433 lines
54 KiB
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433 lines
54 KiB
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<document ID-DOI="10.11646/zootaxa.5278.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7906172" ID-ZooBank="A5A18196-D519-4D43-BF03-5C6D2DEAC842" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683529272315" checkinUser="plazi" docAuthor="Archibald, S. Bruce, Ware, Jessica L., Rasmussen, Jan A., Sylvestersen, René L., Olsen, Kent & Simonsen, Thomas J." docDate="2023" docId="03A5BA5CFFADFFBBFF40EAF7EDD24EB4" docLanguage="en" docName="zootaxa.5278.2.4.pdf" docOrigin="Zootaxa 5278 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Furagrion jutlandicus" docType="treatment" docVersion="3" lastPageNumber="310" masterDocId="FF9CC224FFA2FFAEFFD7EA60E938481D" masterDocTitle="The damselfly genus Furagrion Petrulevičius et al. (Odonata, Zygoptera) from the early Eocene Fur Formation of Denmark and the dysagrionoid grade" masterLastPageNumber="317" masterPageNumber="289" pageNumber="304" updateTime="1683553539095" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>The damselfly genus Furagrion Petrulevičius et al. (Odonata, Zygoptera) from the early Eocene Fur Formation of Denmark and the dysagrionoid grade</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation>Beaty Biodiversity Museum, University of British Columbia, Vancouver, British Columbia, Canada. https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ware, Jessica L.</mods:namePart>
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<mods:affiliation>Division of Invertebrate Zoology, American Museum of Natural History, New York, New York, 10024, United States of America.</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rasmussen, Jan A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-0520-9148</mods:nameIdentifier>
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<mods:affiliation>Museum Mors, Skarrehagevej 8, DK- 7950 NykØbing Mors, Denmark. https: // orcid. org / 0000 - 0003 - 0520 - 9148 & Natural History Museum of Denmark, Øster Voldgade 5 - 7, DK- 1350 Copenhagen K, Denmark. jan. rasmussen @ museummors. dk</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sylvestersen, René L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-1946-4716</mods:nameIdentifier>
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<mods:affiliation>Fur Museum, Nederby 28, DK- 7884 Fur, Denmark. RLSY @ museumsalling. dk; https: // orcid. org / 0000 - 0002 - 1946 - 4716</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Olsen, Kent</mods:namePart>
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<mods:affiliation>Natural History Museum Aarhus, Wilhelm Meyers Allé 10, Aarhus, DK- 8000 Aarhus C, Denmark.</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Simonsen, Thomas J.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-9857-9564</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum Aarhus, Wilhelm Meyers Allé 10, Aarhus, DK- 8000 Aarhus C, Denmark. https: // orcid. org / 0000 - 0001 - 9857 - 9564</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-05</mods:number>
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<mods:detail type="volume">
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<mods:number>5278</mods:number>
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</mods:detail>
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<mods:number>2</mods:number>
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<mods:extent unit="page">
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<mods:start>289</mods:start>
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<mods:end>317</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.2.4</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5278.2.4</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7906172</mods:identifier>
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<mods:identifier type="ZooBank">A5A18196-D519-4D43-BF03-5C6D2DEAC842</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7906162" ID-Zenodo-Dep="7906162" LSID="urn:lsid:plazi:treatment:03A5BA5CFFADFFBBFF40EAF7EDD24EB4" httpUri="http://treatment.plazi.org/id/03A5BA5CFFADFFBBFF40EAF7EDD24EB4" lastPageId="21" lastPageNumber="310" pageId="15" pageNumber="304">
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<subSubSection box="[151,651,151,178]" pageId="15" pageNumber="304" type="nomenclature">
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<paragraph blockId="15.[151,651,151,213]" box="[151,651,151,178]" pageId="15" pageNumber="304">
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<heading bold="true" box="[151,651,151,178]" fontSize="11" level="1" pageId="15" pageNumber="304" reason="1">
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<taxonomicName authority="(Henriksen, 1922)" baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[151,651,151,178]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="jutlandicus">
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<emphasis bold="true" box="[151,651,151,178]" pageId="15" pageNumber="304">
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<emphasis bold="true" box="[151,421,151,178]" italics="true" pageId="15" pageNumber="304">Furagrion jutlandicus</emphasis>
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(
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<bibRefCitation author="Henriksen, K." box="[436,643,151,177]" pageId="15" pageNumber="304" pagination="1 - 36" refId="ref12417" refString="Henriksen, K. (1922) Eocene insects from Denmark. Danmarks Grologiske UndersOgelser, II Raekke, 37, 1 - 36." type="journal article" year="1922">Henriksen, 1922</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection box="[151,402,187,213]" pageId="15" pageNumber="304" type="description">
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||
<paragraph blockId="15.[151,651,151,213]" box="[151,402,187,213]" pageId="15" pageNumber="304">
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<figureCitation box="[151,257,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[152,255,1633,1657]" captionStartId-1="2.[152,255,1807,1831]" captionStartId-2="4.[151,250,1462,1486]" captionStartId-3="5.[151,250,1977,2001]" captionTargetBox-0="[151,1427,525,1601]" captionTargetBox-1="[151,1421,1101,1773]" captionTargetBox-2="[151,1435,489,1438]" captionTargetBox-3="[176,1415,670,1949]" captionTargetId-0="figure-182@1.[151,1436,525,1610]" captionTargetId-1="figure-470@2.[151,1436,1101,1784]" captionTargetId-2="figure-188@4.[151,1436,489,1438]" captionTargetId-3="figure-172@5.[171,1415,662,1954]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. MGUH 1819, Henriksen’s type specimen of Phenacolestes jutlandica: A, photograph; B, drawing; C–E, labelled wings. A, B to scale, C–E to scale, both scales 5 mm." captionText-1="FIGURE 2. FUM-N 13856, with which Petrulevičius et al. (2008) defined Furagrion (as ERK-KL-T1): A, photograph; B, drawing. Both to scale, 5 mm. A forewing. Ax1, Ax2 = antenodal cross veins 1 and 2; arc = arculus; b = brace vein; CuA = anterior branch of the cubitus; IR1, IR2 = intercalar vein 1 and 2; MA, MP = the anterior and posterior branches of the media; nod = nodus; pt = pterostigma; Q = quadrangle; RA = radius anterior; RP1, RP2, RP3-4 = branches of the radius posterior; sn = subnodus; * = origin of RP2." captionText-2="FIGURE 3. Furagrion jutlandicus proposed neotype MM-10752:A, photograph of the whole insect; B, photograph of the head; C, drawing of the head, showing bulging left compound eye. A to scale, 5 mm; B, C to scale, 5 mm." captionText-3="FIGURE 4. Wings of Furagrion jutlandicus proposed neotype MM-10752. The top two are hind wings and the bottom two are forewings. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906174" figureDoi-1="http://doi.org/10.5281/zenodo.7906176" figureDoi-2="http://doi.org/10.5281/zenodo.7906180" figureDoi-3="http://doi.org/10.5281/zenodo.7906182" httpUri-0="https://zenodo.org/record/7906174/files/figure.png" httpUri-1="https://zenodo.org/record/7906176/files/figure.png" httpUri-2="https://zenodo.org/record/7906180/files/figure.png" httpUri-3="https://zenodo.org/record/7906182/files/figure.png" pageId="15" pageNumber="304">Figs. 1–4</figureCitation>
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,
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<figureCitation box="[269,314,187,213]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="9.[151,250,1269,1293]" captionStartId-1="10.[151,250,1802,1826]" captionStartId-2="11.[151,250,1756,1780]" captionTargetBox-0="[158,1436,181,1240]" captionTargetBox-1="[151,1436,181,1772]" captionTargetBox-2="[169,1420,422,1724]" captionTargetId-0="figure-24@9.[151,1436,181,1245]" captionTargetId-1="figure-28@10.[151,1436,181,1778]" captionTargetId-2="figure-144@11.[163,1425,417,1733]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 6. 14M-A2163: A, photograph; B, drawing; C. drawing of the hind wing (red in Fig. 6B). All to scale, 5 mm." captionText-1="FIGURE 7. MM-10750: A, photograph; B, drawing, wing colours as in Fig. 8. Both to scale, 5 mm." captionText-2="FIGURE 8. Wings of MM-10750, colour-coded forewings (FW) and hind wings (HW) to match colours in Fig. 7. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906186" figureDoi-1="http://doi.org/10.5281/zenodo.7906188" figureDoi-2="http://doi.org/10.5281/zenodo.7906190" httpUri-0="https://zenodo.org/record/7906186/files/figure.png" httpUri-1="https://zenodo.org/record/7906188/files/figure.png" httpUri-2="https://zenodo.org/record/7906190/files/figure.png" pageId="15" pageNumber="304">6–8</figureCitation>
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,
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<figureCitation box="[325,397,187,213]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStart-5="FIGURE 16" captionStart-6="FIGURE 17" captionStart-7="FIGURE 18" captionStartId-0="15.[151,250,1992,2016]" captionStartId-1="16.[151,250,1226,1250]" captionStartId-2="17.[151,250,959,983]" captionStartId-3="17.[151,250,1857,1881]" captionStartId-4="18.[151,250,1183,1207]" captionStartId-5="19.[151,250,1943,1967]" captionStartId-6="20.[151,250,913,937]" captionStartId-7="20.[151,250,1716,1740]" captionTargetBox-0="[151,1436,598,1961]" captionTargetBox-1="[193,1399,639,1195]" captionTargetBox-2="[155,1434,181,928]" captionTargetBox-3="[151,1436,1008,1818]" captionTargetBox-4="[190,1397,181,1150]" captionTargetBox-5="[178,1408,186,1919]" captionTargetBox-6="[151,1436,181,875]" captionTargetBox-7="[151,1434,962,1680]" captionTargetId-0="figure-158@15.[151,1436,592,1968]" captionTargetId-1="figure-208@16.[188,1400,633,1203]" captionTargetId-2="figure-24@17.[151,1436,181,935]" captionTargetId-3="figure-48@17.[151,1436,1008,1833]" captionTargetId-4="figure-28@18.[190,1397,181,1159]" captionTargetId-5="figure-24@19.[178,1410,181,1919]" captionTargetId-6="figure-28@20.[151,1436,181,889]" captionTargetId-7="figure-48@20.[151,1436,962,1693]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionTargetPageId-6="20" captionTargetPageId-7="20" captionText-0="FIGURE 11. MM-4785A: A, photograph; B, drawing. Both to scale, 5 mm." captionText-1="FIGURE 12. Wings of MM-4785A: A, top wing in Fig. 9 (red), a forewing; B, wing below (blue), a hind wing. Both to scale, 5 mm." captionText-2="FIGURE 13. FUM-N 11146: A, photograph; B, drawing. Note infuscation (see text). Both to scale, 5 mm. A forewing." captionText-3="FIGURE 14. FUM-N 16192: A, photograph; B, drawing. Both to scale, 5 mm. A hind wing." captionText-4="FIGURE 15. 14M-2765: A, photograph; B, drawing as preserved; C, the fore- or hind wing graphically unfolded. All to scale, 5 mm." captionText-5="FIGURE 16. FUM-N 11696: A, drawing of bottom wing in 17C; B, drawing to top wing in 17C; C, photograph. All to scale, 5 mm." captionText-6="FIGURE 17. FUM-N 14704: A, photograph; B, drawing. A forewing. Both to scale, 5 mm." captionText-7="FIGURE 18. MM-2085: A, photograph; B, drawing. A forewing. Both to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906196" figureDoi-1="http://doi.org/10.5281/zenodo.7906198" figureDoi-2="http://doi.org/10.5281/zenodo.7906200" figureDoi-3="http://doi.org/10.5281/zenodo.7906202" figureDoi-4="http://doi.org/10.5281/zenodo.7906206" figureDoi-5="http://doi.org/10.5281/zenodo.7906208" figureDoi-6="http://doi.org/10.5281/zenodo.7906210" figureDoi-7="http://doi.org/10.5281/zenodo.7906212" httpUri-0="https://zenodo.org/record/7906196/files/figure.png" httpUri-1="https://zenodo.org/record/7906198/files/figure.png" httpUri-2="https://zenodo.org/record/7906200/files/figure.png" httpUri-3="https://zenodo.org/record/7906202/files/figure.png" httpUri-4="https://zenodo.org/record/7906206/files/figure.png" httpUri-5="https://zenodo.org/record/7906208/files/figure.png" httpUri-6="https://zenodo.org/record/7906210/files/figure.png" httpUri-7="https://zenodo.org/record/7906212/files/figure.png" pageId="15" pageNumber="304">11–18</figureCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection box="[151,560,256,280]" pageId="15" pageNumber="304" type="reference_group">
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||
<paragraph blockId="15.[151,560,256,280]" box="[151,560,256,280]" pageId="15" pageNumber="304">
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<taxonomicName authority="Zessin, 2011" authorityName="Zessin" authorityYear="2011" box="[151,458,256,280]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="morsi" status="syn. nov.">
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<emphasis box="[151,321,256,279]" italics="true" pageId="15" pageNumber="304">Furagrion morsi</emphasis>
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<bibRefCitation author="Zessin, W." box="[327,458,256,280]" pageId="15" pageNumber="304" pagination="64 - 73" refId="ref14309" refString="Zessin, W. (2011) Neue Insekten aus dem Moler (Palaozan / Eozan) von Danemark Teil 1 (Odonata: Epallagidae, Megapodagrioniidae), Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg, 14, 64 - 73." type="journal article" year="2011">Zessin, 2011</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[468,560,256,280]" pageId="15" pageNumber="304">syn. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="304" type="diagnosis">
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<paragraph blockId="15.[151,1436,325,568]" pageId="15" pageNumber="304">
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<emphasis bold="true" box="[151,390,325,352]" pageId="15" pageNumber="304">Emended diagnosis.</emphasis>
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The wings of
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<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[553,801,326,352]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="jutlandicus">
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<emphasis box="[553,801,326,352]" italics="true" pageId="15" pageNumber="304">Furagrion jutlandicus</emphasis>
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</taxonomicName>
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may be distinguished from those of
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<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[1215,1436,327,352]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="ansorgei">
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<emphasis box="[1215,1436,327,352]" italics="true" pageId="15" pageNumber="304">Furagrion ansorgei</emphasis>
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</taxonomicName>
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by shape: both fore- and hind wings have smaller length/width ratios in all measurements except RP2 to apex / width (
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<tableCitation box="[231,325,398,424]" captionStart="TABLE 2" captionStartId="5.[152,244,151,175]" captionTargetId="graphics-41@5.[151,1434,180,606]" captionTargetPageId="5" captionText="TABLE 2. Ratios of measurements in Table 1. Abbreviations as in Table 1." pageId="15" pageNumber="304">Tables 2</tableCitation>
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and
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<tableCitation box="[379,394,398,424]" captionStart="TABLE 3" captionStartId="6.[152,244,1052,1076]" captionTargetId="graphics-456@6.[151,1435,1117,1636]" captionTargetPageId="6" captionText="TABLE 3. Measurements in millimeters and ratios of these for wings not known to be fore- or hind wings. Abbreviations as in Table 1." pageId="15" pageNumber="304">3</tableCitation>
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, and see PCA, above); and by the IR1-RP2 space becoming more than one cell wide at about level of basal end of pterostigma or slightly basal to that [
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<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[815,941,435,460]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="ansorgei">
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<emphasis box="[815,941,435,460]" italics="true" pageId="15" pageNumber="304">F. ansorgei</emphasis>
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</taxonomicName>
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: distad distal end of pterostigma (tentative, see below)].
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</paragraph>
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<paragraph blockId="15.[151,1436,325,568]" pageId="15" pageNumber="304">
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<emphasis bold="true" box="[199,460,505,532]" pageId="15" pageNumber="304">Emended description.</emphasis>
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As for genus description and diagnosis, species diagnosis, and see forewing, hind wing measurements and ratios of these for the proposed
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<typeStatus box="[720,809,542,568]" pageId="15" pageNumber="304">neotype</typeStatus>
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and additional specimens in
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<tableCitation box="[1134,1228,542,568]" captionStart="TABLE 1" captionStartId="4.[152,244,1570,1594]" captionTargetId="graphics-268@4.[151,1434,1635,2018]" captionTargetPageId="4" captionText="TABLE 1. Measured distances from known Furagrion fore- and hind wings in millimeters: pt = pterostigma, apx = apex, arc = arculus, nod = nodus, RP2 = the origin of RP2, w = maximum width." pageId="15" pageNumber="304">Tables 1</tableCitation>
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and
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<tableCitation box="[1282,1299,542,568]" captionStart="TABLE 2" captionStartId="5.[152,244,151,175]" captionTargetId="graphics-41@5.[151,1434,180,606]" captionTargetPageId="5" captionText="TABLE 2. Ratios of measurements in Table 1. Abbreviations as in Table 1." pageId="15" pageNumber="304">2</tableCitation>
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.
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</paragraph>
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</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906196" ID-Zenodo-Dep="7906196" box="[151,923,1992,2016]" httpUri="https://zenodo.org/record/7906196/files/figure.png" pageId="15" pageNumber="304" startId="15.[151,250,1992,2016]" targetBox="[151,1436,598,1961]" targetPageId="15" targetType="figure">
|
||
<paragraph blockId="15.[151,923,1992,2016]" box="[151,923,1992,2016]" pageId="15" pageNumber="304">
|
||
<emphasis bold="true" box="[151,286,1992,2016]" pageId="15" pageNumber="304">FIGURE 11.</emphasis>
|
||
MM-4785A: A, photograph; B, drawing. Both to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="18" lastPageNumber="307" pageId="16" pageNumber="305" type="materials_examined">
|
||
<paragraph blockId="16.[151,1437,150,609]" pageId="16" pageNumber="305">
|
||
<emphasis bold="true" box="[199,375,150,177]" pageId="16" pageNumber="305">Type material.</emphasis>
|
||
<emphasis box="[384,490,151,177]" italics="true" pageId="16" pageNumber="305">Proposed</emphasis>
|
||
<materialsCitation box="[500,1026,150,177]" collectingDate="2009-12" country="Denmark" county="Nykobing Mors" location="Fossil" municipality="Museum Mors" pageId="16" pageNumber="305" specimenCode="MM-10752" specimenCount="1" typeStatus="neotype">
|
||
<emphasis box="[500,587,151,176]" italics="true" pageId="16" pageNumber="305">
|
||
<typeStatus box="[500,587,151,176]" pageId="16" pageNumber="305">neotype</typeStatus>
|
||
</emphasis>
|
||
MM-10752 (previously MHM-17-42)
|
||
</materialsCitation>
|
||
(
|
||
<figureCitation box="[1043,1155,151,177]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1462,1486]" captionStartId-1="5.[151,250,1977,2001]" captionTargetBox-0="[151,1435,489,1438]" captionTargetBox-1="[176,1415,670,1949]" captionTargetId-0="figure-188@4.[151,1436,489,1438]" captionTargetId-1="figure-172@5.[171,1415,662,1954]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Furagrion jutlandicus proposed neotype MM-10752:A, photograph of the whole insect; B, photograph of the head; C, drawing of the head, showing bulging left compound eye. A to scale, 5 mm; B, C to scale, 5 mm." captionText-1="FIGURE 4. Wings of Furagrion jutlandicus proposed neotype MM-10752. The top two are hind wings and the bottom two are forewings. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906180" figureDoi-1="http://doi.org/10.5281/zenodo.7906182" httpUri-0="https://zenodo.org/record/7906180/files/figure.png" httpUri-1="https://zenodo.org/record/7906182/files/figure.png" pageId="16" pageNumber="305">Figs. 3–4</figureCitation>
|
||
): head mostly indistinct, with faintly preserved left compound eye, complete thorax and abdomen, all four wings, in soft diatomite, coll.
|
||
<collectorName box="[151,329,223,249]" pageId="16" pageNumber="305">Henrik Madsen</collectorName>
|
||
at
|
||
<collectorName box="[370,496,223,249]" pageId="16" pageNumber="305">Skarrehage</collectorName>
|
||
mo-clay pit,
|
||
<collectorName box="[656,718,223,249]" pageId="16" pageNumber="305">Mors</collectorName>
|
||
,
|
||
<collectorName box="[733,964,223,249]" pageId="16" pageNumber="305">Knudeklint Member</collectorName>
|
||
(informally the Skarrehage bed, 0.6–1.0 m below ash layer -13),
|
||
<collectorName box="[437,603,259,285]" pageId="16" pageNumber="305">Fur Formation</collectorName>
|
||
in
|
||
<date box="[643,827,259,285]" pageId="16" pageNumber="305" value="2009-12">
|
||
<collectingDate box="[643,827,259,285]" pageId="16" pageNumber="305" value="2009-12">December 2009</collectingDate>
|
||
</date>
|
||
, in the
|
||
<location LSID="urn:lsid:plazi:treatment:03A5BA5CFFADFFBBFF40EAF7EDD24EB4:8ED35D91FFB2FFBEFC43EB63EAEF4900" box="[916,983,259,285]" country="Denmark" county="Nykobing Mors" municipality="Museum Mors" name="Fossil" pageId="16" pageNumber="305">Fossil</location>
|
||
and
|
||
<location LSID="urn:lsid:plazi:treatment:03A5BA5CFFADFFBBFF40EAF7EDD24EB4:8ED35D91FFB2FFBEFBC4EB63EDD94900" box="[1043,1249,259,285]" country="Denmark" county="Nykobing Mors" municipality="Museum Mors" name="Mo-clay Museum" pageId="16" pageNumber="305">Mo-clay Museum</location>
|
||
,
|
||
<collectingMunicipality box="[1262,1432,259,285]" pageId="16" pageNumber="305">Museum Mors</collectingMunicipality>
|
||
,
|
||
<collectingCounty box="[151,332,295,321]" pageId="16" pageNumber="305">Nykøbing Mors</collectingCounty>
|
||
,
|
||
<collectingCountry box="[343,451,295,321]" name="Denmark" pageId="16" pageNumber="305">Denmark</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,150,609]" pageId="16" pageNumber="305">
|
||
<emphasis bold="true" box="[199,384,330,356]" pageId="16" pageNumber="305">Other material.</emphasis>
|
||
MM-10750 (previously MHM-17-40) (
|
||
<figureCitation box="[827,936,331,357]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,250,1802,1826]" captionStartId-1="11.[151,250,1756,1780]" captionTargetBox-0="[151,1436,181,1772]" captionTargetBox-1="[169,1420,422,1724]" captionTargetId-0="figure-28@10.[151,1436,181,1778]" captionTargetId-1="figure-144@11.[163,1425,417,1733]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7. MM-10750: A, photograph; B, drawing, wing colours as in Fig. 8. Both to scale, 5 mm." captionText-1="FIGURE 8. Wings of MM-10750, colour-coded forewings (FW) and hind wings (HW) to match colours in Fig. 7. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906188" figureDoi-1="http://doi.org/10.5281/zenodo.7906190" httpUri-0="https://zenodo.org/record/7906188/files/figure.png" httpUri-1="https://zenodo.org/record/7906190/files/figure.png" pageId="16" pageNumber="305">Figs. 7–8</figureCitation>
|
||
): head missing the compound eyes, complete thorax and almost complete abdomen, most of all four wings, parts of five legs, in soft diatomite, coll. Henrik Madsen at Skarrehage mo-clay pit, Mors, Knudeklint Member (informally Skarrehage bed), Fur Formation,
|
||
<date box="[1253,1432,403,429]" pageId="16" pageNumber="305" value="2009-12">December 2009</date>
|
||
, in the Fossil and Mo-clay Museum.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,150,609]" pageId="16" pageNumber="305">
|
||
MM-4785 (previously MHM-5242A) (
|
||
<figureCitation box="[638,776,475,501]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="15.[151,250,1992,2016]" captionStartId-1="16.[151,250,1226,1250]" captionTargetBox-0="[151,1436,598,1961]" captionTargetBox-1="[193,1399,639,1195]" captionTargetId-0="figure-158@15.[151,1436,592,1968]" captionTargetId-1="figure-208@16.[188,1400,633,1203]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 11. MM-4785A: A, photograph; B, drawing. Both to scale, 5 mm." captionText-1="FIGURE 12. Wings of MM-4785A: A, top wing in Fig. 9 (red), a forewing; B, wing below (blue), a hind wing. Both to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906196" figureDoi-1="http://doi.org/10.5281/zenodo.7906198" httpUri-0="https://zenodo.org/record/7906196/files/figure.png" httpUri-1="https://zenodo.org/record/7906198/files/figure.png" pageId="16" pageNumber="305">Figs. 11–12</figureCitation>
|
||
): head missing, almost complete thorax and abdomen, one forewing with damaged posterior margin and two hind wings in varying states of incomplete preservation, in soft diatomite, coll. by unknown museum guest at the Ejerslev mo-clay pit, Mors, Knudeklint Member, Fur Formation,
|
||
<date box="[151,311,583,609]" pageId="16" pageNumber="305" value="2013-06-21">June 21, 2013</date>
|
||
, in the Fossil and Mo-clay Museum.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906198" ID-Zenodo-Dep="7906198" httpUri="https://zenodo.org/record/7906198/files/figure.png" pageId="16" pageNumber="305" startId="16.[151,250,1226,1250]" targetBox="[193,1399,639,1195]" targetPageId="16" targetType="figure">
|
||
<paragraph blockId="16.[151,1436,1226,1286]" pageId="16" pageNumber="305">
|
||
<emphasis bold="true" box="[151,288,1226,1250]" pageId="16" pageNumber="305">FIGURE 12.</emphasis>
|
||
Wings of MM-4785A: A, top wing in Fig. 9 (red), a forewing; B, wing below (blue), a hind wing. Both to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="16.[151,1437,1332,2007]" pageId="16" pageNumber="305">
|
||
<collectionCode box="[199,289,1333,1359]" pageId="16" pageNumber="305">FUM-N</collectionCode>
|
||
13856 (
|
||
<figureCitation box="[382,451,1332,1358]" captionStart="FIGURE 2" captionStartId="2.[152,255,1807,1831]" captionTargetBox="[151,1421,1101,1773]" captionTargetId="figure-470@2.[151,1436,1101,1784]" captionTargetPageId="2" captionText="FIGURE 2. FUM-N 13856, with which Petrulevičius et al. (2008) defined Furagrion (as ERK-KL-T1): A, photograph; B, drawing. Both to scale, 5 mm. A forewing. Ax1, Ax2 = antenodal cross veins 1 and 2; arc = arculus; b = brace vein; CuA = anterior branch of the cubitus; IR1, IR2 = intercalar vein 1 and 2; MA, MP = the anterior and posterior branches of the media; nod = nodus; pt = pterostigma; Q = quadrangle; RA = radius anterior; RP1, RP2, RP3-4 = branches of the radius posterior; sn = subnodus; * = origin of RP2." figureDoi="http://doi.org/10.5281/zenodo.7906176" httpUri="https://zenodo.org/record/7906176/files/figure.png" pageId="16" pageNumber="305">Fig. 2</figureCitation>
|
||
): an almost complete forewing missing a portion of the posterior at about a third length, in a concretion, coll. Erwin Rettig at Klitgaard Cliff, Mors, Fur Formation, date unknown, in the Fur Museum.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,1332,2007]" pageId="16" pageNumber="305">
|
||
<collectionCode box="[199,289,1405,1431]" pageId="16" pageNumber="305">FUM-N</collectionCode>
|
||
11146 (previously
|
||
<collectionCode box="[514,570,1405,1431]" pageId="16" pageNumber="305">ERK</collectionCode>
|
||
GU E7) (
|
||
<figureCitation box="[688,775,1404,1430]" captionStart="FIGURE 13" captionStartId="17.[151,250,959,983]" captionTargetBox="[155,1434,181,928]" captionTargetId="figure-24@17.[151,1436,181,935]" captionTargetPageId="17" captionText="FIGURE 13. FUM-N 11146: A, photograph; B, drawing. Note infuscation (see text). Both to scale, 5 mm. A forewing." figureDoi="http://doi.org/10.5281/zenodo.7906200" httpUri="https://zenodo.org/record/7906200/files/figure.png" pageId="16" pageNumber="305">Fig. 13</figureCitation>
|
||
): an almost complete forewing missing parts of the wing basad the nodus, coll. Erwin Rettig at Gullerup; Mors, Fur Formation, date unknown, in the Fur Museum.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,1332,2007]" pageId="16" pageNumber="305">
|
||
<collectionCode box="[199,289,1477,1503]" pageId="16" pageNumber="305">FUM-N</collectionCode>
|
||
16192 (
|
||
<figureCitation box="[384,470,1476,1503]" captionStart="FIGURE 14" captionStartId="17.[151,250,1857,1881]" captionTargetBox="[151,1436,1008,1818]" captionTargetId="figure-48@17.[151,1436,1008,1833]" captionTargetPageId="17" captionText="FIGURE 14. FUM-N 16192: A, photograph; B, drawing. Both to scale, 5 mm. A hind wing." figureDoi="http://doi.org/10.5281/zenodo.7906202" httpUri="https://zenodo.org/record/7906202/files/figure.png" pageId="16" pageNumber="305">Fig. 14</figureCitation>
|
||
): an almost complete hind wing missing parts basad about Ax2, in a concretion, coll. Jan and Elly Verkleij at the Ejerslev mo-clay pit, Mors, Knudeklint Member, Fur Formation, 2008, in the Fur Museum.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,1332,2007]" pageId="16" pageNumber="305">
|
||
14M-2765 (
|
||
<figureCitation box="[333,416,1584,1611]" captionStart="FIGURE 15" captionStartId="18.[151,250,1183,1207]" captionTargetBox="[190,1397,181,1150]" captionTargetId="figure-28@18.[190,1397,181,1159]" captionTargetPageId="18" captionText="FIGURE 15. 14M-2765: A, photograph; B, drawing as preserved; C, the fore- or hind wing graphically unfolded. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906206" httpUri="https://zenodo.org/record/7906206/files/figure.png" pageId="16" pageNumber="305">Fig. 15</figureCitation>
|
||
): a partial thorax and basal fragment of abdomen with a [fore- or hind?] wing attached, the distal posterior portion of which is folded over the rest of the wing, coll. Henrik Madsen at the Ejerslev mo-clay pit, Mors, Knudeklint Member, Fur Formation,
|
||
<date box="[648,876,1656,1682]" pageId="16" pageNumber="305" value="1991-11-14">November 14, 1991</date>
|
||
, in the negative series, in the Fossil and Mo-clay Museum, Museum Mors.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,1332,2007]" pageId="16" pageNumber="305">
|
||
<collectionCode box="[199,289,1729,1755]" pageId="16" pageNumber="305">FUM-N</collectionCode>
|
||
11616 (
|
||
<figureCitation box="[382,466,1728,1754]" captionStart="FIGURE 16" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[178,1408,186,1919]" captionTargetId="figure-24@19.[178,1410,181,1919]" captionTargetPageId="19" captionText="FIGURE 16. FUM-N 11696: A, drawing of bottom wing in 17C; B, drawing to top wing in 17C; C, photograph. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906208" httpUri="https://zenodo.org/record/7906208/files/figure.png" pageId="16" pageNumber="305">Fig. 16</figureCitation>
|
||
): two very complete hind wings joined by a small fragment of thorax, in a concretion, coll. Jan and Elly Verkleij at Manhøj, new pit, Fur Formation, 2008, in the Fur Museum.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,1332,2007]" pageId="16" pageNumber="305">
|
||
<collectionCode box="[199,289,1801,1827]" pageId="16" pageNumber="305">FUM-N</collectionCode>
|
||
14704 (previously
|
||
<collectionCode box="[514,554,1801,1827]" country="China" name="Department of Nature, Fujian Province Museum" pageId="16" pageNumber="305">FM</collectionCode>
|
||
1106) (
|
||
<figureCitation box="[644,730,1800,1826]" captionStart="FIGURE 17" captionStartId="20.[151,250,913,937]" captionTargetBox="[151,1436,181,875]" captionTargetId="figure-28@20.[151,1436,181,889]" captionTargetPageId="20" captionText="FIGURE 17. FUM-N 14704: A, photograph; B, drawing. A forewing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906210" httpUri="https://zenodo.org/record/7906210/files/figure.png" pageId="16" pageNumber="305">Fig. 17</figureCitation>
|
||
): an almost complete forewing missing a small portion of the apex, coll. Erik Fjeldsø at Stolleklint Cliff, Silstrup Member, between ash layers +25 and +30, Fur Formation, date unknown, in the Fur Museum.
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1437,1332,2007]" pageId="16" pageNumber="305">
|
||
MM-2085 (previously MHM-i1583AB): (
|
||
<figureCitation box="[674,758,1908,1934]" captionStart="FIGURE 18" captionStartId="20.[151,250,1716,1740]" captionTargetBox="[151,1434,962,1680]" captionTargetId="figure-48@20.[151,1436,962,1693]" captionTargetPageId="20" captionText="FIGURE 18. MM-2085: A, photograph; B, drawing. A forewing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906212" httpUri="https://zenodo.org/record/7906212/files/figure.png" pageId="16" pageNumber="305">Fig. 18</figureCitation>
|
||
): an almost complete forewing in a concretion, coll. Henrik Madsen at Ejerslev mo-clay pit, Mors, Knudeklint Member, between ash layers +25 and +30, Fur Formation, 2005, in the Fossil and Mo-clay Museum, Museum Mors.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906200" ID-Zenodo-Dep="7906200" box="[151,1358,959,984]" httpUri="https://zenodo.org/record/7906200/files/figure.png" pageId="17" pageNumber="306" startId="17.[151,250,959,983]" targetBox="[155,1434,181,928]" targetPageId="17" targetType="figure">
|
||
<paragraph blockId="17.[151,1358,959,984]" box="[151,1358,959,984]" pageId="17" pageNumber="306">
|
||
<emphasis bold="true" box="[151,288,959,983]" pageId="17" pageNumber="306">FIGURE 13.</emphasis>
|
||
FUM-N 11146: A, photograph; B, drawing. Note infuscation (see text). Both to scale, 5 mm. A forewing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906202" ID-Zenodo-Dep="7906202" box="[151,1089,1857,1881]" httpUri="https://zenodo.org/record/7906202/files/figure.png" pageId="17" pageNumber="306" startId="17.[151,250,1857,1881]" targetBox="[151,1436,1008,1818]" targetPageId="17" targetType="figure">
|
||
<paragraph blockId="17.[151,1089,1857,1881]" box="[151,1089,1857,1881]" pageId="17" pageNumber="306">
|
||
<emphasis bold="true" box="[151,288,1857,1881]" pageId="17" pageNumber="306">FIGURE 14.</emphasis>
|
||
FUM-N 16192: A, photograph; B, drawing. Both to scale, 5 mm. A hind wing.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="17.[151,1436,1927,2025]" pageId="17" pageNumber="306">
|
||
<collectionCode box="[199,285,1928,1954]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="17" pageNumber="306">MGUH</collectionCode>
|
||
1819 (
|
||
<figureCitation box="[367,435,1927,1954]" captionStart="FIGURE 1" captionStartId="1.[152,255,1633,1657]" captionTargetBox="[151,1427,525,1601]" captionTargetId="figure-182@1.[151,1436,525,1610]" captionTargetPageId="1" captionText="FIGURE 1. MGUH 1819, Henriksen’s type specimen of Phenacolestes jutlandica: A, photograph; B, drawing; C–E, labelled wings. A, B to scale, C–E to scale, both scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906174" httpUri="https://zenodo.org/record/7906174/files/figure.png" pageId="17" pageNumber="306">Fig. 1</figureCitation>
|
||
): the distal portions of three wings and much of the abdomen, coll. and date unknown, Struer, loose slab from an unknown level within the Fur Formation, in the Geological Museum, University of Copenhagen,
|
||
<collectingCountry box="[305,413,1999,2025]" name="Denmark" pageId="17" pageNumber="306">Denmark</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906206" ID-Zenodo-Dep="7906206" httpUri="https://zenodo.org/record/7906206/files/figure.png" pageId="18" pageNumber="307" startId="18.[151,250,1183,1207]" targetBox="[190,1397,181,1150]" targetPageId="18" targetType="figure">
|
||
<paragraph blockId="18.[151,1436,1183,1243]" pageId="18" pageNumber="307">
|
||
<emphasis bold="true" box="[151,288,1183,1207]" pageId="18" pageNumber="307">FIGURE 15.</emphasis>
|
||
14M-2765: A, photograph; B, drawing as preserved; C, the fore- or hind wing graphically unfolded. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="18.[151,1437,1289,2035]" pageId="18" pageNumber="307">
|
||
14M-A2163 (
|
||
<figureCitation box="[357,428,1289,1315]" captionStart="FIGURE 6" captionStartId="9.[151,250,1269,1293]" captionTargetBox="[158,1436,181,1240]" captionTargetId="figure-24@9.[151,1436,181,1245]" captionTargetPageId="9" captionText="FIGURE 6. 14M-A2163: A, photograph; B, drawing; C. drawing of the hind wing (red in Fig. 6B). All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906186" httpUri="https://zenodo.org/record/7906186/files/figure.png" pageId="18" pageNumber="307">Fig. 6</figureCitation>
|
||
) (
|
||
<collectionCode box="[455,507,1289,1315]" country="France" name="University of Montpellier" pageId="18" pageNumber="307">MM</collectionCode>
|
||
14M-A2163 of
|
||
<bibRefCitation author="Rust, J." box="[701,822,1289,1316]" pageId="18" pageNumber="307" refId="ref13654" refString="Rust, J. (1999) Biologie der Insekten aus dem altesten Tertiar Nordeuropas. Habilitationsschrift zur Erlangung der venia legendi fu ¨ r das Fach Zoologie. Postdoctoral thesis, Biologische Fakultat der Georg-August-Universitat Gottingen, Gottingen, Lower Saxony, 482 pp." type="book" year="1999">Rust 1999</bibRefCitation>
|
||
, see his plate 1, fig. b): with a well-preserved head, including compound eyes, thorax, and most of the abdomen, the anterior of a forewing and much of a hind wing, parts of two legs, coll. Henrik Madsen at Ejerslev mo-clay pit, Mors, Knudeklint Member between ash layers -11 and -13, Fur Formation,
|
||
<date box="[427,617,1397,1423]" pageId="18" pageNumber="307" value="1993-08-21">August 21, 1993</date>
|
||
, in the Fossil and Mo-clay Museum. Although the distal IR1-RP2 region is missing and the maximum wing width is not known, the preserved portion of the hind wing is shaped like those of
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[182,333,1469,1495]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis box="[182,333,1469,1495]" italics="true" pageId="18" pageNumber="307">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
, clearly not as slender as the
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[663,788,1470,1495]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[663,788,1470,1495]" italics="true" pageId="18" pageNumber="307">F. ansorgei</emphasis>
|
||
</taxonomicName>
|
||
wing.
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,1289,2035]" pageId="18" pageNumber="307">
|
||
14M-3801 (
|
||
<figureCitation box="[332,414,1505,1531]" captionStart="FIGURE 19" captionStartId="21.[151,250,1429,1453]" captionTargetBox="[151,1436,187,1405]" captionTargetId="figure-24@21.[151,1436,181,1405]" captionTargetPageId="21" captionText="FIGURE 19. 14M-3801: A; drawing of forewing; B, drawing of the hind wing; C, photograph, complete. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906214" httpUri="https://zenodo.org/record/7906214/files/figure.png" pageId="18" pageNumber="307">Fig. 19</figureCitation>
|
||
): missing the head, thorax, and abdomen mostly complete, one forewing and one hind partly preserved, parts of three legs, in a concretion, coll. Henrik Madsen at the Ejerslev mo-clay pit, Mors, Knudeklint Member between ash layers -13 and -14, Fur Formation,
|
||
<date box="[819,1009,1577,1603]" pageId="18" pageNumber="307" value="1992-10-02">October 2, 1992</date>
|
||
, in the Fossil and Mo-clay Museum, Museum Mors. Although the distal IR1-RP2 region is missing, its wing shape is not as slender as the
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[1310,1436,1614,1639]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[1310,1436,1614,1639]" italics="true" pageId="18" pageNumber="307">F. ansorgei</emphasis>
|
||
</taxonomicName>
|
||
wing.
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,1289,2035]" pageId="18" pageNumber="307">
|
||
MOA 769/1, 2 (part, counterpart), “
|
||
<taxonomicName authorityName="Zessin" authorityYear="2011" box="[608,797,1686,1711]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="morsi">
|
||
<emphasis box="[608,797,1686,1711]" italics="true" pageId="18" pageNumber="307">Furagrion morsi</emphasis>
|
||
</taxonomicName>
|
||
” is known by the basal portion of a wing (
|
||
<figureCitation box="[1293,1376,1685,1711]" captionStart="FIGURE 20" captionStartId="22.[151,250,773,797]" captionTargetBox="[157,1431,189,744]" captionTargetId="figure-28@22.[151,1436,181,749]" captionTargetPageId="22" captionText="FIGURE 20. A, “Furagrion morsi” MOA 769 redrawn from part and counterpart photographs (Zessin 2011 Figs. 6 and 7); B–D, F. morsi (red) superimposed on F. jutlandicus wings: B, MHM 1742; C, D, FUM-N 11616. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906216" httpUri="https://zenodo.org/record/7906216/files/figure.png" pageId="18" pageNumber="307">Fig. 20</figureCitation>
|
||
). We could not locate the fossil and based our discussion on the description, photograph and drawing of
|
||
<bibRefCitation author="Zessin, W." box="[1274,1432,1721,1747]" pageId="18" pageNumber="307" pagination="64 - 73" refId="ref14309" refString="Zessin, W. (2011) Neue Insekten aus dem Moler (Palaozan / Eozan) von Danemark Teil 1 (Odonata: Epallagidae, Megapodagrioniidae), Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg, 14, 64 - 73." type="journal article" year="2011">Zessin (2011)</bibRefCitation>
|
||
. We find its putative differences with
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[571,724,1757,1783]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis box="[571,724,1757,1783]" italics="true" pageId="18" pageNumber="307">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[781,906,1758,1783]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[781,906,1758,1783]" italics="true" pageId="18" pageNumber="307">F. ansorgei</emphasis>
|
||
</taxonomicName>
|
||
to be within their intraspecific variations (
|
||
<figureCitation captionStart="FIGURE 20" captionStartId="22.[151,250,773,797]" captionTargetBox="[157,1431,189,744]" captionTargetId="figure-28@22.[151,1436,181,749]" captionTargetPageId="22" captionText="FIGURE 20. A, “Furagrion morsi” MOA 769 redrawn from part and counterpart photographs (Zessin 2011 Figs. 6 and 7); B–D, F. morsi (red) superimposed on F. jutlandicus wings: B, MHM 1742; C, D, FUM-N 11616. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906216" httpUri="https://zenodo.org/record/7906216/files/figure.png" pageId="18" pageNumber="307">Fig. 20B–D</figureCitation>
|
||
). The shape of its preserved portion is like
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[724,876,1793,1819]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis box="[724,876,1793,1819]" italics="true" pageId="18" pageNumber="307">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
not as slender as the
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[1120,1245,1794,1819]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[1120,1245,1794,1819]" italics="true" pageId="18" pageNumber="307">F. ansorgei</emphasis>
|
||
</taxonomicName>
|
||
wing, and so we treat “
|
||
<taxonomicName authorityName="Zessin" authorityYear="2011" box="[220,408,1830,1855]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="morsi">
|
||
<emphasis box="[220,408,1830,1855]" italics="true" pageId="18" pageNumber="307">Furagrion morsi</emphasis>
|
||
</taxonomicName>
|
||
” as a junior synonym of
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[691,842,1829,1855]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis box="[691,842,1829,1855]" italics="true" pageId="18" pageNumber="307">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,1289,2035]" pageId="18" pageNumber="307">
|
||
MM-11044 (
|
||
<figureCitation box="[345,432,1865,1891]" captionStart="FIGURE 21" captionStartId="22.[151,250,1657,1681]" captionTargetBox="[151,1436,858,1619]" captionTargetId="figure-79@22.[151,1436,858,1634]" captionTargetPageId="22" captionText="FIGURE 21. MM-11044: A, photograph; B, drawing.A fore- or hind wing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906218" httpUri="https://zenodo.org/record/7906218/files/figure.png" pageId="18" pageNumber="307">Fig. 21</figureCitation>
|
||
): basal portion of a wing in a concretion, coll. Henrik Madsen, 2018, Ejerslev mo-clay pit, Mors, Knudeklint Member, between ash layers +25 - +30, Fur Formation, in the Fossil and Mo-clay Museum, Museum Mors. Incomplete and lacking the IR1-RP2 space. Although the distal portion of the wing is missing, we tentatively assign this fossil to
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[493,643,1973,1999]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis box="[493,643,1973,1999]" italics="true" pageId="18" pageNumber="307">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
, as its preserved portion appears not as slender as an equivalent portion of the
|
||
<taxonomicName box="[223,335,2010,2035]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="307" phylum="Arthropoda" rank="species" species="ansorgi">
|
||
<emphasis box="[223,335,2010,2035]" italics="true" pageId="18" pageNumber="307">F. ansorgi</emphasis>
|
||
</taxonomicName>
|
||
wing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906208" ID-Zenodo-Dep="7906208" httpUri="https://zenodo.org/record/7906208/files/figure.png" pageId="19" pageNumber="308" startId="19.[151,250,1943,1967]" targetBox="[178,1408,186,1919]" targetPageId="19" targetType="figure">
|
||
<paragraph blockId="19.[151,1436,1943,2003]" pageId="19" pageNumber="308">
|
||
<emphasis bold="true" box="[151,288,1943,1967]" pageId="19" pageNumber="308">FIGURE 16.</emphasis>
|
||
FUM-N 11696: A, drawing of bottom wing in 17C; B, drawing to top wing in 17C; C, photograph. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906210" ID-Zenodo-Dep="7906210" box="[151,1079,913,938]" httpUri="https://zenodo.org/record/7906210/files/figure.png" pageId="20" pageNumber="309" startId="20.[151,250,913,937]" targetBox="[151,1436,181,875]" targetPageId="20" targetType="figure">
|
||
<paragraph blockId="20.[151,1079,913,938]" box="[151,1079,913,938]" pageId="20" pageNumber="309">
|
||
<emphasis bold="true" box="[151,288,913,937]" pageId="20" pageNumber="309">FIGURE 17.</emphasis>
|
||
FUM-N 14704: A, photograph; B, drawing. A forewing. Both to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906212" ID-Zenodo-Dep="7906212" box="[151,1032,1716,1741]" httpUri="https://zenodo.org/record/7906212/files/figure.png" pageId="20" pageNumber="309" startId="20.[151,250,1716,1740]" targetBox="[151,1434,962,1680]" targetPageId="20" targetType="figure">
|
||
<paragraph blockId="20.[151,1032,1716,1741]" box="[151,1032,1716,1741]" pageId="20" pageNumber="309">
|
||
<emphasis bold="true" box="[151,288,1716,1740]" pageId="20" pageNumber="309">FIGURE 18.</emphasis>
|
||
MM-2085: A, photograph; B, drawing. A forewing. Both to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906214" ID-Zenodo-Dep="7906214" box="[151,1421,1429,1453]" httpUri="https://zenodo.org/record/7906214/files/figure.png" pageId="21" pageNumber="310" startId="21.[151,250,1429,1453]" targetBox="[151,1436,187,1405]" targetPageId="21" targetType="figure">
|
||
<paragraph blockId="21.[151,1421,1429,1453]" box="[151,1421,1429,1453]" pageId="21" pageNumber="310">
|
||
<emphasis bold="true" box="[151,288,1429,1453]" pageId="21" pageNumber="310">FIGURE 19.</emphasis>
|
||
14M-3801: A; drawing of forewing; B, drawing of the hind wing; C, photograph, complete. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="21" pageNumber="310" type="discussion">
|
||
<paragraph blockId="21.[151,1437,1499,1706]" pageId="21" pageNumber="310">
|
||
<emphasis bold="true" box="[199,308,1499,1525]" pageId="21" pageNumber="310">Remarks</emphasis>
|
||
. The proposed
|
||
<typeStatus box="[479,568,1500,1526]" pageId="21" pageNumber="310">neotype</typeStatus>
|
||
MM-10752 was figured by
|
||
<bibRefCitation author="Pedersen, G. K. & Pedersen, S. A. S. & Bonde, N. & Heilman-Clausen, C. & Larsen, L. M. & Lindow, B. E. K. & Madsen, H. & Pedersen, A. K. & Rust, J. & Schultz, B. P. & Storey, M." box="[880,1116,1499,1525]" pageId="21" pageNumber="310" pagination="41 - 135" refId="ref13284" refString="Pedersen, G. K., Pedersen, S. A. S., Bonde, N., Heilman-Clausen, C., Larsen, L. M., Lindow, B. E. K., Madsen, H., Pedersen, A. K., Rust, J., Schultz, B. P. & Storey, M. (2012) Moleromradets geologi - sedimenter, fossiler, askelag og glacialtektonik [in Danish]. Geologisk Tidsskrift 2011, 41 - 135." type="journal article" year="2012">
|
||
Pedersen
|
||
<emphasis box="[987,1042,1499,1525]" italics="true" pageId="21" pageNumber="310">et al.</emphasis>
|
||
(2012
|
||
</bibRefCitation>
|
||
, fig. 46D) who characterised it as a damselfly (translated from Danish:
|
||
<emphasis box="[618,739,1535,1561]" italics="true" pageId="21" pageNumber="310">vandnymfe</emphasis>
|
||
) from the level just below ash layer -
|
||
<quantity box="[1155,1212,1535,1561]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="21" pageNumber="310" unit="in" value="13.0">13 in</quantity>
|
||
the Skarrehage mo-clay pit, Mors. The specimen was not named or described in further detail.
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1437,1499,1706]" pageId="21" pageNumber="310">
|
||
<emphasis bold="true" box="[199,394,1607,1633]" pageId="21" pageNumber="310">Range and age.</emphasis>
|
||
<materialsCitation country="Denmark" county="The" latitude="56.944168" location="Earliest Ypresian Fur Formation" longLatPrecision="10" longitude="8.8735" municipality="Jutland" pageId="21" pageNumber="310" specimenCount="1" typeStatus="neotype">
|
||
<location LSID="urn:lsid:plazi:treatment:03A5BA5CFFADFFBBFF40EAF7EDD24EB4:8ED35D91FFB7FFBBFE40EC27EA2C4E7C" box="[407,788,1607,1634]" country="Denmark" county="The" latitude="56.944168" longLatPrecision="10" longitude="8.8735" municipality="Jutland" name="Earliest Ypresian Fur Formation" pageId="21" pageNumber="310">Earliest Ypresian Fur Formation</location>
|
||
,
|
||
<collectingMunicipality box="[805,889,1607,1633]" pageId="21" pageNumber="310">Jutland</collectingMunicipality>
|
||
,
|
||
<collectingCountry box="[905,1013,1607,1633]" name="Denmark" pageId="21" pageNumber="310">Denmark</collectingCountry>
|
||
.
|
||
<collectingCounty box="[1029,1073,1607,1633]" pageId="21" pageNumber="310">The</collectingCounty>
|
||
proposed
|
||
<typeStatus box="[1201,1290,1608,1634]" pageId="21" pageNumber="310">neotype</typeStatus>
|
||
is from the Skarrehage mo-clay pit (
|
||
<geoCoordinate box="[427,560,1643,1670]" degrees="56" direction="north" minutes="56.65" orientation="latitude" pageId="21" pageNumber="310" precision="9" value="56.944168">N 56°56.65</geoCoordinate>
|
||
–,
|
||
<geoCoordinate box="[586,698,1643,1670]" degrees="8" direction="east" minutes="52.41" orientation="longitude" pageId="21" pageNumber="310" precision="9" value="8.8735">E 8°52.41</geoCoordinate>
|
||
–) between 61 and
|
||
<quantity box="[912,995,1643,1670]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="310" unit="cm" value="100.0">100 cm</quantity>
|
||
below ash layer -13, informally named the Skarrehage bed of the Knudeklint Member, Fur Formation. All specimens: as for genus, above
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |