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<document id="DC9C8451DF067C1373728287C9D957DC" ID-DOI="10.11646/zootaxa.5315.5.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8142393" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689236949496" checkinUser="plazi" docAuthor="Tuzovskij, Petr V." docDate="2023" docId="03CF072BFFD0FFA7EAFAFA9D29D36995" docLanguage="en" docName="zootaxa.5315.5.6.pdf" docOrigin="Zootaxa 5315 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.5.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebertia (Pilolebertia) diakonovoensis Tuzovskij 2023, sp.n." docType="treatment" docVersion="2" lastPageNumber="490" masterDocId="FFF67F53FFD0FFA2EA6DFFAD28056E43" masterDocTitle="A new water mite species of the genus Lebertia Neuman, 1880 (Acari, Hydrachnidae: Lebertiidae) from Yaroslavl Province of Russia" masterLastPageNumber="491" masterPageNumber="485" pageNumber="485" updateTime="1689237330206" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C1D9780768600AC821F6635F5BD0E830">A new water mite species of the genus Lebertia Neuman, 1880 (Acari, Hydrachnidae: Lebertiidae) from Yaroslavl Province of Russia</mods:title>
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<mods:namePart id="5D19074B7AD5B5FAF902AFA74064AC22">Tuzovskij, Petr V.</mods:namePart>
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<mods:date id="7508F245055FD0C15FD2EF330F3AAF43">2023</mods:date>
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<mods:identifier id="6F6030C741057BD5170F05046762C363" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CF072BFFD0FFA7EAFAFA9D29D36995" LSID="urn:lsid:plazi:treatment:03CF072BFFD0FFA7EAFAFA9D29D36995" httpUri="http://treatment.plazi.org/id/03CF072BFFD0FFA7EAFAFA9D29D36995" lastPageId="5" lastPageNumber="490" pageId="0" pageNumber="485">
<subSubSection id="C37CE5B6FFD0FFA2EAFAFA9D2AA26B08" box="[151,679,1328,1355]" pageId="0" pageNumber="485" type="nomenclature">
<paragraph id="8BD9B63DFFD0FFA2EAFAFA9D2AA26B08" blockId="0.[151,1437,1328,1998]" box="[151,679,1328,1355]" pageId="0" pageNumber="485">
<heading id="D0910151FFD0FFA2EAFAFA9D2AA26B08" bold="true" box="[151,679,1328,1355]" fontSize="11" level="1" pageId="0" pageNumber="485" reason="1">
<emphasis id="B9126A2FFFD0FFA2EAFAFA9D2AA26B08" bold="true" box="[151,679,1328,1355]" pageId="0" pageNumber="485">
<taxonomicName id="4C66CDBEFFD0FFA2EAFAFA9D2A606B08" authority="Tuzovskij, 2023" authorityName="Tuzovskij" authorityYear="2023" box="[151,613,1328,1355]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="0" pageNumber="485" phylum="Arthropoda" rank="species" species="diakonovoensis" status="sp.n." subGenus="Pilolebertia">
<emphasis id="B9126A2FFFD0FFA2EAFAFA9D28F96B08" bold="true" box="[151,252,1328,1355]" italics="true" pageId="0" pageNumber="485">Lebertia</emphasis>
(
<emphasis id="B9126A2FFFD0FFA2EB60FA9D299C6B08" bold="true" box="[269,409,1328,1355]" italics="true" pageId="0" pageNumber="485">Pilolebertia</emphasis>
)
<emphasis id="B9126A2FFFD0FFA2EBC6FA9D2A606B08" bold="true" box="[427,613,1328,1355]" italics="true" pageId="0" pageNumber="485">diakonovoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D754FFD0FFA2E801FA9C2AA26B08" box="[620,679,1329,1355]" pageId="0" pageNumber="485" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37CE5B6FFD0FFA7EAFAFAFC29D36995" lastPageId="5" lastPageNumber="490" pageId="0" pageNumber="485" type="description">
<paragraph id="8BD9B63DFFD0FFA2EAFAFAFC29196B2A" blockId="0.[151,1437,1328,1998]" box="[151,284,1361,1385]" pageId="0" pageNumber="485">
<figureCitation id="135DAAB8FFD0FFA2EAFAFAFC29196B2A" box="[151,284,1361,1385]" captionStart-0="FIGURES 17" captionStart-1="FIGURES 812" captionStart-2="FIGURE 1315" captionStart-3="FIGURES 1619" captionStartId-0="1.[151,264,1935,1959]" captionStartId-1="2.[151,264,1943,1967]" captionStartId-2="3.[151,250,1927,1951]" captionStartId-3="4.[151,264,1502,1526]" captionTargetBox-0="[168,1436,199,1899]" captionTargetBox-1="[177,1414,201,1914]" captionTargetBox-2="[156,1427,186,1899]" captionTargetBox-3="[156,1431,186,1473]" captionTargetId-0="figure-15@1.[151,1436,180,1910]" captionTargetId-1="figure-29@2.[166,1422,181,1919]" captionTargetId-2="figure-15@3.[151,1436,181,1904]" captionTargetId-3="figure-29@4.[151,1436,181,1478]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." captionText-1="FIGURES 812. Lebertia diakonovoensis sp. n., male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm." captionText-2="FIGURE 1315. Lebertia diakonovoensis sp. n., deutonymph: 13—seta Fch; 14—coxal shield; 15—pedipalp. Scale bar: 13, 15 = 50 μm, 14 = 100 μm." captionText-3="FIGURES 1619. Lebertia diakonovoensis sp. n., deutonymph: 16—chelicera; 17—capitulum, lateral view, 18—genital field, 19—IV-Leg-46. Scale bars: 1618 = 50 μm; 19 = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.8142395" figureDoi-1="http://doi.org/10.5281/zenodo.8142397" figureDoi-2="http://doi.org/10.5281/zenodo.8142399" figureDoi-3="http://doi.org/10.5281/zenodo.8142401" httpUri-0="https://zenodo.org/record/8142395/files/figure.png" httpUri-1="https://zenodo.org/record/8142397/files/figure.png" httpUri-2="https://zenodo.org/record/8142399/files/figure.png" httpUri-3="https://zenodo.org/record/8142401/files/figure.png" pageId="0" pageNumber="485">Figures 119</figureCitation>
</paragraph>
<paragraph id="8BD9B63DFFD0FFA2EAFAFADB2D6B6B96" blockId="0.[151,1437,1328,1998]" pageId="0" pageNumber="485">
Type series:
<materialsCitation id="3B0EBC60FFD0FFA2EB71FAD82BC06BF2" collectingDate="2002-06-01" collectorName="P. Tuzovskij" country="Russia" county="Europe" location="Diakonovo" municipality="Nekouz District" pageId="0" pageNumber="485" specimenCount="1" specimenCount-female="1" stateProvince="Yaroslavl'" typeStatus="holotype">
<typeStatus id="54DD089FFFD0FFA2EB71FAD8297A6BCE" box="[284,383,1397,1421]" pageId="0" pageNumber="485">Holotype</typeStatus>
<specimenCount id="9D607DB4FFD0FFA2EBE8FADB299C6BCE" box="[389,409,1398,1421]" pageId="0" pageNumber="485" type="female"></specimenCount>
, slide 8544,
<collectingCounty id="62B8CEB1FFD0FFA2E84DFADB2A756BCE" box="[544,624,1398,1421]" pageId="0" pageNumber="485">Europe</collectingCounty>
,
<collectingCountry id="F371F6ADFFD0FFA2E817FAD82AC16BCE" box="[634,708,1397,1421]" name="Russia" pageId="0" pageNumber="485">Russia</collectingCountry>
,
<collectingRegion id="49A278DFFFD0FFA2E8A0FAD82B9F6BCE" box="[717,922,1397,1421]" country="Russia" name="Yaroslavl'" pageId="0" pageNumber="485">Yaroslavl Province</collectingRegion>
,
<collectingMunicipality id="6BBD2C47FFD0FFA2E9CEFAD82C4A6BCE" box="[931,1103,1397,1421]" pageId="0" pageNumber="485">Nekouz District</collectingMunicipality>
, brook near village
<location id="8EB9E0E6FFD0FFA2EF72FAD82D9D6BCE" LSID="urn:lsid:plazi:treatment:03CF072BFFD0FFA7EAFAFA9D29D36995:8EB9E0E6FFD0FFA2EF72FAD82D9D6BCE" box="[1311,1432,1397,1421]" country="Russia" county="Europe" municipality="Nekouz District" name="Diakonovo" pageId="0" pageNumber="485" stateProvince="Yaroslavl'">Diakonovo</location>
, depth 0.20.5 m, substrate: slit, detritus, stones,
<date id="FFD890FDFFD0FFA2E8F4FA342B086BF2" box="[665,781,1433,1457]" pageId="0" pageNumber="485" value="2002-06-01">
<collectingDate id="EF9C6915FFD0FFA2E8F4FA342B086BF2" box="[665,781,1433,1457]" pageId="0" pageNumber="485" value="2002-06-01">1.06. 2002</collectingDate>
</date>
, leg
<collectorName id="2693D3EBFFD0FFA2E952FA372BC06BF2" box="[831,965,1433,1457]" pageId="0" pageNumber="485">P. Tuzovskij</collectorName>
</materialsCitation>
.
<materialsCitation id="3B0EBC60FFD0FFA2E9A2FA372A7F6B96" collectingDate="2002-06-16" collectingDateMax="2005-08-08" collectingDateMin="2002-06-16" collectorName="P. Tuzovskij" country="Russia" county="Europe" location="Diakonovo" municipality="Nekouz District" pageId="0" pageNumber="485" specimenCount="4" specimenCount-female="4" stateProvince="Yaroslavl'" typeStatus="paratype">
<typeStatus id="54DD089FFFD0FFA2E9A2FA372C3E6BF1" box="[975,1083,1434,1458]" pageId="0" pageNumber="485">Paratypes</typeStatus>
: same locality as holotype:
<specimenCount id="9D607DB4FFD0FFA2EF0DFA342D996BF2" box="[1376,1436,1433,1457]" pageId="0" pageNumber="485" type="female">2 ♀♀</specimenCount>
<date id="FFD890FDFFD0FFA2EAFAFA10291A6B96" box="[151,287,1469,1493]" pageId="0" pageNumber="485" value="2002-06-16">
<collectingDate id="EF9C6915FFD0FFA2EAFAFA10291A6B96" box="[151,287,1469,1493]" pageId="0" pageNumber="485" value="2002-06-16">16. 06. 2002</collectingDate>
</date>
,
<specimenCount id="9D607DB4FFD0FFA2EB47FA1029576B96" box="[298,338,1469,1493]" pageId="0" pageNumber="485" type="female">1 ♀</specimenCount>
<date id="FFD890FDFFD0FFA2EB34FA1029D66B96" box="[345,467,1469,1493]" pageId="0" pageNumber="485" value="2002-07-02">
<collectingDate id="EF9C6915FFD0FFA2EB34FA1029D66B96" box="[345,467,1469,1493]" pageId="0" pageNumber="485" value="2002-07-02">2. 07. 2002</collectingDate>
</date>
,
<specimenCount id="9D607DB4FFD0FFA2EBB3FA102A036B96" box="[478,518,1469,1493]" pageId="0" pageNumber="485" type="female">1 ♀</specimenCount>
<date id="FFD890FDFFD0FFA2E860FA102A7F6B96" box="[525,634,1469,1493]" pageId="0" pageNumber="485" value="2005-08-08">
<collectingDate id="EF9C6915FFD0FFA2E860FA102A7F6B96" box="[525,634,1469,1493]" pageId="0" pageNumber="485" value="2005-08-08">8.08.2005</collectingDate>
</date>
</materialsCitation>
, 1 ♁
<date id="FFD890FDFFD0FFA2E8DEFA102B246B96" box="[691,801,1469,1493]" pageId="0" pageNumber="485" value="2005-08-09">9.08.2005</date>
, 4 dn
<date id="FFD890FDFFD0FFA2E90CFA102BD96B96" box="[865,988,1469,1493]" pageId="0" pageNumber="485" value="2002-05-26">26.05.2002</date>
, 2 dn
<date id="FFD890FDFFD0FFA2EE76FA102C836B96" box="[1051,1158,1469,1493]" pageId="0" pageNumber="485" value="2002-06-01">1.06.2002</date>
and 1 dn
<date id="FFD890FDFFD0FFA2EE82FA102D6F6B96" box="[1263,1386,1469,1493]" pageId="0" pageNumber="485" value="2002-12-07">12.07.2002</date>
.
</paragraph>
<paragraph id="8BD9B63DFFD0FFA2EAAAFA4F2A8968CA" blockId="0.[151,1437,1328,1998]" pageId="0" pageNumber="485">
<emphasis id="B9126A2FFFD0FFA2EAAAFA4F29366BBA" box="[199,307,1506,1529]" italics="true" pageId="0" pageNumber="485">Diagnosis</emphasis>
. Integument thin and smooth, with thin lines locally visible, setae
<emphasis id="B9126A2FFFD0FFA2EE61FA4C2C306BBA" box="[1036,1077,1505,1529]" italics="true" pageId="0" pageNumber="485">Fch</emphasis>
with short proximal furca; coxal shield wider than long; P-2 ventrodistal seta thin; P-3 mediodistal seta halfway between ventro- and dorsodistal setae or slightly towards dorsodistal seta; P-4 slender, tapering distally, both ventral setae shifted to the distal half of segment, ventral margin divided by seta insertions into sectors 3: 2: 1, mediodistal peg-like seta short with rounded tip; IV-Leg-6 with four to five short, spine-like ventral setae.
</paragraph>
<paragraph id="8BD9B63DFFD0FFA2EAAAF93B2D95698E" blockId="0.[151,1437,1328,1998]" pageId="0" pageNumber="485">
<emphasis id="B9126A2FFFD0FFA2EAAAF93B29CD68EE" box="[199,456,1685,1709]" italics="true" pageId="0" pageNumber="485">Description. Both sexes.</emphasis>
Colour brown. Integument soft and smooth, with short, thin lines which are especially clearly visible along the idiosoma lateral margins (
<figureCitation id="135DAAB8FFD0FFA2E809F9142AA76892" box="[612,674,1721,1745]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="0" pageNumber="485">Fig. 1</figureCitation>
). Setae
<emphasis id="B9126A2FFFD0FFA2E894F9142B276892" box="[761,802,1721,1745]" italics="true" pageId="0" pageNumber="485">Fch</emphasis>
(
<figureCitation id="135DAAB8FFD0FFA2E95CF9142B746892" box="[817,881,1721,1745]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="0" pageNumber="485">Fig. 2</figureCitation>
) longer and thicker than other idiosomal setae, with short proximal furca. Trichobothria
<emphasis id="B9126A2FFFD0FFA2E873F9732A6368B7" box="[542,614,1757,1781]" italics="true" pageId="0" pageNumber="485">Fp, Oi</emphasis>
and setae
<emphasis id="B9126A2FFFD0FFA2E8B6F9732AF668B6" box="[731,755,1758,1781]" italics="true" pageId="0" pageNumber="485">Pi</emphasis>
not associated with glandularia (
<figureCitation id="135DAAB8FFD0FFA2EE30F9702C9B68B6" box="[1117,1182,1757,1781]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="0" pageNumber="485">Fig. 3</figureCitation>
), other idiosomal setae associated with glandularia (
<figureCitation id="135DAAB8FFD0FFA2EBA3F8AC2A17695A" box="[462,530,1793,1817]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="0" pageNumber="485">Fig. 4</figureCitation>
). Coxal shield (
<figureCitation id="135DAAB8FFD0FFA2E8ADF8AC2B07695A" box="[704,770,1793,1817]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="0" pageNumber="485">Fig. 5</figureCitation>
) wider than long (L/W ratio 0.83-0.85). Coxal plates I fused to each other nearly complete, fragment of suture line present only in their posterior portion. Suture line between coxal plates I longer than median length of coxal plates II (Cx-I/Cx II ratio mL 1.36-1.95). Coxal shield bearing two pairs of glandularia (
<emphasis id="B9126A2FFFD0FFA2EB73F8C3293869C6" box="[286,317,1902,1925]" italics="true" pageId="0" pageNumber="485">Hv</emphasis>
and
<emphasis id="B9126A2FFFD0FFA2EB1DF8C3298969C6" box="[368,396,1902,1925]" italics="true" pageId="0" pageNumber="485">Pe</emphasis>
),
<emphasis id="B9126A2FFFD0FFA2EBCFF8C329BB69C6" box="[418,446,1902,1925]" italics="true" pageId="0" pageNumber="485">Pe</emphasis>
located near tips of coxal plates I. Coxal plates II with narrow posterior margin. Genital field with three pairs of unequal acetabula, posterior pair of acetabula smaller than both anterior pairs of acetabula. All coxal plates porous, in additional, coxal plates IV with several relatively large oval cells each. Excretory pore not sclerotized.
</paragraph>
<caption id="DF19E6B5FFD1FFA3EAFAF8222CBD698F" ID-DOI="http://doi.org/10.5281/zenodo.8142395" ID-Zenodo-Dep="8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="1" pageNumber="486" startId="1.[151,264,1935,1959]" targetBox="[168,1436,199,1899]" targetPageId="1" targetType="figure">
<paragraph id="8BD9B63DFFD1FFA3EAFAF8222CBD698F" blockId="1.[151,1436,1935,1996]" pageId="1" pageNumber="486">
<emphasis id="B9126A2FFFD1FFA3EAFAF822293C69E4" bold="true" box="[151,313,1935,1959]" pageId="1" pageNumber="486">FIGURES 17.</emphasis>
<taxonomicName id="4C66CDBEFFD1FFA3EB52F8222A3369E4" authority="Tuzovskij, 2023" authorityName="Tuzovskij" authorityYear="2023" box="[319,566,1935,1959]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="486" phylum="Arthropoda" rank="species" species="diakonovoensis" status="sp. n.">
<emphasis id="B9126A2FFFD1FFA3EB52F8222A3369E4" box="[319,566,1935,1959]" italics="true" pageId="1" pageNumber="486">Lebertia diakonovoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D754FFD1FFA3E856F83D2A7769E4" box="[571,626,1936,1959]" pageId="1" pageNumber="486" rank="species">sp. n.</taxonomicNameLabel>
, male: 1—detail of integument; 2—seta
<emphasis id="B9126A2FFFD1FFA3EE65F8222C2A69E4" box="[1032,1071,1935,1959]" italics="true" pageId="1" pageNumber="486">Fch</emphasis>
; 3—seta
<emphasis id="B9126A2FFFD1FFA3EEE3F83D2CA269E4" box="[1166,1191,1936,1959]" italics="true" pageId="1" pageNumber="486">Oi</emphasis>
; 4—seta
<emphasis id="B9126A2FFFD1FFA3EF6BF83D2D2169E4" box="[1286,1316,1936,1959]" italics="true" pageId="1" pageNumber="486">Oe</emphasis>
; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm.
</paragraph>
</caption>
<caption id="DF19E6B5FFD2FFA0EAFAF83A2C806997" ID-DOI="http://doi.org/10.5281/zenodo.8142397" ID-Zenodo-Dep="8142397" httpUri="https://zenodo.org/record/8142397/files/figure.png" pageId="2" pageNumber="487" startId="2.[151,264,1943,1967]" targetBox="[177,1414,201,1914]" targetPageId="2" targetType="figure">
<paragraph id="8BD9B63DFFD2FFA0EAFAF83A2C806997" blockId="2.[151,1436,1943,2004]" pageId="2" pageNumber="487">
<emphasis id="B9126A2FFFD2FFA0EAFAF83A294D69EC" bold="true" box="[151,328,1943,1967]" pageId="2" pageNumber="487">FIGURES 812.</emphasis>
<taxonomicName id="4C66CDBEFFD2FFA0EB23F8352A4269F3" authority="Tuzovskij, 2023" authorityName="Tuzovskij" authorityYear="2023" box="[334,583,1944,1968]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="487" phylum="Arthropoda" rank="species" species="diakonovoensis" status="sp. n.">
<emphasis id="B9126A2FFFD2FFA0EB23F8352A4269F3" box="[334,583,1944,1968]" italics="true" pageId="2" pageNumber="487">Lebertia diakonovoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D754FFD2FFA0E823F8352A8369EC" box="[590,646,1944,1967]" pageId="2" pageNumber="487" rank="species">sp. n.</taxonomicNameLabel>
, male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm.
</paragraph>
</caption>
<caption id="DF19E6B5FFD3FFA1EAFAF82A29A76987" ID-DOI="http://doi.org/10.5281/zenodo.8142399" ID-Zenodo-Dep="8142399" httpUri="https://zenodo.org/record/8142399/files/figure.png" pageId="3" pageNumber="488" startId="3.[151,250,1927,1951]" targetBox="[156,1427,186,1899]" targetPageId="3" targetType="figure">
<paragraph id="8BD9B63DFFD3FFA1EAFAF82A29A76987" blockId="3.[151,1437,1927,1988]" pageId="3" pageNumber="488">
<emphasis id="B9126A2FFFD3FFA1EAFAF82A294269E3" bold="true" box="[151,327,1927,1952]" pageId="3" pageNumber="488">FIGURE 1315.</emphasis>
<taxonomicName id="4C66CDBEFFD3FFA1EB22F8252A4D69E3" authority="Tuzovskij, 2023" authorityName="Tuzovskij" authorityYear="2023" box="[335,584,1928,1952]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="diakonovoensis" status="sp. n.">
<emphasis id="B9126A2FFFD3FFA1EB22F8252A4D69E3" box="[335,584,1928,1952]" italics="true" pageId="3" pageNumber="488">Lebertia diakonovoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D754FFD3FFA1E83DF8252A8C69DC" box="[592,649,1928,1951]" pageId="3" pageNumber="488" rank="species">sp. n.</taxonomicNameLabel>
, deutonymph: 13—seta
<emphasis id="B9126A2FFFD3FFA1E9E8F8252BA969E3" box="[901,940,1928,1952]" italics="true" pageId="3" pageNumber="488">Fch</emphasis>
; 14—coxal shield; 15—pedipalp. Scale bar: 13, 15 = 50 μm, 14 = 100 μm.
</paragraph>
</caption>
<caption id="DF19E6B5FFD4FFA6EAFAFA732AF16858" ID-DOI="http://doi.org/10.5281/zenodo.8142401" ID-Zenodo-Dep="8142401" httpUri="https://zenodo.org/record/8142401/files/figure.png" pageId="4" pageNumber="489" startId="4.[151,264,1502,1526]" targetBox="[156,1431,186,1473]" targetPageId="4" targetType="figure">
<paragraph id="8BD9B63DFFD4FFA6EAFAFA732AF16858" blockId="4.[151,1436,1502,1563]" pageId="4" pageNumber="489">
<emphasis id="B9126A2FFFD4FFA6EAFAFA7329516BB5" bold="true" box="[151,340,1502,1526]" pageId="4" pageNumber="489">FIGURES 1619.</emphasis>
<taxonomicName id="4C66CDBEFFD4FFA6EB37FA732A576BB5" authority="Tuzovskij, 2023" authorityName="Tuzovskij" authorityYear="2023" box="[346,594,1502,1526]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="diakonovoensis" status="sp. n.">
<emphasis id="B9126A2FFFD4FFA6EB37FA732A576BB5" box="[346,594,1502,1526]" italics="true" pageId="4" pageNumber="489">Lebertia diakonovoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D754FFD4FFA6E834FA722A956BB5" box="[601,656,1503,1526]" pageId="4" pageNumber="489" rank="species">sp. n.</taxonomicNameLabel>
, deutonymph: 16—chelicera; 17—capitulum, lateral view, 18—genital field, 19—IV-Leg-46. Scale bars: 1618 = 50 μm; 19 = 100 μm.
</paragraph>
</caption>
<paragraph id="8BD9B63DFFD4FFA6EAAAF9E7296A6979" blockId="4.[151,1437,1610,2030]" pageId="4" pageNumber="489">
Pedipalp (
<figureCitation id="135DAAB8FFD4FFA6EB59F9E729726821" box="[308,375,1610,1634]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="4" pageNumber="489">Fig. 6</figureCitation>
): P-1 short, with single short dorsodistal setae; P-2 large, with concave ventral margin, three short unequal dorsoproximal and two long subequal dorsodistal setae, ventrodistal seta thin, longer than ventral margin but shorter than dorsal one (occasionally longer of segment), inserted slightly away from ventrodistal segment edge; P-3 rather long, with five long setae, bases of two proximal a little separated, mediodistal seta halfway between ventro- and dorsodistal setae or slightly towards dorsodistal seta; P-4 slender, tapering distally, both ventral setae shifted to the distal half of the segment, ventral margin divided by seta insertions into sectors 3: 2: 1, mediodistal peg-like seta short with rounded tip (
<figureCitation id="135DAAB8FFD4FFA6EB4DF88F29676979" box="[288,354,1826,1850]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="4" pageNumber="489">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BD9B63DFFD4FFA6EAAAF8EB2C0269E5" blockId="4.[151,1437,1610,2030]" pageId="4" pageNumber="489">
Gnathosoma ventral margin strong convex with posterior indentation, posterodorsal projections well developed and pointed, rostrum short, mouth opening flanking by two pairs of short equal setae (
<figureCitation id="135DAAB8FFD4FFA6EE6BF8C72C4269C1" box="[1030,1095,1898,1922]" captionStart="FIGURES 812" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[177,1414,201,1914]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 812. Lebertia diakonovoensis sp. n., male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.8142397" httpUri="https://zenodo.org/record/8142397/files/figure.png" pageId="4" pageNumber="489">Fig. 8</figureCitation>
). Chelicera basal segment large and thickened distally, chela short with two rows of fine teeth on the concave side.
</paragraph>
<paragraph id="8BD9B63DFFD4FFA7EAAAF81E2AA36E90" blockId="4.[151,1437,1610,2030]" lastBlockId="5.[151,1437,151,2007]" lastPageId="5" lastPageNumber="490" pageId="4" pageNumber="489">
Legs II-IV with the following number of swimming setae: four to six on II-Leg-5, five to six on III-Leg-4, 6 to 10 on III-Leg-5, four to eight on IV-Leg-4, 8 to 12 on IV-Leg-5 (
<figureCitation id="135DAAB8FFD4FFA6E946F87B2B6869AD" box="[811,877,2006,2030]" captionStart="FIGURES 812" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[177,1414,201,1914]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 812. Lebertia diakonovoensis sp. n., male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.8142397" httpUri="https://zenodo.org/record/8142397/files/figure.png" pageId="4" pageNumber="489">Fig. 9</figureCitation>
). IV-Leg-6 with four to five (occasionally with two) short, spine-like ventral setae. Leg claws with short internal and comparatively long external clawlets, lamella moderately developed with concave ventral margin (
<figureCitation id="135DAAB8FFD5FFA7E827FF162A9F6E90" box="[586,666,187,211]" captionStart="FIGURES 812" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[177,1414,201,1914]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 812. Lebertia diakonovoensis sp. n., male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.8142397" httpUri="https://zenodo.org/record/8142397/files/figure.png" pageId="5" pageNumber="490">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFF732A476F7F" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAFF73291A6EB5" box="[199,287,222,246]" italics="true" pageId="5" pageNumber="490">Female.</emphasis>
Posterior margin of coxal plates IV not extending to the posterior margin of genital field, with widely rounded posteromedial edges. Genital field with 13-15 medial and three to four lateral setae, anterior genital sclerite with short posteromedian projection (
<figureCitation id="135DAAB8FFD5FFA7EB8AFE892A306F7F" box="[487,565,292,316]" captionStart="FIGURES 812" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[177,1414,201,1914]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 812. Lebertia diakonovoensis sp. n., male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.8142397" httpUri="https://zenodo.org/record/8142397/files/figure.png" pageId="5" pageNumber="490">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFEE52B766C4C" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">Measurements (n=5). Idiosoma L 1160-1250; coxal shield L 665-740, W 800-875; coxal plate I mL 155-185, coxal plate II mL 95-115; genital flap L 185-195, W 72-85, genital acetabula (ac-1-3) L 65-70, 55-65, 45-55; cheliceral segments L: base 210-228, chela 36-42; pedipalp segments (P15) L/H: 30-36/0.48-0.50, 100-120/60-75, 100-115/42-54, 140-145/42-45, 20-25/15-18; leg segments L: I-Leg-16: 60-70, 78-100, 85-95, 125-145, 145-165, 130-145; IILeg-16: 65- 80, 95-120, 98-110, 150-165, 180-205, 180-195; IIILeg-16: 70-85, 115-180, 125-150-150, 200-240, 240-270, 220-240; IVLeg-16: 140-175, 130-150, 180-210, 260-285, 270-380, 235-270.</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFDB629F86C15" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAFDB628F96C70" box="[199,252,539,563]" italics="true" pageId="5" pageNumber="490">Male</emphasis>
. Genital field with about 30 pairs of medial and six to seven pairs of lateral setae, anterior genital sclerite with straight posterior margin (
<figureCitation id="135DAAB8FFD5FFA7EBC9FD9329F46C15" box="[420,497,574,598]" captionStart="FIGURES 812" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[177,1414,201,1914]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 812. Lebertia diakonovoensis sp. n., male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.8142397" httpUri="https://zenodo.org/record/8142397/files/figure.png" pageId="5" pageNumber="490">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFDCF29CD6D45" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">Measurements (n=1). Idiosoma L 1125; coxal shield L 725, W 775; coxal plate I mL 185, coxal plate II mL 135; genital flap L 200, W 87, genital acetabula (ac-1-3) L 50-60, 50-60, 30-37; cheliceral segments L: base 235, chela 62; pedip[alp total L 425; pedipalp segments (P15) L/H,: 37/50, 112/70, 100/45, 150/35, 25/15, leg segments L: Leg-16: 74, 100, 95, 150, 150, 150; IILeg-16: 75, 110, 115, 170, 200, 195; IIILeg-16: 85, 125, 115, 225, 275, 250; IVLeg-16: 165, 150, 200, 275, 290, 270.</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFCBC29D56A00" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAFCBC29526D6A" box="[199,343,785,809]" italics="true" pageId="5" pageNumber="490">Deutonymph.</emphasis>
Dorsal surface, structure of integument and capitulum similar to that of adults. Setae
<emphasis id="B9126A2FFFD5FFA7EF6FFCBC2D2E6D6A" box="[1282,1323,785,809]" italics="true" pageId="5" pageNumber="490">Fch</emphasis>
long, thin with short proximal furca (
<figureCitation id="135DAAB8FFD5FFA7EBD7FC992A0E6D0F" box="[442,523,820,844]" captionStart="FIGURE 1315" captionStartId="3.[151,250,1927,1951]" captionTargetBox="[156,1427,186,1899]" captionTargetId="figure-15@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1315. Lebertia diakonovoensis sp. n., deutonymph: 13—seta Fch; 14—coxal shield; 15—pedipalp. Scale bar: 13, 15 = 50 μm, 14 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142399" httpUri="https://zenodo.org/record/8142399/files/figure.png" pageId="5" pageNumber="490">Fig. 13</figureCitation>
). Coxal shield wider than long (
<figureCitation id="135DAAB8FFD5FFA7E904FC992BBF6D0F" box="[873,954,820,844]" captionStart="FIGURE 1315" captionStartId="3.[151,250,1927,1951]" captionTargetBox="[156,1427,186,1899]" captionTargetId="figure-15@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1315. Lebertia diakonovoensis sp. n., deutonymph: 13—seta Fch; 14—coxal shield; 15—pedipalp. Scale bar: 13, 15 = 50 μm, 14 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142399" httpUri="https://zenodo.org/record/8142399/files/figure.png" pageId="5" pageNumber="490">Fig. 14</figureCitation>
). Medial length of coxal plates I longer than suture line between coxal plates II (L median Cx-I/Cx-II ratio 1.28-1.45). Medial margin of coxal plates III+IV concave, posteromedial corners of coxal plates IV widely rounded. Pregenital sclerite free and very small. Pedipalp (
<figureCitation id="135DAAB8FFD5FFA7EF7EFCD62D666DD0" box="[1299,1379,891,915]" captionStart="FIGURE 1315" captionStartId="3.[151,250,1927,1951]" captionTargetBox="[156,1427,186,1899]" captionTargetId="figure-15@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1315. Lebertia diakonovoensis sp. n., deutonymph: 13—seta Fch; 14—coxal shield; 15—pedipalp. Scale bar: 13, 15 = 50 μm, 14 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142399" httpUri="https://zenodo.org/record/8142399/files/figure.png" pageId="5" pageNumber="490">Fig. 15</figureCitation>
): P-1 short without seta; P-2 ventral margin concave, with relatively short dorsoproximal seta and two rather long dorsodistal ones; P-3 with two long distal setae, dorsal setae rather far away from distal edge, P-4 long tapering distally, with concave ventral margin and convex dorsal margin in distal half, ventral margin divided by seta insertions into sectors: 3:2:1. Capitulum ventral margin strong convex (
<figureCitation id="135DAAB8FFD5FFA7E823FBAA2A996A5C" box="[590,668,1031,1055]" captionStart="FIGURES 1619" captionStartId="4.[151,264,1502,1526]" captionTargetBox="[156,1431,186,1473]" captionTargetId="figure-29@4.[151,1436,181,1478]" captionTargetPageId="4" captionText="FIGURES 1619. Lebertia diakonovoensis sp. n., deutonymph: 16—chelicera; 17—capitulum, lateral view, 18—genital field, 19—IV-Leg-46. Scale bars: 1618 = 50 μm; 19 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142401" httpUri="https://zenodo.org/record/8142401/files/figure.png" pageId="5" pageNumber="490">Fig. 16</figureCitation>
). Chelicera basal segment large with convex dorsal margin and thickened distally, chela short (
<figureCitation id="135DAAB8FFD5FFA7EB19FB8629C66A00" box="[372,451,1067,1091]" captionStart="FIGURES 1619" captionStartId="4.[151,264,1502,1526]" captionTargetBox="[156,1431,186,1473]" captionTargetId="figure-29@4.[151,1436,181,1478]" captionTargetPageId="4" captionText="FIGURES 1619. Lebertia diakonovoensis sp. n., deutonymph: 16—chelicera; 17—capitulum, lateral view, 18—genital field, 19—IV-Leg-46. Scale bars: 1618 = 50 μm; 19 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142401" httpUri="https://zenodo.org/record/8142401/files/figure.png" pageId="5" pageNumber="490">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFBE32C086ACA" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
Genital flaps very narrow, fused to each other and forming ring bearing three fine setae on each side (
<figureCitation id="135DAAB8FFD5FFA7EF68FBE32D516A25" box="[1285,1364,1102,1126]" captionStart="FIGURES 1619" captionStartId="4.[151,264,1502,1526]" captionTargetBox="[156,1431,186,1473]" captionTargetId="figure-29@4.[151,1436,181,1478]" captionTargetPageId="4" captionText="FIGURES 1619. Lebertia diakonovoensis sp. n., deutonymph: 16—chelicera; 17—capitulum, lateral view, 18—genital field, 19—IV-Leg-46. Scale bars: 1618 = 50 μm; 19 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142401" httpUri="https://zenodo.org/record/8142401/files/figure.png" pageId="5" pageNumber="490">Fig. 18</figureCitation>
). Both pairs of genital acetabula subequal in shape and size. Excretory pore unsclerotized.
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFB382CE76A93" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
Legs III-IV with the following number of swimming setae: one to two on II-Leg-5, two on III/IV-Leg-4, four to five on III-Leg-5, four to six on IV-Leg-5; IV-Leg-6 with one to two short spine-like ventral setae (
<figureCitation id="135DAAB8FFD5FFA7EEEBFB1A2CD06A93" box="[1158,1237,1207,1232]" captionStart="FIGURES 1619" captionStartId="4.[151,264,1502,1526]" captionTargetBox="[156,1431,186,1473]" captionTargetId="figure-29@4.[151,1436,181,1478]" captionTargetPageId="4" captionText="FIGURES 1619. Lebertia diakonovoensis sp. n., deutonymph: 16—chelicera; 17—capitulum, lateral view, 18—genital field, 19—IV-Leg-46. Scale bars: 1618 = 50 μm; 19 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142401" httpUri="https://zenodo.org/record/8142401/files/figure.png" pageId="5" pageNumber="490">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFB7628F06BE0" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
Measurements (n=7). Idiosoma L 600-660; seta
<emphasis id="B9126A2FFFD5FFA7E8A5FB762AF76AB0" box="[712,754,1243,1267]" italics="true" pageId="5" pageNumber="490">Fch</emphasis>
L 95-110; coxal shield L 320-330, W 380-390; coxal plate I mL 95-110, coxal plate II mL 65-85; genital ring L 60-65, W 60-70, genital acetabula (ac-1-2) L 23-25, 20-25; capitulum L 120-125; cheliceral segments L base 120, chela 20-24; pedipalp segments (P15) L: 18-22, 63-66, 57-60, 68-85, 17-19; leg segments L: I-Leg-16: 40-45, 42-50, 47-50, 65-72, 75-80, 80-85; IILeg-16: 40-45, 54-60, 54-58, 75-80, 100-105, 105- 110; IIILeg-16: 42-48, 60-65, 70-75, 98-108, 135-140, 130-140; IVLeg-16: 78-85, 65-72, 95-100, 125-140, 150-160, 140-160.
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFA022AE96B86" blockId="5.[151,1437,151,2007]" box="[199,748,1454,1478]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAFA02290B6B85" box="[199,270,1455,1478]" italics="true" pageId="5" pageNumber="490">Larva.</emphasis>
See
<bibRefCitation id="EFF7CBCCFFD5FFA7EB2CFA0329FE6B85" author="Tuzovskij, P. V." box="[321,507,1454,1478]" pageId="5" pageNumber="490" pagination="569 - 580" refId="ref3082" refString="Tuzovskij, P. V. (2013) Larval morphology of Lebertia longiseta Bader, 1955, L. dubia Thor, 1899 and Oxus nodigerus Koenike, 1898 (Acari, Hydrachnidia: Lebertiidae, Oxidae). Zootaxa, 3619 (5), 569 - 580. https: // doi. org / 10.11646 / zootaxa. 3619.5.6" type="journal article" year="2013">Tuzovskij (2013)</bibRefCitation>
, as
<taxonomicName id="4C66CDBEFFD5FFA7E84EFA032AE36B85" authorityName="Bader" authorityYear="1955" box="[547,742,1454,1478]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="B9126A2FFFD5FFA7E84EFA032AE36B85" box="[547,742,1454,1478]" italics="true" pageId="5" pageNumber="490">Lebertia longiseta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAFA7F2ABC6810" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAFA7F29416BAA" box="[199,324,1490,1513]" italics="true" pageId="5" pageNumber="490">Discussion.</emphasis>
The new species is similar to
<taxonomicName id="4C66CDBEFFD5FFA7E8F6FA7C2C0F6BAA" authority="(Koch, 1837)" baseAuthorityName="Koch" baseAuthorityYear="1837" box="[667,1034,1489,1513]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B9126A2FFFD5FFA7E8F6FA7C2B6B6BAA" box="[667,878,1489,1513]" italics="true" pageId="5" pageNumber="490">Lebertia inaequalis</emphasis>
(Koch, 1837)
</taxonomicName>
and
<taxonomicName id="4C66CDBEFFD5FFA7EE29FA7F2D566BAA" authority="Bader, 1955" authorityName="Bader" authorityYear="1955" box="[1092,1363,1489,1513]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="B9126A2FFFD5FFA7EE29FA7F2CC06BAA" box="[1092,1221,1489,1513]" italics="true" pageId="5" pageNumber="490">L. longiseta</emphasis>
<bibRefCitation id="EFF7CBCCFFD5FFA7EEA2FA7C2D566BAA" author="Bader, C." box="[1231,1363,1489,1513]" pageId="5" pageNumber="490" pagination="61 - 84" refId="ref2853" refString="Bader, C. (1955) Hydracarinen-Diagnosen aus dem Nachlass von Dr. C. Walter. Verhandlungen der Naturforschunden Gesellschaft Basel, 66 (1), 61 - 84." type="journal article" year="1955">
Bader,
<quantity id="4C9E1BD8FFD5FFA7EF73FA7C2D566BAA" box="[1310,1363,1489,1513]" metricMagnitude="1" metricUnit="m" metricValue="4.9657" pageId="5" pageNumber="490" unit="in" value="1955.0">1955</quantity>
</bibRefCitation>
</taxonomicName>
in the structure of the idiosoma and pedipalp. The new species is more similar to
<taxonomicName id="4C66CDBEFFD5FFA7E98BFA582C70684F" baseAuthorityName="Koch" baseAuthorityYear="1837" box="[998,1141,1524,1548]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B9126A2FFFD5FFA7E98BFA582C70684F" box="[998,1141,1524,1548]" italics="true" pageId="5" pageNumber="490">L. inaequalis</emphasis>
</taxonomicName>
and differs from it by the following characters (characters states of
<taxonomicName id="4C66CDBEFFD5FFA7E831F9B52AEC686C" baseAuthorityName="Koch" baseAuthorityYear="1837" box="[604,745,1559,1583]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B9126A2FFFD5FFA7E831F9B52A6E686C" box="[604,619,1560,1583]" italics="true" pageId="5" pageNumber="490">L</emphasis>
.
<emphasis id="B9126A2FFFD5FFA7E817F9BA2AEC686C" box="[634,745,1559,1583]" italics="true" pageId="5" pageNumber="490">inaequalis</emphasis>
</taxonomicName>
are given in parenthesis after
<bibRefCitation id="EFF7CBCCFFD5FFA7EE58F9BA2CCB6873" author="Gerecke, R." box="[1077,1230,1559,1584]" pageId="5" pageNumber="490" pagination="1 - 144" refId="ref2967" refString="Gerecke, R. (2009) Revisional studies on the European species of the water mite genus Lebertia Neuman, 1880 (Acari: Hydrachnidia: Lebertiidae). Abhandlungen der Senckenbergischen Gesellschaft fur Naturforschung, 566, 1 - 144." type="journal article" year="2009">Gerecke 2009</bibRefCitation>
,
<bibRefCitation id="EFF7CBCCFFD5FFA7EEB6F9B528C96810" author="Di Sabatino, A. &amp; Gerecke, R. &amp; Gledhill, T. &amp; Smit, H." pageId="5" pageNumber="490" pagination="1 - 234" refId="ref2886" refString="Di Sabatino, A., Gerecke, R., Gledhill, T. &amp; Smit, H. (2010) 8. Acari, Hydrachnidia II. In: Gerecke, R. (Ed.), Suβwasserfauna von Mitteleuropa. 7 / 2 - 2. Spektrum Akademischer Verlag Heidelberg, pp. 1 - 234 https: // doi. org / 10.1007 / 978 - 3 - 8274 - 2266 - 8 _ 1" type="book chapter" year="2010">
<emphasis id="B9126A2FFFD5FFA7EEB6F9B52D93686C" box="[1243,1430,1559,1583]" italics="true" pageId="5" pageNumber="490">Di Sabatino et al</emphasis>
. 2010
</bibRefCitation>
for adults,
<bibRefCitation id="EFF7CBCCFFD5FFA7EB2BF99629EE6810" author="Tuzovskij, P. V." box="[326,491,1595,1619]" pageId="5" pageNumber="490" refId="ref3048" refString="Tuzovskij, P. V. (1990) Opredelitel' deutonymphs vodyanykh kleschey [Key to water mite deutonymphs]. Nauka Publ., Moscow, 238 pp. [In Russian]" type="book" year="1990">Tuzovskij 1990</bibRefCitation>
for deutonymphs):
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAF9F32D38690A" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAF9F3290D6835" box="[199,264,1630,1654]" italics="true" pageId="5" pageNumber="490">adults</emphasis>
: integument with fine lines,
<figureCitation id="135DAAB8FFD5FFA7E827F9F32A8E6835" box="[586,651,1630,1654]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="5" pageNumber="490">Fig. 1</figureCitation>
(smooth), capitulum ventral margin strongly convex near middle, with concave distal part,
<figureCitation id="135DAAB8FFD5FFA7EB1DF92C29AA68DA" box="[368,431,1665,1689]" captionStart="FIGURES 812" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[177,1414,201,1914]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 812. Lebertia diakonovoensis sp. n., male: 8—gnathosoma, lateral view; 9—IV-Leg-46; 10—leg claw; 1112— genital field; 811—female, 12—male. Scale bars: 8, 10 = 50 μm, 9, 1112 = 100 μm, 8 = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.8142397" httpUri="https://zenodo.org/record/8142397/files/figure.png" pageId="5" pageNumber="490">Fig. 8</figureCitation>
(equally convex); coxal field wider than long, (coxal field almost as long as wide or slightly longer than wide); setae
<emphasis id="B9126A2FFFD5FFA7EBF2F90929CD68FF" box="[415,456,1700,1724]" italics="true" pageId="5" pageNumber="490">Fch</emphasis>
with proximal furca,
<figureCitation id="135DAAB8FFD5FFA7E8DFF9092AF468FF" box="[690,753,1700,1724]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="5" pageNumber="490">Fig. 2</figureCitation>
(without furca); P-2 ventral seta thin,
<figureCitation id="135DAAB8FFD5FFA7EEE0F9092CC968FF" box="[1165,1228,1700,1724]" captionStart="FIGURES 17" captionStartId="1.[151,264,1935,1959]" captionTargetBox="[168,1436,199,1899]" captionTargetId="figure-15@1.[151,1436,180,1910]" captionTargetPageId="1" captionText="FIGURES 17. Lebertia diakonovoensis sp. n., male: 1—detail of integument; 2—seta Fch; 3—seta Oi; 4—seta Oe; 5—ventral view; 6—pedipalp; 7—P-4, distal part. Scale bars: 1, 24 = 100 μm, 5 = 200 μm, 6 = 50 μm, 7 = 25 μm." figureDoi="http://doi.org/10.5281/zenodo.8142395" httpUri="https://zenodo.org/record/8142395/files/figure.png" pageId="5" pageNumber="490">Fig. 6</figureCitation>
(rather strong,
<figureCitation id="135DAAB8FFD5FFA7EF1FF90928B068A3" captionStart="FIGURES 2021" captionStartId="6.[151,264,1118,1142]" captionTargetBox="[154,1425,183,1090]" captionTargetId="figure-29@6.[151,1436,181,1094]" captionTargetPageId="6" captionText="FIGURES 2021. Lebertia, subgenus Pilolebertia: 20—L. inaequalis, pedipalp (from Gerecke 2009); 21—L. longiseta, pedipalp (from K.O. Viets 1976)." figureDoi="http://doi.org/10.5281/zenodo.8142405" httpUri="https://zenodo.org/record/8142405/files/figure.png" pageId="5" pageNumber="490">Fig. 20</figureCitation>
);
<emphasis id="B9126A2FFFD5FFA7EAA7F96A2949689C" box="[202,332,1735,1759]" italics="true" pageId="5" pageNumber="490">deutonymph</emphasis>
: setae
<emphasis id="B9126A2FFFD5FFA7EBF5F96A29C4689C" box="[408,449,1735,1759]" italics="true" pageId="5" pageNumber="490">Fch</emphasis>
with proximal furca,
<figureCitation id="135DAAB8FFD5FFA7E8C3F96A2AFE68A3" box="[686,763,1735,1760]" captionStart="FIGURE 1315" captionStartId="3.[151,250,1927,1951]" captionTargetBox="[156,1427,186,1899]" captionTargetId="figure-15@3.[151,1436,181,1904]" captionTargetPageId="3" captionText="FIGURE 1315. Lebertia diakonovoensis sp. n., deutonymph: 13—seta Fch; 14—coxal shield; 15—pedipalp. Scale bar: 13, 15 = 50 μm, 14 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142399" httpUri="https://zenodo.org/record/8142399/files/figure.png" pageId="5" pageNumber="490">Fig. 13</figureCitation>
(without furca); genital acetabula rather large and subequal in shape and size,
<figureCitation id="135DAAB8FFD5FFA7EB53F946298C6940" box="[318,393,1771,1795]" captionStart="FIGURES 1619" captionStartId="4.[151,264,1502,1526]" captionTargetBox="[156,1431,186,1473]" captionTargetId="figure-29@4.[151,1436,181,1478]" captionTargetPageId="4" captionText="FIGURES 1619. Lebertia diakonovoensis sp. n., deutonymph: 16—chelicera; 17—capitulum, lateral view, 18—genital field, 19—IV-Leg-46. Scale bars: 1618 = 50 μm; 19 = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8142401" httpUri="https://zenodo.org/record/8142401/files/figure.png" pageId="5" pageNumber="490">Fig. 18</figureCitation>
(posterior pair of acetabula large than anterior pair acetabula); coxal plate IV posteromedial angle widely rounded (with a right angle).
<taxonomicName id="4C66CDBEFFD5FFA7E849F8A32AED6965" authorityName="Bader" authorityYear="1955" box="[548,744,1806,1830]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="B9126A2FFFD5FFA7E849F8A32AED6965" box="[548,744,1806,1830]" italics="true" pageId="5" pageNumber="490">Lebertia longiseta</emphasis>
</taxonomicName>
differs especially from
<taxonomicName id="4C66CDBEFFD5FFA7E984F8A22CAF6965" authority="Tuzovskij, 2023" authorityName="Tuzovskij" authorityYear="2023" box="[1001,1194,1806,1830]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="diakonovoensis" status="sp.n.">
<emphasis id="B9126A2FFFD5FFA7E984F8A22CAF6965" box="[1001,1194,1806,1830]" italics="true" pageId="5" pageNumber="490">L. diakonovoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221D754FFD5FFA7EEDFF8A32CE06966" box="[1202,1253,1806,1829]" pageId="5" pageNumber="490" rank="species">sp.n.</taxonomicNameLabel>
by the very long ventrodistal seta on P-2 (
<figureCitation id="135DAAB8FFD5FFA7EBCCF89C29F5690A" box="[417,496,1841,1865]" captionStart="FIGURES 2021" captionStartId="6.[151,264,1118,1142]" captionTargetBox="[154,1425,183,1090]" captionTargetId="figure-29@6.[151,1436,181,1094]" captionTargetPageId="6" captionText="FIGURES 2021. Lebertia, subgenus Pilolebertia: 20—L. inaequalis, pedipalp (from Gerecke 2009); 21—L. longiseta, pedipalp (from K.O. Viets 1976)." figureDoi="http://doi.org/10.5281/zenodo.8142405" httpUri="https://zenodo.org/record/8142405/files/figure.png" pageId="5" pageNumber="490">Fig. 21</figureCitation>
), much longer than the segment (
<bibRefCitation id="EFF7CBCCFFD5FFA7E93EF89C2BD6690A" author="Bader, C." box="[851,979,1841,1865]" pageId="5" pageNumber="490" pagination="61 - 84" refId="ref2853" refString="Bader, C. (1955) Hydracarinen-Diagnosen aus dem Nachlass von Dr. C. Walter. Verhandlungen der Naturforschunden Gesellschaft Basel, 66 (1), 61 - 84." type="journal article" year="1955">Bader 1955</bibRefCitation>
, K.O.
<bibRefCitation id="EFF7CBCCFFD5FFA7EE75F89C2C88690A" author="Viets, K. O." box="[1048,1165,1841,1865]" pageId="5" pageNumber="490" pagination="17 - 24" refId="ref3145" refString="Viets, K. O. (1976) Uber Lebertia (Pilolebertia) longiseta (Walter) Bader, 1955 eine seltene Wassermilbenart aus dem Bodensee (Acari, Hydrachnellae). Nachrichten Naturwissenschaftlichen Museums der Stadt Aschaffenburg, 84, 17 - 24." type="journal article" year="1976">Viets 1976</bibRefCitation>
,
<bibRefCitation id="EFF7CBCCFFD5FFA7EEF5F89C2D34690A" author="Gerecke, R." box="[1176,1329,1841,1865]" pageId="5" pageNumber="490" pagination="1 - 144" refId="ref2967" refString="Gerecke, R. (2009) Revisional studies on the European species of the water mite genus Lebertia Neuman, 1880 (Acari: Hydrachnidia: Lebertiidae). Abhandlungen der Senckenbergischen Gesellschaft fur Naturforschung, 566, 1 - 144." type="journal article" year="2009">Gerecke 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAF8F9292269CC" blockId="5.[151,1437,151,2007]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAF8F9293C692F" box="[199,313,1876,1900]" italics="true" pageId="5" pageNumber="490">Etymology</emphasis>
. The species epithet
<taxonomicName id="4C66CDBEFFD5FFA7E845F8F92AC9692F" authorityName="Tuzovskij" authorityYear="2023" box="[552,716,1876,1900]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="diakonovoensis">
<emphasis id="B9126A2FFFD5FFA7E845F8F92AC9692F" box="[552,716,1876,1900]" italics="true" pageId="5" pageNumber="490">diakonovoensis</emphasis>
</taxonomicName>
is named after the village in the vicinity where it was collected (Diakonovo).
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAF8362AE669F0" blockId="5.[151,1437,151,2007]" box="[199,739,1947,1971]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAF836295469F0" box="[199,337,1947,1971]" italics="true" pageId="5" pageNumber="490">Distribution.</emphasis>
Europea,
<collectingCountry id="F371F6ADFFD5FFA7EBD0F8362A0D69F0" box="[445,520,1947,1971]" name="Russia" pageId="5" pageNumber="490">Russia</collectingCountry>
:
<collectingRegion id="49A278DFFFD5FFA7E87CF8362ADA69F0" box="[529,735,1947,1971]" country="Russia" name="Yaroslavl'" pageId="5" pageNumber="490">Yaroslavl Province</collectingRegion>
.
</paragraph>
<paragraph id="8BD9B63DFFD5FFA7EAAAF81229D36995" blockId="5.[151,1437,151,2007]" box="[199,470,1983,2007]" pageId="5" pageNumber="490">
<emphasis id="B9126A2FFFD5FFA7EAAAF812291D6994" box="[199,280,1983,2007]" italics="true" pageId="5" pageNumber="490">Habitat</emphasis>
. Running waters.
</paragraph>
</subSubSection>
</treatment>
</document>