">T. depressiceps</emphasis>
</taxonomicName>
. He did not examine the
<typeStatus id="C71BDBB8127DFFF5FEEFF99D8EFCF987" box="[378,479,1591,1616]" pageId="4" pageNumber="79" type="holotype">holotype</typeStatus>
, although he presumed it to reside in the ZMB collection.
<bibRefCitation id="7C3118EB127DFFF5FBFCF9938BCCF987" author="Vogt" box="[1129,1263,1592,1617]" pageId="4" pageNumber="79" refString="Vogt, T. (1932) Beitrag zur Reptilienfauna der ehemaligen Kolonie Deutsch-Neuguinea. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1932, 281 - 294." type="journal article" year="1932">Vogt (1932)</bibRefCitation>
did not give an exact
<typeStatus id="C71BDBB8127DFFF5FF4EF9F78E28F9A2" box="[219,267,1629,1653]" pageId="4" pageNumber="79">type</typeStatus>
locality for his species either, but noted that the vast majority of the collection he treated in the paper describing
<taxonomicName id="DFA01E99127DFFF5FE82F92B8E90F94F" authorityName="Vogt" authorityYear="1932" box="[279,435,1663,1688]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127DFFF5FE82F92B8E90F94F" box="[279,435,1663,1688]" italics="true" pageId="4" pageNumber="79">T. monochrus</emphasis>
</taxonomicName>
came from an expedition along the
<collectingRegion id="DA64ABF8127DFFF5FCF1F9D58C87F94F" box="[868,932,1663,1688]" country="Papua New Guinea" name="West Sepik" pageId="4" pageNumber="79">Sepik</collectingRegion>
River (
<figureCitation id="809B799F127DFFF5FB94F9D58B69F94F" box="[1025,1098,1663,1688]" captionStart="FIGURE 2" captionStartId="3.[151,249,1021,1043]" captionTargetBox="[229,1360,193,999]" captionTargetId="figure@3.[227,1360,193,999]" captionTargetPageId="3" captionText="FIGURE 2. Map of Papua New Guinea, showing the two known localities for Gerrhopilus depressiceps (circles) and type localities for Gerrhopilus addisoni (triangle), Gerrhopilus eurydice (square), Gerrhopilus persephone (star), and Gerrhopilus lestes (diamond). The demarcated northern portion of Papua New Guinea is the former German New Guinea from which the holotype of Typhlops depressiceps presumably came; the irregular solid line running through northwestern Papua New Guinea shows the course of the Sepik River, along which the holotype of Typhlops monochrus was presumably obtained." httpUri="https://zenodo.org/record/835107/files/figure.png" pageId="4" pageNumber="79">Fig. 2</figureCitation>
), and it has been reasonably presumed that the
<typeStatus id="C71BDBB8127DFFF5FEF0F90F8EB6F96A" box="[357,405,1701,1725]" pageId="4" pageNumber="79">type</typeStatus>
specimen did as well (
<bibRefCitation id="7C3118EB127DFFF5FD03F90E8C70F96B" author="McDowell" box="[662,851,1700,1725]" pageId="4" pageNumber="79" refString="McDowell, S. B. (1974) A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part I. Scolecophidia. Journal of Herpetology, 8, 1 - 57." type="journal article" year="1974">McDowell 1974</bibRefCitation>
). The
<typeStatus id="C71BDBB8127DFFF5FC0CF90E8CD8F96A" box="[921,1019,1700,1725]" pageId="4" pageNumber="79" type="holotype">holotype</typeStatus>
has not subsequently been located in the ZMB (
<bibRefCitation id="7C3118EB127DFFF5FE8FF9628ED6F937" author="Bauer" box="[282,501,1735,1760]" pageId="4" pageNumber="79" refString="Bauer, A. M., Wallach, V. &amp; Gunther, R. (2002) An annotated type catalogue of the scolecophidian, alethinophidian, and macrostomatan snakes in the collection of the Museum fur Naturkunde der Humboldt-Universitat zu Berlin. Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 78, 157 - 176." type="journal article" year="2002">
Bauer
<emphasis id="2AD4B908127DFFF5FEF9F9638E88F937" box="[364,427,1735,1760]" italics="true" pageId="4" pageNumber="79">et al.</emphasis>
2002
</bibRefCitation>
), and it remains uncertain whether it still exists and, if so, where it resides. Consequently, it appears to have remained unexamined since Vogt’s original description, the brevity of which does not allow for useful taxonomic discrimination using modern diagnostic characters for the group.
</paragraph>
<paragraph id="181F651A127DFFF4FF52F89E8D25FD1B" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,1112]" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79">
McDowell’s (1974) synonymization of
<taxonomicName id="DFA01E99127DFFF5FD13F89E8C4CF89A" authorityName="Vogt" authorityYear="1932" box="[646,879,1844,1869]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127DFFF5FD13F89E8C4CF89A" box="[646,879,1844,1869]" italics="true" pageId="4" pageNumber="79">Typhlops monochrus</emphasis>
</taxonomicName>
with
<taxonomicName id="DFA01E99127DFFF5FC24F89F8B78F89A" authorityName="Sternfeld" authorityYear="1913" box="[945,1115,1844,1869]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127DFFF5FC24F89F8B78F89A" box="[945,1115,1844,1869]" italics="true" pageId="4" pageNumber="79">T. depressiceps</emphasis>
</taxonomicName>
may have been based on an error in Sternfeld’s (1913) original description of the latter, in which he claimed the
<typeStatus id="C71BDBB8127DFFF5FBCEF8FD8B9EF8A7" box="[1115,1213,1879,1904]" pageId="4" pageNumber="79" type="holotype">holotype</typeStatus>
to have 24 rows of scales. Presumably this count was taken at midbody, although that is not clearly specified. Sternfeld’s report was repeated by de
<bibRefCitation id="7C3118EB127DFFF5FED8F80A8EC3F86F" author="Rooij" box="[333,480,1952,1977]" pageId="4" pageNumber="79" refString="de Rooij, N. (1917) The reptiles of the Indo-Australian Archipelago, II: Ophidia. E. J. Brill, Leiden, 334 pp." type="book" year="1917">Rooij (1917)</bibRefCitation>
and
<bibRefCitation id="7C3118EB127DFFF5FD8BF80A8DCDF86F" author="McDowell" box="[542,750,1952,1977]" pageId="4" pageNumber="79" refString="McDowell, S. B. (1974) A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part I. Scolecophidia. Journal of Herpetology, 8, 1 - 57." type="journal article" year="1974">McDowell (1974)</bibRefCitation>
, neither of whom examined the
<typeStatus id="C71BDBB8127DFFF5FBFBF80B8BBDF86E" box="[1134,1182,1953,1977]" pageId="4" pageNumber="79">type</typeStatus>
specimen.
<bibRefCitation id="7C3118EB127DFFF5FAB0F80A8FC2F80B" author="McDowell" pageId="4" pageNumber="79" refString="McDowell, S. B. (1974) A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part I. Scolecophidia. Journal of Herpetology, 8, 1 - 57." type="journal article" year="1974">McDowell (1974)</bibRefCitation>
did remark that the single complete specimen of presumptive
<taxonomicName id="DFA01E99127DFFF5FC27F86F8B42F80A" authorityName="Sternfeld" authorityYear="1913" box="[946,1121,1988,2013]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127DFFF5FC27F86F8B42F80A" box="[946,1121,1988,2013]" italics="true" pageId="4" pageNumber="79">G. depressiceps</emphasis>
</taxonomicName>
that he did examine (YPM 9066) had 22 mid-body scale rows, which he thought served to connect Sternfeld’s
<taxonomicName id="DFA01E99127CFFF4FBAAFF338BCAFF67" authorityName="Sternfeld" authorityYear="1913" box="[1087,1257,151,176]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FBAAFF338BCAFF67" box="[1087,1257,151,176]" italics="true" pageId="5" pageNumber="80">T. depressiceps</emphasis>
</taxonomicName>
(24 scale rows) with Vogt’s
<taxonomicName id="DFA01E99127CFFF4FEB8FF178EEDFF02" authorityName="Vogt" authorityYear="1932" box="[301,462,188,213]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127CFFF4FEB8FF178EEDFF02" box="[301,462,188,213]" italics="true" pageId="5" pageNumber="80">T. monochrus</emphasis>
</taxonomicName>
(20 scale rows). It was partly this presumed annectance that led McDowell to synonymize the latter with the former. Based presumably on this history,
<bibRefCitation id="7C3118EB127CFFF4FC40FF758BB3FF2F" author="Wallach" box="[981,1168,223,248]" pageId="5" pageNumber="80" refString="Wallach, V. (1996 b) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996" yearSuffix="b">Wallach (1996b)</bibRefCitation>
reported the number of longitudinal scale rows (LSR) in
<taxonomicName id="DFA01E99127CFFF4FD83FEAF8DE6FECA" authorityName="Sternfeld" authorityYear="1913" box="[534,709,260,285]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FD83FEAF8DE6FECA" box="[534,709,260,285]" italics="true" pageId="5" pageNumber="80">G. depressiceps</emphasis>
</taxonomicName>
to range from 20–24. He did not define LSR, but based on his reported values (and the contrast in values between what he reports for
<taxonomicName id="DFA01E99127CFFF4FC53FE828B7AFE97" authorityName="Wallach" authorityYear="1996" box="[966,1113,295,320]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="mcdowelli">
<emphasis id="2AD4B908127CFFF4FC53FE828B7AFE97" box="[966,1113,295,320]" italics="true" pageId="5" pageNumber="80">G. mcdowelli</emphasis>
</taxonomicName>
in his tables 1 and 2), it too evidently refers to a count taken at midbody, which is standard for snakes unless otherwise noted. This interpretation is consistent too with his use of mid-body scale counts in his dichotomous key provided in the same paper. In contrast to these reports, I find the
<typeStatus id="C71BDBB8127CFFF4FD20FE3E8C34FE7A" box="[693,791,404,429]" pageId="5" pageNumber="80" type="holotype">holotype</typeStatus>
of
<taxonomicName id="DFA01E99127CFFF4FCD3FE3F8CD6FE7A" authorityName="Sternfeld" authorityYear="1913" box="[838,1013,404,429]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FCD3FE3F8CD6FE7A" box="[838,1013,404,429]" italics="true" pageId="5" pageNumber="80">T. depressiceps</emphasis>
</taxonomicName>
to have 22 scale rows at midbody, consistent with what McDowell and I have found for the Omuru specimen. Hence, there is no annectance in longitudinal scale rows between the assorted specimens I assign to
<taxonomicName id="DFA01E99127CFFF4FC3CFE778B7AFE22" authorityName="Sternfeld" authorityYear="1913" box="[937,1113,476,501]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FC3CFE778B7AFE22" box="[937,1113,476,501]" italics="true" pageId="5" pageNumber="80">G. depressiceps</emphasis>
</taxonomicName>
and the
<typeStatus id="C71BDBB8127CFFF4FB56FE778BD0FE22" box="[1219,1267,477,501]" pageId="5" pageNumber="80">type</typeStatus>
description of
<taxonomicName id="DFA01E99127CFFF4FF02FE558EA1FDCF" authorityName="Vogt" authorityYear="1932" box="[151,386,511,536]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127CFFF4FF02FE558EA1FDCF" box="[151,386,511,536]" italics="true" pageId="5" pageNumber="80">Typhlops monochrus</emphasis>
</taxonomicName>
. Given the generally conserved nature of scale-row patterns among blindsnakes, there is, therefore, reason to entertain the notion that
<taxonomicName id="DFA01E99127CFFF4FD3AFD8F8C6EFDEA" authorityName="Vogt" authorityYear="1932" box="[687,845,548,573]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127CFFF4FD3AFD8F8C6EFDEA" box="[687,845,548,573]" italics="true" pageId="5" pageNumber="80">T. monochrus</emphasis>
</taxonomicName>
may prove to be a valid taxon differing from
<taxonomicName id="DFA01E99127CFFF4FA11FD8F8E06FDB7" authorityName="Sternfeld" authorityYear="1913" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FA11FD8F8E06FDB7" italics="true" pageId="5" pageNumber="80">G. depressiceps</emphasis>
</taxonomicName>
in having 20 instead of 22 mid-body scale rows. Determining whether that possibility is valid will require collection of new material from northern
<collectingCountry id="60B7258A127CFFF4FD50FDC78CBCFD52" box="[709,927,620,645]" name="Papua New Guinea" pageId="5" pageNumber="80">
<collectingRegion id="DA64ABF8127CFFF4FD50FDC78C29FD53" box="[709,778,621,644]" country="Indonesia" name="Papua Barat" pageId="5" pageNumber="80">Papua</collectingRegion>
New Guinea
</collectingCountry>
inasmuch as the
<typeStatus id="C71BDBB8127CFFF4FBF1FDC68BE5FD52" box="[1124,1222,620,645]" pageId="5" pageNumber="80" type="holotype">holotype</typeStatus>
remains unlocated and possibly lost. Nonetheless, future workers on these snakes should be aware that present evidence does not yet settle the status of
<taxonomicName id="DFA01E99127CFFF4FEF2FD1F8EDCFD1A" authorityName="Vogt" authorityYear="1932" box="[359,511,692,717]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="monochrus">
<emphasis id="2AD4B908127CFFF4FEF2FD1F8EDCFD1A" box="[359,511,692,717]" italics="true" pageId="5" pageNumber="80">T. monochrus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="181F651A127CFFF4FF52FD728E7CFB8F" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="80">
An additional morphological discrepancy in the literature is that
<bibRefCitation id="7C3118EB127CFFF4FC09FD7D8B74FD27" author="Wallach" box="[924,1111,727,752]" pageId="5" pageNumber="80" refString="Wallach, V. (1996 b) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996" yearSuffix="b">Wallach (1996b)</bibRefCitation>
reported the number of middorsal scale rows in
<taxonomicName id="DFA01E99127CFFF4FE1FFD568D88FCC2" authorityName="Sternfeld" authorityYear="1913" box="[394,683,764,789]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FE1FFD568D88FCC2" box="[394,683,764,789]" italics="true" pageId="5" pageNumber="80">Gerrhopilus depressiceps</emphasis>
</taxonomicName>
to vary from 631–713. Of the three specimens
<bibRefCitation id="7C3118EB127CFFF4FB4BFD568ABFFCC2" author="Wallach" box="[1246,1436,764,789]" pageId="5" pageNumber="80" refString="Wallach, V. (1996 b) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996" yearSuffix="b">Wallach (1996b)</bibRefCitation>
examined, I obtain values of 627 (vs. his 631) for USNM 195953, 647 (vs. his 648) for MCZ 145954, and 649 (vs. his 713) for ZMB 23986 (
<tableCitation id="552250A1127CFFF4FE5CFCEE8D03FC8B" box="[457,544,836,861]" captionStart="TABLE 1" captionStartId="8.[119,197,488,507]" captionText="TABLE 1. Variation in qualitative anđ quantitative attributes among specimens of Gerrhopilus treateđ in this paper. Length measurements are given in millimeters. SVA = supraocular when vieweđ from above." pageId="5" pageNumber="80">Table 1</tableCitation>
). Count differences for the first two specimens are trivial, but that for ZMB 23986 is significant. This large divergence between our observations for ZMB 23986 is important because Wallach’s value would imply a much smaller difference in mid-dorsal scale rows between
<taxonomicName id="DFA01E99127CFFF4FB1BFC278A18FC72" authorityName="Sternfeld" authorityYear="1913" box="[1166,1339,908,933]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FB1BFC278A18FC72" box="[1166,1339,908,933]" italics="true" pageId="5" pageNumber="80">G. depressiceps</emphasis>
</taxonomicName>
and two of the species described below than appears to be the case. I have repeated my counts for these specimens two or three times for each and I am confident that
<taxonomicName id="DFA01E99127CFFF4FD17FC7F8C0DFC3A" authorityName="Sternfeld" authorityYear="1913" box="[642,814,980,1005]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FD17FC7F8C0DFC3A" box="[642,814,980,1005]" italics="true" pageId="5" pageNumber="80">G. depressiceps</emphasis>
</taxonomicName>
, on the basis of available specimens, does not attain the large number of mid-dorsal scale rows attributed to it by
<bibRefCitation id="7C3118EB127CFFF4FC83FC5D8CF0FBC7" author="Wallach" box="[790,979,1015,1040]" pageId="5" pageNumber="80" refString="Wallach, V. (1996 b) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996" yearSuffix="b">Wallach (1996b)</bibRefCitation>
. Consequently, I report a much narrower range in this character for
<taxonomicName id="DFA01E99127CFFF4FE55FBB78D4EFBE2" authorityName="Sternfeld" authorityYear="1913" box="[448,621,1052,1077]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="2AD4B908127CFFF4FE55FBB78D4EFBE2" box="[448,621,1052,1077]" italics="true" pageId="5" pageNumber="80">G. depressiceps</emphasis>
</taxonomicName>
and the new taxa I remove from that species than appears from the data presented by him.
</paragraph>
</subSubSection>
</treatment>
</document>
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