599 lines
80 KiB
XML
599 lines
80 KiB
XML
<document id="7A41C40C69E7F48A476C30AD268645E0" ID-CLB-Dataset="8907" ID-DOI="10.11646/zootaxa.4819.3.1" ID-GBIF-Dataset="c4ab39b0-7ded-4e6e-ae61-496887fb12b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4397003" ID-ZooBank="406EA604-84B1-4ABC-8416-95ED0AA93C46" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609203341047" checkinUser="plazi" docAuthor="Bonomo, Lynn J. & Gosliner, Terrence M." docDate="2020" docId="03CFFA0FFFFAFF9699D0FF663C50AD38" docLanguage="en" docName="zootaxa.4819.3.1.pdf" docOrigin="Zootaxa 4819 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chromodoris kalawakan Bonomo & Gosliner 2020, sp. nov." docType="treatment" docUuid="F18F8F3B-E46E-426D-BD29-9C65185B9B16" docUuidSource="ZooBank" docVersion="8" lastPageNumber="423" masterDocId="FFF68277FFEEFF819947FFF1396EAC25" masterDocTitle="Adding stars to the Chromodoris (Nudibranchia, Chromodorididae) galaxy with the description of four new species" masterLastPageNumber="435" masterPageNumber="401" pageNumber="421" updateTime="1698848172372" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="6FE118048FB35D529EC0DCFCFD277012">
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<mods:title id="69EA026138CC6646B162D2153482C79F">Adding stars to the Chromodoris (Nudibranchia, Chromodorididae) galaxy with the description of four new species</mods:title>
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<mods:name id="002A5A896529E1D9411782E03A3AAE64" type="personal">
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<mods:roleTerm id="C519EEF015819AA64DA942670E8B83B3">Author</mods:roleTerm>
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<mods:namePart id="ECE7428A25DC784E9C1349362BF6BDD9">Bonomo, Lynn J.</mods:namePart>
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<mods:nameIdentifier id="56883FE019A31B9430051AF98DFB5080" type="ORCID">0000-0002-3950-0243</mods:nameIdentifier>
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<mods:affiliation id="D83673FE728CF76E7870E32BD6344ECF">Department of Invertebrate Zoology and Geology, California Academy of Sciences, San Francisco, CA, 94118, USA tgosliner @ calacademy. org, https: // orcid. org / 0000 - 0002 - 3950 - 0243 & Department of Biology, San Francisco State University, San Francisco, CA, 94132, USA.</mods:affiliation>
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<mods:roleTerm id="C8AF55B2D33DAC6E08B3475D73A4BB95">Author</mods:roleTerm>
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<mods:namePart id="D9FA923C71DA93BBAC4695237F9550CC">Gosliner, Terrence M.</mods:namePart>
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<mods:nameIdentifier id="0D9A6479B72E60F98EB576927E949D7A" type="ORCID">0000-0002-3950-0243</mods:nameIdentifier>
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<mods:affiliation id="93FA6756FE4B4B6D6ABEEA6062A5F382">Department of Invertebrate Zoology and Geology, California Academy of Sciences, San Francisco, CA, 94118, USA tgosliner @ calacademy. org, https: // orcid. org / 0000 - 0002 - 3950 - 0243</mods:affiliation>
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<mods:typeOfResource id="106AE2494CFEF1AAFA8C53AF07F21C0F">text</mods:typeOfResource>
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<mods:title id="B4351A12CBA5C05CF811E3AE58746CDD">Zootaxa</mods:title>
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<mods:part id="36448C169365D76ECACD7A69A080AADA">
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<mods:date id="F3B833D6A35EC69660BD8B24AE01828B">2020</mods:date>
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<mods:number id="DACEF56C095B988726E646DF0A865847">2020-07-24</mods:number>
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<mods:detail id="5ED13E361BC0A613EC6F4F86BEBFBDD0" type="volume">
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<mods:number id="7F0FBAF251461A442F98C7E1DF75B588">4819</mods:number>
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<mods:detail id="EB01BB82DA3FA8193E0ACE4A674DE848" type="issue">
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<mods:number id="549EA553E80070166EDC3BFFD313A0A3">3</mods:number>
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<treatment id="03CFFA0FFFFAFF9699D0FF663C50AD38" ID-DOI="http://doi.org/10.5281/zenodo.4438816" ID-GBIF-Taxon="176580875" ID-Zenodo-Dep="4438816" LSID="urn:lsid:zoobank.org:act:F18F8F3B-E46E-426D-BD29-9C65185B9B16" httpUri="http://treatment.plazi.org/id/03CFFA0FFFFAFF9699D0FF663C50AD38" lastPageId="23" lastPageNumber="423" pageId="20" pageNumber="421">
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<subSubSection id="C37C1892FFFAFF9599D0FF6638D5ACF0" pageId="20" pageNumber="421" type="nomenclature">
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<paragraph id="8BD94B19FFFAFF9599D0FF663A47AC94" blockId="20.[151,963,151,250]" box="[151,809,151,178]" pageId="20" pageNumber="421">
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<heading id="D091FC75FFFAFF9599D0FF663A47AC94" bold="true" box="[151,809,151,178]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
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<emphasis id="B912970BFFFAFF9599D0FF663A47AC94" bold="true" box="[151,809,151,178]" pageId="20" pageNumber="421">
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<taxonomicName id="4C66309AFFFAFF9599D0FF663BAAAC94" ID-CoL="7QP37" authority="Bonomo & Gosliner" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[151,708,151,178]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="421" phylum="Mollusca" rank="species" species="kalawakan" status="sp. nov.">
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<emphasis id="B912970BFFFAFF9599D0FF6638ACAC97" bold="true" box="[151,450,151,178]" italics="true" pageId="20" pageNumber="421">Chromodoris kalawakan</emphasis>
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Bonomo & Gosliner
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</taxonomicName>
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<taxonomicNameLabel id="A2212A70FFFAFF959B8CFF663A47AC94" box="[715,809,151,177]" pageId="20" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD94B19FFFAFF9599D0FF4A38D5ACF0" blockId="20.[151,963,151,250]" box="[151,443,187,214]" pageId="20" pageNumber="421">Figures (2C, 2D, 2E, 7, 8)</paragraph>
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</subSubSection>
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<subSubSection id="C37C1892FFFAFF9599D0FF2E3AA0AD19" pageId="20" pageNumber="421" type="description">
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<paragraph id="8BD94B19FFFAFF9599D0FF2E3AADACDF" blockId="20.[151,963,151,250]" box="[151,963,223,250]" pageId="20" pageNumber="421">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BD94B19FFFAFF9599D0FED53AA0AD19" blockId="20.[151,974,292,377]" box="[151,974,292,316]" pageId="20" pageNumber="421">
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<taxonomicName id="4C66309AFFFAFF9599D0FED53873AD19" authorityName="Alder & Hancock" authorityYear="1855" box="[151,285,292,316]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="421" phylum="Mollusca" rank="genus">
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<emphasis id="B912970BFFFAFF9599D0FED53873AD19" box="[151,285,292,316]" italics="true" pageId="20" pageNumber="421">Chromodoris</emphasis>
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</taxonomicName>
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sp. 23—Gosliner
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<emphasis id="B912970BFFFAFF959890FED53B64AD19" box="[471,522,292,316]" italics="true" pageId="20" pageNumber="421">et al.</emphasis>
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2008: 229, second from the top photograph.
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</paragraph>
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<subSubSection id="C37C1892FFFAFF9599D0FEB33A19AD5C" pageId="20" pageNumber="421" type="reference_group">
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<paragraph id="8BD94B19FFFAFF9599D0FEB33AFDAD7F" blockId="20.[151,974,292,377]" box="[151,915,322,346]" pageId="20" pageNumber="421">
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<treatmentCitationGroup id="AB766C37FFFAFF9599D0FEB33AFDAD7F" box="[151,915,322,346]" pageId="20" pageNumber="421">
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<taxonomicName id="4C66309AFFFAFF9599D0FEB3385BAD7F" ID-CoL="7P4GC" box="[151,309,322,346]" class="Gastropoda" family="Chromodorididae" genus="Goniobranchus" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="421" phylum="Mollusca" rank="genus">
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<emphasis id="B912970BFFFAFF9599D0FEB3385BAD7F" box="[151,309,322,346]" italics="true" pageId="20" pageNumber="421">Goniobranchus</emphasis>
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</taxonomicName>
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sp. 40—
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<treatmentCitation id="0AC76D08FFFAFF9598D6FEB33BE7AD7F" author="Gosliner, T. & Behrens, D. & Valdes, A." box="[401,649,322,346]" page="230" pageId="20" pageNumber="421" year="2015">
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<bibRefCitation id="EFF736E8FFFAFF9598D6FEB33BE7AD7F" author="Gosliner, T. & Behrens, D. & Valdes, A." box="[401,649,322,346]" pageId="20" pageNumber="421" refId="ref15951" refString="Gosliner, T., Behrens, D. & Valdes, A. (2015) Indo-Pacific nudibranchs and sea slugs: A field guide to the world's most diverse fauna. New World Publications, Inc., Jacksonville, Florida, 426 pp." type="book" year="2015">
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Gosliner
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<emphasis id="B912970BFFFAFF9598A8FEB23B75AD7F" box="[495,539,322,346]" italics="true" pageId="20" pageNumber="421">et al</emphasis>
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2015: 230
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</bibRefCitation>
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</treatmentCitation>
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, bottom right photograph.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD94B19FFFAFF9599D0FE903A19AD5C" blockId="20.[151,974,292,377]" box="[151,887,353,377]" pageId="20" pageNumber="421">
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<treatmentCitationGroup id="AB766C37FFFAFF9599D0FE903A19AD5C" box="[151,887,353,377]" pageId="20" pageNumber="421">
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<taxonomicName id="4C66309AFFFAFF9599D0FE90385BAD5C" ID-CoL="7P4GC" box="[151,309,353,377]" class="Gastropoda" family="Chromodorididae" genus="Goniobranchus" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="421" phylum="Mollusca" rank="genus">
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<emphasis id="B912970BFFFAFF9599D0FE90385BAD5C" box="[151,309,353,377]" italics="true" pageId="20" pageNumber="421">Goniobranchus</emphasis>
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</taxonomicName>
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sp. 43—
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<treatmentCitation id="0AC76D08FFFAFF9598D6FE903BE7AD5C" author="Gosliner, T. & Valdeiz, A. & Behrens, D. W." box="[401,649,353,377]" page="160" pageId="20" pageNumber="421" year="2018">
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<bibRefCitation id="EFF736E8FFFAFF9598D6FE903BE7AD5C" author="Gosliner, T. & Valdeiz, A. & Behrens, D. W." box="[401,649,353,377]" pageId="20" pageNumber="421" refId="ref15998" refString="Gosliner, T., Valdeiz, A. & Behrens, D. W. (2018) Indo-Pacific nudibranchs and sea slugs: A field guide to the world's most diverse fauna. New World Publications, Inc., Jacksonville, Florida, 451 pp. Available from: https: // www. kogan. com / au / buy / nudibranch-and-sea-slug-identification-indo-pacific-terrence-gosliner- 9781878348678 / (accessed 19 July 2019)" type="book" year="2018">
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Gosliner
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<emphasis id="B912970BFFFAFF9598A8FE933B75AD5C" box="[495,539,353,377]" italics="true" pageId="20" pageNumber="421">et al</emphasis>
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2018: 160
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</bibRefCitation>
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</treatmentCitation>
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, upper left photograph.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C37C1892FFFAFF9599D0FE563B35ABE8" pageId="20" pageNumber="421" type="materials_examined">
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<emphasis id="B912970BFFFAFF9599D0FE563826ADE4" bold="true" box="[151,328,423,449]" pageId="20" pageNumber="421">Type material.</emphasis>
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<materialsCitation id="3B0E4144FFFAFF959815FE563B43ADC0" ID-GBIF-Occurrence="3015405301" collectingDate="1996-09-21" collectionCode="T" collectorName="M. Gosliner" country="Palau" elevation="24" location="Uchelbeluu Reef" pageId="20" pageNumber="421" specimenCode="CASIZ 109740" specimenCount="1" typeStatus="holotype">
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<emphasis id="B912970BFFFAFF959815FE5638A6ADE4" bold="true" box="[338,456,423,449]" pageId="20" pageNumber="421">
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<typeStatus id="54DDF5BBFFFAFF959815FE5638ADADE4" box="[338,451,423,449]" pageId="20" pageNumber="421" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCode id="DBC0E362FFFAFF959895FE563BEAADE4" box="[466,644,423,449]" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 109740</specimenCode>
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, one specimen.
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<location id="8EB91DC2FFFAFF959A78FE563D6CADE4" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959A78FE563D6CADE4" box="[831,1026,423,449]" country="Palau" name="Uchelbeluu Reef" pageId="20" pageNumber="421">Uchelbeluu Reef</location>
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,
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,
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depth
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</elevation>
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,
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<quantity id="4C9EE6FCFFFAFF959DB3FE563C28ADE7" box="[1268,1350,423,450]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="421" unit="mm" value="10.0">10 mm</quantity>
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length,
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<collectingDate id="EF9C9431FFFAFF9599D0FE3A3815ADC0" box="[151,379,459,485]" pageId="20" pageNumber="421" value="1996-09-21">21 September 1996</collectingDate>
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</date>
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,
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<collectionCode id="ED77D3DCFFFAFF9598CEFE3D38F2ADC3" box="[393,412,460,486]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="421">T</collectionCode>
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.
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<collectorName id="26932ECFFFFAFF9598E7FE3D3B43ADC0" box="[416,557,459,486]" pageId="20" pageNumber="421">M. Gosliner</collectorName>
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</materialsCitation>
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.
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<materialsCitation id="3B0E4144FFFAFF959B7DFE3A3A79AE2C" ID-GBIF-Occurrence="3015405315" collectingDate="2006-09-10" collectionCode="CASIZ" collectorName="NW Tutuba Island" country="Vanuatu" latitude="-15.55" location="Tutuba Island" longLatPrecision="7" longitude="167.28" pageId="20" pageNumber="421" specimenCode="CASIZ 178286" specimenCount="1" typeStatus="paratype">
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<emphasis id="B912970BFFFAFF959B7DFE3A3BD5ADC0" bold="true" box="[570,699,459,485]" pageId="20" pageNumber="421">
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<typeStatus id="54DDF5BBFFFAFF959B7DFE3A3BD8ADC0" box="[570,694,459,485]" pageId="20" pageNumber="421" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCode id="DBC0E362FFFAFF959B81FE3A3A16ADC0" box="[710,888,459,485]" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 178286</specimenCode>
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, one specimen.
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<collectorName id="26932ECFFFFAFF959D72FE3D3C63ADC0" box="[1077,1293,459,486]" pageId="20" pageNumber="421">NW Tutuba Island</collectorName>
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,
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,
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<location id="8EB91DC2FFFAFF959868FE1E38A4AE2C" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959868FE1E38A4AE2C" box="[303,458,495,521]" country="Vanuatu" latitude="-15.55" longLatPrecision="7" longitude="167.28" name="Tutuba Island" pageId="20" pageNumber="421">Tutuba Island</location>
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,
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<collectingCountry id="F3710B89FFFAFF959893FE013B5DAE2F" box="[468,563,496,522]" name="Vanuatu" pageId="20" pageNumber="421">Vanuatu</collectingCountry>
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,
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<date id="FFD86DD9FFFAFF959B7AFE1E3A79AE2C" box="[573,791,495,521]" pageId="20" pageNumber="421" value="2006-09-10">
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<collectingDate id="EF9C9431FFFAFF959B7AFE1E3A79AE2C" box="[573,791,495,521]" pageId="20" pageNumber="421" value="2006-09-10">10 September 2006</collectingDate>
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,
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<materialsCitation id="3B0E4144FFFAFF959A66FE013D8FAE2C" ID-GBIF-Occurrence="3015405311" box="[801,1249,495,522]" collectorName="M. Pola-Perez" country="Vanuatu" location="Camacho-Garcia" pageId="20" pageNumber="421" specimenCount="1" typeStatus="paratype">
|
||
<collectorName id="26932ECFFFFAFF959A66FE013AADAE2C" box="[801,963,495,522]" pageId="20" pageNumber="421">M. Pola-Perez</collectorName>
|
||
, Y.
|
||
<location id="8EB91DC2FFFAFF959AADFE1E3DC6AE2C" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959AADFE1E3DC6AE2C" box="[1002,1192,495,521]" country="Vanuatu" name="Camacho-Garcia" pageId="20" pageNumber="421">Camacho-Garcia</location>
|
||
<emphasis id="B912970BFFFAFF959DEAFE013D8FAE2C" box="[1197,1249,495,521]" italics="true" pageId="20" pageNumber="421">et al</emphasis>
|
||
</materialsCitation>
|
||
<emphasis id="B912970BFFFAFF959DA6FE1E3D8BAE2C" box="[1249,1253,495,521]" italics="true" pageId="20" pageNumber="421">.</emphasis>
|
||
<materialsCitation id="3B0E4144FFFAFF959DADFE1E39BCAE74" ID-GBIF-Occurrence="3015405304" collectingDate="2006-09-28" collectionCode="CASIZ" collectorName="SE Matewulu" country="Vanuatu" latitude="-15.4983" location="Espiritu Santo Island" longLatPrecision="7" longitude="167.1983" pageId="20" pageNumber="421" specimenCode="CASIZ 178300" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBC0E362FFFAFF959DADFE1E3CF6AE2C" box="[1258,1432,495,521]" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 178300</specimenCode>
|
||
, one specimen, dissected,
|
||
<collectorName id="26932ECFFFFAFF9598FCFDE23B34AE08" box="[443,602,531,557]" pageId="20" pageNumber="421">SE Matewulu</collectorName>
|
||
,
|
||
<geoCoordinate id="EE522DDEFFFAFF959B2FFDE23B86AE08" box="[616,744,531,558]" degrees="15.4983" direction="south" orientation="latitude" pageId="20" pageNumber="421" precision="5" value="-15.4983">15.4983° S</geoCoordinate>
|
||
<geoCoordinate id="EE522DDEFFFAFF959BB5FDE23AEDAE0B" box="[754,899,531,558]" degrees="167.1983" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="5" value="167.1983">167.1983° E</geoCoordinate>
|
||
,
|
||
<location id="8EB91DC2FFFAFF959AD6FDE23DEAAE08" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959AD6FDE23DEAAE08" box="[913,1156,531,557]" country="Vanuatu" latitude="-15.4983" longLatPrecision="7" longitude="167.1983" name="Espiritu Santo Island" pageId="20" pageNumber="421">Espiritu Santo Island</location>
|
||
,
|
||
<collectingCountry id="F3710B89FFFAFF959DD5FDE53D9FAE0B" box="[1170,1265,532,558]" name="Vanuatu" pageId="20" pageNumber="421">Vanuatu</collectingCountry>
|
||
,
|
||
<date id="FFD86DD9FFFAFF959DB8FDE239BCAE74" pageId="20" pageNumber="421" value="2006-09-28">
|
||
<collectingDate id="EF9C9431FFFAFF959DB8FDE239BCAE74" pageId="20" pageNumber="421" value="2006-09-28">28 September 2006</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B0E4144FFFAFF9599A7FDC93BDDAE74" ID-GBIF-Occurrence="3015405319" box="[224,691,567,594]" collectorName="M. Pola-Perez" country="Vanuatu" location="Camacho-Garcia" pageId="20" pageNumber="421" specimenCount="1" typeStatus="paratype">
|
||
<collectorName id="26932ECFFFFAFF9599A7FDC938E8AE74" box="[224,390,567,594]" pageId="20" pageNumber="421">M. Pola-Perez</collectorName>
|
||
, Y.
|
||
<location id="8EB91DC2FFFAFF9598F3FDC63B1CAE74" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF9598F3FDC63B1CAE74" box="[436,626,567,593]" country="Vanuatu" name="Camacho-Garcia" pageId="20" pageNumber="421">Camacho-Garcia</location>
|
||
<emphasis id="B912970BFFFAFF959B3BFDC93BDDAE74" box="[636,691,567,593]" italics="true" pageId="20" pageNumber="421">et al</emphasis>
|
||
</materialsCitation>
|
||
<emphasis id="B912970BFFFAFF959BF4FDC63BD9AE74" box="[691,695,567,593]" italics="true" pageId="20" pageNumber="421">.</emphasis>
|
||
<materialsCitation id="3B0E4144FFFAFF959B86FDC63A5BAEBC" ID-GBIF-Occurrence="3015405320" collectingDate="2017-04-03" collectionCode="CASIZ" collectorName="South" country="Philippines" county="Island" latitude="12.58355" location="Lugbung Island" longLatPrecision="1" longitude="122.2438" municipality="Lugbung Island" pageId="20" pageNumber="421" specimenCode="CASIZ 222016" specimenCount="1" stateProvince="Romblon" typeStatus="paratype">
|
||
<specimenCode id="DBC0E362FFFAFF959B86FDC63A1DAE74" box="[705,883,567,593]" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 222016</specimenCode>
|
||
, one specimen, subsampled.
|
||
<collectorName id="26932ECFFFFAFF959D85FDC63C6DAE74" box="[1218,1283,567,593]" pageId="20" pageNumber="421">South</collectorName>
|
||
side of
|
||
<location id="8EB91DC2FFFAFF959C21FDC93872AE50" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959C21FDC93872AE50" country="Philippines" county="Island" latitude="12.58355" longLatPrecision="1" longitude="122.2438" municipality="Lugbung Island" name="Lugbung Island" pageId="20" pageNumber="421" stateProvince="Romblon">Lugbung Island</location>
|
||
<locationDeviation id="9A6B523BFFFAFF959863FDAA3A35AE50" box="[292,859,603,629]" firstLocation="Nabagbagan Rocks" pageId="20" pageNumber="421" secondLocation="Logbon Sandbar">between Nabagbagan Rocks and Logbon Sandbar</locationDeviation>
|
||
,
|
||
<geoCoordinate id="EE522DDEFFFAFF959A20FDAA3A9EAE53" box="[871,1008,603,630]" degrees="12.58355" direction="north" orientation="latitude" pageId="20" pageNumber="421" precision="1" value="12.58355">12.58355°N</geoCoordinate>
|
||
<geoCoordinate id="EE522DDEFFFAFF959ABFFDAA3DE1AE50" box="[1016,1167,603,629]" degrees="122.24380" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="1" value="122.2438">122.24380°E</geoCoordinate>
|
||
,
|
||
<collectingMunicipality id="6BBDD163FFFAFF959DDCFDAA3C3EAE50" box="[1179,1360,603,629]" pageId="20" pageNumber="421">Lugbung Island</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49A285FBFFFAFF959C1BFDAD39A7AEBC" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon</collectingRegion>
|
||
<collectingCounty id="62B83395FFFAFF959994FD8E3874AEBC" box="[211,282,639,665]" pageId="20" pageNumber="421">Island</collectingCounty>
|
||
,
|
||
<collectingRegion id="49A285FBFFFAFF95986FFD8E3B6FAEBC" box="[296,513,639,665]" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3710B89FFFAFF959B48FD8E3BFEAEBC" box="[527,656,639,665]" name="Philippines" pageId="20" pageNumber="421">Philippines</collectingCountry>
|
||
,
|
||
<date id="FFD86DD9FFFAFF959BD9FD8E3A5BAEBC" box="[670,821,639,665]" pageId="20" pageNumber="421" value="2017-04-03">
|
||
<collectingDate id="EF9C9431FFFAFF959BD9FD8E3A5BAEBC" box="[670,821,639,665]" pageId="20" pageNumber="421" value="2017-04-03">3 April 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B0E4144FFFAFF959A05FD713A8BAEBC" ID-GBIF-Occurrence="3015405323" box="[834,997,639,666]" collectionCode="T" country="Philippines" location="Gosliner" pageId="20" pageNumber="421" specimenCount="1" stateProvince="Romblon" typeStatus="paratype">
|
||
<collectionCode id="ED77D3DCFFFAFF959A05FD713A3BAEBF" box="[834,853,640,666]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="421">T</collectionCode>
|
||
.M.
|
||
<location id="8EB91DC2FFFAFF959AC5FD8E3A8BAEBC" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959AC5FD8E3A8BAEBC" box="[898,997,639,665]" country="Philippines" name="Gosliner" pageId="20" pageNumber="421" stateProvince="Romblon">Gosliner</location>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0E4144FFFAFF959AB5FD713B9FAEC4" ID-GBIF-Occurrence="3015405312" collectingDate="2017-04-06" collectionCode="NMP, CASIZ" collectorName="Cobrador Rocks" country="Philippines" county="Naguso Island" latitude="12.6518" location="Cabugaan Island" longLatPrecision="1" longitude="122.23113" municipality="Cobrador Island" pageId="20" pageNumber="421" specimenCode="NMP 041293, CASIZ 222019" specimenCount="1" stateProvince="Romblon" typeStatus="paratype">
|
||
<specimenCode id="DBC0E362FFFAFF959AB5FD713DE2AEBC" box="[1010,1164,639,666]" collectionCode="NMP" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="20" pageNumber="421">NMP 041293</specimenCode>
|
||
(formerly from
|
||
<specimenCode id="DBC0E362FFFAFF959C0CFD8E399EAE98" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 222019</specimenCode>
|
||
a), one specimen,
|
||
<collectorName id="26932ECFFFFAFF9598FDFD523B1BAE98" box="[442,629,675,701]" pageId="20" pageNumber="421">Cobrador Rocks</collectorName>
|
||
,
|
||
<geoCoordinate id="EE522DDEFFFAFF959BC6FD523A64AE9B" box="[641,778,675,702]" degrees="12.65180" direction="north" orientation="latitude" pageId="20" pageNumber="421" precision="1" value="12.6518">12.65180°N</geoCoordinate>
|
||
<geoCoordinate id="EE522DDEFFFAFF959A56FD523AC9AE98" box="[785,935,675,701]" degrees="122.23113" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="1" value="122.23113">122.23113°E</geoCoordinate>
|
||
,
|
||
<location id="8EB91DC2FFFAFF959AF4FD523D1CAE98" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959AF4FD523D1CAE98" box="[947,1138,675,701]" country="Philippines" county="Naguso Island" latitude="12.6518" longLatPrecision="1" longitude="122.23113" municipality="Cobrador Island" name="Cabugaan Island" pageId="20" pageNumber="421" stateProvince="Romblon">Cabugaan Island</location>
|
||
,
|
||
<collectingMunicipality id="6BBDD163FFFAFF959D38FD523C5BAE98" box="[1151,1333,675,701]" pageId="20" pageNumber="421">Cobrador Island</collectingMunicipality>
|
||
(
|
||
<collectingCounty id="62B83395FFFAFF959C02FD52398EAEC7" pageId="20" pageNumber="421">Naguso Island</collectingCounty>
|
||
),
|
||
<collectingRegion id="49A285FBFFFAFF9599B4FD3638A6AEC4" box="[243,456,711,737]" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3710B89FFFAFF959893FD363B3BAEC4" box="[468,597,711,737]" name="Philippines" pageId="20" pageNumber="421">Philippines</collectingCountry>
|
||
,
|
||
<date id="FFD86DD9FFFAFF959B27FD363B9FAEC4" box="[608,753,711,737]" pageId="20" pageNumber="421" value="2017-04-06">
|
||
<collectingDate id="EF9C9431FFFAFF959B27FD363B9FAEC4" box="[608,753,711,737]" pageId="20" pageNumber="421" value="2017-04-06">6 April 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, J. Maestro.
|
||
<materialsCitation id="3B0E4144FFFAFF959A38FD3638D5AF0C" ID-GBIF-Occurrence="3015405305" collectingDate="2017-04-06" collectionCode="CASIZ" collectorName="Cobrador Rocks" country="Philippines" county="Naguso Island" latitude="12.6518" location="Cabugaan Island" longLatPrecision="1" longitude="122.23113" municipality="Cobrador Island" pageId="20" pageNumber="421" specimenCode="CASIZ 222019" specimenCount="1" stateProvince="Romblon" typeStatus="paratype">
|
||
<specimenCode id="DBC0E362FFFAFF959A38FD363D40AEC4" box="[895,1070,711,737]" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 222019</specimenCode>
|
||
, one specimen, subsampled.
|
||
<collectorName id="26932ECFFFFAFF959C35FD36385CAF20" pageId="20" pageNumber="421">Cobrador Rocks</collectorName>
|
||
,
|
||
<geoCoordinate id="EE522DDEFFFAFF959879FD1A38A9AF23" box="[318,455,747,774]" degrees="12.65180" direction="north" orientation="latitude" pageId="20" pageNumber="421" precision="1" value="12.6518">12.65180°N</geoCoordinate>
|
||
<geoCoordinate id="EE522DDEFFFAFF959888FD1A3B0BAF20" box="[463,613,747,773]" degrees="122.23113" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="1" value="122.23113">122.23113°E</geoCoordinate>
|
||
,
|
||
<location id="8EB91DC2FFFAFF959B36FD1A3A5EAF20" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959B36FD1A3A5EAF20" box="[625,816,747,773]" country="Philippines" county="Naguso Island" latitude="12.6518" longLatPrecision="1" longitude="122.23113" municipality="Cobrador Island" name="Cabugaan Island" pageId="20" pageNumber="421" stateProvince="Romblon">Cabugaan Island</location>
|
||
,
|
||
<collectingMunicipality id="6BBDD163FFFAFF959A7AFD1A3A9DAF20" box="[829,1011,747,773]" pageId="20" pageNumber="421">Cobrador Island</collectingMunicipality>
|
||
(
|
||
<collectingCounty id="62B83395FFFAFF959D43FD1A3DC3AF23" box="[1028,1197,747,774]" pageId="20" pageNumber="421">Naguso Island</collectingCounty>
|
||
),
|
||
<collectingRegion id="49A285FBFFFAFF959D86FD1A3CF6AF20" box="[1217,1432,747,773]" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3710B89FFFAFF9599D0FCFE3876AF0C" box="[151,280,783,809]" name="Philippines" pageId="20" pageNumber="421">Philippines</collectingCountry>
|
||
,
|
||
<date id="FFD86DD9FFFAFF959861FCFE38D5AF0C" box="[294,443,783,809]" pageId="20" pageNumber="421" value="2017-04-06">
|
||
<collectingDate id="EF9C9431FFFAFF959861FCFE38D5AF0C" box="[294,443,783,809]" pageId="20" pageNumber="421" value="2017-04-06">6 April 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, J. Maestro.
|
||
<materialsCitation id="3B0E4144FFFAFF959B17FCFE3DC5AF68" ID-GBIF-Occurrence="3015405303" collectingDate="2018-03-19" collectionCode="CASIZ" country="Philippines" latitude="12.65212" location="Bangug Island" longLatPrecision="1" longitude="122.23082" municipality="Island" pageId="20" pageNumber="421" specimenCode="CASIZ 224653" specimenCount="1" stateProvince="Romblon" typeStatus="paratype">
|
||
<specimenCode id="DBC0E362FFFAFF959B17FCFE3A6FAF0F" box="[592,769,783,810]" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 224653</specimenCode>
|
||
, one specimen, subsampled and dissected.
|
||
<location id="8EB91DC2FFFAFF959DB7FCE13CF6AF0C" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959DB7FCE13CF6AF0C" box="[1264,1432,783,810]" country="Philippines" latitude="12.65212" longLatPrecision="1" longitude="122.23082" municipality="Island" name="Bangug Island" pageId="20" pageNumber="421" stateProvince="Romblon">Bangug Island</location>
|
||
,
|
||
<geoCoordinate id="EE522DDEFFFAFF9599D0FCC2384EAF6B" box="[151,288,819,846]" degrees="12.65212" direction="north" orientation="latitude" pageId="20" pageNumber="421" precision="1" value="12.65212">12.65212°N</geoCoordinate>
|
||
<geoCoordinate id="EE522DDEFFFAFF959860FCC238D0AF68" box="[295,446,819,845]" degrees="122.23082" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="1" value="122.23082">122.23082°E</geoCoordinate>
|
||
,
|
||
<collectingRegion id="49A285FBFFFAFF95988EFCC23B5FAF68" box="[457,561,819,845]" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon</collectingRegion>
|
||
<collectingMunicipality id="6BBDD163FFFAFF959B7EFCC23BEEAF68" box="[569,640,819,845]" pageId="20" pageNumber="421">Island</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49A285FBFFFAFF959BCAFCC23A0CAF68" box="[653,866,819,845]" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3710B89FFFAFF959A28FCC23A9EAF68" box="[879,1008,819,845]" name="Philippines" pageId="20" pageNumber="421">Philippines</collectingCountry>
|
||
,
|
||
<date id="FFD86DD9FFFAFF959ABBFCC23DC5AF68" box="[1020,1195,819,845]" pageId="20" pageNumber="421" value="2018-03-19">
|
||
<collectingDate id="EF9C9431FFFAFF959ABBFCC23DC5AF68" box="[1020,1195,819,845]" pageId="20" pageNumber="421" value="2018-03-19">19 March 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, K.L. Larkin.
|
||
<materialsCitation id="3B0E4144FFFAFF959C0CFCC23B60AFB0" ID-GBIF-Occurrence="3015405310" collectingDate="2018-03-18" collectionCode="CASIZ" country="Philippines" latitude="12.58418" location="Lugbong Island" longLatPrecision="1" longitude="122.24429" municipality="Island" pageId="20" pageNumber="421" specimenCode="CASIZ 224654" specimenCount="1" stateProvince="Romblon" typeStatus="paratype">
|
||
<specimenCode id="DBC0E362FFFAFF959C0CFCC23980AF57" collectionCode="CASIZ" pageId="20" pageNumber="421">CASIZ 224654</specimenCode>
|
||
, one specimen, subsampled.
|
||
<location id="8EB91DC2FFFAFF959B7EFCA63B85AF54" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF959B7EFCA63B85AF54" box="[569,747,855,881]" country="Philippines" latitude="12.58418" longLatPrecision="1" longitude="122.24429" municipality="Island" name="Lugbong Island" pageId="20" pageNumber="421" stateProvince="Romblon">Lugbong Island</location>
|
||
<geoCoordinate id="EE522DDEFFFAFF959BB3FCA63A13AF57" box="[756,893,855,882]" degrees="12.58418" direction="north" orientation="latitude" pageId="20" pageNumber="421" precision="1" value="12.58418">12.58418°N</geoCoordinate>
|
||
<geoCoordinate id="EE522DDEFFFAFF959AC2FCA63D72AF54" box="[901,1052,855,881]" degrees="122.24429" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="1" value="122.24429">122.24429°E</geoCoordinate>
|
||
,
|
||
<collectingRegion id="49A285FBFFFAFF959D6FFCA63DFEAF54" box="[1064,1168,855,881]" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon</collectingRegion>
|
||
<collectingMunicipality id="6BBDD163FFFAFF959DDEFCA63D8EAF54" box="[1177,1248,855,881]" pageId="20" pageNumber="421">Island</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49A285FBFFFAFF959DAAFCA639A6AFB0" country="Philippines" name="Romblon" pageId="20" pageNumber="421">Romblon Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3710B89FFFAFF959993FC8A383BAFB0" box="[212,341,891,917]" name="Philippines" pageId="20" pageNumber="421">Philippines</collectingCountry>
|
||
,
|
||
<date id="FFD86DD9FFFAFF959827FC8A3B60AFB0" box="[352,526,891,917]" pageId="20" pageNumber="421" value="2018-03-18">
|
||
<collectingDate id="EF9C9431FFFAFF959827FC8A3B60AFB0" box="[352,526,891,917]" pageId="20" pageNumber="421" value="2018-03-18">18 March 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, C. Alahado.
|
||
</paragraph>
|
||
<paragraph id="8BD94B19FFFAFF959980FC6E3B13AF9C" blockId="20.[151,1437,423,2033]" box="[199,637,927,953]" pageId="20" pageNumber="421">
|
||
<materialsCitation id="3B0E4144FFFAFF959980FC6E3B17AF9C" ID-GBIF-Occurrence="3015405306" box="[199,633,927,953]" country="Palau" location="Uchelbeluu Reef" pageId="20" pageNumber="421" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B912970BFFFAFF959980FC6E3808AF9C" bold="true" box="[199,358,927,953]" pageId="20" pageNumber="421">
|
||
<typeStatus id="54DDF5BBFFFAFF959980FC6E386CAF9C" box="[199,258,927,953]" pageId="20" pageNumber="421">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EB91DC2FFFAFF95982BFC6E3B42AF9C" LSID="urn:lsid:plazi:treatment:03CFFA0FFFFAFF9699D0FF663C50AD38:8EB91DC2FFFAFF95982BFC6E3B42AF9C" box="[364,556,927,953]" country="Palau" name="Uchelbeluu Reef" pageId="20" pageNumber="421">Uchelbeluu Reef</location>
|
||
,
|
||
<collectingCountry id="F3710B89FFFAFF959B70FC6E3B17AF9C" box="[567,633,927,953]" name="Palau" pageId="20" pageNumber="421">Palau</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD94B19FFFAFF959980FC323CE2A8FC" blockId="20.[151,1437,423,2033]" pageId="20" pageNumber="421">
|
||
<emphasis id="B912970BFFFAFF959980FC3238A1AFF8" bold="true" box="[199,463,963,989]" pageId="20" pageNumber="421">External morphology.</emphasis>
|
||
Living animals (
|
||
<figureCitation id="135D579CFFFAFF959BD3FC323BB9AFF8" box="[660,727,963,989]" captionStart="FIGURE 2" captionStartId="14.[151,250,1794,1818]" captionTargetBox="[148,1439,181,1770]" captionTargetId="figure-24@14.[148,1439,181,1770]" captionTargetPageId="14" captionText="FIGURE 2. A. Chromodoris balat sp. nov. NMP 041292, Philippines; B. Chromodoris baqe sp. nov. CASIZ 192285, Red Sea; C–E. Chromodoris kalawakan sp. nov. C. CASIZ 109740, Palau; D. CASIZ 178300, Vanuatu; E. CASIZ 222016, Philippines; F. Chromodoris aff. mandapamensis, CASIZ 181260, Philippines; G–H. Chromodoris quagga sp. nov. G. CASIZ 184320, Philippines; H. CASIZ 229987, Philippines. Photos: T.M. Gosliner." figureDoi="http://doi.org/10.5281/zenodo.4397009" httpUri="https://zenodo.org/record/4397009/files/figure.png" pageId="20" pageNumber="421">Fig. 2</figureCitation>
|
||
C–E) are small, with a maximum length of
|
||
<quantity id="4C9EE6FCFFFAFF959D89FC323C4CAFFB" box="[1230,1314,963,990]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="421" unit="mm" value="20.0">20 mm</quantity>
|
||
. Body is a translucent white or light gray color with bright opaque white spots across the body. The white spots are found on the tips of short conical projections. Some of the bright white spots are interconnected by diffuse opaque white lines. The marginal band includes small orange dots around the whole nudibranch. Nine to ten unipinnate gill branches that are white in color with bright opaque white spots and orange brown markings (
|
||
<figureCitation id="135D579CFFFAFF959D03FBA23DF0A848" box="[1092,1182,1107,1133]" captionStart="FIGURE 7" captionStartId="21.[151,250,1837,1861]" captionTargetBox="[164,1428,569,1804]" captionTargetId="figure-178@21.[149,1438,561,1814]" captionTargetPageId="21" captionText="FIGURE 7. Chromodoris kalawakan sp. nov., CASIZ 178300. A. Gill branches (scale bar = 0.17 mm); B. Rhinophore (scale bar = 0.1 mm) and C. Mantle glands. CASIZ 224653: D. Buccal mass (scale bar = 0.18 mm) and E. Reproductive system (scale bar = 0.25 mm).Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; ej, ejaculatory duct; fgm, female gland mass; ot, oral tube; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; v, vagina; vg, vestibular gland." figureDoi="http://doi.org/10.5281/zenodo.4397021" httpUri="https://zenodo.org/record/4397021/files/figure.png" pageId="20" pageNumber="421">Fig. 7A</figureCitation>
|
||
). The perfoliate rhinophores are white and have eight distinct lamellae with an orange accent at the top and the bright white opaque spots (
|
||
<figureCitation id="135D579CFFFAFF9599E7FB6A3999A893" box="[160,247,1179,1206]" captionStart="FIGURE 7" captionStartId="21.[151,250,1837,1861]" captionTargetBox="[164,1428,569,1804]" captionTargetId="figure-178@21.[149,1438,561,1814]" captionTargetPageId="21" captionText="FIGURE 7. Chromodoris kalawakan sp. nov., CASIZ 178300. A. Gill branches (scale bar = 0.17 mm); B. Rhinophore (scale bar = 0.1 mm) and C. Mantle glands. CASIZ 224653: D. Buccal mass (scale bar = 0.18 mm) and E. Reproductive system (scale bar = 0.25 mm).Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; ej, ejaculatory duct; fgm, female gland mass; ot, oral tube; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; v, vagina; vg, vestibular gland." figureDoi="http://doi.org/10.5281/zenodo.4397021" httpUri="https://zenodo.org/record/4397021/files/figure.png" pageId="20" pageNumber="421">Fig. 7B</figureCitation>
|
||
). The posterior end of the foot extends well beyond the posterior end of the mantle and is ornamented with a network of diffuse opaque white markings. On either side of the mouth there is a pair of digitiform oral tentacles.
|
||
</paragraph>
|
||
<paragraph id="8BD94B19FFFAFF959980FB123C7AA960" blockId="20.[151,1437,423,2033]" pageId="20" pageNumber="421">
|
||
<emphasis id="B912970BFFFAFF959980FB1238A6A8D8" bold="true" box="[199,456,1251,1277]" pageId="20" pageNumber="421">Internal morphology.</emphasis>
|
||
<emphasis id="B912970BFFFAFF959896FB123B1DA8D8" box="[465,627,1251,1277]" italics="true" pageId="20" pageNumber="421">Mantle glands</emphasis>
|
||
: The mantle glands are subcutaneous around the entire mantle margin, except for the anterior margin. There are approximately six mantle glands per side of the mantle (
|
||
<figureCitation id="135D579CFFFAFF959D97FAF63C49A904" box="[1232,1319,1287,1313]" captionStart="FIGURE 7" captionStartId="21.[151,250,1837,1861]" captionTargetBox="[164,1428,569,1804]" captionTargetId="figure-178@21.[149,1438,561,1814]" captionTargetPageId="21" captionText="FIGURE 7. Chromodoris kalawakan sp. nov., CASIZ 178300. A. Gill branches (scale bar = 0.17 mm); B. Rhinophore (scale bar = 0.1 mm) and C. Mantle glands. CASIZ 224653: D. Buccal mass (scale bar = 0.18 mm) and E. Reproductive system (scale bar = 0.25 mm).Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; ej, ejaculatory duct; fgm, female gland mass; ot, oral tube; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; v, vagina; vg, vestibular gland." figureDoi="http://doi.org/10.5281/zenodo.4397021" httpUri="https://zenodo.org/record/4397021/files/figure.png" pageId="20" pageNumber="421">Fig. 7C</figureCitation>
|
||
). They are slightly yellower in color on the preserved
|
||
<typeStatus id="54DDF5BBFFFAFF959B3EFADA3BB1A960" box="[633,735,1323,1349]" pageId="20" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
. The mantle glands in
|
||
<taxonomicName id="4C66309AFFFAFF959A9CFADD3D1AA960" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[987,1140,1323,1350]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="421" phylum="Mollusca" rank="species" species="kalawakan">
|
||
<emphasis id="B912970BFFFAFF959A9CFADD3D1AA960" box="[987,1140,1323,1350]" italics="true" pageId="20" pageNumber="421">C. kalawakan</emphasis>
|
||
</taxonomicName>
|
||
are Y-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BD94B19FFFAFF959980FABE38A1AAAC" blockId="20.[151,1437,423,2033]" pageId="20" pageNumber="421">
|
||
<emphasis id="B912970BFFFAFF959980FABE38B9A94C" box="[199,471,1359,1385]" italics="true" pageId="20" pageNumber="421">Buccal mass and radula</emphasis>
|
||
: The muscular portion of the buccal mass is slightly smaller than the oral tube length. A chitinous labial cuticle is found at the anterior end of the muscular portion (
|
||
<figureCitation id="135D579CFFFAFF959D4AFA823D08A9AB" box="[1037,1126,1395,1422]" captionStart="FIGURE 7" captionStartId="21.[151,250,1837,1861]" captionTargetBox="[164,1428,569,1804]" captionTargetId="figure-178@21.[149,1438,561,1814]" captionTargetPageId="21" captionText="FIGURE 7. Chromodoris kalawakan sp. nov., CASIZ 178300. A. Gill branches (scale bar = 0.17 mm); B. Rhinophore (scale bar = 0.1 mm) and C. Mantle glands. CASIZ 224653: D. Buccal mass (scale bar = 0.18 mm) and E. Reproductive system (scale bar = 0.25 mm).Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; ej, ejaculatory duct; fgm, female gland mass; ot, oral tube; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; v, vagina; vg, vestibular gland." figureDoi="http://doi.org/10.5281/zenodo.4397021" httpUri="https://zenodo.org/record/4397021/files/figure.png" pageId="20" pageNumber="421">Fig. 7D</figureCitation>
|
||
) with widely-forked, bifurcated jaw elements (
|
||
<figureCitation id="135D579CFFFAFF95983AFA6638A7A994" box="[381,457,1431,1457]" captionStart="FIGURE 8" captionStartId="22.[151,250,1580,1604]" captionTargetBox="[151,1436,181,1556]" captionTargetId="figure-24@22.[151,1436,181,1556]" captionTargetPageId="22" captionText="FIGURE 8. Scanning electron micrographs. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. A. Jaw. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. B. Jaw rodlets. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. C. Radula. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. D. Central teeth; E. Mid-lateral teeth; F. Outer lateral teeth." figureDoi="http://doi.org/10.5281/zenodo.4397023" httpUri="https://zenodo.org/record/4397023/files/figure.png" pageId="20" pageNumber="421">Figs. 8</figureCitation>
|
||
A–B). The radular formula in two
|
||
<typeStatus id="54DDF5BBFFFAFF959A09FA693AD2A997" box="[846,956,1432,1458]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<collectionCode id="ED77D3DCFFFAFF959A8FFA663D77A994" box="[968,1049,1431,1457]" pageId="20" pageNumber="421">CASIZ</collectionCode>
|
||
224653 and
|
||
<collectionCode id="ED77D3DCFFFAFF959DECFA663D92A994" box="[1195,1276,1431,1457]" pageId="20" pageNumber="421">CASIZ</collectionCode>
|
||
178300, is 43 x 32.1.32 and 43 x 29–31.1.29–31, respectively (
|
||
<figureCitation id="135D579CFFFAFF959B86FA4A3A74A9F0" box="[705,794,1467,1493]" captionStart="FIGURE 8" captionStartId="22.[151,250,1580,1604]" captionTargetBox="[151,1436,181,1556]" captionTargetId="figure-24@22.[151,1436,181,1556]" captionTargetPageId="22" captionText="FIGURE 8. Scanning electron micrographs. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. A. Jaw. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. B. Jaw rodlets. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. C. Radula. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. D. Central teeth; E. Mid-lateral teeth; F. Outer lateral teeth." figureDoi="http://doi.org/10.5281/zenodo.4397023" httpUri="https://zenodo.org/record/4397023/files/figure.png" pageId="20" pageNumber="421">Fig. 8C</figureCitation>
|
||
). The rachidian tooth is thin and linear without a distinct cusp. On either side of it, the inner lateral teeth have 1–3 denticles on the inner side and 2–3 on the outer side (
|
||
<figureCitation id="135D579CFFFAFF959C28FA2E39D3AA38" captionStart="FIGURE 8" captionStartId="22.[151,250,1580,1604]" captionTargetBox="[151,1436,181,1556]" captionTargetId="figure-24@22.[151,1436,181,1556]" captionTargetPageId="22" captionText="FIGURE 8. Scanning electron micrographs. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. A. Jaw. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. B. Jaw rodlets. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. C. Radula. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. D. Central teeth; E. Mid-lateral teeth; F. Outer lateral teeth." figureDoi="http://doi.org/10.5281/zenodo.4397023" httpUri="https://zenodo.org/record/4397023/files/figure.png" pageId="20" pageNumber="421">Fig. 8D</figureCitation>
|
||
). The inner lateral tooth has an elongate central cusp that is about twice the length of the adjacent denticles. The remaining laterals only have denticles on the outer side of the central cusp. The middle lateral teeth have an elongate cutting edge with 3 to 7 acutely pointed denticles (
|
||
<figureCitation id="135D579CFFFAFF959B9DF9BA3A5DAA40" box="[730,819,1611,1637]" captionStart="FIGURE 8" captionStartId="22.[151,250,1580,1604]" captionTargetBox="[151,1436,181,1556]" captionTargetId="figure-24@22.[151,1436,181,1556]" captionTargetPageId="22" captionText="FIGURE 8. Scanning electron micrographs. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. A. Jaw. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. B. Jaw rodlets. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. C. Radula. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. D. Central teeth; E. Mid-lateral teeth; F. Outer lateral teeth." figureDoi="http://doi.org/10.5281/zenodo.4397023" httpUri="https://zenodo.org/record/4397023/files/figure.png" pageId="20" pageNumber="421">Fig. 8E</figureCitation>
|
||
). The outer lateral teeth are elongate with 2 to 6 more rounded denticles (
|
||
<figureCitation id="135D579CFFFAFF95982AF99E38ACAAAC" box="[365,450,1647,1673]" captionStart="FIGURE 8" captionStartId="22.[151,250,1580,1604]" captionTargetBox="[151,1436,181,1556]" captionTargetId="figure-24@22.[151,1436,181,1556]" captionTargetPageId="22" captionText="FIGURE 8. Scanning electron micrographs. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. A. Jaw. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. B. Jaw rodlets. Chromodoris kalawakan sp. nov., CASIZ 178300, Vanuatu. C. Radula. Chromodoris kalawakan sp. nov., CASIZ 224653, Philippines. D. Central teeth; E. Mid-lateral teeth; F. Outer lateral teeth." figureDoi="http://doi.org/10.5281/zenodo.4397023" httpUri="https://zenodo.org/record/4397023/files/figure.png" pageId="20" pageNumber="421">Fig. 8F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD94B19FFFAFF959980F9623B35ABE8" blockId="20.[151,1437,423,2033]" pageId="20" pageNumber="421">
|
||
<emphasis id="B912970BFFFAFF959980F96238DFAA88" box="[199,433,1683,1709]" italics="true" pageId="20" pageNumber="421">Reproductive system</emphasis>
|
||
(
|
||
<figureCitation id="135D579CFFFAFF959884F9623B70AA8B" box="[451,542,1683,1710]" captionStart="FIGURE 7" captionStartId="21.[151,250,1837,1861]" captionTargetBox="[164,1428,569,1804]" captionTargetId="figure-178@21.[149,1438,561,1814]" captionTargetPageId="21" captionText="FIGURE 7. Chromodoris kalawakan sp. nov., CASIZ 178300. A. Gill branches (scale bar = 0.17 mm); B. Rhinophore (scale bar = 0.1 mm) and C. Mantle glands. CASIZ 224653: D. Buccal mass (scale bar = 0.18 mm) and E. Reproductive system (scale bar = 0.25 mm).Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; ej, ejaculatory duct; fgm, female gland mass; ot, oral tube; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; v, vagina; vg, vestibular gland." figureDoi="http://doi.org/10.5281/zenodo.4397021" httpUri="https://zenodo.org/record/4397021/files/figure.png" pageId="20" pageNumber="421">Fig. 7E</figureCitation>
|
||
): The thin pre-ampullary duct connects the ovotestis with the wider elongate ampulla. The ampulla narrows proximally and divides into a short oviduct and an elongate vas deferens. The distal portion of the vas deferens is wide, convoluted, and prostatic. The prostatic portion narrows proximally and enters the short muscular ejaculatory segment. This segment expands into the wider penial bulb, which also joins with the distal end of the vagina. The vagina is moderately long and curved and enters the base of the thin-walled, spherical bursa copulatrix. Also, joining at the base of the bursa is the receptaculum seminis duct, which connects with the pyriform curved receptaculum seminis. Along the length of the receptaculum seminis duct is a short branch of the uterine duct that enters the female gland mass. The female gland mass is composed of a small albumen and membrane glands and a larger mucous gland.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C1892FFFAFF949980F8263C4BACDC" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421" type="etymology">
|
||
<paragraph id="8BD94B19FFFAFF949980F8263AC0ACF0" blockId="20.[151,1437,423,2033]" lastBlockId="21.[151,1437,151,537]" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
|
||
<emphasis id="B912970BFFFAFF959980F8263820ABD4" bold="true" box="[199,334,2007,2033]" pageId="20" pageNumber="421">Etymology.</emphasis>
|
||
This species is named
|
||
<taxonomicName id="4C66309AFFFAFF959B1EF8263A01ABD4" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[601,879,2007,2033]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="421" phylum="Mollusca" rank="species" species="kalawakan">
|
||
<emphasis id="B912970BFFFAFF959B1EF8263A01ABD4" box="[601,879,2007,2033]" italics="true" pageId="20" pageNumber="421">Chromodoris kalawakan</emphasis>
|
||
</taxonomicName>
|
||
after the Filipino word for galaxy, since the coloration pattern looks like stars floating in a space cloud. The lines connecting some of the white dots appear as if astronomers were drawing the constellation patterns between the stars.
|
||
</paragraph>
|
||
<paragraph id="8BD94B19FFFBFF949980FF2F3C4BACDC" blockId="21.[151,1437,151,537]" box="[199,1317,222,249]" pageId="21" pageNumber="422">
|
||
<emphasis id="B912970BFFFBFF949980FF2F3B6BACDD" bold="true" box="[199,517,222,248]" pageId="21" pageNumber="422">Geographical distribution.</emphasis>
|
||
This species is known from
|
||
<collectingCountry id="F3710B89FFFBFF949A01FF2E3ACBACDC" box="[838,933,223,249]" name="Vanuatu" pageId="21" pageNumber="422">Vanuatu</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3710B89FFFBFF949AF7FF2E3D4EACDC" box="[944,1056,223,249]" name="Indonesia" pageId="21" pageNumber="422">Indonesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3710B89FFFBFF949D6CFF2E3DC7ACDC" box="[1067,1193,223,249]" name="Philippines" pageId="21" pageNumber="422">Philippines</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3710B89FFFBFF949D98FF2E3C4FACDC" box="[1247,1313,223,249]" name="Palau" pageId="21" pageNumber="422">Palau</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C1892FFFBFF969980FEF23C50AD38" lastPageId="23" lastPageNumber="424" pageId="21" pageNumber="422" type="discussion">
|
||
<paragraph id="8BD94B19FFFBFF949980FEF239BCAE3C" blockId="21.[151,1437,151,537]" pageId="21" pageNumber="422">
|
||
<emphasis id="B912970BFFFBFF949980FEF23855AD38" bold="true" box="[199,315,259,285]" pageId="21" pageNumber="422">Remarks.</emphasis>
|
||
In our molecular phylogeny,
|
||
<taxonomicName id="4C66309AFFFBFF949BC2FEF23AF7AD38" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[645,921,259,285]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="kalawakan">
|
||
<emphasis id="B912970BFFFBFF949BC2FEF23AF7AD38" box="[645,921,259,285]" italics="true" pageId="21" pageNumber="422">Chromodoris kalawakan</emphasis>
|
||
</taxonomicName>
|
||
is a member of a large, polytomous clade that includes:
|
||
<taxonomicName id="4C66309AFFFBFF949845FED63B1FAD64" authority="Bergh, 1877" authorityName="Bergh" authorityYear="1877" box="[258,625,295,321]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="annae">
|
||
<emphasis id="B912970BFFFBFF949845FED6388EAD65" box="[258,480,295,321]" italics="true" pageId="21" pageNumber="422">Chromodoris annae</emphasis>
|
||
<bibRefCitation id="EFF736E8FFFBFF9498A2FED63B1FAD64" author="Bergh, L. S. R." box="[485,625,295,321]" pageId="21" pageNumber="422" pagination="429 - 546" refId="ref15402" refString="Bergh, L. S. R. (1877) Malacologische Untersuchungen 11. Reisen im Archipel Philippinen. Wissenschaftliche Resultate, 2, 429 - 546." type="journal article" year="1877">Bergh, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF949B3CFED63D41AD64" authority="Bergh, 1877" authorityName="Bergh" authorityYear="1877" box="[635,1071,295,321]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="elisabethina">
|
||
<emphasis id="B912970BFFFBFF949B3CFED63AF0AD64" box="[635,926,295,321]" italics="true" pageId="21" pageNumber="422">Chromodoris elisabethina</emphasis>
|
||
<bibRefCitation id="EFF736E8FFFBFF949AE4FED63D41AD64" author="Bergh, L. S. R." box="[931,1071,295,321]" pageId="21" pageNumber="422" pagination="429 - 546" refId="ref15402" refString="Bergh, L. S. R. (1877) Malacologische Untersuchungen 11. Reisen im Archipel Philippinen. Wissenschaftliche Resultate, 2, 429 - 546." type="journal article" year="1877">Bergh, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF949D7EFED63DD2AD64" baseAuthorityName="van Hasselt" baseAuthorityYear="1824" box="[1081,1212,295,321]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="lineolata">
|
||
<emphasis id="B912970BFFFBFF949D7EFED63DD2AD64" box="[1081,1212,295,321]" italics="true" pageId="21" pageNumber="422">C. lineolata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF949D8EFED6389EAD40" authority="Valdes et al., 1999" authorityName="Valdes" authorityYear="1999" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="mandapamensis">
|
||
<emphasis id="B912970BFFFBFF949D8EFED63875AD40" italics="true" pageId="21" pageNumber="422">Chromodoris mandapamensis</emphasis>
|
||
<bibRefCitation id="EFF736E8FFFBFF949866FEBA389EAD40" author="Valdes, A. & Mollo, E. & Ortea, J." box="[289,496,331,357]" pageId="21" pageNumber="422" pagination="461 - 472" refId="ref18062" refString="Valdes, A., Mollo, E. & Ortea, J. (1999) Two new species of Chromodoris (Mollusca, Nudibranchia, Chromodorididae) from Southern India, with a redescription of Chromodoris trimarginata (Winckworth, 1946). Proceedings of the California Academy of Sciences, 51, 461 - 472." type="journal article" year="1999">
|
||
Valdés
|
||
<emphasis id="B912970BFFFBFF949836FEBA38C1AD40" box="[369,431,331,357]" italics="true" pageId="21" pageNumber="422">et al.,</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF9498BDFEBA3D4EAD40" authority="Gosliner & Behrens, 1998" authorityName="Gosliner & Behrens" authorityYear="1998" box="[506,1056,331,357]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="michaeli">
|
||
<emphasis id="B912970BFFFBFF9498BDFEBA3B9BAD40" box="[506,757,331,357]" italics="true" pageId="21" pageNumber="422">Chromodoris michaeli</emphasis>
|
||
<bibRefCitation id="EFF736E8FFFBFF949BBCFEBA3D4EAD40" author="Gosliner, T. M. & Behrens, D. W." box="[763,1056,331,357]" pageId="21" pageNumber="422" pagination="139 - 165" refId="ref15903" refString="Gosliner, T. M. & Behrens, D. W. (1998) Five new species of Chromodoris (Mollusca: Nudibranchia: Chromodorididae) from the tropical Indo-Pacific Ocean. Proceedings of the California Academy of Sciences, 50, 139 - 165." type="journal article" year="1998">Gosliner & Behrens, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF949D6CFEBA39BCADAC" authority="Rudman, 1983" authorityName="Rudman" authorityYear="1983" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="orientalis">
|
||
<emphasis id="B912970BFFFBFF949D6CFEBA3C5EAD40" box="[1067,1328,331,357]" italics="true" pageId="21" pageNumber="422">Chromodoris orientalis</emphasis>
|
||
<bibRefCitation id="EFF736E8FFFBFF949C71FEBA39BCADAC" author="Rudman, W. B." pageId="21" pageNumber="422" pagination="105 - 173" refId="ref17646" refString="Rudman, W. B. (1983) The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris splendida, C. aspersa and Hypselodoris placida colour groups. Zoological Journal of the Linnean Society, 78, 105 - 173. https: // doi. org / 10.1111 / j. 1096 - 3642.1983. tb 00864. x" type="journal article" year="1983">Rudman, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF949998FE9E3B6FADAC" baseAuthorityName="Ruppell & Leuckart" baseAuthorityYear="1830" box="[223,513,367,393]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="quadricolor">
|
||
<emphasis id="B912970BFFFBFF949998FE9E3B6FADAC" box="[223,513,367,393]" italics="true" pageId="21" pageNumber="422">Chromodoris quadricolor</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF949B57FE9E3BF9ADAC" authorityName="Bergh" authorityYear="1877" box="[528,663,367,393]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="striatella">
|
||
<emphasis id="B912970BFFFBFF949B57FE9E3BF9ADAC" box="[528,663,367,393]" italics="true" pageId="21" pageNumber="422">C. striatella</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFFBFF949BE0FE9E3D24ADAC" authority="Rudman, 1982" authorityName="Rudman" authorityYear="1982" box="[679,1098,367,393]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="strigata">
|
||
<emphasis id="B912970BFFFBFF949BE0FE9E3AF4ADAD" box="[679,922,367,393]" italics="true" pageId="21" pageNumber="422">Chromodoris strigata</emphasis>
|
||
<bibRefCitation id="EFF736E8FFFBFF949AE5FE9E3D24ADAC" author="Rudman, W. B." box="[930,1098,367,393]" pageId="21" pageNumber="422" pagination="183 - 241" refId="ref17581" refString="Rudman, W. B. (1982) The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris quadricolor, C. lineolata and Hypselodoris nigrolineata colour groups. Zoological Journal of the Linnean Society, 76, 183 - 241. https: // doi. org / 10.1111 / j. 1096 - 3642.1982. tb 02182. x" type="journal article" year="1982">Rudman, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
; and several undescribed species. The shortest uncorrected pairwise genetic distances for COI are of 10.07%, 10.07%, 10.77%, 9.20%, 10.94%, 9.72%, 9.97%, 8.86%, and 9.89% respectively to the described species stated in the previous sentence. This species was not included in the study by
|
||
<bibRefCitation id="EFF736E8FFFBFF949B51FE2A3B92ADD0" author="Layton, K. K. S. & Gosliner, T. M. & Wilson, N. G." box="[534,764,475,501]" pageId="21" pageNumber="422" pagination="27 - 36" refId="ref16812" refString="Layton, K. K. S., Gosliner, T. M. & Wilson, N. G. (2018) Flexible colour patterns obscure identification and mimicry in Indo-Pacific Chromodoris nudibranchs (Gastropoda: Chromodorididae). Molecular Phylogenetics and Evolution, 124, 27 - 36. https: // doi. org / 10.1016 / j. ympev. 2018.02.008" type="journal article" year="2018">
|
||
Layton
|
||
<emphasis id="B912970BFFFBFF949B29FE2A3BC7ADD0" box="[622,681,475,501]" italics="true" pageId="21" pageNumber="422">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
as it was originally thought to be a species of
|
||
<taxonomicName id="4C66309AFFFBFF949C5FFE2A39A4AE3C" class="Gastropoda" family="Chromodorididae" genus="Goniobranchus" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B912970BFFFBFF949C5FFE2A39A4AE3C" italics="true" pageId="21" pageNumber="422">Goniobranchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF191B91FFFBFF9499D0F8DC3DB8AB94" ID-DOI="http://doi.org/10.5281/zenodo.4397021" ID-Zenodo-Dep="4397021" httpUri="https://zenodo.org/record/4397021/files/figure.png" pageId="21" pageNumber="422" startId="21.[151,250,1837,1861]" targetBox="[164,1428,569,1804]" targetPageId="21">
|
||
<paragraph id="8BD94B19FFFBFF9499D0F8DC3DB8AB94" blockId="21.[151,1436,1837,1970]" pageId="21" pageNumber="422">
|
||
<emphasis id="B912970BFFFBFF9499D0F8DC387AAB60" bold="true" box="[151,276,1837,1861]" pageId="21" pageNumber="422">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C66309AFFFBFF94985CF8DC3B78AB60" authority="Bonomo & Gosliner, 2020" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[283,534,1837,1861]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="422" phylum="Mollusca" rank="species" species="kalawakan" status="sp. nov.">
|
||
<emphasis id="B912970BFFFBFF94985CF8DC3B78AB60" box="[283,534,1837,1861]" italics="true" pageId="21" pageNumber="422">Chromodoris kalawakan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912970BFFFBFF949B5AF8DC3B00AB60" bold="true" box="[541,622,1837,1861]" pageId="21" pageNumber="422">
|
||
<taxonomicNameLabel id="A2212A70FFFBFF949B5AF8DC3B00AB60" box="[541,622,1837,1861]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, CASIZ 178300. A. Gill branches (scale bar = 0.17 mm); B. Rhinophore (scale bar = 0.1 mm) and C. Mantle glands. CASIZ 224653: D. Buccal mass (scale bar = 0.18 mm) and E. Reproductive system (scale bar = 0.25 mm). Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; ej, ejaculatory duct; fgm, female gland mass; ot, oral tube; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; v, vagina; vg, vestibular gland.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF191B91FFF8FF9799D0F9DD39BEAA95" ID-DOI="http://doi.org/10.5281/zenodo.4397023" ID-Zenodo-Dep="4397023" httpUri="https://zenodo.org/record/4397023/files/figure.png" pageId="22" pageNumber="423" startId="22.[151,250,1580,1604]" targetBox="[151,1436,181,1556]" targetPageId="22">
|
||
<paragraph id="8BD94B19FFF8FF9799D0F9DD39BEAA95" blockId="22.[151,1437,1580,1712]" pageId="22" pageNumber="423">
|
||
<emphasis id="B912970BFFF8FF9799D0F9DD387DAA61" bold="true" box="[151,275,1580,1604]" pageId="22" pageNumber="423">FIGURE 8.</emphasis>
|
||
Scanning electron micrographs.
|
||
<taxonomicName id="4C66309AFFF8FF979B26F9DD3A35AA61" authority="Bonomo & Gosliner, 2020" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[609,859,1580,1604]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="kalawakan" status="sp. nov.">
|
||
<emphasis id="B912970BFFF8FF979B26F9DD3A35AA61" box="[609,859,1580,1604]" italics="true" pageId="22" pageNumber="423">Chromodoris kalawakan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912970BFFF8FF979A26F9DD3ADCAA61" bold="true" box="[865,946,1580,1604]" pageId="22" pageNumber="423">
|
||
<taxonomicNameLabel id="A2212A70FFF8FF979A26F9DD3ADCAA61" box="[865,946,1580,1604]" pageId="22" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, CASIZ 178300, Vanuatu. A. Jaw.
|
||
<taxonomicName id="4C66309AFFF8FF979C51F9DD386BAA4D" authority="Bonomo & Gosliner, 2020" authorityName="Bonomo & Gosliner" authorityYear="2020" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="kalawakan" status="sp. nov.">
|
||
<emphasis id="B912970BFFF8FF979C51F9DD386BAA4D" italics="true" pageId="22" pageNumber="423">Chromodoris kalawakan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912970BFFF8FF97984BF9A13830AA4D" bold="true" box="[268,350,1616,1640]" pageId="22" pageNumber="423">
|
||
<taxonomicNameLabel id="A2212A70FFF8FF97984BF9A13830AA4D" box="[268,350,1616,1640]" pageId="22" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, CASIZ 224653, Philippines. B. Jaw rodlets.
|
||
<taxonomicName id="4C66309AFFF8FF979A71F9A13D5CAA4D" authority="Bonomo & Gosliner, 2020" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[822,1074,1616,1640]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="kalawakan" status="sp. nov.">
|
||
<emphasis id="B912970BFFF8FF979A71F9A13D5CAA4D" box="[822,1074,1616,1640]" italics="true" pageId="22" pageNumber="423">Chromodoris kalawakan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912970BFFF8FF979D7EF9A13DE5AA4D" bold="true" box="[1081,1163,1616,1640]" pageId="22" pageNumber="423">
|
||
<taxonomicNameLabel id="A2212A70FFF8FF979D7EF9A13DE5AA4D" box="[1081,1163,1616,1640]" pageId="22" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, CASIZ 178300, Vanuatu. C. Radula.
|
||
<taxonomicName id="4C66309AFFF8FF97984FF9853B6CAAA9" authority="Bonomo & Gosliner, 2020" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[264,514,1652,1676]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="kalawakan" status="sp. nov.">
|
||
<emphasis id="B912970BFFF8FF97984FF9853B6CAAA9" box="[264,514,1652,1676]" italics="true" pageId="22" pageNumber="423">Chromodoris kalawakan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912970BFFF8FF979B40F9853B39AAA9" bold="true" box="[519,599,1652,1676]" pageId="22" pageNumber="423">
|
||
<taxonomicNameLabel id="A2212A70FFF8FF979B40F9853B39AAA9" box="[519,599,1652,1676]" pageId="22" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, CASIZ 224653, Philippines. D. Central teeth; E. Mid-lateral teeth; F. Outer lateral teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD94B19FFF8FF969980F92F3C50AD38" blockId="22.[151,1437,1758,2037]" lastBlockId="23.[151,1437,151,285]" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423">
|
||
Externally,
|
||
<taxonomicName id="4C66309AFFF8FF979800F92E388EAADD" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[327,480,1758,1785]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="kalawakan">
|
||
<emphasis id="B912970BFFF8FF979800F92E388EAADD" box="[327,480,1758,1785]" italics="true" pageId="22" pageNumber="423">C. kalawakan</emphasis>
|
||
</taxonomicName>
|
||
is the only species of
|
||
<taxonomicName id="4C66309AFFF8FF979B94F92F3A09AADD" authorityName="Alder & Hancock" authorityYear="1855" box="[723,871,1758,1784]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B912970BFFF8FF979B94F92F3A09AADD" box="[723,871,1758,1784]" italics="true" pageId="22" pageNumber="423">Chromodoris</emphasis>
|
||
</taxonomicName>
|
||
that is entirely devoid of black lines or dark brown to black spots. It shares having white spots on the rhinophores and gill with several other species including,
|
||
<taxonomicName id="4C66309AFFF8FF979C7FF8F338BAAB65" authority="Rudman, 1982" authorityName="Rudman" authorityYear="1982" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="willani">
|
||
<emphasis id="B912970BFFF8FF979C7FF8F33848AB65" italics="true" pageId="22" pageNumber="423">Chromodoris willani</emphasis>
|
||
<bibRefCitation id="EFF736E8FFF8FF97986BF8D738BAAB65" author="Rudman, W. B." box="[300,468,1830,1856]" pageId="22" pageNumber="423" pagination="183 - 241" refId="ref17581" refString="Rudman, W. B. (1982) The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris quadricolor, C. lineolata and Hypselodoris nigrolineata colour groups. Zoological Journal of the Linnean Society, 76, 183 - 241. https: // doi. org / 10.1111 / j. 1096 - 3642.1982. tb 02182. x" type="journal article" year="1982">Rudman, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66309AFFF8FF979898F8D63B0DAB65" baseAuthorityName="van Hasselt" baseAuthorityYear="1824" box="[479,611,1830,1857]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="lineolata">
|
||
<emphasis id="B912970BFFF8FF979898F8D63B0DAB65" box="[479,611,1830,1857]" italics="true" pageId="22" pageNumber="423">C. lineolata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66309AFFF8FF979B36F8D63B98AB65" authorityName="Bergh" authorityYear="1877" box="[625,758,1830,1857]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="striatella">
|
||
<emphasis id="B912970BFFF8FF979B36F8D63B98AB65" box="[625,758,1830,1857]" italics="true" pageId="22" pageNumber="423">C. striatella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66309AFFF8FF979A43F8D63AB6AB65" box="[772,984,1830,1857]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="mandapamensis">
|
||
<emphasis id="B912970BFFF8FF979A43F8D63AB6AB65" box="[772,984,1830,1857]" italics="true" pageId="22" pageNumber="423">C. mandapamensis</emphasis>
|
||
</taxonomicName>
|
||
, and six undescribed species. Internally, the Y-shaped mantle glands in
|
||
<taxonomicName id="4C66309AFFF8FF9798AEF8BA3BEFAB41" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[489,641,1866,1893]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="kalawakan">
|
||
<emphasis id="B912970BFFF8FF9798AEF8BA3BEFAB41" box="[489,641,1866,1893]" italics="true" pageId="22" pageNumber="423">C. kalawakan</emphasis>
|
||
</taxonomicName>
|
||
are distinct from the more highly ramified mantle glands of other
|
||
<taxonomicName id="4C66309AFFF8FF979C1DF8BB399DABAD" authorityName="Alder & Hancock" authorityYear="1855" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B912970BFFF8FF979C1DF8BB399DABAD" italics="true" pageId="22" pageNumber="423">Chromodoris</emphasis>
|
||
</taxonomicName>
|
||
species that have been examined (
|
||
<bibRefCitation id="EFF736E8FFF8FF979B28F89F3AFEABAD" author="Gosliner, T. M. & Behrens, D. W." box="[623,912,1902,1928]" pageId="22" pageNumber="423" pagination="139 - 165" refId="ref15903" refString="Gosliner, T. M. & Behrens, D. W. (1998) Five new species of Chromodoris (Mollusca: Nudibranchia: Chromodorididae) from the tropical Indo-Pacific Ocean. Proceedings of the California Academy of Sciences, 50, 139 - 165." type="journal article" year="1998">Gosliner & Behrens 1998</bibRefCitation>
|
||
). The radula is smaller and contains fewer rows of radular teeth than do other species of
|
||
<emphasis id="B912970BFFF8FF979B10F8633B9CAB89" box="[599,754,1938,1964]" italics="true" pageId="22" pageNumber="423">
|
||
<taxonomicName id="4C66309AFFF8FF979B10F8633B80AB89" authorityName="Alder & Hancock" authorityYear="1855" box="[599,750,1938,1964]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="genus">Chromodoris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with a maximum of 43 rows of teeth. The rachidian teeth are short and narrow with a linear shape as in most other species of
|
||
<taxonomicName id="4C66309AFFF8FF979A2EF8473A93ABF5" authorityName="Alder & Hancock" authorityYear="1855" box="[873,1021,1974,2000]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B912970BFFF8FF979A2EF8473A93ABF5" box="[873,1021,1974,2000]" italics="true" pageId="22" pageNumber="423">Chromodoris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFF736E8FFF8FF979D4BF8473DC1ABF5" author="Rudman, W. B." box="[1036,1199,1974,2000]" pageId="22" pageNumber="423" pagination="183 - 241" refId="ref17581" refString="Rudman, W. B. (1982) The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris quadricolor, C. lineolata and Hypselodoris nigrolineata colour groups. Zoological Journal of the Linnean Society, 76, 183 - 241. https: // doi. org / 10.1111 / j. 1096 - 3642.1982. tb 02182. x" type="journal article" year="1982">Rudman 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF736E8FFF8FF979DFBF84739BCABD1" author="Gosliner, T. M. & Behrens, D. W." pageId="22" pageNumber="423" pagination="139 - 165" refId="ref15903" refString="Gosliner, T. M. & Behrens, D. W. (1998) Five new species of Chromodoris (Mollusca: Nudibranchia: Chromodorididae) from the tropical Indo-Pacific Ocean. Proceedings of the California Academy of Sciences, 50, 139 - 165." type="journal article" year="1998">Gosliner & Behrens 1998</bibRefCitation>
|
||
), whereas a rachidian row of teeth appears to be absent in
|
||
<taxonomicName id="4C66309AFFF8FF979A32F82B3D94ABD1" authority="Eliot, 1904" authorityName="Eliot" authorityYear="1904" box="[885,1274,2010,2036]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="africana">
|
||
<emphasis id="B912970BFFF8FF979A32F82B3D1FABD1" box="[885,1137,2010,2036]" italics="true" pageId="22" pageNumber="423">Chromodoris africana</emphasis>
|
||
<bibRefCitation id="EFF736E8FFF8FF979D3EF82B3D94ABD1" author="Eliot, C. N. E." box="[1145,1274,2010,2036]" pageId="22" pageNumber="423" pagination="380 - 406" refId="ref15510" refString="Eliot, C. N. E. (1904) On some nudibranchs from East Africa and Zanzibar. Part IV. Proceedings of the Zoological Society of London, 1904, 380 - 406. https: // doi. org / 10.1111 / j. 1469 - 7998.1904. tb 08295. x" type="journal article" year="1904">Eliot, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C66309AFFF8FF969C4FF82B38FEAC94" authority="Rudman, 1982" authorityName="Rudman" authorityYear="1982" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" lastPageId="23" lastPageNumber="424" order="Nudibranchia" pageId="22" pageNumber="423" phylum="Mollusca" rank="species" species="kuiteri">
|
||
<emphasis id="B912970BFFF8FF969C4FF82B398EAC94" italics="true" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423">Chromodoris kuiteri</emphasis>
|
||
<bibRefCitation id="EFF736E8FFF9FF9699AFFF6638FEAC94" author="Rudman, W. B." box="[232,400,151,177]" pageId="23" pageNumber="424" pagination="183 - 241" refId="ref17581" refString="Rudman, W. B. (1982) The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris quadricolor, C. lineolata and Hypselodoris nigrolineata colour groups. Zoological Journal of the Linnean Society, 76, 183 - 241. https: // doi. org / 10.1111 / j. 1096 - 3642.1982. tb 02182. x" type="journal article" year="1982">Rudman, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C66309AFFF9FF96988AFF663ABFAC94" authority="Gosliner & Behrens, 1998" authorityName="Gosliner & Behrens" authorityYear="1998" box="[461,977,151,177]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="424" phylum="Mollusca" rank="species" species="joshi">
|
||
<emphasis id="B912970BFFF9FF96988AFF663BF1AC94" box="[461,671,151,177]" italics="true" pageId="23" pageNumber="424">Chromodoris joshi</emphasis>
|
||
<bibRefCitation id="EFF736E8FFF9FF969BE0FF663ABFAC94" author="Gosliner, T. M. & Behrens, D. W." box="[679,977,151,177]" pageId="23" pageNumber="424" pagination="139 - 165" refId="ref15903" refString="Gosliner, T. M. & Behrens, D. W. (1998) Five new species of Chromodoris (Mollusca: Nudibranchia: Chromodorididae) from the tropical Indo-Pacific Ocean. Proceedings of the California Academy of Sciences, 50, 139 - 165." type="journal article" year="1998">Gosliner & Behrens, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C66309AFFF9FF969A9AFF663DA9AC94" authorityName="Rudman" authorityYear="1982" box="[989,1223,151,177]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="424" phylum="Mollusca" rank="species" species="willani">
|
||
<emphasis id="B912970BFFF9FF969A9AFF663DA9AC94" box="[989,1223,151,177]" italics="true" pageId="23" pageNumber="424">Chromodoris willani</emphasis>
|
||
</taxonomicName>
|
||
is the only species that has a well-developed rachidian row of teeth that are large and triangular (
|
||
<bibRefCitation id="EFF736E8FFF9FF969D52FF4A3DD5ACF0" author="Rudman, W. B." box="[1045,1211,187,213]" pageId="23" pageNumber="424" pagination="183 - 241" refId="ref17581" refString="Rudman, W. B. (1982) The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris quadricolor, C. lineolata and Hypselodoris nigrolineata colour groups. Zoological Journal of the Linnean Society, 76, 183 - 241. https: // doi. org / 10.1111 / j. 1096 - 3642.1982. tb 02182. x" type="journal article" year="1982">Rudman 1982</bibRefCitation>
|
||
). Like most species of
|
||
<taxonomicName id="4C66309AFFF9FF9699F0FF2E3825ACDC" authorityName="Alder & Hancock" authorityYear="1855" box="[183,331,223,249]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="424" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B912970BFFF9FF9699F0FF2E3825ACDC" box="[183,331,223,249]" italics="true" pageId="23" pageNumber="424">Chromodoris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66309AFFF9FF96981DFF2E389BACDC" authorityName="Bonomo & Gosliner" authorityYear="2020" box="[346,501,223,249]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="424" phylum="Mollusca" rank="species" species="kalawakan">
|
||
<emphasis id="B912970BFFF9FF96981DFF2E389BACDC" box="[346,501,223,249]" italics="true" pageId="23" pageNumber="424">C. kalawakan</emphasis>
|
||
</taxonomicName>
|
||
has a long prostatic portion of the vas deferens and short ejaculatory portion. The ABGD and bPTP analyses distinctive coloration of this species clearly differentiate it as a distinct species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |