308 lines
35 KiB
XML
308 lines
35 KiB
XML
<document ID-CLB-Dataset="22826" ID-DOI="http://dx.doi.org/10.3897/alpento.4.50703" ID-GBIF-Dataset="a9e0336c-85fe-4d7c-8998-fadb8b1e4a29" ID-Pensoft-Pub="2535-0889-4-39" ID-Pensoft-UUID="73E3CBB8841D53E98BB78BA05DDC69C9" ID-ZooBank="D9502F1AAEC04B0F845C87D86632AF17" ModsDocID="2535-0889-4-39" checkinTime="1587713590471" checkinUser="pensoft" docAuthor="Huemer, Peter" docDate="2020" docId="62782D16A86A5ED891798A210F2DEAE9" docLanguage="en" docName="AlpEntomol 4: 39-63" docOrigin="Alpine Entomology 4" docSource="http://dx.doi.org/10.3897/alpento.4.50703" docTitle="Caryocolum dianthella Huemer 2020, stat. rev." docType="treatment" docVersion="6" id="73E3CBB8841D53E98BB78BA05DDC69C9" lastPageNumber="39" masterDocId="73E3CBB8841D53E98BB78BA05DDC69C9" masterDocTitle="Integrative revision of the Caryocolum schleichi species group - a striking example of a temporally changing species concept (Lepidoptera, Gelechiidae)" masterLastPageNumber="63" masterPageNumber="39" pageNumber="39" updateTime="1732751505677" updateUser="ExternalLinkService">
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<mods:titleInfo id="E548438B74CD713694E51098BE07D609">
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<mods:title id="A23262FF46E740B66C719061A5A5B487">Integrative revision of the Caryocolum schleichi species group - a striking example of a temporally changing species concept (Lepidoptera, Gelechiidae)</mods:title>
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<mods:namePart id="369DBDE125990D1942CAC726D09B3461">Huemer, Peter</mods:namePart>
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<mods:affiliation id="0FCC0E99252C6D03780E05DC3C8E29DE">Tiroler Landesmuseen Betriebgsges. m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria</mods:affiliation>
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<mods:nameIdentifier id="871E2368855512FBB1653331ACD56B59" type="orcid">https://orcid.org/0000-0002-0630-545X</mods:nameIdentifier>
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<mods:nameIdentifier id="07E40AA81AE8C640464235F6AEAC2AD9" type="email">p.huemer@tiroler-landesmuseen.at</mods:nameIdentifier>
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<mods:typeOfResource id="DBF62395EA32AB8DBC969198749BACEB">text</mods:typeOfResource>
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<mods:title id="DB10661AE791889A3736BFFD0D5F058D">Alpine Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="9868BAA584B0E336F63B322F90AFAD6D">
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<mods:date id="FE53BCA3AA4BE8784AA5D313AB76B2DC">2020</mods:date>
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<mods:number id="BE762F94C70A49B9ADA40DB754D50E79">4</mods:number>
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<mods:start id="6052D48FB11B3A25A31B8843518B46EB">39</mods:start>
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<mods:end id="5B9E20D4918CBB6CDE2D88C19DA85093">63</mods:end>
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<mods:url id="C7F15BE5AF58109B4EE8745EEF9971BB">http://dx.doi.org/10.3897/alpento.4.50703</mods:url>
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<mods:classification id="524F415378FA409609B65D5C377550AF">journal article</mods:classification>
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<mods:identifier id="3B6407AC74B9F3D6C9D498253C44307E" type="DOI">http://dx.doi.org/10.3897/alpento.4.50703</mods:identifier>
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<treatment id="62782D16A86A5ED891798A210F2DEAE9" ID-GBIF-Taxon="163737597" LSID="urn:lsid:plazi:treatment:62782D16A86A5ED891798A210F2DEAE9" httpUri="http://treatment.plazi.org/id/62782D16A86A5ED891798A210F2DEAE9" lastPageNumber="39" pageId="0" pageNumber="39" scope_family="Gelechiidae" scope_order="Lepidoptera">
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<subSubSection id="518E20540289E1F2B3B89F14A0C180FC" pageId="0" pageNumber="39" type="nomenclature">
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<paragraph id="0E6E0E988920F3DDFDBF76E3BA3412BF" pageId="0" pageNumber="39">
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<taxonomicName id="DAA45FFC46D0E3E364A7F4FA3B00EB34" ID-CoL="98ZMY" LSID="62782D16-A86A-5ED8-9179-8A210F2DEAE9" authority="(Chretien, 1925)" authorityName="Huemer" authorityYear="2020" baseAuthorityName="Chretien" baseAuthorityYear="1925" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum dianthella" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="dianthella" status="stat. rev.">
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Caryocolum dianthella (
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<normalizedToken id="B131C031810F962DC9C673C0E4D34383" originalValue="Chrétien">Chretien</normalizedToken>
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, 1925)
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</taxonomicName>
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<taxonomicNameLabel id="56D1B6F593B8BA7735627A82E31D7D73" pageId="0" pageNumber="39">stat. rev.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="8778533D2CC7BB8884769871EBF69628" pageId="0" pageNumber="39" type="reference_group">
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<paragraph id="E47C568F7F3F349CFC5E3B6EDCE4E6CE" pageId="0" pageNumber="39">
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<taxonomicName id="541D24A4DD32F643CF57C5B9D81917A3" authorityName="Chretien" authorityYear="1925" class="Insecta" family="Gelechiidae" genus="Lita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lita dianthella" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="dianthella">Lita dianthella</taxonomicName>
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<bibRefCitation id="C461BF4FB8CBB0B2133D2E6D097A2E33" author="Chretien, P" journalOrPublisher="L'Amateur de Papillons" pageId="0" pageNumber="39" pagination="241 - 247" refId="B7" refString="Chretien, P, 1925. La legende de Graellsia isabellae. Appendice. L'Amateur de Papillons 2: 241 - 247" title="La legende de Graellsia isabellae. Appendice." volume="2" year="1925">
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<normalizedToken id="16324761D3EA896E8A8BC484B48251B0" originalValue="Chrétien">Chretien</normalizedToken>
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1925
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</bibRefCitation>
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: 246. Syntypes, Spain: Segovia, La Granja de San Ildefonso (MNHN) [not examined].
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</paragraph>
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<paragraph id="3005C1BAD0678456ECCAADB3BB5F788A" pageId="0" pageNumber="39">
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<taxonomicName id="5EF75FAD8D1F4D2CC30AF781C312CD8F" authorityName="Derra" authorityYear="1985" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum hackeri" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="hackeri">Caryocolum hackeri</taxonomicName>
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<bibRefCitation id="80B163A4F2D3D34584D22CBDA4DD604F" author="Derra, G" journalOrPublisher="Entomofauna, Linz" pageId="0" pageNumber="39" pagination="373 - 379" refId="B9" refString="Derra, G, 1985. Caryocolum hackeri sp. n., eine neue Gelechiidae aus den Pyrenaeen Frankreichs und Spaniens (Lepidoptera, Gelechiidae). Entomofauna, Linz 6: 373 - 379" title="Caryocolum hackeri sp. n., eine neue Gelechiidae aus den Pyrenaeen Frankreichs und Spaniens (Lepidoptera, Gelechiidae)." volume="6" year="1985">Derra 1985</bibRefCitation>
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: 373, figs 1, 4-6. Holotype ♂, Spain (coll. Derra, Bamberg). Synonymized by
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<bibRefCitation id="4BDC4030D80F467089D81370E4198B99" author="Huemer, P" journalOrPublisher="Bulletin of the British Museum of Natural History (Entomology)" pageId="0" pageNumber="39" pagination="439 - 571" refId="B15" refString="Huemer, P, 1988. A taxonomic revision of Caryocolum (Lepidoptera: Gelechiidae). Bulletin of the British Museum of Natural History (Entomology) 57: 439 - 571" title="A taxonomic revision of Caryocolum (Lepidoptera: Gelechiidae)." volume="57" year="1988">Huemer 1988</bibRefCitation>
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: 489.
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</paragraph>
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</subSubSection>
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<subSubSection id="B65310AF0D6DDA4B1E1DBAF5CC6309A0" pageId="0" pageNumber="39" type="materials_examined">
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<paragraph id="F4B4D50A2BB514E6239CA8E1754925F7" pageId="0" pageNumber="39">Material examined.</paragraph>
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<paragraph id="765D0EA0C1E145B7607D7119508D451B" pageId="0" pageNumber="39">
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<materialsCitation id="825653B6F4FD5899DA98E1EA3B1A7BB9" ID-GBIF-Occurrence="2609799212" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectorName="Derra & Bamberg" elevation="0" location="Pyrenees" municipality="Spain" specimenCode="86/248" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="CD92154CC51C44E31B312BFCBC2DD54C" bold="true" pageId="0" pageNumber="39">
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<typeStatus id="C0014C1E8F844329A8683C961E3A10FB">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="76DC4CF8A2CC188D29E34973545566B1" type="male">♂</specimenCount>
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(
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<taxonomicName id="E7C4E014BB4F59B2D1B1215CC3FC76EE" authorityName="Derra" authorityYear="1985" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="hackeri" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="hackeri">
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<emphasis id="C692437B277B47324D6524F9F0A96843" italics="true" pageId="0" pageNumber="39">hackeri</emphasis>
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</taxonomicName>
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),
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<collectingMunicipality id="9265F317AF34547A97B431DB6BB70D19">Spain</collectingMunicipality>
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:
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<location id="AB69EB81DBC3BDE71FFB42E6A6101C22" LSID="urn:lsid:plazi:treatment:62782D16A86A5ED891798A210F2DEAE9:AB69EB81DBC3BDE71FFB42E6A6101C22" municipality="Spain" name="Pyrenees">Pyrenees</location>
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,
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<location id="3C78A81E35399D479B60FC5E263D7F62" LSID="urn:lsid:plazi:treatment:62782D16A86A5ED891798A210F2DEAE9:3C78A81E35399D479B60FC5E263D7F62" municipality="Spain" name="Lerida province, Bellver de Cerdana">
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||
Lerida province, Bellver de
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<normalizedToken id="FC3831A8967ADE024E5D79BF6939431A" originalValue="Cerdaña">Cerdana</normalizedToken>
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</location>
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,
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<elevation id="DE5943FDC36A26F6C4D850A880065605">900 m,</elevation>
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<collectingDate id="C4D344F234D5D3EAEAD7543DF01C9DF8">20 Sep. 1981</collectingDate>
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, leg.
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<collectorName id="CDF825C8CB0C0DFF1D37729DCCF94B5E">Derra</collectorName>
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, genitalia slide no.
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<specimenCode id="CA66F39A40AA2683335F51E173885287">86/248</specimenCode>
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P. Huemer (coll.
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<collectorName id="4134501DF7D506D1268CC95CACB3F7D4">Derra</collectorName>
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,
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<collectorName id="D674D9E0DE8E6C18243FB2EF4B3AEE5D">Bamberg</collectorName>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="313DCE150CF348C47E56DE12FD611134" pageId="0" pageNumber="39">
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Andorra: 1 ♀, La Massana, 2 Jun. 2002 e.l. (
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<taxonomicName id="E863291329D70D7DFF50CBAD74542574" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus pungens" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="species" species="pungens">
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<emphasis id="3CC8BE362D8F6A05DFE515D53EC29450" italics="true" pageId="0" pageNumber="39">Dianthus pungens</emphasis>
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</taxonomicName>
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), leg. Mazel (TLMF). Spain: 1 ♂, 1 ♀ (paratypes
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<taxonomicName id="78E5712C2B003AA2123EBDED58BF4892" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="hackeri" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="species" species="hackeri">
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<emphasis id="9137FBCF0A40FFB8D2169426746002A5" italics="true" pageId="0" pageNumber="39">hackeri</emphasis>
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</taxonomicName>
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), same data as holotype but 7 Oct 1981, leg. Derra (coll. Derra, Bamberg); 1 ♀, Zaragoza province, 3 km E Cerveruela, Rio del Huerva, 800 m, 6 Sep. 2001, leg. Skule & Skou (TLMF); 1 ♂, Alicante province, Alcoj, Font Roja, W El Menejador, S slope, 1300 m, 4 Sep. 2005, leg. Huemer (TLMF); 2 ♂♂, 6 ♀♀, Granada province, Camino de Capileira, 1250 m, 13 Jul. 1985, leg. Baldizzone & Traugott-Olsen; 2 ♂♂, Granada province, Sierra Nevada, Puerto de la Ragua, 1000 m, 25 Jun. 1968, 20 Jul. 1969, leg. Sattler & Carter; 2 ♂♂, 1 ♀, Granada province, Sierra de Alfacar, 1500 m, 3 Jul. l962, 13 Sep. 1972, leg. Sattler; 1 ♂, Huesca province, Jaca, 12 Aug. 1933, leg. Fassnidge (all NHM); 5 ♂♂, 1 ♀, Granada province, Sierra Nevada, Cam. de Veleta, 2000-2300 m, 24 Jul. 1983, 19 Aug. 1984, leg. Traugott-Olsen (ZMUC, TLMF); 1 ♂, 1 ♀, Granada province, Sierra Nevada, Cam. de Veleta, 2250-2300 m, 1 Aug. 1986, leg. Traugott-Olsen; 1 ♂, Granada province, Sierra Nevada, Ruta del Veleta, 1600 m, 19 Sep. 1987 (all TLMF). France: 2 ♂♂, 1 ♀,
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<normalizedToken id="FCE1E37FA93CB02CB4F2F6B40C96DE39" originalValue="Pyrénées-Orientales">Pyrenees-Orientales</normalizedToken>
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,
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<normalizedToken id="32B5612F83063ADEE3BB73F8D2D80410" originalValue="Thuès-les-Bains">Thues-les-Bains</normalizedToken>
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, larvae 24 Jun. 1900 on
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<taxonomicName id="508FD339CE4532DE7DDF37937354A8A0" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="genus">
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<emphasis id="949FC1FE7D58AF5515AF4346A3E8D015" italics="true" pageId="0" pageNumber="39">Dianthus</emphasis>
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</taxonomicName>
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, moths emerged 26 Jul. 1900, leg. Walsingham (NHM). Morocco: 1 ♂, Haut Atlas,
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<normalizedToken id="31F1201CF6EB8307CC77694596D4776C" originalValue="Oukaïmeden">Oukaimeden</normalizedToken>
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, 2600 m, 9 & 11 Jul. l975, leg. Kasy (NHMW).
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</paragraph>
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<caption id="A6274099EF803954FDCFCAAFC4495E16" doi="10.3897/alpento.4.50703.figures13-16" httpUri="https://binary.pensoft.net/fig/403190" pageId="0" pageNumber="39" start="Figures 13–16" startId="F5">
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<paragraph id="5975F7A4A9D91721B3436DBFC392B0A0" pageId="0" pageNumber="39">
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<emphasis id="A1244D20993C35A6F97167A5AD00F43E" bold="true" pageId="0" pageNumber="39">Figures 13-16.</emphasis>
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Male genitalia.
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<emphasis id="E7BADCF13F87D6FCB11E2681D9830007" bold="true" pageId="0" pageNumber="39">13</emphasis>
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<taxonomicName id="C23B7DB8DFBB67E0FFCBC259C13F4920" baseAuthorityName="Christoph" baseAuthorityYear="1872" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum schleichi" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="schleichi">
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<emphasis id="117797E0FC6921816A7D45E87B7BB2C2" italics="true" pageId="0" pageNumber="39">Caryocolum schleichi</emphasis>
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</taxonomicName>
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, Syria, slide GU 86/273 P. Huemer;
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<emphasis id="FA51E2BAD6CAEC5E50BAC1040E69E740" bold="true" pageId="0" pageNumber="39">14</emphasis>
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<taxonomicName id="32050CCB3CCFD3FF5236CB1B5339EF49" lsidName="C. dianthella" pageId="0" pageNumber="39" rank="species" species="dianthella">
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<emphasis id="BC63C96F8F30539768E7EC0D9BC033FD" italics="true" pageId="0" pageNumber="39">C. dianthella</emphasis>
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</taxonomicName>
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, Spain, slide GEL 1284 P. Huemer;
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<emphasis id="DBF70554532698A167D2F4DE593C975E" bold="true" pageId="0" pageNumber="39">15</emphasis>
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<taxonomicName id="82C0F9A518D00A3216ACD0ECF0F20997" lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
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<emphasis id="744B6DB74453A21AC4DD804B0CFFAD06" italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
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</taxonomicName>
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, Austria, slide GEL 1256 P. Huemer;
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<emphasis id="D93B19D63D3177EEA2EB7A5DDDC4AC4B" bold="true" pageId="0" pageNumber="39">16</emphasis>
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||
<taxonomicName id="01B847198907ECCF35EB8D8C7D95BF68" lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
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<emphasis id="EFD3C81FFFBD99F2A3B212D92A612519" italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
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</taxonomicName>
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, Italy, slide GEL 1290 P. Huemer.
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</paragraph>
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</caption>
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<caption id="9E4495ED5949CD2F4B342453BCE19338" doi="10.3897/alpento.4.50703.figures17-20" httpUri="https://binary.pensoft.net/fig/403191" pageId="0" pageNumber="39" start="Figures 17–20" startId="F6">
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||
<paragraph id="593C426EBA2E3B7AD311BDE0DBA24708" pageId="0" pageNumber="39">
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<emphasis id="B447426E5BC670549BEEE2E37F6BEA82" bold="true" pageId="0" pageNumber="39">Figures 17-20.</emphasis>
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Male genitalia.
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<emphasis id="5D1B83358BB2A106F5240BB29EA6DE54" bold="true" pageId="0" pageNumber="39">17</emphasis>
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<taxonomicName id="8EE2E8B3581406BD43D6677A2990C369" baseAuthorityName="Benander" baseAuthorityYear="1937" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum arenariella" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="arenariella">
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||
<emphasis id="48C86AF7ADBE52840ED0475E1BE2429E" italics="true" pageId="0" pageNumber="39">Caryocolum arenariella</emphasis>
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</taxonomicName>
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, Russia, slide GEL 1260 P. Huemer;
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<emphasis id="1556D60DA36EB81E2AF2156ADDE76655" bold="true" pageId="0" pageNumber="39">18</emphasis>
|
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<taxonomicName id="0FC1B8D920D2C4CC2B848AB361273648" lsidName="C. messneri" pageId="0" pageNumber="39" rank="species" species="messneri">
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||
<emphasis id="3E982C6CEE36FF3E4234EAE589A5E059" italics="true" pageId="0" pageNumber="39">C. messneri</emphasis>
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</taxonomicName>
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, holotype, Italy, slide GEL 1255;
|
||
<emphasis id="EC6080947EFB5082971AB6B32293D588" bold="true" pageId="0" pageNumber="39">19</emphasis>
|
||
<taxonomicName id="9C2529425AA215578618F88C7B147372" lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
|
||
<emphasis id="0A56295084BE771BC017DA5A2FA1D8D0" italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Italy, slide GEL 1258 P. Huemer;
|
||
<emphasis id="CC6287747C8424C81523766FA8119205" bold="true" pageId="0" pageNumber="39">21</emphasis>
|
||
<taxonomicName id="60FB6E47BA727F3C3CC8861EAEE90CB2" lsidName="C. habeleri" pageId="0" pageNumber="39" rank="species" species="habeleri">
|
||
<emphasis id="D4813BDC9D3FCE711710EF5AC381CBEC" italics="true" pageId="0" pageNumber="39">C. habeleri</emphasis>
|
||
</taxonomicName>
|
||
, paratype, France, slide GEL 1289 P. Huemer.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="64C08742218AD54E6B24E4BD6D773455" pageId="0" pageNumber="39" type="diagnosis">
|
||
<paragraph id="138EAE0649706A4A7EF2863B158FC2F8" pageId="0" pageNumber="39">Diagnosis.</paragraph>
|
||
<paragraph id="017F763C748C4CBC95395AA044D595CF" pageId="0" pageNumber="39">
|
||
<taxonomicName id="5ABE70E9239F70262E55CEAA9B3BE88E" lsidName="C. dianthella" pageId="0" pageNumber="39" rank="species" species="dianthella">
|
||
<emphasis id="265522DBF7E64ED4DAD0D9C9A12771F1" italics="true" pageId="0" pageNumber="39">C. dianthella</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="48EC050CBFC851EA3103EEA46200F9D3" lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
|
||
<emphasis id="C3EBADF4871BD3AD84AE7F9AAB007B67" italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
|
||
</taxonomicName>
|
||
by the brown rather than white head, thorax and tegulae but cannot be reliably separated from the other species of the complex by external characters. However, the male genitalia are characterized by the shape of the valva, particularly the convex dorsal margin with a sub-apical hump, the shortly pointed sub-dorsal process and the small digitate ventral process. A similarly short but broader dorso-apical process is only found in
|
||
<taxonomicName id="5193DA54D143F246ACBFFE858B52E4DD" lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
|
||
<emphasis id="3B4FF04C236F1F5A63B3D7186088BFCA" italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
|
||
</taxonomicName>
|
||
, whereas
|
||
<taxonomicName id="5707B693FE0D0EA1FE2A2E585380E102" lsidName="C. habeleri" pageId="0" pageNumber="39" rank="species" species="habeleri">
|
||
<emphasis id="EDB6749C699063AEFA599B38279C3F6E" italics="true" pageId="0" pageNumber="39">C. habeleri</emphasis>
|
||
</taxonomicName>
|
||
with a similar dorso-apical process differs in the large ventro-apical projection and the weakly curved dorsal margin of the valva. In all other species of the
|
||
<taxonomicName id="D0E2746E6FAED42153CD7323A505685A" lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
|
||
<emphasis id="79D623EDB84838E11F56019FB6B5C70C" italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
|
||
</taxonomicName>
|
||
species group, the valva is more slender with a much longer dorso-apical process and a more slender dorso-ventral process. The female genitalia are almolst indistinguishable from other species of the
|
||
<taxonomicName id="C643AA15B90032ADD1F4A6BF50F278E1" lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
|
||
<emphasis id="D0E1D00E8F0CDA5428313A399210F69B" italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
|
||
</taxonomicName>
|
||
species group except for the characteristic signum with a broad hook and a largely reduced base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="507CDF05F985EC5168E11FC765D66138" pageId="0" pageNumber="39" type="description">
|
||
<paragraph id="6D0190F468FBD8EFAB87F8AAC45B0E94" pageId="0" pageNumber="39">Description.</paragraph>
|
||
<paragraph id="2101726B881578AC5EDDF48ECF75752E" pageId="0" pageNumber="39">
|
||
Adult (Fig.
|
||
<figureCitation id="4436BE299DCC216525F498960584378E" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Adults. 3 Caryocolum schleichi, male, Syria; 4 C. dianthella, male, Spain; 5 C. improvisella, male, Austria; 6 C. improvisella, female, Austria; 7 C. arenariella, male, Sweden; 8 C. arenariella, female, Sweden." figureDoi="10.3897/alpento.4.50703.figures3-8" httpUri="https://binary.pensoft.net/fig/403188" pageId="0" pageNumber="39">4</figureCitation>
|
||
). Forewing length ♂ 4.0-5.5 mm, ♀ 4.0-5.0 mm. Head dark brown, frons cream-white to white; labial palpus dark brown, second segment cream-white on inner and upper surface, mottled with grey-brown on outer surface, third segment dark brown mottled with few white scales; antenna black, weakly ringed paler. Thorax and tegula rusty brown, anterior part dark brown. Abdomen predominantly grey on ventral surface, medially cream-coloured. Forewing dark brown with weak light mottling, distinct white markings: transverse fascia from fold to costa at one-fifth, distinct white medial spot, separate costal and tornal spots; fringes basally dark brown, distal part lighter. Hindwing light grey.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61BD977E88164ACD88679973BD4B4607" pageId="0" pageNumber="39" type="variation">
|
||
<paragraph id="EDEAAE5FCA7E5952AFE72FD2BABD569A" pageId="0" pageNumber="39">Variation.</paragraph>
|
||
<paragraph id="2DA27EC1055C96EA964C41F75E0AB899" pageId="0" pageNumber="39">Weak variation in the extent of the white markings.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0991AD1695C41A06CC56D19BE39EC66A" pageId="0" pageNumber="39" type="male genitalia">
|
||
<paragraph id="1D208E40556C612D2A622D6296F36212" pageId="0" pageNumber="39">Male genitalia</paragraph>
|
||
<paragraph id="CDFBBD4F511CF80BCDC13D716D691B62" pageId="0" pageNumber="39">
|
||
(Figs
|
||
<figureCitation id="1E87BDC8D36290EABE7CE3063989D8E9" captionStart="Figures 13–16" captionStartId="F5" captionText="Figures 13 - 16. Male genitalia. 13 Caryocolum schleichi, Syria, slide GU 86 / 273 P. Huemer; 14 C. dianthella, Spain, slide GEL 1284 P. Huemer; 15 C. improvisella, Austria, slide GEL 1256 P. Huemer; 16 C. improvisella, Italy, slide GEL 1290 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures13-16" httpUri="https://binary.pensoft.net/fig/403190" pageId="0" pageNumber="39">14</figureCitation>
|
||
,
|
||
<figureCitation id="3AA40D6B50AE9BD77AACCB158A5B1F1A" captionStart="Figures 21–24" captionStartId="F7" captionText="Figures 21 - 24. Male genitalia (details vinculum-valva-complex). 21 Caryocolum schleichi, Syria, slide GU 86 / 273 P. Huemer; 22 C. dianthella, Spain, slide GEL 1284 P. Huemer; 23 C. improvisella, Austria, slide GEL 1256 P. Huemer; 24 C. improvisella, Italy, slide GEL 120 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures21-24" httpUri="https://binary.pensoft.net/fig/403192" pageId="0" pageNumber="39">22</figureCitation>
|
||
). Uncus broadly sub-quadrangular, posterior corners rounded; lateral sclerites of gnathos distinct, medial part with large minutely spined culcitula; tegumen weakly widened anteriorly, with slightly emarginated anterior margin; transtilla sclerotized, longitudinally folded; pedunculus large, sub-triangular, with sclerotized inner edge; valva nearly straight, moderately long, distal part moderately broad, shovel-shaped, dorsal edge with distinct convex hump, apex with two processes, short pointed sub-dorsal and small digitate ventral process, medially separated by small excavation; sacculus moderately long, knife-shaped; posterior margin of vinculum deeply incised medially, with pair of long digitate processes, pair of latero-medial processes broadly projected; saccus slightly longer than valva, slender, gradually tapered to apex; phallus long, slender, nearly straight, apically with area of small cornuti.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7386B4A199048B046DB0444455BF5B2F" pageId="0" pageNumber="39" type="female genitalia">
|
||
<paragraph id="7C9A590832DFA2160AF1A0D7BFF1E9C3" pageId="0" pageNumber="39">Female genitalia</paragraph>
|
||
<paragraph id="40B211D4EA0E646FDCF36D5AAD99EB3F" pageId="0" pageNumber="39">
|
||
(Figs
|
||
<figureCitation id="93F0C1E96C9074AA5AE64D2051225A6A" captionStart="Figures 29, 30" captionStartId="F9" captionText="Figures 29, 30. Female genitalia. 29 Caryocolum schleichi, Syria, slide GU 86 / 277 P. Huemer; 30 C. dianthella, Spain, slide GEL 1285 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures29-30" httpUri="https://binary.pensoft.net/fig/403194" pageId="0" pageNumber="39">30</figureCitation>
|
||
,
|
||
<figureCitation id="5C2D87F046B3FDD8C0059DEA8F3843DE" captionStart="Figures 36–39" captionStartId="F12" captionText="Figures 36 - 39. Female genitalia (segment VIII). 36 Caryocolum schleichi, Syria, slide GU 86 / 277 P. Huemer; 37 C. dianthella, Spain, slide GEL 1285 P. Huemer; 38 C. improvisella, Italy, slide GEL 1291 P. Huemer; 39 C. arenariella, Sweden, slide GEL 1292 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures36-39" httpUri="https://binary.pensoft.net/fig/403197" pageId="0" pageNumber="39">37</figureCitation>
|
||
,
|
||
<figureCitation id="86D0E8D5AF1A8D3A71551C41E98E6F25" captionStart="Figures 43–46" captionStartId="F14" captionText="Figures 43 - 46. Female genitalia (signum). 43 Caryocolum schleichi, Syria, slide GU 86 / 277 P. Huemer; 44 C. dianthella, Spain, slide GEL 1285 P. Huemer; 45 C. improvisella, Italy, slide GEL 1291 P. Huemer; 46 C. arenariella, Sweden, slide GEL 1292 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures43-46" httpUri="https://binary.pensoft.net/fig/403199" pageId="0" pageNumber="39">44</figureCitation>
|
||
). Apophysis posterior about four times length of apophysis anterior; segment VIII without processes, smooth; ostium bursae with short lateral folds; apophysis anterior 1.3 times length of segment VIII; antrum short, about one-fifth length of apophysis anterior, funnel-shaped; posterior part of ductus bursae with pair of long lateral sclerites extending to about apex of apophysis anterior, membranous part of ductus bursae about length of segment VIII including apophysis anterior; signum on right side of entrance to sub-oval corpus bursae, with broad base lacking crescent-shaped lateral projections, and stout hook.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="485AB5525A4A9972B7B7A2579A6B44B4" pageId="0" pageNumber="39" type="molecular data">
|
||
<paragraph id="AF164F8DFCF9D27BB548C38BAE9A3E22" pageId="0" pageNumber="39">Molecular data.</paragraph>
|
||
<paragraph id="87A8E1F23B38BBE3868276D336A4B82F" pageId="0" pageNumber="39">
|
||
BIN: BOLD:AAU1854. The intraspecific average distance of the barcode region is 1.43%, the maximum distance 1.83% (p-dist) (n = 3). The minimum distance to the nearest neighbor,
|
||
<taxonomicName id="B66E0BB7B109695ED86F1D3EBE6993C6" lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
|
||
<emphasis id="EF854CB4822D010970AB3AA71B8C909A" italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
|
||
</taxonomicName>
|
||
, is 3.05%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2CB6CB090655C8E535682691A4B4BB09" pageId="0" pageNumber="39" type="distribution">
|
||
<paragraph id="8308D372A750AF57247672381CD752B0" pageId="0" pageNumber="39">Distribution</paragraph>
|
||
<paragraph id="E86F2598E015EFEE1041B637DEE1C5AC" pageId="0" pageNumber="39">
|
||
(Fig.
|
||
<figureCitation id="DCFBA8F1DE681193A053C04DFF324009" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of successfully sequenced material of the C. schleichi species group. Map created with SimpleMappr (http: // www. simplemappr. net)." figureDoi="10.3897/alpento.4.50703.figure1" httpUri="https://binary.pensoft.net/fig/403186" pageId="0" pageNumber="39">1</figureCitation>
|
||
).
|
||
<taxonomicName id="8CEC38E39F45217989F93E6E018612D2" lsidName="C. dianthella" pageId="0" pageNumber="39" rank="species" species="dianthella">
|
||
<emphasis id="49B3BF53816FCEFF749651E911B6EC48" italics="true" pageId="0" pageNumber="39">C. dianthella</emphasis>
|
||
</taxonomicName>
|
||
is restricted to the Western Mediterranean, with country records from Morocco, Portugal, Spain and France (Pyrenees).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AE5789D91FC6605B2CA86EC9CA92F545" pageId="0" pageNumber="39" type="bionomics">
|
||
<paragraph id="4DFD4A456E34531B4211A91721792110" pageId="0" pageNumber="39">Bionomics.</paragraph>
|
||
<paragraph id="CEDA128F1151A9C52E6A74D6D0EA82DD" pageId="0" pageNumber="39">
|
||
The larva has been recorded in May and June feeding on
|
||
<taxonomicName id="0B4240611BDE99D2DB5C3955AD037521" authorityName="Tutin" authorityYear="1963" baseAuthorityName="Boiss. & Reuter" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus scaber subsp. toletanus" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="subSpecies" species="scaber" subSpecies="toletanus">
|
||
<emphasis id="BF05A061E9D6DBBBAADAE52AE7E682E0" italics="true" pageId="0" pageNumber="39">Dianthus scaber toletanus</emphasis>
|
||
</taxonomicName>
|
||
(Boiss and Reuter) Tutin and
|
||
<taxonomicName id="02C371DAFD3720ECA495C65C84344E5B" lsidName="D. deltoides" pageId="0" pageNumber="39" rank="species" species="deltoides">
|
||
<emphasis id="EB149FDE933B7E12A88C928323E80D59" italics="true" pageId="0" pageNumber="39">D. deltoides</emphasis>
|
||
</taxonomicName>
|
||
L. and moths emerged in July (
|
||
<bibRefCitation id="68A5BF26882E34A306116BF43BB14EE2" author="Chretien, P" journalOrPublisher="L'Amateur de Papillons" pageId="0" pageNumber="39" pagination="241 - 247" refId="B7" refString="Chretien, P, 1925. La legende de Graellsia isabellae. Appendice. L'Amateur de Papillons 2: 241 - 247" title="La legende de Graellsia isabellae. Appendice." volume="2" year="1925">
|
||
<normalizedToken id="88888F4AE7FF06326510B7849865BC58" originalValue="Chrétien">Chretien</normalizedToken>
|
||
1925
|
||
</bibRefCitation>
|
||
). In the Pyrenees it was also bred from
|
||
<taxonomicName id="6FEAE5C0ED2FC8090F8BCB161B625DD4" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus pungens" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="species" species="pungens">
|
||
<emphasis id="10346F65F8054841B0B23157FAFB563D" italics="true" pageId="0" pageNumber="39">Dianthus pungens</emphasis>
|
||
</taxonomicName>
|
||
L. (see material examined). In the field moths have been collected from late June to October up to altitudes of about 2600 m in Morocco.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="54407677EE58C8E769FCC4A6215A35AA" pageId="0" pageNumber="39" type="remarks">
|
||
<paragraph id="156BACBA546CC1362CD08DE88E34B8C0" pageId="0" pageNumber="39">Remarks.</paragraph>
|
||
<paragraph id="3232F37673C38700CFB3EAD0B9816536" pageId="0" pageNumber="39">
|
||
<taxonomicName id="529B0CD652D92539BEA95CDD9B4AAFDE" authorityName="Chretien" authorityYear="1925" class="Insecta" family="Gelechiidae" genus="Lita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lita dianthella" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="dianthella">
|
||
<emphasis id="C1E4D0F9B1C8081B9D71F7600CE95CF2" italics="true" pageId="0" pageNumber="39">Lita dianthella</emphasis>
|
||
</taxonomicName>
|
||
was described from an unspecified number of specimens from central Spain (
|
||
<bibRefCitation id="126D88D3E922A15B318A984B94BA6CFD" author="Chretien, P" journalOrPublisher="L'Amateur de Papillons" pageId="0" pageNumber="39" pagination="241 - 247" refId="B7" refString="Chretien, P, 1925. La legende de Graellsia isabellae. Appendice. L'Amateur de Papillons 2: 241 - 247" title="La legende de Graellsia isabellae. Appendice." volume="2" year="1925">
|
||
<normalizedToken id="05BC66FD20FB6AC45EC26FF4C9F47022" originalValue="Chrétien">Chretien</normalizedToken>
|
||
1925
|
||
</bibRefCitation>
|
||
) and the genitalia of a syntype figured by
|
||
<bibRefCitation id="0DF746D41896475AAD75AC0C9AF36AE7" author="Agenjo, R" journalOrPublisher="Revista Espanola de Entomologia" pageId="0" pageNumber="39" pagination="147 - 189" refId="B2" refString="Agenjo, R, 1962. Resultados cientificos de una Pension de Estudios en el 'Museum National d'Histoire Naturelle' de Paris, con la descripcion de un genero y otra especie nuevos de lepidopteros espanoles, dedicados al Excmo. Sr. D. Jesus Rubio y Garcia-Mina, Ministro de Educacion Nacional. Eos. Revista Espanola de Entomologia 38: 147 - 189" title="Resultados cientificos de una Pension de Estudios en el ' Museum National d'Histoire Naturelle' de Paris, con la descripcion de un genero y otra especie nuevos de lepidopteros espanoles, dedicados al Excmo. Sr. D. Jesus Rubio y Garcia-Mina, Ministro de Educacion Nacional. Eos." volume="38" year="1962">Agenjo (1962)</bibRefCitation>
|
||
leave no doubt as to the identity.
|
||
<taxonomicName id="E3046F3721D55B2211F9D32F10F1B4E6" authorityName="Derra" authorityYear="1985" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum hackeri" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="hackeri">
|
||
<emphasis id="CD7EB22139AF030791B0CB9989B37E7B" italics="true" pageId="0" pageNumber="39">Caryocolum hackeri</emphasis>
|
||
</taxonomicName>
|
||
was described from northern Spain (Pyrenees) with the holotype fixed in the original description (
|
||
<bibRefCitation id="7DE5B714BD2BD6A4720CF19F2F55F63B" author="Derra, G" journalOrPublisher="Entomofauna, Linz" pageId="0" pageNumber="39" pagination="373 - 379" refId="B9" refString="Derra, G, 1985. Caryocolum hackeri sp. n., eine neue Gelechiidae aus den Pyrenaeen Frankreichs und Spaniens (Lepidoptera, Gelechiidae). Entomofauna, Linz 6: 373 - 379" title="Caryocolum hackeri sp. n., eine neue Gelechiidae aus den Pyrenaeen Frankreichs und Spaniens (Lepidoptera, Gelechiidae)." volume="6" year="1985">Derra 1985</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |