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<document id="C931D904D011F85889CF7BEB116E4AE6" ID-CLB-Dataset="121335" ID-DOI="10.11646/zootaxa.5174.4.2" ID-GBIF-Dataset="d6b7a54e-37b1-44c0-963d-c68720fa9e97" ID-ISSN="1175-5326" ID-Zenodo-Dep="6986320" ID-ZooBank="B09A5F4E-31FA-44CE-BF50-007D1DEF43D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660326602367" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2022" docId="03B0B202FFE2BF61C0AAF6BAFDE6FDC5" docLanguage="en" docName="zootaxa.5174.4.2.pdf" docOrigin="Zootaxa 5174 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stenus hessei L. Benick 1933" docType="treatment" docVersion="7" lastPageNumber="339" masterDocId="FF89CA7AFFEABF69C03DF62DFF81FFDC" masterDocTitle="Four new species and additional records of the genus Stenus of southern Africa (Coleoptera: Staphylinidae: Steninae)" masterLastPageNumber="356" masterPageNumber="331" pageNumber="339" updateTime="1699366806495" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2E58CB2FCCE15E46A0A04F702A554A53">Four new species and additional records of the genus Stenus of southern Africa (Coleoptera: Staphylinidae: Steninae)</mods:title>
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<mods:namePart id="E1EA0E334A1EA502A2E666C0B534ADB8">Janák, Jiří</mods:namePart>
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<mods:title id="FA022C2963C0F6715B462DF6F8727C6A">Zootaxa</mods:title>
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<mods:date id="8F1E7BE36E100C7C0886A16A0DC1781E">2022</mods:date>
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<treatment id="03B0B202FFE2BF61C0AAF6BAFDE6FDC5" ID-DOI="http://doi.org/10.5281/zenodo.6986708" ID-GBIF-Taxon="199056757" ID-Zenodo-Dep="6986708" LSID="urn:lsid:plazi:treatment:03B0B202FFE2BF61C0AAF6BAFDE6FDC5" httpUri="http://treatment.plazi.org/id/03B0B202FFE2BF61C0AAF6BAFDE6FDC5" lastPageNumber="339" pageId="8" pageNumber="339">
<subSubSection id="C303509FFFE2BF61C0AAF6BAFD82FF6D" box="[151,515,151,178]" pageId="8" pageNumber="339" type="nomenclature">
<paragraph id="8BA60314FFE2BF61C0AAF6BAFD82FF6D" blockId="8.[151,515,151,178]" box="[151,515,151,178]" pageId="8" pageNumber="339">
<heading id="D0EEB478FFE2BF61C0AAF6BAFD82FF6D" bold="true" box="[151,515,151,178]" fontSize="11" level="1" pageId="8" pageNumber="339" reason="1">
<taxonomicName id="4C197897FFE2BF61C0AAF6BAFD82FF6D" ID-CoL="529CJ" authority="L. Benick, 1933" authorityName="L. Benick" authorityYear="1933" box="[151,515,151,178]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="hessei">
<emphasis id="B96DDF06FFE2BF61C0AAF6BAFD82FF6D" bold="true" box="[151,515,151,178]" pageId="8" pageNumber="339">
<emphasis id="B96DDF06FFE2BF61C0AAF6BAFEB8FF6E" bold="true" box="[151,313,151,178]" italics="true" pageId="8" pageNumber="339">Stenus hessei</emphasis>
L. Benick, 1933
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFE2BF61C0AAF6F2FE32FE71" pageId="8" pageNumber="339" type="materials_examined">
<paragraph id="8BA60314FFE2BF61C0AAF6F2FC05FF26" blockId="8.[151,1436,223,538]" box="[151,900,223,250]" pageId="8" pageNumber="339">
<materialsCitation id="3B710949FFE2BF61C0AAF6F2FC01FF26" ID-GBIF-Occurrence="3888061388" box="[151,896,223,250]" country="South Africa" location="Town" municipality="Ilati" pageId="8" pageNumber="339" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="B96DDF06FFE2BF61C0AAF6F2FEB7FF25" bold="true" box="[151,310,223,249]" pageId="8" pageNumber="339">
<typeStatus id="54A2BDB6FFE2BF61C0AAF6F2FF53FF25" box="[151,210,223,249]" pageId="8" pageNumber="339">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F30E4384FFE2BF61C101F6F2FE4CFF25" box="[316,461,223,249]" name="South Africa" pageId="8" pageNumber="339">South Africa</collectingCountry>
,
<collectingRegion id="49DDCDF6FFE2BF61C1E5F6CDFDF7FF25" box="[472,630,223,250]" country="South Africa" name="Western Cape" pageId="8" pageNumber="339">Western Cape</collectingRegion>
,
<collectingRegion id="49DDCDF6FFE2BF61C2BFF6F2FD3DFF25" box="[642,700,223,249]" country="South Africa" name="Western Cape" pageId="8" pageNumber="339">Cape</collectingRegion>
<collectingMunicipality id="6BC2996EFFE2BF61C2FFF6F2FD73FF25" box="[706,754,223,249]" pageId="8" pageNumber="339">Ilati</collectingMunicipality>
,
<collectingRegion id="49DDCDF6FFE2BF61C2C0F6F2FCB6FF25" box="[765,823,223,249]" country="South Africa" name="Western Cape" pageId="8" pageNumber="339">Cape</collectingRegion>
<location id="8EC655CFFFE2BF61C300F6CDFC01FF26" LSID="urn:lsid:plazi:treatment:03B0B202FFE2BF61C0AAF6BAFDE6FDC5:8EC655CFFFE2BF61C300F6CDFC01FF26" box="[829,896,224,250]" country="South Africa" municipality="Ilati" name="Town" pageId="8" pageNumber="339" stateProvince="Western Cape">Town</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA60314FFE2BF61C0FAF72EFE32FE71" blockId="8.[151,1436,223,538]" pageId="8" pageNumber="339">
<emphasis id="B96DDF06FFE2BF61C0FAF72EFE34FEC1" bold="true" box="[199,437,259,285]" pageId="8" pageNumber="339">Material examined.</emphasis>
<materialsCitation id="3B710949FFE2BF61C1FDF72EFC9EFE9D" ID-GBIF-Occurrence="3888061438" collectingDate="2019-11-05" collectionCode="JJRC" collectorName="Bulirsch" country="South Africa" latitude="-34.315" location="Bordjiesrif" longLatPrecision="119" longitude="18.46" municipality="Table Mts." pageId="8" pageNumber="339" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape">
<emphasis id="B96DDF06FFE2BF61C1FDF72EFD23FEC1" bold="true" box="[448,674,259,285]" pageId="8" pageNumber="339">
<collectingCountry id="F30E4384FFE2BF61C1FDF72EFD1CFEC1" box="[448,669,259,285]" name="South Africa" pageId="8" pageNumber="339">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DDCDF6FFE2BF61C293F729FCD3FEC1" box="[686,850,259,286]" country="South Africa" name="Western Cape" pageId="8" pageNumber="339">Western Cape</collectingRegion>
:
<specimenCount id="9D1FC89DFFE2BF61C35CF72EFC10FEC2" box="[865,913,259,286]" pageId="8" pageNumber="339" type="female">4 ♀</specimenCount>
:
<collectingMunicipality id="6BC2996EFFE2BF61C39FF72EFB9FFEC2" box="[930,1054,259,286]" pageId="8" pageNumber="339">Table Mts.</collectingMunicipality>
NP,
<location id="8EC655CFFFE2BF61C461F72EFB59FEC1" LSID="urn:lsid:plazi:treatment:03B0B202FFE2BF61C0AAF6BAFDE6FDC5:8EC655CFFFE2BF61C461F72EFB59FEC1" box="[1116,1240,259,285]" country="South Africa" latitude="-34.315" longLatPrecision="119" longitude="18.46" municipality="Table Mts." name="Bordjiesrif" pageId="8" pageNumber="339" stateProvince="Western Cape">Bordjiesrif</location>
, creek banks, ca
<geoCoordinate id="EE2D65D3FFE2BF61C0AAF70AFE8BFE9D" box="[151,266,295,321]" degrees="34" direction="south" minutes="18.9" orientation="latitude" pageId="8" pageNumber="339" precision="92" value="-34.315">34°18.9S</geoCoordinate>
,
<geoCoordinate id="EE2D65D3FFE2BF61C124F70AFE0FFE9D" box="[281,398,295,321]" degrees="18" direction="east" minutes="27.6" orientation="longitude" pageId="8" pageNumber="339" precision="92" value="18.46">18°27.6E</geoCoordinate>
,
<date id="FFA725D4FFE2BF61C1A0F70AFD8BFE9E" box="[413,522,295,322]" pageId="8" pageNumber="339" value="2019-11-05">
<collectingDate id="EFE3DC3CFFE2BF61C1A0F70AFD8BFE9E" box="[413,522,295,322]" pageId="8" pageNumber="339" value="2019-11-05">5.xi.2019</collectingDate>
</date>
, P.
<collectorName id="26EC66C2FFE2BF61C20BF70AFD14FE9D" box="[566,661,295,321]" pageId="8" pageNumber="339">Bulirsch</collectorName>
lgt. (
<collectionCode id="ED089BD1FFE2BF61C2EBF70AFC96FE9D" box="[726,791,295,321]" pageId="8" pageNumber="339">JJRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B710949FFE2BF61C313F70AFA01FE9D" ID-GBIF-Occurrence="3888061495" box="[814,1408,295,322]" collectingDate="2021-11-05" collectionCode="JJRC" collectorName="Janak" country="South Africa" latitude="-34.315" location="Bordjiesrif" longLatPrecision="119" longitude="18.46" municipality="Table Mts." pageId="8" pageNumber="339" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape">
<specimenCount id="9D1FC89DFFE2BF61C313F70AFCDCFE9E" box="[814,861,295,322]" pageId="8" pageNumber="339" type="female">4 ♀</specimenCount>
: same data, but
<date id="FFA725D4FFE2BF61C422F70AFB0AFE9E" box="[1055,1163,295,322]" pageId="8" pageNumber="339" value="2021-11-05">
<collectingDate id="EFE3DC3CFFE2BF61C422F70AFB0AFE9E" box="[1055,1163,295,322]" pageId="8" pageNumber="339" value="2021-11-05">5.xi.2021</collectingDate>
</date>
, J.
<collectorName id="26EC66C2FFE2BF61C48BF70AFB77FE9D" box="[1206,1270,295,321]" pageId="8" pageNumber="339">Janák</collectorName>
lgt. (
<collectionCode id="ED089BD1FFE2BF61C50BF70AFAF6FE9D" box="[1334,1399,295,321]" pageId="8" pageNumber="339">JJRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B710949FFE2BF61C5B3F70AFE22FE55" ID-GBIF-Occurrence="3888061472" collectingDate="2021-11-04" collectionCode="JJRC" collectorName="Janak" country="South Africa" county="Table Mts." latitude="-34.255" location="Olifantsbos" longLatPrecision="119" longitude="18.385" municipality="Peninsula" pageId="8" pageNumber="339" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
<specimenCount id="9D1FC89DFFE2BF61C5B3F70AFF2EFEBA" pageId="8" pageNumber="339" type="male">1 ♂</specimenCount>
:
<collectingCounty id="62C77B98FFE2BF61C083F766FEB8FEBA" box="[190,313,331,358]" pageId="8" pageNumber="339">Table Mts.</collectingCounty>
NP,
<collectingRegion id="49DDCDF6FFE2BF61C148F766FE2EFEB9" box="[373,431,331,357]" country="South Africa" name="Western Cape" pageId="8" pageNumber="339">Cape</collectingRegion>
<collectingMunicipality id="6BC2996EFFE2BF61C184F766FDA8FEB9" box="[441,553,331,357]" pageId="8" pageNumber="339">Peninsula</collectingMunicipality>
,
<location id="8EC655CFFFE2BF61C20AF766FD3AFEB9" LSID="urn:lsid:plazi:treatment:03B0B202FFE2BF61C0AAF6BAFDE6FDC5:8EC655CFFFE2BF61C20AF766FD3AFEB9" box="[567,699,331,357]" country="South Africa" county="Table Mts." latitude="-34.255" longLatPrecision="119" longitude="18.385" municipality="Peninsula" name="Olifantsbos" pageId="8" pageNumber="339" stateProvince="Western Cape">Olifantsbos</location>
,
<geoCoordinate id="EE2D65D3FFE2BF61C2F7F766FCBCFEBA" box="[714,829,331,358]" degrees="34" direction="south" minutes="15.3" orientation="latitude" pageId="8" pageNumber="339" precision="92" value="-34.255">34°15.3S</geoCoordinate>
,
<geoCoordinate id="EE2D65D3FFE2BF61C371F766FC43FEB9" box="[844,962,331,357]" degrees="18" direction="east" minutes="23.1" orientation="longitude" pageId="8" pageNumber="339" precision="92" value="18.385">18°23.1E</geoCoordinate>
,
<date id="FFA725D4FFE2BF61C3EDF766FBBCFEB9" box="[976,1085,331,357]" pageId="8" pageNumber="339" value="2021-11-04">
<collectingDate id="EFE3DC3CFFE2BF61C3EDF766FBBCFEB9" box="[976,1085,331,357]" pageId="8" pageNumber="339" value="2021-11-04">4.xi.2021</collectingDate>
</date>
, J.
<collectorName id="26EC66C2FFE2BF61C455F766FB29FEB9" box="[1128,1192,331,357]" pageId="8" pageNumber="339">Janák</collectorName>
lgt., wetland, in low fynbos, treading (
<collectionCode id="ED089BD1FFE2BF61C167F742FE1AFE55" box="[346,411,367,393]" pageId="8" pageNumber="339">JJRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B710949FFE2BF61C190F742FE2EFE71" ID-GBIF-Occurrence="3888061413" collectingDate="1983-10-28" collectionCode="JJRC" collectorName="Endrody-Younga" country="South Africa" latitude="-34.42" location="Heuningnes Riv." longLatPrecision="720" longitude="20.02" pageId="8" pageNumber="339" specimenCount="1" specimenCount-male="1" stateProvince="Cape">
<specimenCount id="9D1FC89DFFE2BF61C190F742FE58FE56" box="[429,473,367,394]" pageId="8" pageNumber="339" type="male">1 ♂</specimenCount>
:
<location id="8EC655CFFFE2BF61C1D9F742FD21FE55" LSID="urn:lsid:plazi:treatment:03B0B202FFE2BF61C0AAF6BAFDE6FDC5:8EC655CFFFE2BF61C1D9F742FD21FE55" box="[484,672,367,393]" country="South Africa" latitude="-34.42" longLatPrecision="720" longitude="20.02" name="Heuningnes Riv." pageId="8" pageNumber="339" stateProvince="Cape">Heuningnes Riv.</location>
,
<geoCoordinate id="EE2D65D3FFE2BF61C297F742FD7DFE55" box="[682,764,367,393]" degrees="34.42" direction="south" orientation="latitude" pageId="8" pageNumber="339" precision="555" value="-34.42">34.42 S</geoCoordinate>
<geoCoordinate id="EE2D65D3FFE2BF61C336F742FCE4FE56" box="[779,869,367,394]" degrees="20.02" direction="east" orientation="longitude" pageId="8" pageNumber="339" precision="555" value="20.02">20.02 E</geoCoordinate>
,
<date id="FFA725D4FFE2BF61C34DF742FC63FE55" box="[880,994,367,393]" pageId="8" pageNumber="339" value="1983-10-28">
<collectingDate id="EFE3DC3CFFE2BF61C34DF742FC63FE55" box="[880,994,367,393]" pageId="8" pageNumber="339" value="1983-10-28">28.x.1983</collectingDate>
</date>
, leg.
<collectorName id="26EC66C2FFE2BF61C421F742FB5DFE55" box="[1052,1244,367,393]" pageId="8" pageNumber="339">Endrödy-Younga</collectorName>
(
<collectionCode id="ED089BD1FFE2BF61C4D4F742FAA9FE55" box="[1257,1320,367,393]" pageId="8" pageNumber="339">JJRC</collectionCode>
, specimen recorded by
<bibRefCitation id="EF887EE5FFE2BF61C11EF7BEFE26FE71" author="Puthz, V." box="[291,423,403,429]" pageId="8" pageNumber="339" pagination="29 - 67" refId="ref16758" refString="Puthz, V. (2006) On the genus Stenus Latreille, mostly from southern Africa, with a revision of the Stenus mendicus speciesgroup (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 43, 29 - 67." type="journal article" year="2006">Puthz 2006</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFE2BF61C0FAF79AFDE6FDC5" pageId="8" pageNumber="339" type="distribution">
<paragraph id="8BA60314FFE2BF61C0FAF79AFBD1FE0D" blockId="8.[151,1436,223,538]" box="[199,1104,439,466]" pageId="8" pageNumber="339">
<emphasis id="B96DDF06FFE2BF61C0FAF79AFEE0FE0D" bold="true" box="[199,353,439,465]" pageId="8" pageNumber="339">Distribution.</emphasis>
The species is distributed in
<collectingRegion id="49DDCDF6FFE2BF61C29BF795FC2EFE0D" box="[678,943,439,466]" country="South Africa" name="Western Cape" pageId="8" pageNumber="339">Western Cape Province</collectingRegion>
,
<collectingCountry id="F30E4384FFE2BF61C387F79AFBCDFE0D" box="[954,1100,439,465]" name="South Africa" pageId="8" pageNumber="339">South Africa</collectingCountry>
.
</paragraph>
<paragraph id="8BA60314FFE2BF61C0FAF7F6FDE6FDC5" blockId="8.[151,1436,223,538]" pageId="8" pageNumber="339">
<emphasis id="B96DDF06FFE2BF61C0FAF7F6FE87FE29" bold="true" box="[199,262,475,501]" pageId="8" pageNumber="339">Note.</emphasis>
Aedeagus as in
<figureCitation id="13221F91FFE2BF61C186F7F6FDC3FE29" box="[443,578,475,501]" captionStart="FIGURES 2537" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[172,1408,321,1890]" captionTargetId="figure-74@9.[159,1429,309,1903]" captionTargetPageId="9" captionText="FIGURES 2537. 2531: Stenus rugosellus Janák, sp. nov., 2530, holotype; 2930, paratype. 3237: Stenus hessei L. Benick, 32, Heuningnes river, 3337, Bordjiersrif. 25, habitus; 26, abdomen ventral; 27, 32, 33, aedeagus, dorsal; 28, 34, sternite 9; 29, 35, male tergite 10; 31, 37, female tergite 10; 30, 36, valves. Scale 1 mm: 2526; scale 0.5 mm: 2737." figureDoi="http://doi.org/10.5281/zenodo.6986326" httpUri="https://zenodo.org/record/6986326/files/figure.png" pageId="8" pageNumber="339">Figs. 3233</figureCitation>
, male sternite 9 as in
<figureCitation id="13221F91FFE2BF61C30FF7F6FC02FE29" box="[818,899,475,501]" captionStart="FIGURES 2537" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[172,1408,321,1890]" captionTargetId="figure-74@9.[159,1429,309,1903]" captionTargetPageId="9" captionText="FIGURES 2537. 2531: Stenus rugosellus Janák, sp. nov., 2530, holotype; 2930, paratype. 3237: Stenus hessei L. Benick, 32, Heuningnes river, 3337, Bordjiersrif. 25, habitus; 26, abdomen ventral; 27, 32, 33, aedeagus, dorsal; 28, 34, sternite 9; 29, 35, male tergite 10; 31, 37, female tergite 10; 30, 36, valves. Scale 1 mm: 2526; scale 0.5 mm: 2737." figureDoi="http://doi.org/10.5281/zenodo.6986326" httpUri="https://zenodo.org/record/6986326/files/figure.png" pageId="8" pageNumber="339">Fig. 34</figureCitation>
, male tergite 10 as in
<figureCitation id="13221F91FFE2BF61C44AF7F6FB49FE2A" box="[1143,1224,475,502]" captionStart="FIGURES 2537" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[172,1408,321,1890]" captionTargetId="figure-74@9.[159,1429,309,1903]" captionTargetPageId="9" captionText="FIGURES 2537. 2531: Stenus rugosellus Janák, sp. nov., 2530, holotype; 2930, paratype. 3237: Stenus hessei L. Benick, 32, Heuningnes river, 3337, Bordjiersrif. 25, habitus; 26, abdomen ventral; 27, 32, 33, aedeagus, dorsal; 28, 34, sternite 9; 29, 35, male tergite 10; 31, 37, female tergite 10; 30, 36, valves. Scale 1 mm: 2526; scale 0.5 mm: 2737." figureDoi="http://doi.org/10.5281/zenodo.6986326" httpUri="https://zenodo.org/record/6986326/files/figure.png" pageId="8" pageNumber="339">Fig. 35</figureCitation>
, female valvifer as in
<figureCitation id="13221F91FFE2BF61C089F7D2FE84FDC5" box="[180,261,511,537]" captionStart="FIGURES 2537" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[172,1408,321,1890]" captionTargetId="figure-74@9.[159,1429,309,1903]" captionTargetPageId="9" captionText="FIGURES 2537. 2531: Stenus rugosellus Janák, sp. nov., 2530, holotype; 2930, paratype. 3237: Stenus hessei L. Benick, 32, Heuningnes river, 3337, Bordjiersrif. 25, habitus; 26, abdomen ventral; 27, 32, 33, aedeagus, dorsal; 28, 34, sternite 9; 29, 35, male tergite 10; 31, 37, female tergite 10; 30, 36, valves. Scale 1 mm: 2526; scale 0.5 mm: 2737." figureDoi="http://doi.org/10.5281/zenodo.6986326" httpUri="https://zenodo.org/record/6986326/files/figure.png" pageId="8" pageNumber="339">Fig. 36</figureCitation>
, female tergite 10 as in
<figureCitation id="13221F91FFE2BF61C22CF7D2FDE2FDC5" box="[529,611,511,537]" captionStart="FIGURES 2537" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[172,1408,321,1890]" captionTargetId="figure-74@9.[159,1429,309,1903]" captionTargetPageId="9" captionText="FIGURES 2537. 2531: Stenus rugosellus Janák, sp. nov., 2530, holotype; 2930, paratype. 3237: Stenus hessei L. Benick, 32, Heuningnes river, 3337, Bordjiersrif. 25, habitus; 26, abdomen ventral; 27, 32, 33, aedeagus, dorsal; 28, 34, sternite 9; 29, 35, male tergite 10; 31, 37, female tergite 10; 30, 36, valves. Scale 1 mm: 2526; scale 0.5 mm: 2737." figureDoi="http://doi.org/10.5281/zenodo.6986326" httpUri="https://zenodo.org/record/6986326/files/figure.png" pageId="8" pageNumber="339">Fig. 37</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>