267 lines
35 KiB
XML
267 lines
35 KiB
XML
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<mods:title id="9D17CB05D60EFC2C2ED11E3217A5C510">Systematics and palaeobiology of kangaroos of the late Cenozoic genus Protemnodon (Marsupialia, Macropodidae)</mods:title>
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<mods:namePart id="86BDA1B6284A85A2A3D63C65F8978D2C">Kerr, Isaac A. R.</mods:namePart>
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<mods:affiliation id="23DDE3E70309C9BD7E859E419C336E90">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:namePart id="B9A7A0B32BA9F0D1D1B218017258D41B">Camens, Aaron B.</mods:namePart>
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<mods:affiliation id="990325AEB1FBCAC85227120E7D760273">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:namePart id="437C63BDA4B34E045EFC572EAE22D6B9">Van Zoelen, Jacob D.</mods:namePart>
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<mods:affiliation id="65CF2D8FAB7BFA731CD18B939F838DFA">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:namePart id="76380C1F880BD53E59437431F0C4B6CB">Worthy, Trevor H.</mods:namePart>
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<mods:nameIdentifier id="FCEEF41F4CA9F2EE80409E8D16B7264F" type="ORCID">0000-0001-7047-4680</mods:nameIdentifier>
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<mods:namePart id="6754A91C14428B5DAA70B2575D8D4B19">Prideaux, Gavin J.</mods:namePart>
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<mods:affiliation id="DEA457FA60C40DF4B4D84FBE78A09F46">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:title id="C9C3420052A4F266A2C1647C1FC24288">Megataxa</mods:title>
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<mods:part id="26143DBACE38D709AD7C3669DF6A90FE">
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<mods:date id="C1B08F203BF18E197273EF479FDE8B5E">2024</mods:date>
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<mods:number id="0EA40193CD1AB565E639BEA729D95C71">2024-04-15</mods:number>
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<treatment id="03E587FDFF2CD5F4FCA276CDFD75FBC9" LSID="urn:lsid:plazi:treatment:03E587FDFF2CD5F4FCA276CDFD75FBC9" httpUri="http://treatment.plazi.org/id/03E587FDFF2CD5F4FCA276CDFD75FBC9" lastPageId="193" lastPageNumber="194" pageId="192" pageNumber="193">
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<subSubSection id="C3566560FF2CD5F5FCA276CDFBEAFD49" box="[810,1082,651,677]" pageId="192" pageNumber="193" type="nomenclature">
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<paragraph id="8BF336EBFF2CD5F5FCA276CDFBEAFD49" blockId="192.[810,1452,651,2086]" box="[810,1082,651,677]" pageId="192" pageNumber="193">
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<emphasis id="B938EAF9FF2CD5F5FCA276CDFC02FD49" box="[810,978,651,677]" italics="true" pageId="192" pageNumber="193">
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‘
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<taxonomicName id="4C4C4D68FF2CD5F5FCA676CDFC02FD49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[814,978,651,677]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus" status="n. sp.">Protemnodon</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A20B5782FF2CD5F5FC5176CDFBC8FD49" box="[985,1048,651,677]" pageId="192" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
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A’
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</paragraph>
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</subSubSection>
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<subSubSection id="C3566560FF2CD5F4FCA276EDFD75FBC9" lastPageId="193" lastPageNumber="194" pageId="192" pageNumber="193" type="discussion">
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<paragraph id="8BF336EBFF2CD5F5FCA276EDFA42FB49" blockId="192.[810,1452,651,2086]" pageId="192" pageNumber="193">
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Several specimens were referred to ‘
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<taxonomicName id="4C4C4D68FF2CD5F5FB4576EDFAB4FD29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1229,1380,683,709]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus" status="n. sp.">
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<emphasis id="B938EAF9FF2CD5F5FB4576EDFAB4FD29" box="[1229,1380,683,709]" italics="true" pageId="192" pageNumber="193">Protemnodon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B5782FF2CD5F5FAE676EDFA7BFD29" box="[1390,1451,683,709]" pageId="192" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
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A’ by
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<bibRefCitation id="EFDD4B1AFF2CD5F5FCFA768DFBD4FD0A" author="Piper, K. J." box="[882,1028,715,742]" pageId="192" pageNumber="193" pagination="233 - 253" refId="ref153263" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper (2016)</bibRefCitation>
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, who described but did not formally name a taxon from mostly isolated upper cheek teeth from the early Pliocene Hamilton LF (4.46 ± 0.1 Ma) (
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<bibRefCitation id="EFDD4B1AFF2CD5F5FCB9776DFBC8FCA9" author="Turnbull, W. D. & Lundelius Jr, E. L. & Archer, M." box="[817,1048,811,837]" pageId="192" pageNumber="193" pagination="513 - 540" refId="ref155756" refString="Turnbull, W. D., Lundelius Jr, E. L. & Archer, M. (2003) Chapter 18: dasyurids, perameloids, phalangeroids, and vombatoids from the Early Pliocene Hamilton Fauna, Victoria, Australia. Bulletin of the American Museum of Natural History, 279, 513 - 540. https: // doi. org / 10.1206 / 0003 - 0090 (2003) 279 % 3 C 0513:" type="journal article" year="2003">
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Turnbull
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<emphasis id="B938EAF9FF2CD5F5FC14776AFC05FCA9" box="[924,981,811,837]" italics="true" pageId="192" pageNumber="193">et al.</emphasis>
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2003
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</bibRefCitation>
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), early mid Pleistocene
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<collectingRegion id="4988F809FF2CD5F5FAAF776DFAA6FCA9" box="[1319,1398,811,837]" country="New Zealand" name="Nelson" pageId="192" pageNumber="193">Nelson</collectingRegion>
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Bay LF (0.78–1.77 Ma) (
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||
<bibRefCitation id="EFDD4B1AFF2CD5F5FB81770DFA77FC89" author="Aziz-ur-Rahman, A. & McDougall, I." box="[1033,1447,843,869]" pageId="192" pageNumber="193" pagination="61 - 69" refId="ref142981" refString="Aziz-ur-Rahman, A. & McDougall, I. (1972) Potassium-Argon ages on the newer volcanics of Victoria. Proceedings of the Royal Society of Victoria, 85, 61 - 69." type="journal article" year="1972">Aziz-ur-Rahman & McDougall 1972</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFDD4B1AFF2CD5F5FCA2772DFBEAFC69" author="MacFadden, B. J. & Whitelaw, M. J. & McFadden, P. & Rich, T. H." box="[810,1082,875,901]" pageId="192" pageNumber="193" pagination="364 - 373" refId="ref150809" refString="MacFadden, B. J., Whitelaw, M. J., McFadden, P. & Rich, T. H. (1987) Magnetic polarity stratigraphy of the Pleistocene section at Portland (Victoria), Australia. Quaternary Research, 28 (3), 364 - 373. https: // doi. org / 10.1016 / 0033 - 5894 (87) 90004 - 4" type="journal article" year="1987">
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MacFadden
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||
<emphasis id="B938EAF9FF2CD5F5FC30772AFC24FC69" box="[952,1012,875,901]" italics="true" pageId="192" pageNumber="193">et al.</emphasis>
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||
1987
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||
</bibRefCitation>
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||
), and the early Pleistocene Dog Rocks LF (2.06–2.58 Ma) (
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||
<bibRefCitation id="EFDD4B1AFF2CD5F5FBDE77CDFAD5FC49" author="Whitelaw, M. J." box="[1110,1285,907,933]" pageId="192" pageNumber="193" pagination="417 - 423" refId="ref156853" refString="Whitelaw, M. J. (1991) Magnetic polarity stratigraphy of the Fisherman's Cliff and Bone Gulch vertebrate fossil faunas from the Murray Basin, New South Wales, Australia. Earth and Planetary Science Letters, 104, 417 - 423. https: // doi. org / 10.1016 / 0012 - 821 X (91) 90219 - 8" type="journal article" year="1991">Whitelaw 1991</bibRefCitation>
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||
), southwestern
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<collectingRegion id="4988F809FF2CD5F5FCA277EDFC58FC29" box="[810,904,939,965]" country="Australia" name="Victoria" pageId="192" pageNumber="193">Victoria</collectingRegion>
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||
.
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||
<bibRefCitation id="EFDD4B1AFF2CD5F5FC1877EDFBCBFC2A" author="Piper, K. J." box="[912,1051,939,966]" pageId="192" pageNumber="193" pagination="233 - 253" refId="ref153263" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper (2016)</bibRefCitation>
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included a complete description and diagnosis but refrained from assigning a formal name or
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<typeStatus id="54F78849FF2CD5F5FCA277ADFC8AFBE9" box="[810,858,1003,1029]" pageId="192" pageNumber="193">type</typeStatus>
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specimen due to the need for more complete material and for a taxonomic review of the genus. All specimens listed under
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<taxonomicName id="4C4C4D68FF2CD5F5FC38706DFB97FBA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[944,1095,1067,1093]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus" status="n. sp.">
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<emphasis id="B938EAF9FF2CD5F5FC38706DFB97FBA9" box="[944,1095,1067,1093]" italics="true" pageId="192" pageNumber="193">Protemnodon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B5782FF2CD5F5FBC5706DFB57FBA9" box="[1101,1159,1067,1093]" pageId="192" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
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A are isolated upper cheek teeth with the exception of NMV P216005, an associated DP2, unerupted P3 and M1 from
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<collectingRegion id="4988F809FF2CD5F5FB23702DFB2AFB69" box="[1195,1274,1131,1157]" country="New Zealand" name="Nelson" pageId="192" pageNumber="193">Nelson</collectingRegion>
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Bay and NMV P201862b, an associated M1 and M3 from Dog Rocks.
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</paragraph>
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||
<paragraph id="8BF336EBFF2CD5F5FCD270EDFC36F909" blockId="192.[810,1452,651,2086]" pageId="192" pageNumber="193">
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The following is an extract from the specific diagnosis: ‘…differs from all other known species of
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<taxonomicName id="4C4C4D68FF2CD5F5FA85708DFA74FB09" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1293,1444,1227,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus">
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||
<emphasis id="B938EAF9FF2CD5F5FA85708DFA74FB09" box="[1293,1444,1227,1253]" italics="true" pageId="192" pageNumber="193">Protemnodon</emphasis>
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</taxonomicName>
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, except
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<taxonomicName id="4C4C4D68FF2CD5F5FCF370AAFBD6FAE9" authorityName="Plane" authorityYear="1967" box="[891,1030,1259,1285]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="species" species="otibandus">
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||
<emphasis id="B938EAF9FF2CD5F5FCF370AAFBD6FAE9" box="[891,1030,1259,1285]" italics="true" pageId="192" pageNumber="193">P. otibandus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4C4D68FF2CD5F5FBC970AAFAD6FAE9" authorityName="Bartholomai" authorityYear="1973" box="[1089,1286,1259,1285]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="species" species="chinchillaensis">
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<emphasis id="B938EAF9FF2CD5F5FBC970AAFAD6FAE9" box="[1089,1286,1259,1285]" italics="true" pageId="192" pageNumber="193">P. chinchillaensis</emphasis>
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</taxonomicName>
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, in possessing a P3 that is relatively more elongate compared to upper molars (see table 2), and from all species in having upper molars that are more rounded in occlusal outline (
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<emphasis id="B938EAF9FF2CD5F5FA01710AFA7CFA89" box="[1417,1452,1356,1381]" italics="true" pageId="192" pageNumber="193">i.e.</emphasis>
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unconstricted across the [interloph] valley—base of lophs expanded lingually and swollen in the [buccal] moiety of the [interloph] valley, forming a convex [buccal] margin in occlusal view). Differs from all species except
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<taxonomicName id="4C4C4D68FF2CD5F5FCA271AAFC69F9E9" authorityName="Plane" authorityYear="1967" box="[810,953,1515,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="species" species="otibandus">
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<emphasis id="B938EAF9FF2CD5F5FCA271AAFC69F9E9" box="[810,953,1515,1541]" italics="true" pageId="192" pageNumber="193">P. otibandus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4C4D68FF2CD5F5FC7371AAFBAEF9E9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1019,1150,1515,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="species" species="tumbuna">
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<emphasis id="B938EAF9FF2CD5F5FC7371AAFBAEF9E9" box="[1019,1150,1515,1541]" italics="true" pageId="192" pageNumber="193">P. tumbuna</emphasis>
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</taxonomicName>
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in having gently sloping lingual loph margins and a variably developed postlink on anterior molars in some individuals’ (
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<bibRefCitation id="EFDD4B1AFF2CD5F5FA8D726DFA58F9A9" author="Piper, K. J." box="[1285,1416,1579,1605]" pageId="192" pageNumber="193" pagination="233 - 253" refId="ref153263" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper 2016</bibRefCitation>
|
||
, p. 243). Our morphological comparisons suggest that the dental material allocated by
|
||
<bibRefCitation id="EFDD4B1AFF2CD5F5FBED722DFB21F96A" author="Piper, K. J." box="[1125,1265,1643,1670]" pageId="192" pageNumber="193" pagination="233 - 253" refId="ref153263" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper (2016)</bibRefCitation>
|
||
to
|
||
<taxonomicName id="4C4C4D68FF2CD5F5FA9C722DFA7BF969" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1300,1451,1643,1669]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus" status="n. sp.">
|
||
<emphasis id="B938EAF9FF2CD5F5FA9C722DFA7BF969" box="[1300,1451,1643,1669]" italics="true" pageId="192" pageNumber="193">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B5782FF2CD5F5FCA272CDFCBAF949" box="[810,874,1675,1701]" pageId="192" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
|
||
A most likely represents two or three species of
|
||
<taxonomicName id="4C4C4D68FF2CD5F5FCA272EDFC11F929" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[810,961,1707,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF2CD5F5FCA272EDFC11F929" box="[810,961,1707,1733]" italics="true" pageId="192" pageNumber="193">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
from the Pliocene and Pleistocene, as discussed below.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF2CD5F4FCD272ADFD65FEA9" blockId="192.[810,1452,651,2086]" lastBlockId="193.[136,778,171,1061]" lastPageId="193" lastPageNumber="194" pageId="192" pageNumber="193">
|
||
<bibRefCitation id="EFDD4B1AFF2CD5F5FCD272ADFC24F8EA" author="Piper, K. J." box="[858,1012,1771,1798]" pageId="192" pageNumber="193" pagination="233 - 253" refId="ref153263" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper (2016)</bibRefCitation>
|
||
identified the mean ratio of M1 length to P3 length in
|
||
<taxonomicName id="4C4C4D68FF2CD5F5FBA7734DFB16F8C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1071,1222,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus" status="n. sp.">
|
||
<emphasis id="B938EAF9FF2CD5F5FBA7734DFB16F8C9" box="[1071,1222,1803,1829]" italics="true" pageId="192" pageNumber="193">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B5782FF2CD5F5FB58734DFADFF8C9" box="[1232,1295,1803,1829]" pageId="192" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
|
||
A as 0.54, in contrast to the more similar lengths in all taxa except
|
||
<taxonomicName id="4C4C4D68FF2CD5F5FCA2730AFC67F889" authorityName="Plane" authorityYear="1967" box="[810,951,1867,1893]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF2CD5F5FCA2730AFC67F889" box="[810,951,1867,1893]" italics="true" pageId="192" pageNumber="193">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF2CD5F5FC7E730AFB6EF889" authorityName="Bartholomai" authorityYear="1973" box="[1014,1214,1867,1893]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="species" species="chinchillaensis">
|
||
<emphasis id="B938EAF9FF2CD5F5FC7E730AFB6EF889" box="[1014,1214,1867,1893]" italics="true" pageId="192" pageNumber="193">P. chinchillaensis</emphasis>
|
||
</taxonomicName>
|
||
. However, the teeth identified as M1s have narrow protoloph bases relative to the metalophs, very narrow anterior cingula, anterior moieties rotated distinctly toward the lingual side and overall anteroposterior constriction, consistent with the morphology of the DP
|
||
<quantity id="4CB49B0EFF2CD5F5FBD473ADFB57F7E9" box="[1116,1159,2027,2053]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="192" pageNumber="193" unit="in" value="3.0">3 in</quantity>
|
||
species of
|
||
<taxonomicName id="4C4C4D68FF2CD5F5FA8573ADFA74F7E9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1293,1444,2027,2053]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF2CD5F5FA8573ADFA74F7E9" box="[1293,1444,2027,2053]" italics="true" pageId="192" pageNumber="193">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
, particularly that of
|
||
<taxonomicName id="4C4C4D68FF2CD5F5FC767C4AFB8BF7C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1022,1115,2060,2085]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="192" pageNumber="193" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF2CD5F5FC767C4AFB8BF7C9" box="[1022,1115,2060,2085]" italics="true" pageId="192" pageNumber="193">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF2CD5F5FBE97C4DFB6AF7C9" bold="true" box="[1121,1210,2059,2085]" pageId="192" pageNumber="193">
|
||
<taxonomicNameLabel id="A20B5782FF2CD5F5FBE97C4DFB6AF7C9" box="[1121,1210,2059,2085]" pageId="192" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="B938EAF9FF2CD5F5FB417C4AFB21F7C9" box="[1225,1265,2060,2085]" italics="true" pageId="192" pageNumber="193">e.g.</emphasis>
|
||
SAMA P53836) and
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FF3D74EAFE92FF29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[181,322,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF2DD5F4FF3D74EAFE92FF29" box="[181,322,171,197]" italics="true" pageId="193" pageNumber="194">P.mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF2DD5F4FECE74EDFE4DFF29" bold="true" box="[326,413,171,197]" pageId="193" pageNumber="194">
|
||
<taxonomicNameLabel id="A20B5782FF2DD5F4FECE74EDFE4DFF29" box="[326,413,171,197]" pageId="193" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="B938EAF9FF2DD5F4FE2374EAFE03FF29" box="[427,467,172,197]" italics="true" pageId="193" pageNumber="194">e.g.</emphasis>
|
||
SAMAP21863).Therefore, the apparent great difference in relative size between the P3 and M
|
||
<quantity id="4CB49B0EFF2DD5F4FEA174ADFE82FEE9" box="[297,338,235,261]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="193" pageNumber="194" unit="in" value="1.0">1 in</quantity>
|
||
this taxon is a result of the accidental comparison of the dimensions of the P3 with those of the DP3 rather than the larger and more elongate M1.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF2DD5F4FF30750DFD75FBC9" blockId="193.[136,778,171,1061]" pageId="193" pageNumber="194">
|
||
Two DP2s from
|
||
<collectingRegion id="4988F809FF2DD5F4FE11750DFE38FE89" box="[409,488,331,357]" country="New Zealand" name="Nelson" pageId="193" pageNumber="194">Nelson</collectingRegion>
|
||
Bay LF with distinct morphologies were allocated to this taxon; one was included due to its association with diagnostic material, the other by morphological association (
|
||
<bibRefCitation id="EFDD4B1AFF2DD5F4FE3175EDFDE6FE29" author="Piper, K. J." box="[441,566,427,453]" pageId="193" pageNumber="194" pagination="233 - 253" refId="ref153263" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper 2016</bibRefCitation>
|
||
, fig. 8, a–c and d–f respectively). The former of the DP2s, NMV P216005, is associated with an unerupted P3 and what was listed as an M3 (
|
||
<bibRefCitation id="EFDD4B1AFF2DD5F4FE89764DFE50FDC9" author="Piper, K. J." box="[257,384,523,549]" pageId="193" pageNumber="194" pagination="233 - 253" refId="ref153263" refString="Piper, K. J. (2016) The Macropodidae (Marsupialia) of the early Pleistocene Nelson Bay Local Fauna, Victoria, Australia. Memoirs of Museum Victoria, 74, 233 - 253. https: // doi. org / 10.24199 / j. mmv. 2016.74.18" type="journal article" year="2016">Piper 2016</bibRefCitation>
|
||
, fig.
|
||
<quantity id="4CB49B0EFF2DD5F4FE33764DFE0FFDC9" box="[443,479,523,549]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="193" pageNumber="194" unit="m" value="8.0">8m</quantity>
|
||
–o, fig. 9p–r respectively). This DP2, which is broad and rounded in occlusal view, falls within the morphological definitions of
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FDF3760AFCD9FD89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[635,777,587,613]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF2DD5F4FDF3760AFCD9FD89" box="[635,777,587,613]" italics="true" pageId="193" pageNumber="194">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF2DD5F4FF00762DFF33FD69" bold="true" box="[136,227,619,645]" pageId="193" pageNumber="194">
|
||
<taxonomicNameLabel id="A20B5782FF2DD5F4FF00762DFF33FD69" box="[136,227,619,645]" pageId="193" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FE94762AFEACFD69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[284,380,620,645]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF2DD5F4FE94762AFEACFD69" box="[284,380,620,645]" italics="true" pageId="193" pageNumber="194">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF2DD5F4FE0D762DFE0FFD69" bold="true" box="[389,479,619,645]" pageId="193" pageNumber="194">
|
||
<taxonomicNameLabel id="A20B5782FF2DD5F4FE0D762DFE0FFD69" box="[389,479,619,645]" pageId="193" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Based on its relative loph widths and the slight rotation of its anterior moiety towards the lingual side, the ‘M3’ of this specimen is in fact an M1. The associated unerupted P3 and the DP2 are within the morphological bounds of both
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FE6D76AAFDA5FCE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[485,629,747,773]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF2DD5F4FE6D76AAFDA5FCE9" box="[485,629,747,773]" italics="true" pageId="193" pageNumber="194">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF2DD5F4FDF676ADFD09FCE9" bold="true" box="[638,729,747,773]" pageId="193" pageNumber="194">
|
||
<taxonomicNameLabel id="A20B5782FF2DD5F4FDF676ADFD09FCE9" box="[638,729,747,773]" pageId="193" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FF00774AFF36FCC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,230,780,805]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF2DD5F4FF00774AFF36FCC9" box="[136,230,780,805]" italics="true" pageId="193" pageNumber="194">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF2DD5F4FF66774DFE98FCC9" bold="true" box="[238,328,779,805]" pageId="193" pageNumber="194">
|
||
<taxonomicNameLabel id="A20B5782FF2DD5F4FF66774DFE98FCC9" box="[238,328,779,805]" pageId="193" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The second DP2 is attributable to
|
||
<emphasis id="B938EAF9FF2DD5F4FD46774AFD33FCC9" box="[718,739,780,805]" italics="true" pageId="193" pageNumber="194">P.</emphasis>
|
||
sp. cf.
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FF25776AFF2FFCA9" authorityName=", NHMUK PVM" authorityYear="1895" box="[173,255,811,837]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF2DD5F4FF25776AFF2FFCA9" box="[173,255,811,837]" italics="true" pageId="193" pageNumber="194">P. anak</emphasis>
|
||
</taxonomicName>
|
||
due to its small size, narrow posterior relative to total length, and anteroposteriorly short lingual valley. Thus, although the material used to describe this species is definitely of the genus
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FE5B77CDFDB9FC49" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[467,617,907,933]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF2DD5F4FE5B77CDFDB9FC49" box="[467,617,907,933]" italics="true" pageId="193" pageNumber="194">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
, the isolated nature of all but five of the teeth has been a factor in its misidentification as a new species. We conclude that none of the material listed under ‘
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FE3777ADFD85FBE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[447,597,1003,1029]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="genus" status="n. sp.">
|
||
<emphasis id="B938EAF9FF2DD5F4FE3777ADFD85FBE9" box="[447,597,1003,1029]" italics="true" pageId="193" pageNumber="194">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B5782FF2DD5F4FDD377ADFD46FBE9" box="[603,662,1003,1029]" pageId="193" pageNumber="194" rank="species">n. sp.</taxonomicNameLabel>
|
||
A’ reflects a hitherto unrecognised species of
|
||
<taxonomicName id="4C4C4D68FF2DD5F4FD80704DFD4EFBC9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[520,670,1035,1061]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="193" pageNumber="194" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF2DD5F4FD80704DFD4EFBC9" box="[520,670,1035,1061]" italics="true" pageId="193" pageNumber="194">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |