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<document id="824CFC1731EE24D30E77DD29F54B14E4" ID-CLB-Dataset="29291" ID-DOI="10.11646/zootaxa.4493.1.1" ID-GBIF-Dataset="ac55c1cd-0227-47c0-a130-15d18759b313" ID-ISSN="1175-5326" ID-Zenodo-Dep="1445162" ID-ZooBank="BA78044B-2C16-4F64-AA20-D1838D423CCC" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1538637090652" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2018" docId="03A587B82A44795129B7DE17F7842EAC" docLanguage="en" docName="zootaxa.4493.1.1.pdf" docOrigin="Zootaxa 4493 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Guyalna woldai Sanborn 2018, n. sp." docType="treatment" docVersion="18" lastPageNumber="32" masterDocId="FF9CFFC02A5F794E2920DB44F6142811" masterDocTitle="The cicadas (Hemiptera: Cicadidae) of Panama including the description of six new species, three new combinations, one new synonymy, and nine new records" masterLastPageNumber="69" masterPageNumber="1" pageNumber="28" updateTime="1698566846198" updateUser="ExternalLinkService">
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<mods:title id="9CA7E1E94CC246954C475A4C59EC1437">The cicadas (Hemiptera: Cicadidae) of Panama including the description of six new species, three new combinations, one new synonymy, and nine new records</mods:title>
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<mods:namePart id="90D43593EF8BD38D52DD79C8DD8166C5">Sanborn, Allen F.</mods:namePart>
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<mods:title id="7BA26845AF4C55FBBD983247FA023D31">Zootaxa</mods:title>
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<mods:date id="6A9F5B9459FA79CA4AE7B83A4F063887">2018</mods:date>
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<mods:number id="493D7D76381949972A91183484403B11">2018-10-04</mods:number>
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<mods:classification id="FFF4ED76926B8ACD276F5771CF0FEFFA">journal article</mods:classification>
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<treatment id="03A587B82A44795129B7DE17F7842EAC" ID-DOI="http://doi.org/10.5281/zenodo.4480604" ID-GBIF-Taxon="148462739" ID-Zenodo-Dep="4480604" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC" httpUri="http://treatment.plazi.org/id/03A587B82A44795129B7DE17F7842EAC" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28">
<subSubSection id="C31665252A44795529B7DE17F7B42D7C" box="[151,416,1363,1389]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BB336AE2A44795529B7DE17F7B42D7C" blockId="27.[151,416,1363,1422]" box="[151,416,1363,1389]" pageId="27" pageNumber="28">
<heading id="D0FB81C22A44795529B7DE17F7B42D7C" bold="true" box="[151,416,1363,1389]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B978EABC2A44795529B7DE17F7B42D7C" bold="true" box="[151,416,1363,1389]" pageId="27" pageNumber="28">
<taxonomicName id="4C0C4D2D2A44795529B7DE17F7422D7C" ID-CoL="8JST5" authority="Sanborn, 2018" authorityName="Sanborn" authorityYear="2018" box="[151,342,1363,1389]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="woldai" status="sp. nov.">
<emphasis id="B978EABC2A44795529B7DE17F7422D7C" bold="true" box="[151,342,1363,1389]" italics="true" pageId="27" pageNumber="28">Guyalna woldai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C72A447955287DDE17F7B42D7C" box="[349,416,1363,1389]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31665252A44795529B7DE31F6FF2D9F" box="[151,235,1397,1422]" pageId="27" pageNumber="28" type="description">
<paragraph id="8BB336AE2A44795529B7DE31F6FF2D9F" blockId="27.[151,416,1363,1422]" box="[151,235,1397,1422]" pageId="27" pageNumber="28">
(
<figureCitation id="13372A2B2A44795529BFDE31F6F72D9F" box="[159,227,1397,1422]" captionStart="FIGURE 4" captionStartId="29.[151,250,1917,1939]" captionTargetBox="[214,1372,193,1896]" captionTargetId="figure@29.[214,1372,193,1896]" captionTargetPageId="29" captionText="FIGURE 4. Guyalna woldai n. sp.: A, holotype male and paratype female habitus; B, holotype male dorsum; C, holotype male timbal cover; D, paratype male operculum; E, paratype female operculum; F, holotype male lateral view of genitalia; G, holotype male posterior view of genitalia; H, paratype female lateral view of genitalia; I, paratype female ventral view of genitalia. Scale bar: A, 2 cm; B, 5 mm; C–E, 2 mm; F–G, 1 mm; H–I, 2 mm." httpUri="https://zenodo.org/record/1445170/files/figure.png" pageId="27" pageNumber="28">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31665252A44795529B7DEFAF57E2DE3" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BB336AE2A44795529B7DEFAF51F2DC2" blockId="27.[151,874,1469,1522]" box="[151,779,1469,1491]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB1C11802A44795529B7DEFAF51F2DC2" box="[151,779,1469,1491]" pageId="27" pageNumber="28">
<taxonomicName id="4C0C4D2D2A44795529B7DEFAF4352DC2" ID-CoL="8HFWG" authorityName="Metcalf" authorityYear="1952" box="[151,545,1469,1491]" class="Insecta" family="Cicadidae" genus="Dorisiana" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sp. b">
<emphasis id="B978EABC2A44795529B7DEFAF6EA2DC2" box="[151,254,1470,1491]" italics="true" pageId="27" pageNumber="28">Dorisiana</emphasis>
sp. B
<bibRefCitation id="EF9D4B5F2A447955281EDEF9F4352DC2" author="Wolda, H." box="[318,545,1469,1491]" journalOrPublisher="Oecologia, Berlin" pageId="27" pageNumber="28" pagination="437 - 442" part="80" refId="ref65636" refString="Wolda, H. (1989) Seasonal cues in tropical organisms: rainfall? Not necessarily. Oecologia, Berlin, 80, 437 - 442. https: // doi. org / 10.1007 / BF 00380064" title="Seasonal cues in tropical organisms: rainfall? Not necessarily" type="journal article" year="1989">
<treatmentCitation id="0AAD10BF2A447955281EDEF9F7F92DC2" author="Wolda" box="[318,493,1469,1491]" page="438" pageId="27" pageNumber="28" year="1989">Wolda 1989: 438</treatmentCitation>
–440
</bibRefCitation>
</taxonomicName>
,
<figureCitation id="13372A2B2A4479552B0BDEF9F49F2DC2" box="[555,651,1469,1491]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="14.[151,250,1911,1933]" captionStartId-1="16.[151,250,1914,1936]" captionStartId-2="22.[151,250,1850,1872]" captionTargetBox-0="[220,1378,193,1896]" captionTargetBox-1="[214,1372,193,1892]" captionTargetBox-2="[151,1433,517,1828]" captionTargetId-0="figure@14.[220,1378,193,1896]" captionTargetId-1="figure@16.[214,1372,193,1892]" captionTargetId-2="figure@22.[151,1436,517,1828]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionTargetPageId-2="22" captionText-0="FIGURE 1. Procollina quadrimaculata n. sp.: A, holotype male and paratype female habitus; B, holotype male dorsum; C, holotype male timbal; D, holotype male operculum; E, paratype female operculum; F, holotype male lateral view of genitalia; G, holotype male posterior view of genitalia; H, paratype female lateral view of genitalia; I, paratype female ventral view of genitalia. Scale bar: A, 2 cm; B, 5 mm; C–E, 2 mm; F–G, 1 mm; H–I, 2 mm." captionText-1="FIGURE 2. Procollina stigmosa n. sp.: A, holotype male and paratype female habitus; B, holotype male dorsum; C, paratype male timbal; D, paratype male operculum; E, paratype female operculum; F, holotype male lateral view of genitalia; G, holotype male posterior view of genitalia; H, paratype female lateral view of genitalia; I, paratype female ventral view of genitalia. Scale bar: A, 2 cm; B, 5 mm; C–E, 2 mm; F–G, 1 mm; H–I, 2 mm." captionText-2="FIGURE 3. Comparison of the dorsum and timbal covers of Proarna championi n. comb. to the type species of the genera Proarna and Pacarina. A, Proarna championi n. comb. dorsum; B, Proarna hilaris dorsum; C, Pacarina puella dorsum; D, Proarna championi n. comb. timbal cover; E, Proarna hilaris timbal cover; F, Pacarina puella timbal cover. Scale bar: A–C, 5 mm; D–F, 2 mm." httpUri-0="https://zenodo.org/record/1445164/files/figure.png" httpUri-1="https://zenodo.org/record/1445166/files/figure.png" httpUri-2="https://zenodo.org/record/1445168/files/figure.png" pageId="27" pageNumber="28">Figs. 1–3</figureCitation>
, Tables 1–2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB336AE2A44795529B7DE99F57E2DE3" blockId="27.[151,874,1469,1522]" box="[151,874,1500,1522]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB1C11802A44795529B7DE99F57E2DE3" box="[151,874,1500,1522]" pageId="27" pageNumber="28">
<taxonomicName id="4C0C4D2D2A44795529B7DE99F72C2DE3" ID-CoL="8HFWG" authorityName="Metcalf" authorityYear="1952" box="[151,312,1501,1522]" class="Insecta" family="Cicadidae" genus="Dorisiana" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A44795529B7DE99F6EA2DE3" box="[151,254,1501,1522]" italics="true" pageId="27" pageNumber="28">Dorisiana</emphasis>
sp. B
</taxonomicName>
<bibRefCitation id="EF9D4B5F2A447955281EDE98F4932DE3" author="Wolda, H. &amp; Ramos, J. A." box="[318,647,1500,1522]" editor="Quintero, D. &amp; Aiello, A." journalOrPublisher="Oxford University Press, New York" pageId="27" pageNumber="28" pagination="271 - 279" refId="ref65723" refString="Wolda, H. &amp; Ramos, J. A. (1992) Cicadas in Panama, their distribution, seasonality and diversity. In: Quintero, D. &amp; Aiello, A. (Eds.), The insects of Panama and Mesoamerica. Selected studies. Oxford University Press, New York, pp. 271 - 279." title="Cicadas in Panama, their distribution, seasonality and diversity" type="book chapter" volumeTitle="The insects of Panama and Mesoamerica. Selected studies" year="1992">
<treatmentCitation id="0AAD10BF2A447955281EDE98F4472DE3" author="Wolda &amp; Ramos" box="[318,595,1500,1522]" page="272" pageId="27" pageNumber="28" year="1992">Wolda &amp; Ramos 1992: 272</treatmentCitation>
–273
</bibRefCitation>
, Fig. 17.1, Table 17.1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31665252A44795229B7DD5AF4932C01" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BB336AE2A44795229B7DD5AF6E328E9" blockId="27.[151,1437,1566,2023]" lastBlockId="28.[151,1437,151,1977]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B978EABC2A44795529B7DD5AF75F2E26" bold="true" box="[151,331,1566,1591]" pageId="27" pageNumber="28">
<typeStatus id="54B7880C2A44795529B7DD5AF6C62E26" box="[151,210,1566,1591]" pageId="27" pageNumber="28">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="3B643CF32A4479552879DD5AF2992E4A" ID-GBIF-Occurrence="4127563328" collectingDate="2015-06-01" collectingDateMax="2015-06-09" collectingDateMin="2015-06-01" collectionCode="TAMU" collectorName="E. Riley" country="Panama" elevation="90" latitude="8.0198" location="PANAMA" longLatPrecision="7" longitude="-77.7322" pageId="27" pageNumber="28" specimenCount="2" stateProvince="PANAMA" typeStatus="holotype">
<typeStatus id="54B7880C2A4479552879DD5AF7D12E26" box="[345,453,1566,1591]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
. “
<collectingCountry id="F31B763E2A44795528C0DD5BF44A2E26" box="[480,606,1567,1591]" name="Panama" pageId="27" pageNumber="28">PANAMA</collectingCountry>
:
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Prov. / P.N. Darién Rancho /
<location id="8ED360752A4479552D16DD5AF2D02E26" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4479552D16DD5AF2D02E26" box="[1078,1220,1566,1591]" country="Panama" latitude="8.0198" longLatPrecision="7" longitude="-77.7322" name="Frio Station" pageId="27" pageNumber="28" stateProvince="Darien">Frio Station</location>
,
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,
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</collectingDate>
,
<collectorName id="26F953782A44795528F8DD07F4222E4A" box="[472,566,1602,1627]" pageId="27" pageNumber="28">E. Riley</collectorName>
, UV // TAMU—ENTO / X1229169” male (
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).
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<typeStatus id="54B7880C2A4479552DB5DD07F3122E4A" box="[1173,1286,1603,1627]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
. “
<collectingCountry id="F31B763E2A4479552C3BDD07F38D2E4A" box="[1307,1433,1603,1627]" name="Panama" pageId="27" pageNumber="28">PANAMA</collectingCountry>
:
<collectingRegion id="49C8F84C2A44795529B7DD22F6F72E6E" box="[151,227,1638,1663]" country="Panama" name="Darien" pageId="27" pageNumber="28">Darién</collectingRegion>
Prov. / P.N. Darién Rancho /
<location id="8ED360752A4479552B64DD22F4DB2E6E" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4479552B64DD22F4DB2E6E" box="[580,719,1638,1663]" country="Panama" latitude="8.0198" longLatPrecision="7" longitude="-77.7322" name="Frio Station" pageId="27" pageNumber="28" stateProvince="Darien">Frio Station</location>
,
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<quantity id="4CF49B4B2A4479552BFCDD23F51A2E6E" box="[732,782,1639,1663]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">90m</quantity>
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/
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,
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/
<collectingDate id="EFF6E9862A4479552D1FDD22F2CA2E6E" box="[1087,1246,1638,1663]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">
<date id="FFB2106E2A4479552D1FDD22F2CA2E6E" box="[1087,1246,1638,1663]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">VI–1–9–2015</date>
</collectingDate>
,
<collectorName id="26F953782A4479552DCCDD23F3582E6E" box="[1260,1356,1638,1663]" pageId="27" pageNumber="28">E. Riley</collectorName>
, UV // TAMU—ENTO / X1230490” one female (
<collectionCode id="ED1DAE6B2A4479552B59DDCFF4C42EB2" box="[633,720,1675,1699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="27" pageNumber="28">TAMU</collectionCode>
);
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<materialsCitation id="3B643CF32A4479552BC3DDCFF3882ED6" ID-GBIF-Occurrence="4127563377" collectingDate="2015-06-01" collectingDateMax="2015-06-09" collectingDateMin="2015-06-01" collectionCode="TAMU" collectorName="E. Riley" country="Panama" elevation="90" latitude="8.0198" location="Frio Station" longLatPrecision="7" longitude="-77.7322" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Darien" typeStatus="paratype">
“
<collectingCountry id="F31B763E2A4479552BCDDDCFF57F2EB2" box="[749,875,1675,1699]" name="Panama" pageId="27" pageNumber="28">PANAMA</collectingCountry>
:
<collectingRegion id="49C8F84C2A4479552A56DDCEF5D62EB2" box="[886,962,1674,1699]" country="Panama" name="Darien" pageId="27" pageNumber="28">Darién</collectingRegion>
Prov. / P.N. Darién Rancho /
<location id="8ED360752A4479552C30DDCEF38C2EB2" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4479552C30DDCEF38C2EB2" box="[1296,1432,1674,1699]" country="Panama" latitude="8.0198" longLatPrecision="7" longitude="-77.7322" name="Frio Station" pageId="27" pageNumber="28" stateProvince="Darien">Frio Station</location>
,
<elevation id="0021D19D2A44795529B7DDEBF6DC2ED6" box="[151,200,1711,1735]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">
<quantity id="4CF49B4B2A44795529B7DDEBF6DC2ED6" box="[151,200,1711,1735]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">90m</quantity>
</elevation>
/
<geoCoordinate id="EE3850692A44795529C2DDEBF75A2ED6" box="[226,334,1711,1735]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="8.0198">8.0198°N</geoCoordinate>
,
<geoCoordinate id="EE3850692A447955287BDDEBF7CD2ED6" box="[347,473,1711,1735]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-77.7322">77.7322°W</geoCoordinate>
/
<collectingDate id="EFF6E9862A44795528D2DDEAF4842ED6" box="[498,656,1710,1735]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">
<date id="FFB2106E2A44795528D2DDEAF4842ED6" box="[498,656,1710,1735]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">VI–1–9–2015</date>
</collectingDate>
,
<collectorName id="26F953782A4479552BBDDDEBF4E82ED6" box="[669,764,1710,1735]" pageId="27" pageNumber="28">E. Riley</collectorName>
, UV // TAMU—ENTO / X1229392” one female (
<collectionCode id="ED1DAE6B2A4479552C19DDEBF3842ED6" box="[1337,1424,1711,1735]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="27" pageNumber="28">TAMU</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A44795529B7DD97F57C2F1E" ID-GBIF-Occurrence="4412493302" collectingDate="2015-06-01" collectingDateMax="2015-06-09" collectingDateMin="2015-06-01" collectionCode="TAMU" collectorName="E. Riley" country="Panama" elevation="90" latitude="8.0198" location="Frio Station" longLatPrecision="7" longitude="-77.7322" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Darien" typeStatus="paratype">
“
<collectingCountry id="F31B763E2A4479552981DD97F70B2EFA" box="[161,287,1747,1771]" name="Panama" pageId="27" pageNumber="28">PANAMA</collectingCountry>
:
<collectingRegion id="49C8F84C2A447955280CDD96F76C2EFA" box="[300,376,1746,1771]" country="Panama" name="Darien" pageId="27" pageNumber="28">Darién</collectingRegion>
Prov. / P.N. Darién Rancho /
<location id="8ED360752A4479552BFADD96F5712EFA" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4479552BFADD96F5712EFA" box="[730,869,1746,1771]" country="Panama" latitude="8.0198" longLatPrecision="7" longitude="-77.7322" name="Frio Station" pageId="27" pageNumber="28" stateProvince="Darien">Frio Station</location>
,
<elevation id="0021D19D2A4479552A53DD97F5B12EFA" box="[883,933,1747,1771]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">
<quantity id="4CF49B4B2A4479552A53DD97F5B12EFA" box="[883,933,1747,1771]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">90m</quantity>
</elevation>
/
<geoCoordinate id="EE3850692A4479552AE2DD97F23B2EFA" box="[962,1071,1747,1771]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="8.0198">8.0198°N</geoCoordinate>
,
<geoCoordinate id="EE3850692A4479552D1DDD97F2AE2EFA" box="[1085,1210,1747,1771]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-77.7322">77.7322°W</geoCoordinate>
/
<collectingDate id="EFF6E9862A4479552DF7DD96F3622EFA" box="[1239,1398,1746,1771]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">
<date id="FFB2106E2A4479552DF7DD96F3622EFA" box="[1239,1398,1746,1771]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">VI–1–9–2015</date>
</collectingDate>
,
<collectorName id="26F953782A4479552CA4DD97F6C12F1E" pageId="27" pageNumber="28">E. Riley</collectorName>
, UV // TAMU—ENTO / X1229374” one female (
<collectionCode id="ED1DAE6B2A4479552A30DDB3F5482F1E" box="[784,860,1783,1807]" pageId="27" pageNumber="28">AFSC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A4479552A51DDB3F7152F46" ID-GBIF-Occurrence="4126471338" collectingDate="2015-06-01" collectingDateMax="2015-06-09" collectingDateMin="2015-06-01" collectionCode="AFSC" collectorName="E. Riley" country="Panama" elevation="90" latitude="8.0198" location="Frio Station" longLatPrecision="7" longitude="-77.7322" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Darien" typeStatus="paratype">
“
<collectingCountry id="F31B763E2A4479552A5BDDB3F5ED2F1E" box="[891,1017,1783,1807]" name="Panama" pageId="27" pageNumber="28">PANAMA</collectingCountry>
:
<collectingRegion id="49C8F84C2A4479552D24DDB2F2442F1E" box="[1028,1104,1782,1807]" country="Panama" name="Darien" pageId="27" pageNumber="28">Darién</collectingRegion>
Prov. / P.N. Darién Rancho /
<location id="8ED360752A44795529B7DC5EF70A2F22" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A44795529B7DC5EF70A2F22" box="[151,286,1818,1843]" country="Panama" latitude="8.0198" longLatPrecision="7" longitude="-77.7322" name="Frio Station" pageId="27" pageNumber="28" stateProvince="Darien">Frio Station</location>
,
<elevation id="0021D19D2A4479552808DC5FF74E2F22" box="[296,346,1819,1843]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">
<quantity id="4CF49B4B2A4479552808DC5FF74E2F22" box="[296,346,1819,1843]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">90m</quantity>
</elevation>
/
<geoCoordinate id="EE3850692A447955284EDC5FF7CF2F22" box="[366,475,1819,1843]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="8.0198">8.0198°N</geoCoordinate>
,
<geoCoordinate id="EE3850692A44795528C6DC5FF4772F22" box="[486,611,1819,1843]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-77.7322">77.7322°W</geoCoordinate>
/
<date id="FFB2106E2A4479552B58DC5EF5032F22" box="[632,791,1818,1843]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">
<collectingDate id="EFF6E9862A4479552B58DC5EF5032F22" box="[632,791,1818,1843]" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">VI–1–9–2015</collectingDate>
</date>
,
<collectorName id="26F953782A4479552A01DC5FF56A2F22" box="[801,894,1818,1843]" pageId="27" pageNumber="28">E. Riley</collectorName>
, UV // TAMU—ENTO / X1230108” one female (
<collectionCode id="ED1DAE6B2A44795529BEDC7BF6E12F46" box="[158,245,1855,1879]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="27" pageNumber="28">TAMU</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A447955282BDC7BF5D52F6A" ID-GBIF-Occurrence="4126471331" collectingDate="2015-06-01" collectingDateMax="2015-06-09" collectingDateMin="2015-06-01" collectionCode="AFSC" collectorName="E. Riley" country="Panama" elevation="90" latitude="8.0198" location="Frio Station" longLatPrecision="7" longitude="-77.7322" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Darien" typeStatus="paratype">
“
<collectingCountry id="F31B763E2A4479552834DC7BF7862F46" box="[276,402,1855,1879]" name="Panama" pageId="27" pageNumber="28">PANAMA</collectingCountry>
: Darién Prov. / P.N.
<collectingRegion id="49C8F84C2A4479552BA0DC7AF4D82F46" box="[640,716,1854,1879]" country="Panama" name="Darien" pageId="27" pageNumber="28">Darién</collectingRegion>
Rancho /
<location id="8ED360752A4479552A64DC7AF5DA2F46" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4479552A64DC7AF5DA2F46" box="[836,974,1854,1879]" country="Panama" latitude="8.0198" longLatPrecision="7" longitude="-77.7322" name="Frio Station" pageId="27" pageNumber="28" stateProvince="Darien">Frio Station</location>
,
<elevation id="0021D19D2A4479552AFBDC7BF2192F46" box="[987,1037,1855,1879]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">
<quantity id="4CF49B4B2A4479552AFBDC7BF2192F46" box="[987,1037,1855,1879]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="90.0">90m</quantity>
</elevation>
/
<geoCoordinate id="EE3850692A4479552D07DC7BF2802F46" box="[1063,1172,1855,1879]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="8.0198">8.0198°N</geoCoordinate>
,
<geoCoordinate id="EE3850692A4479552D81DC7BF30A2F46" box="[1185,1310,1855,1879]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-77.7322">77.7322°W</geoCoordinate>
/
<collectingDate id="EFF6E9862A4479552C19DC7BF6C62F6A" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">
<date id="FFB2106E2A4479552C19DC7BF6C62F6A" pageId="27" pageNumber="28" value="2015-06-01" valueMax="2015-06-09" valueMin="2015-06-01">VI–1–9– 2015</date>
</collectingDate>
,
<collectorName id="26F953782A44795529FFDC27F72A2F6A" box="[223,318,1890,1915]" pageId="27" pageNumber="28">E. Riley</collectorName>
, UV // TAMU—ENTO / X1229755” one male (
<collectionCode id="ED1DAE6B2A4479552A49DC27F5A12F6A" box="[873,949,1891,1915]" pageId="27" pageNumber="28">AFSC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A4479552AEADC26F4352FD2" ID-GBIF-Occurrence="4126471353" collectingDate="2015-06-01" collectingDateMax="2015-06-09" collectingDateMin="2015-06-01" collectionCode="AFSC" collectorName="E. Riley" country="Panama" elevation="90" latitude="8.0198" location="Frio Station" longLatPrecision="7" longitude="-77.7322" municipality="Barro Colorado Isl." pageId="27" pageNumber="28" specimenCode="SDNHM094780" specimenCount="1" stateProvince="Darien" typeStatus="paratype">
“
<collectorName id="26F953782A4479552AF4DC26F24F2F6B" box="[980,1115,1890,1915]" pageId="27" pageNumber="28">Canal Zone</collectorName>
/
<collectorName id="26F953782A4479552D53DC27F2B52F6B" box="[1139,1185,1891,1914]" pageId="27" pageNumber="28">Pan</collectorName>
.
<collectingDate id="EFF6E9862A4479552D8DDC26F32F2F6A" box="[1197,1339,1890,1915]" pageId="27" pageNumber="28" value="1958-09-12">
<date id="FFB2106E2A4479552D8DDC26F32F2F6A" box="[1197,1339,1890,1915]" pageId="27" pageNumber="28" value="1958-09-12">IX–12–1958</date>
</collectingDate>
//
<collectingMunicipality id="6BD7ACD42A4479552C7CDC27F73B2F8E" pageId="27" pageNumber="28">Barro Colorado Isl.</collectingMunicipality>
/
<location id="8ED360752A4479552870DCC3F7CE2F8E" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4479552870DCC3F7CE2F8E" box="[336,474,1926,1951]" country="Panama" municipality="Barro Colorado Isl." name="Gatun Lake" pageId="27" pageNumber="28" stateProvince="Darien">Gatun Lake</location>
//
<collectorName id="26F953782A4479552B23DCC2F46E2F8F" box="[515,634,1926,1951]" pageId="27" pageNumber="28">Charles F.</collectorName>
Harbison /
<collectorName id="26F953782A4479552A2EDCC2F5622F8E" box="[782,886,1926,1951]" pageId="27" pageNumber="28">Collector</collectorName>
//
<collectionCode id="ED1DAE6B2A4479552A80DCC3F2112F8E" box="[928,1029,1927,1951]" country="USA" httpUri="http://grbio.org/cool/4ksk-xsw5" name="San Diego Natural History Museum" pageId="27" pageNumber="28">SDNHM</collectionCode>
/ COLLECTION / DATABASE /
<specimenCode id="DBAA9ED52A44795529B7DCEFF7472FD2" box="[151,339,1963,1987]" collectionCode="SDNHM" country="USA" httpUri="http://grbio.org/cool/4ksk-xsw5" name="San Diego Natural History Museum" pageId="27" pageNumber="28">SDNHM094780</specimenCode>
” female (
<collectionCode id="ED1DAE6B2A447955289FDCEFF4012FD2" box="[447,533,1963,1987]" pageId="27" pageNumber="28">SDMC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A4479552B09DCEEF48C2FF6" ID-GBIF-Occurrence="4127563403" collectingDate="2015-06-01" collectingDateMax="2015-06-09" collectingDateMin="2015-06-01" collectionCode="AFSC" collectorName="E. Riley" country="Panama" elevation="90" latitude="8.0198" location="Frio Station" longLatPrecision="7" longitude="-77.7322" municipality="Barro Colorado Isl." pageId="27" pageNumber="28" specimenCode="SDNHM094780" specimenCount="1" stateProvince="Darien" typeStatus="paratype">
“
<collectorName id="26F953782A4479552B13DCEEF4AC2FD3" box="[563,696,1962,1987]" pageId="27" pageNumber="28">Canal Zone</collectorName>
/
<collectorName id="26F953782A4479552BEFDCEFF4E92FD3" box="[719,765,1963,1986]" pageId="27" pageNumber="28">Pan</collectorName>
.
<collectingDate id="EFF6E9862A4479552A27DCEEF5812FD2" box="[775,917,1962,1987]" pageId="27" pageNumber="28" value="1958-09-12">
<date id="FFB2106E2A4479552A27DCEEF5812FD2" box="[775,917,1962,1987]" pageId="27" pageNumber="28" value="1958-09-12">IX–12–1958</date>
</collectingDate>
//
<collectingMunicipality id="6BD7ACD42A4479552A93DCEFF29A2FD2" box="[947,1166,1962,1987]" pageId="27" pageNumber="28">Barro Colorado Isl.</collectingMunicipality>
/
<location id="8ED360752A4479552D84DCEFF33C2FD2" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4479552D84DCEFF33C2FD2" box="[1188,1320,1962,1987]" country="Panama" municipality="Barro Colorado Isl." name="Gatun Lake" pageId="27" pageNumber="28">Gatun Lake</location>
//
<collectorName id="26F953782A4479552C67DCEEF6B82FF7" pageId="27" pageNumber="28">Charles F.</collectorName>
Harbison /
<collectorName id="26F953782A4479552814DC8AF78B2FF6" box="[308,415,1998,2023]" pageId="27" pageNumber="28">Collector</collectorName>
” one female (
<collectionCode id="ED1DAE6B2A4479552B60DC8BF4982FF6" box="[576,652,1999,2023]" pageId="27" pageNumber="28">AFSC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A4479522B81DC8BF7DC28A1" ID-GBIF-Occurrence="4126471350" collectingDate="1958-09-12" collectingDateMax="1986-08-22" collectingDateMin="1958-09-12" collectionCode="AFSC" collectorName="C. Riley Nelson" country="Panama" lastPageId="28" lastPageNumber="29" location="Gatun Lake" municipality="Barro Colorado Isl." pageId="27" pageNumber="28" specimenCount="1" typeStatus="paratype">
“
<collectingCountry id="F31B763E2A4479552B8BDC8BF5302FF6" box="[683,804,1999,2023]" name="Panama" pageId="27" pageNumber="28">PANAMA</collectingCountry>
Gamboa /
<collectingDate id="EFF6E9862A4479552A86DC8BF2482FF6" box="[934,1116,1998,2023]" pageId="27" pageNumber="28" value="1986-08-22">
<date id="FFB2106E2A4479552A86DC8BF2482FF6" box="[934,1116,1998,2023]" pageId="27" pageNumber="28" value="1986-08-22">22 August 1986</date>
</collectingDate>
/
<collectorName id="26F953782A4479552D56DC8BF3392FF6" box="[1142,1325,1998,2023]" pageId="27" pageNumber="28">C. Riley Nelson</collectorName>
/ malaise trap” one female (AFSC);
</materialsCitation>
<materialsCitation id="3B643CF32A43795228F5DBDCF55528C5" ID-GBIF-Occurrence="1914127211" collectingDate="1989-05-17" collectingDateMax="1989-05-18" collectingDateMin="1989-05-17" collectionCode="UMRM" collectorName="E. Riley &amp; LaDoux" country="Panama" determinerName="Det. L. &amp; Burgess" location="Cardenas Village" pageId="28" pageNumber="29" specimenCount="1">
“
<collectingCountry id="F31B763E2A43795228FFDBDCF44928A1" box="[479,605,152,176]" name="Panama" pageId="28" pageNumber="29">PANAMA</collectingCountry>
:
<collectorName id="26F953782A4379522B4CDBDCF48A28A1" box="[620,670,152,176]" name="International Commission on Zoological Nomenclature" pageId="28" pageNumber="29">C.Z.</collectorName>
/
<location id="8ED360752A4379522B9EDBD3F59028A1" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4379522B9EDBD3F59028A1" box="[702,900,151,176]" country="Panama" name="Cardenas Village" pageId="28" pageNumber="29">Cardenas Village</location>
/
<collectingDate id="EFF6E9862A4379522A84DBDCF26128A1" box="[932,1141,152,176]" pageId="28" pageNumber="29" value="1989-05-17" valueMax="1989-05-18" valueMin="1989-05-17">
<date id="FFB2106E2A4379522A84DBDCF26128A1" box="[932,1141,152,176]" pageId="28" pageNumber="29" value="1989-05-17" valueMax="1989-05-18" valueMin="1989-05-17">May 17–18, 1989</date>
</collectingDate>
/
<collectorName id="26F953782A4379522DB5DBDCF2E228A1" box="[1173,1270,151,176]" pageId="28" pageNumber="29">E. Riley</collectorName>
&amp;
<collectorName id="26F953782A4379522C04DBDCF39428BE" box="[1316,1408,152,175]" pageId="28" pageNumber="29">LaDoux</collectorName>
//
<taxonomicName id="4C0C4D2D2A43795229B7DBF8F73D28C4" authorityName="Stal" authorityYear="1870" box="[151,297,188,213]" class="Insecta" family="Cicadidae" genus="Diceroprocta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="63" phylum="Arthropoda" rank="genus">Diceroprocta</taxonomicName>
spp. /
<determinerName id="7E6FDA6A2A437952285DDBF9F42128C5" box="[381,565,188,212]" pageId="28" pageNumber="29">Det. L. Burgess</determinerName>
” one female (
<collectionCode id="ED1DAE6B2A4379522BFADBF8F52D28C5" box="[730,825,188,212]" country="USA" httpUri="http://grbio.org/cool/w4qt-0wzz" name="W.R. Enns Entomology Museum" pageId="28" pageNumber="29">UMRM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CF32A4379522A6FDBF8F6E328E9" ID-GBIF-Occurrence="4127563415" collectingDate="1967-09" collectingDateMax="1967-10-31" collectingDateMin="1967-09" collectionCode="FSCA" country="Panama" location="Curiche" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Choco">
“
<location id="8ED360752A4379522A79DBF8F5A328C4" LSID="urn:lsid:plazi:treatment:03A587B82A44795129B7DE17F7842EAC:8ED360752A4379522A79DBF8F5A328C4" box="[857,951,188,213]" country="Panama" name="Curiche" pageId="28" pageNumber="29" stateProvince="Choco">Curiche</location>
,
<collectingRegion id="49C8F84C2A4379522AE5DBF8F21A28C4" box="[965,1038,188,213]" country="Panama" name="Embera-Wounaan" pageId="28" pageNumber="29">Choco</collectingRegion>
/ Col.,
<collectingDate id="EFF6E9862A4379522D4BDBF8F31D28C4" box="[1131,1289,188,213]" pageId="28" pageNumber="29" value="1967-09" valueMax="1967-10-31" valueMin="1967-09-00">
<date id="FFB2106E2A4379522D4BDBF8F31D28C4" box="[1131,1289,188,213]" pageId="28" pageNumber="29" value="1967-09" valueMax="1967-10-31" valueMin="1967-09-00">Sept.–Oct. 67</date>
</collectingDate>
” one female (
<collectionCode id="ED1DAE6B2A43795229BFDBA4F6FF28E9" box="[159,235,224,248]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="28" pageNumber="29">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB336AE2A43795229E7DA40F4932C01" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="29">
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,
<collectingRegion id="49C8F84C2A4379522DD8D864F3452B26" box="[1272,1361,800,823]" country="Panama" name="Panama" pageId="28" pageNumber="29">Panama</collectingRegion>
Prov., Canal Zone, BCI, Lab Clearing, 9.1634 -79.8382,
<date id="FFB2106E2A4379522BF0D800F54F2B4D" box="[720,859,836,860]" pageId="28" pageNumber="29" value="1972-06-12">12 Jun 1972</date>
to
<date id="FFB2106E2A4379522AA0D800F20E2B4D" box="[896,1050,836,860]" pageId="28" pageNumber="29" value="1972-06-17">17 June 1972</date>
, R.H. Pine,
<collectionCode id="ED1DAE6B2A4379522D83D800F2CB2B4D" box="[1187,1247,836,860]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
_
<collectionCode id="ED1DAE6B2A4379522DCDD800F3302B4D" box="[1261,1316,836,860]" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="28" pageNumber="29">ENT</collectionCode>
_ 0 0 0 0 451 (
<collectionCode id="ED1DAE6B2A43795229BFD82CF6F52B91" box="[159,225,872,896]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A43795229DCD82CF7742BB5" ID-GBIF-Occurrence="4126471394" pageId="28" pageNumber="29">
<collectingCountry id="F31B763E2A43795229DCD82CF74C2B6E" box="[252,344,872,895]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
,
<collectingRegion id="49C8F84C2A4379522849D82CF7D62B6E" box="[361,450,872,895]" country="Panama" name="Panama" pageId="28" pageNumber="29">Panama</collectingRegion>
Prov., Canal Zone, BCI, 9.1667 -79.8333,
<date id="FFB2106E2A4379522AF4D82CF2472B91" box="[980,1107,872,896]" pageId="28" pageNumber="29" value="1978-10-19">19–Oct–78</date>
, Henk Wolda,
<collectionCode id="ED1DAE6B2A4379522C2DD82CF35D2B91" box="[1293,1353,872,896]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
_
<collectionCode id="ED1DAE6B2A4379522C77D82CF39A2B91" box="[1367,1422,872,896]" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="28" pageNumber="29">ENT</collectionCode>
_ 0 0 0 0 452 (
<collectionCode id="ED1DAE6B2A4379522832D8C8F7402BB5" box="[274,340,908,932]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A4379522851D8C9F7E22BD9" ID-GBIF-Occurrence="4126471422" pageId="28" pageNumber="29">
<collectingCountry id="F31B763E2A4379522851D8C9F7D92BB5" box="[369,461,909,932]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
,
<collectingRegion id="49C8F84C2A43795228C2D8C9F42F2BB5" box="[482,571,909,932]" country="Panama" name="Panama" pageId="28" pageNumber="29">Panama</collectingRegion>
Prov., Canal Zone, BCI, 9.1667 -79.8333,
<date id="FFB2106E2A4379522D40D8C8F2FE2BB4" box="[1120,1258,908,933]" pageId="28" pageNumber="29" value="1992-05-26">26–May–92</date>
, Henk Wolda,
<collectionCode id="ED1DAE6B2A43795229B7D8F4F6C72BD9" box="[151,211,944,968]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
_
<collectionCode id="ED1DAE6B2A43795229C1D8F4F70C2BD9" box="[225,280,944,968]" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="28" pageNumber="29">ENT</collectionCode>
_ 0 0 0 0 453 (
<collectionCode id="ED1DAE6B2A4379522887D8F4F7FE2BD9" box="[423,490,944,968]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A4379522B22D8F4F45B2BFD" ID-GBIF-Occurrence="4126471429" pageId="28" pageNumber="29">
<collectingCountry id="F31B763E2A4379522B22D8F4F44A2BD6" box="[514,606,944,967]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
,
<collectingRegion id="49C8F84C2A4379522B4DD8F4F4D22BD6" box="[621,710,944,967]" country="Panama" name="Panama" pageId="28" pageNumber="29">Panama</collectingRegion>
Prov., Canal Zone, BCI, 9.1667 -79.8333,
<date id="FFB2106E2A4379522DE8D8F4F3442BD9" box="[1224,1360,944,968]" pageId="28" pageNumber="29" value="1977-08-23">23–Aug–77</date>
, Henk Wolda,
<collectionCode id="ED1DAE6B2A43795229D1D890F7392BFD" box="[241,301,980,1004]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
_
<collectionCode id="ED1DAE6B2A437952281BD890F7662BFD" box="[315,370,980,1004]" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="28" pageNumber="29">ENT</collectionCode>
_ 0 0 0 0 454 (
<collectionCode id="ED1DAE6B2A4379522B20D890F4562BFD" box="[512,578,980,1004]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B643CF32A4379522B7AD891F4932C01" ID-GBIF-Occurrence="4126471333" pageId="28" pageNumber="29">
<collectingCountry id="F31B763E2A4379522B7AD891F4A22BFD" box="[602,694,981,1004]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
,
<collectingRegion id="49C8F84C2A4379522BE5D891F50A2BFD" box="[709,798,981,1004]" country="Panama" name="Panama" pageId="28" pageNumber="29">Panama</collectingRegion>
Prov., Canal Zone, BCI, 9.1667 -79.8333,
<date id="FFB2106E2A4379522C3CD890F38C2BFD" box="[1308,1432,980,1004]" pageId="28" pageNumber="29" value="1992-05-08">8–May–92</date>
, Henk Wolda,
<collectionCode id="ED1DAE6B2A4379522811D8BCF7792C01" box="[305,365,1016,1040]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
_
<collectionCode id="ED1DAE6B2A437952285BD8BCF7A62C01" box="[379,434,1016,1040]" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="28" pageNumber="29">ENT</collectionCode>
_ 0 0 0 0 455 (
<collectionCode id="ED1DAE6B2A4379522B18D8BCF46E2C01" box="[568,634,1016,1040]" country="Panama" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="28" pageNumber="29">STRI</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31665252A43795229E7DF58F25C2D69" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BB336AE2A43795229E7DF58F25C2D69" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="29">
<emphasis id="B978EABC2A43795229E7DF58F72F2C24" bold="true" box="[199,315,1052,1077]" pageId="28" pageNumber="29">Remarks.</emphasis>
This new species is considered to be the undetermined new species of
<taxonomicName id="4C0C4D2D2A4379522DACDF59F2EA2C25" authorityName="Metcalf" authorityYear="1952" box="[1164,1278,1053,1076]" class="Insecta" family="Cicadidae" genus="Dorisiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A4379522DACDF59F2EA2C25" box="[1164,1278,1053,1076]" italics="true" pageId="28" pageNumber="29">Dorisiana</emphasis>
</taxonomicName>
discussed by
<bibRefCitation id="EF9D4B5F2A43795229B7DF7BF7252C49" author="Wolda, H." box="[151,305,1087,1112]" firstAuthor="Wolda" journalOrPublisher="Oecologia, Berlin" pageId="28" pageNumber="29" pagination="437 - 442" part="80" refId="ref65636" refString="Wolda, H. (1989) Seasonal cues in tropical organisms: rainfall? Not necessarily. Oecologia, Berlin, 80, 437 - 442. https: // doi. org / 10.1007 / BF 00380064" title="Seasonal cues in tropical organisms: rainfall? Not necessarily" type="journal article" year="1989">Wolda (1989)</bibRefCitation>
&amp;
<bibRefCitation id="EF9D4B5F2A4379522877DF7BF4732C49" author="Wolda, H. &amp; Ramos, J. A." box="[343,615,1087,1112]" editor="Quintero, D. &amp; Aiello, A." firstAuthor="Wolda" journalOrPublisher="Oxford University Press, New York" pageId="28" pageNumber="29" pagination="271 - 279" refId="ref65723" refString="Wolda, H. &amp; Ramos, J. A. (1992) Cicadas in Panama, their distribution, seasonality and diversity. In: Quintero, D. &amp; Aiello, A. (Eds.), The insects of Panama and Mesoamerica. Selected studies. Oxford University Press, New York, pp. 271 - 279." title="Cicadas in Panama, their distribution, seasonality and diversity" type="book chapter" volumeTitle="The insects of Panama and Mesoamerica. Selected studies" year="1992">Wolda &amp; Ramos (1992)</bibRefCitation>
. The
<typeStatus id="54B7880C2A4379522B86DF04F4C22C49" box="[678,726,1088,1112]" pageId="28" pageNumber="29">type</typeStatus>
series was collected at one of the localities mentioned in these works and is the only unidentifiable species of a
<taxonomicName id="4C0C4D2D2A4379522B9DDF21F53B2C6D" authorityName="Metcalf" authorityYear="1952" box="[701,815,1125,1148]" class="Insecta" family="Cicadidae" genus="Dorisiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A4379522B9DDF21F53B2C6D" box="[701,815,1125,1148]" italics="true" pageId="28" pageNumber="29">Dorisiana</emphasis>
</taxonomicName>
-like species that I found in collections. In addition, the image described in the previous paragraph that matches the females in the
<typeStatus id="54B7880C2A4379522AD3DFCCF2372CB1" box="[1011,1059,1160,1184]" pageId="28" pageNumber="29">type</typeStatus>
series is evidence to support the synonymy of the new species identified in
<bibRefCitation id="EF9D4B5F2A4379522BA8DFE8F5302CD5" author="Wolda, H." box="[648,804,1196,1221]" firstAuthor="Wolda" journalOrPublisher="Oecologia, Berlin" pageId="28" pageNumber="29" pagination="437 - 442" part="80" refId="ref65636" refString="Wolda, H. (1989) Seasonal cues in tropical organisms: rainfall? Not necessarily. Oecologia, Berlin, 80, 437 - 442. https: // doi. org / 10.1007 / BF 00380064" title="Seasonal cues in tropical organisms: rainfall? Not necessarily" type="journal article" year="1989">Wolda (1989)</bibRefCitation>
&amp;
<bibRefCitation id="EF9D4B5F2A4379522A6DDFE8F2742CD5" author="Wolda, H. &amp; Ramos, J. A." box="[845,1120,1196,1221]" editor="Quintero, D. &amp; Aiello, A." firstAuthor="Wolda" journalOrPublisher="Oxford University Press, New York" pageId="28" pageNumber="29" pagination="271 - 279" refId="ref65723" refString="Wolda, H. &amp; Ramos, J. A. (1992) Cicadas in Panama, their distribution, seasonality and diversity. In: Quintero, D. &amp; Aiello, A. (Eds.), The insects of Panama and Mesoamerica. Selected studies. Oxford University Press, New York, pp. 271 - 279." title="Cicadas in Panama, their distribution, seasonality and diversity" type="book chapter" volumeTitle="The insects of Panama and Mesoamerica. Selected studies" year="1992">Wolda &amp; Ramos (1992)</bibRefCitation>
with this new species. The genus
<taxonomicName id="4C0C4D2D2A43795229C5DF8BF7512CF9" baseAuthorityName="Sanborn" baseAuthorityYear="2016" box="[229,325,1231,1256]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A43795229C5DF8BF7512CF9" box="[229,325,1231,1256]" italics="true" pageId="28" pageNumber="29">Guyalna</emphasis>
</taxonomicName>
did not exist when the species was discussed by
<bibRefCitation id="EF9D4B5F2A4379522A86DF8BF2522CF9" author="Wolda, H." box="[934,1094,1231,1256]" firstAuthor="Wolda" journalOrPublisher="Oecologia, Berlin" pageId="28" pageNumber="29" pagination="437 - 442" part="80" refId="ref65636" refString="Wolda, H. (1989) Seasonal cues in tropical organisms: rainfall? Not necessarily. Oecologia, Berlin, 80, 437 - 442. https: // doi. org / 10.1007 / BF 00380064" title="Seasonal cues in tropical organisms: rainfall? Not necessarily" type="journal article" year="1989">Wolda (1989)</bibRefCitation>
or
<bibRefCitation id="EF9D4B5F2A4379522D57DF8BF38C2CF9" author="Wolda, H. &amp; Ramos, J. A." box="[1143,1432,1231,1256]" editor="Quintero, D. &amp; Aiello, A." firstAuthor="Wolda" journalOrPublisher="Oxford University Press, New York" pageId="28" pageNumber="29" pagination="271 - 279" refId="ref65723" refString="Wolda, H. &amp; Ramos, J. A. (1992) Cicadas in Panama, their distribution, seasonality and diversity. In: Quintero, D. &amp; Aiello, A. (Eds.), The insects of Panama and Mesoamerica. Selected studies. Oxford University Press, New York, pp. 271 - 279." title="Cicadas in Panama, their distribution, seasonality and diversity" type="book chapter" volumeTitle="The insects of Panama and Mesoamerica. Selected studies" year="1992">Wolda &amp; Ramos (1992)</bibRefCitation>
.
<taxonomicName id="4C0C4D2D2A43795229B7DFB1F71D2D1D" authorityName="Metcalf" authorityYear="1952" box="[151,265,1269,1292]" class="Insecta" family="Cicadidae" genus="Dorisiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A43795229B7DFB1F71D2D1D" box="[151,265,1269,1292]" italics="true" pageId="28" pageNumber="29">Dorisiana</emphasis>
</taxonomicName>
is structurally similar to
<taxonomicName id="4C0C4D2D2A4379522B05DFB0F4912D1C" baseAuthorityName="Sanborn" baseAuthorityYear="2016" box="[549,645,1268,1293]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A4379522B05DFB0F4912D1C" box="[549,645,1268,1293]" italics="true" pageId="28" pageNumber="29">Guyalna</emphasis>
</taxonomicName>
and many species have been classified in either genera at one time or another (
<bibRefCitation id="EF9D4B5F2A43795229DDDE53F7A52D21" author="Sanborn, A. F." box="[253,433,1303,1328]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="28" pageNumber="29" pagination="430 - 454" part="4105" refId="ref63606" refString="Sanborn, A. F. (2016 a) The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species. Zootaxa, 4105 (5), 430 - 454. https: // doi. org / 10.11646 / zootaxa. 4105.5.2" title="The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species" type="journal article" year="2016" yearSuffix="a">Sanborn 2016a</bibRefCitation>
). The lobate timbal covers classify this new species in the genus
<taxonomicName id="4C0C4D2D2A4379522D94DE53F3002D21" baseAuthorityName="Sanborn" baseAuthorityYear="2016" box="[1204,1300,1303,1328]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A4379522D94DE53F3002D21" box="[1204,1300,1303,1328]" italics="true" pageId="28" pageNumber="29">Guyalna</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C0C4D2D2A43795229B7DE79F7C22D44" authority="(Sanborn 2016 a)" baseAuthorityName="Sanborn" baseAuthorityYear="2016" box="[151,470,1340,1365]" class="Insecta" family="Cicadidae" genus="Dorisiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A43795229B7DE79F71D2D45" box="[151,265,1341,1364]" italics="true" pageId="28" pageNumber="29">Dorisiana</emphasis>
(
<bibRefCitation id="EF9D4B5F2A437952283BDE78F7DA2D44" author="Sanborn, A. F." box="[283,462,1340,1365]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="28" pageNumber="29" pagination="430 - 454" part="4105" refId="ref63606" refString="Sanborn, A. F. (2016 a) The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species. Zootaxa, 4105 (5), 430 - 454. https: // doi. org / 10.11646 / zootaxa. 4105.5.2" title="The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species" type="journal article" year="2016" yearSuffix="a">Sanborn 2016a</bibRefCitation>
)
</taxonomicName>
. The species is a rainy season
<taxonomicName id="4C0C4D2D2A4379522A65DE78F5982D44" authorityName="Linnaeus" authorityYear="1758" box="[837,908,1340,1365]" class="Insecta" family="Cicadidae" genus="Cicada" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="genus">cicada</taxonomicName>
with emergences between late April to early November (
<bibRefCitation id="EF9D4B5F2A437952283BDE1BF7BD2D69" author="Wolda, H." box="[283,425,1375,1400]" firstAuthor="Wolda" journalOrPublisher="Oecologia, Berlin" pageId="28" pageNumber="29" pagination="437 - 442" part="80" refId="ref65636" refString="Wolda, H. (1989) Seasonal cues in tropical organisms: rainfall? Not necessarily. Oecologia, Berlin, 80, 437 - 442. https: // doi. org / 10.1007 / BF 00380064" title="Seasonal cues in tropical organisms: rainfall? Not necessarily" type="journal article" year="1989">Wolda 1989</bibRefCitation>
). The species sang during the day in
<collectingRegion id="49C8F84C2A4379522A66DE24F58B2D66" box="[838,927,1376,1399]" country="Panama" name="Panama" pageId="28" pageNumber="29">
<collectingCountry id="F31B763E2A4379522A66DE24F58B2D66" box="[838,927,1376,1399]" name="Panama" pageId="28" pageNumber="29">Panama</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="EF9D4B5F2A4379522A8DDE1BF22F2D69" author="Wolda, H." box="[941,1083,1375,1400]" firstAuthor="Wolda" journalOrPublisher="Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen. Biological, Chemical, Geological, Physical and Medical Sciences" pageId="28" pageNumber="29" pagination="369 - 381" part="96" refId="ref65675" refString="Wolda, H. (1993) Diel and seasonal patterns of mating calls in some neotropical cicadas. Acoustic interference? Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen. Biological, Chemical, Geological, Physical and Medical Sciences, 96, 369 - 381." title="Diel and seasonal patterns of mating calls in some neotropical cicadas. Acoustic interference?" type="journal article" year="1993">Wolda 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31665252A43795229E7DEC0F49F2DD1" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BB336AE2A43795229E7DEC0F49F2DD1" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="29">
<emphasis id="B978EABC2A43795229E7DEC0F75A2D8C" bold="true" box="[199,334,1412,1437]" pageId="28" pageNumber="29">Etymology.</emphasis>
The name is in honor of Henk Wolda who referenced the new species and provided ecological data on multiple Panamanian
<taxonomicName id="4C0C4D2D2A43795228C4DEE3F4382DD1" authorityName="Linnaeus" authorityYear="1758" box="[484,556,1447,1472]" class="Insecta" family="Cicadidae" genus="Cicada" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="genus">cicada</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C31665252A43795129E7DE88F78F2A2C" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="description">
<paragraph id="8BB336AE2A43795229E7DE88F3102E19" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="29">
<emphasis id="B978EABC2A43795229E7DE88F74D2DF4" bold="true" box="[199,345,1484,1509]" pageId="28" pageNumber="29">Description.</emphasis>
Ground color of head and thorax castaneous marked with piceous, abdomen piceous marked with castaneous. The Cardenas specimen has the piceous very reduced or absent from many body regions.
</paragraph>
<paragraph id="8BB336AE2A43795229E7DD50F4DC2FA8" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="29">
Head. Head wider than mesonotum, castaneous with transverse piceous fascia through ocelli and frons expanding anteriorly and posteriorly along margin of eye continuing to piceous posterior of eye and ventral head, piceous mark on anteromedial supra-antennal plate. Head covered with short silvery pile dorsally, longer and denser posterior to eye. Ocelli rosaceous, ochraceous in some
<typeStatus id="54B7880C2A4379522A52DDC4F5F42E89" box="[882,992,1664,1688]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
, eyes ochraceous, castaneous in some
<typeStatus id="54B7880C2A43795229B7DDE1F7122EAC" box="[151,262,1701,1725]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
. Ventral head piceous except castaneous mark on gena next to anterolateral angle of lorum and anteromedial margin of lorum, mark on gena expanded to transverse fascia in one
<typeStatus id="54B7880C2A4379522D1DDD8CF2B52EF1" box="[1085,1185,1736,1760]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
. Postclypeus centrally sulcate, piceous except castaneous ten transverse ridges, dorsal surface, spot near apex and mark on ventroposterior margin that splits around posterior central sulcus, ochraceous in some
<typeStatus id="54B7880C2A4379522ACDDC54F2482F39" box="[1005,1116,1808,1832]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
. Anteclypeus piceous with castaneous spots on anterior, middle and posterior of carina. Long white pile on lorum, gena, lateral postclypeus and anteclypeus, some golden pile laterally in some
<typeStatus id="54B7880C2A4379522A3EDC1CF5982F61" box="[798,908,1880,1904]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
, white pubescence in central sulcus, lateral postclypeus and medial gena. Mentum tawny, labium castaneous becoming piceous distally. Rostrum reaching to abdominal sternite II. Antennal segments piceous.
</paragraph>
<caption id="DF7366262A42795329B7DC39F58A2FFE" httpUri="https://zenodo.org/record/1445170/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1917,1939]" targetBox="[214,1372,193,1896]" targetPageId="29">
<paragraph id="8BB336AE2A42795329B7DC39F58A2FFE" blockId="29.[151,1436,1917,2031]" pageId="29" pageNumber="30">
<emphasis id="B978EABC2A42795329B7DC39F7062F83" bold="true" box="[151,274,1917,1939]" pageId="29" pageNumber="30">FIGURE 4.</emphasis>
<taxonomicName id="4C0C4D2D2A4279532838DC39F7AE2F82" authority="Sanborn, 2018" authorityName="Sanborn" authorityYear="2018" box="[280,442,1917,1939]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="woldai" status="sp. nov.">
<emphasis id="B978EABC2A4279532838DC39F7AE2F82" box="[280,442,1917,1939]" italics="true" pageId="29" pageNumber="30">Guyalna woldai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C72A42795328E0DC39F7ED2F83" box="[448,505,1917,1938]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B978EABC2A42795328E0DC39F7ED2F83" bold="true" box="[448,505,1917,1938]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
: A, holotype male and paratype female habitus; B, holotype male dorsum; C, holotype male timbal cover; D, paratype male operculum; E, paratype female operculum; F, holotype male lateral view of genitalia; G, holotype male posterior view of genitalia; H, paratype female lateral view of genitalia; I, paratype female ventral view of genitalia. Scale bar: A, 2 cm; B, 5 mm; C–E, 2 mm; F–G, 1 mm; H–I, 2 mm.
</paragraph>
</caption>
<paragraph id="8BB336AE2A41795029E7DBD3F71A2ADC" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
Thorax. Dorsal thorax castaneous marked with piceous. Prothorax with transverse piceous fascia on anterior margin, piceous within paramedfian fissure, and a pair of medially fused triangular piceous marks on either side of midline anterior to ambient fissure, mark in paramedian fissure continuous to triangular marks in some
<typeStatus id="54B7880C2A4179502C0ADBA4F38C28E9" box="[1322,1432,224,248]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
, silvery pile anteriorly between fissures, black pile on disc. Pronotal collar ochraceous with piceous posterior and lateral margins, piceous pile on dorsal pronotal collar, more dense laterally, silvery pile across lateral angle, and dense piceous pile on lateral part of pronotal collar. Mesothorax castaneous with piceous submedian sigillae, anterior lateral sigillae, anterior wing groove, posterior lateral to cruciform elevation, scutal depressions, and shield-shaped piceous mark on dorsal midline between submedian sigillae and scutal depressions, dark castaneous across lateral sigillae posterior ro piceous mark, posterior margin of wing groove ochraceous. Dense silvery pile on anterolateral and lateral mesonotum, in medial wing groove and on posterolateral cruciform elevation, dense piceous pile between lateral silvery pile in anterior and posterior wing groove, between anterior arms of cruciform elevation, anterolateral cruciform elevation and along posterior margin of cruciform elevation, silvery pile anterior to piceous pile between anterior arms of cruciform elevation in some
<typeStatus id="54B7880C2A4179502A9CD90CF23F2A71" box="[956,1067,584,608]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
. Metanotum piceous with dense piceous pile. Ventral thoracic segments piceous except ochraceous middle of anepisternum 2, and castaneous anterolateral anepimeron 2 and anterior katepimeron 2. Thoracic sternites covered with white pubescence and white pile.
</paragraph>
<paragraph id="8BB336AE2A41795029E7D993F4192BFC" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
Wings. Fore wing and wings hyaline, slight milky appearance when viewed at an angle. Venation castaneousochraceous at base becoming piceous distally, costal margin piceous anteriorly, piceous beyond node. Basal cell piceous anteriorly with ochraceous spot at base, piceous reduced or replaced with ochraceous in some
<typeStatus id="54B7880C2A4179502C0AD864F38C2B29" box="[1322,1432,800,824]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
, pterostigma extending to about radial crossvein, proximal third and posterior of clavus infuscated, basal membrane of fore wing grayish piceous. Hind wing infuscated proximally, venation proximally castaneous-opchraceous becoming piceous distally except ochraceous cubitus anterior. Anal cell 3 and anal cell 2 along anal veins 2 and 3, anal cell 1 along anal vein 2, proximal half of cubital cell 2 along cubitus posterior vein, and basal cubital, radial and medial cells grayish-piceous.
</paragraph>
<paragraph id="8BB336AE2A41795029E7D8BCF52D2CB1" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
Legs. Legs castaneous, piceous along medial and lateral coxae, proximal, distal and ventral trochanters, ventral femora, tarsi and pretarsal claws, distal tibiae piceous in some
<typeStatus id="54B7880C2A4179502A6FDF59F5AA2C24" box="[847,958,1053,1077]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
. Fore femora with dark ochraceous stripes, proximal spine small, angled, secondary spine slightly angled and slightly larger than primary spine, tertiary spine very small, upright, all spines piceous. Tibial spurs castaneous with piceous tips, tibial combs piceous. Legs with long piceous pile, coxae and femora with white pubescence.
</paragraph>
<paragraph id="8BB336AE2A41795029E7DFE8F2892D45" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
Operculum. Male operculum piceous with short golden pile covered with white pubescence, slightly expanded at lateral base, lateral margin straight, angled to auditory capsule, rounded posterolateral margin, posterior margin straight to rounded medial margin, not meeting medially, barely covering
<taxonomicName id="4C0C4D2D2A4179502AECDFB1F2502D1C" authorityName="Fabricius" authorityYear="1803" box="[972,1092,1269,1293]" class="Insecta" family="Cicadidae" genus="Tettigonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tympanum">tympanum</taxonomicName>
reaching to anterior of sternite II, domed laterally. Meracathus pointed, piceous with ochraceous lateral margin not reaching posterior opercular margin. Female operculum similarly shaped and colored, reaching to posterior of sternite II.
</paragraph>
<paragraph id="8BB336AE2A41795029E7DE1BF7732F61" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
Abdomen. Abdomen tergites piceous, castaneous posterior margins on tergites 2–8 castaneous expanding anteriorly onto dorsolateral tergite 2, lateral tergite 3, and dorsolateral tergites 5–7, tergites covered with piceous pile, silvery pile on dorsolateral tergite 2, lateral tergites 3 and 4, these regions connected in some female
<typeStatus id="54B7880C2A4179502C11DEECF3882DD1" box="[1329,1436,1448,1472]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
forming large silvery dorsolateral spot, posterior dorsolateral tergite 4, dorsoposterior tergites 5–6, posterolateral tergite 7, and dorsoposterior of tergite 8. Timbal cover piceous, incomplete exposing timbal dorsally, lateroanterior margin rounded, medial margin with a notch forming an approximate right angle. Timbal white with piceous markings visible through opening in timbal cover. Male sternite I, anterior of II, sternite VII and posterior sternite VIII piceous, sternite VIII with V-shaped notch posteriorly, remaining sternites castaneous, sternites III–VI translucent, long black pile radiating from sternites, epipleurites piceous, sternites covered with white pubescence, denser laterally. Female tergites colored similar to male, female sternites piceous except castaneous mediolateral posterior sternite III and posterolateral sternites IV–V, white pubescence. Female sternite VII with arching posterior margin and single medial notch. Female abdominal segment 9 piceous with castaneous or ochraceous anterodorsal and dorsolateral marks and spots on posterior angle and castaneous spot on ventroposterior curve, radiating long piceous pile. Dorsal beak extending beyond piceous anal styles. Posterior margin of abdominal segment 9 sinuate.
</paragraph>
<paragraph id="8BB336AE2A41795129E7DC38F51228E9" blockId="30.[151,1437,151,2013]" lastBlockId="31.[151,1437,151,1725]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">Genitalia. Male pygofer castaneous with piceous dorsal outside, distal basal lobe, uncus, base of anal styles, and tip of dorsal beak. Dorsal beak narrow, extending from deep depression on posterior margin. Pygofer basal lobe extended, medial margin bent inward at approximale right angle with squared apex. Uncal dorsal crest finger- like emerging at approximate right angle with rounded at terminus. Lateral branch of uncus curving ventrally to rounded apex, posteromedial margin straight meeting under the median uncus lobe to surround aedeagus. Aedeagus castaneous with a tawny terminal membrane.</paragraph>
<paragraph id="8BB336AE2A40795129E7DA40F59C2974" blockId="31.[151,1437,151,1725]" pageId="31" pageNumber="32">Female gonocoxite IX piceous or castaneous. Gonapophysis IX and X piceous. Ovipositor sheath extends beyond dorsal beak. Dorsal beak extending just beyond piceous-gray anal styles. Long golden pile radiating from ovipositor sheath, ovipositor sheath covered with short golden pile.</paragraph>
<paragraph id="8BB336AE2A40795129E7DA2BF78F2A2C" blockId="31.[151,1437,151,1725]" pageId="31" pageNumber="32">
<emphasis id="B978EABC2A40795129E7DA2BF7DC2999" bold="true" box="[199,456,367,392]" pageId="31" pageNumber="32">Measurements (mm).</emphasis>
N =
<specimenCount id="9D0AFD272A4079512B2BDA34F4702999" box="[523,612,367,392]" pageId="31" pageNumber="32" type="male">2 males</specimenCount>
or
<specimenCount id="9D0AFD272A4079512BB0DA34F5162999" box="[656,770,367,392]" pageId="31" pageNumber="32" type="female">9 females</specimenCount>
, mean (range). Length of body: males 22.15 (21.5–22.8), females 19.46 (18.1–20.4); length of fore wing: males 32.60 (32.4–32.8), females 29.56 (28.1–31.0); width of fore wing: males 9.85 (9.8–9.9), females 9.39 (8.9–10.0); length of head: males 3.60 (3.5–3.7), females 3.33 (2.8–3.6); width of head including eyes: males 9.45 (9.3–9.6), females 8.72 (8.1–9.2); width of pronotum including suprahumeral plates: males 9.35 (9.3–9.4), females 8.54 (7.8–9.3); width of mesonotum: males 8.00 (7.8–8.2), females 7.39 (6.9–7.8).
</paragraph>
</subSubSection>
<subSubSection id="C31665252A40795129E7D903F5622E19" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BB336AE2A40795129E7D903F4172BB5" blockId="31.[151,1437,151,1725]" pageId="31" pageNumber="32">
<emphasis id="B978EABC2A40795129E7D903F7552A71" bold="true" box="[199,321,583,608]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
The new species can be distinguished easily from the other Panamanian species of
<taxonomicName id="4C0C4D2D2A4079512C14D903F3802A71" baseAuthorityName="Sanborn" baseAuthorityYear="2016" box="[1332,1428,583,608]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A4079512C14D903F3802A71" box="[1332,1428,583,608]" italics="true" pageId="31" pageNumber="32">Guyalna</emphasis>
</taxonomicName>
.
<taxonomicName id="4C0C4D2D2A40795129B7D928F7442A94" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[151,336,620,645]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B978EABC2A40795129B7D928F7442A94" box="[151,336,620,645]" italics="true" pageId="31" pageNumber="32">Guyalna bicolor</emphasis>
</taxonomicName>
is a short, stocky species being about half as wide as long with a green (fading to ochraceous in some specimens) head and thorax and castaneous or testaceous abdomen and
<taxonomicName id="4C0C4D2D2A4079512D27D9D4F2852AB9" authorityName="Distant" authorityYear="1892" box="[1031,1169,655,680]" class="Insecta" family="Cicadidae" genus="Fidicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bogotana">
<emphasis id="B978EABC2A4079512D27D9D4F20C2AB6" box="[1031,1048,656,679]" italics="true" pageId="31" pageNumber="32">G</emphasis>
.
<emphasis id="B978EABC2A4079512D07D9CBF2852AB9" box="[1063,1169,655,680]" italics="true" pageId="31" pageNumber="32">bogotana</emphasis>
</taxonomicName>
is greenish tawny head and thorax with an ochraceous abdomen rather than the dark brown and piceous new species. In addition, the abdominal tergites of male
<taxonomicName id="4C0C4D2D2A40795128FBD99CF47D2AE1" authorityName="Distant" authorityYear="1892" box="[475,617,727,752]" class="Insecta" family="Cicadidae" genus="Fidicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bogotana">
<emphasis id="B978EABC2A40795128FBD99CF7F82AFE" box="[475,492,728,751]" italics="true" pageId="31" pageNumber="32">G</emphasis>
.
<emphasis id="B978EABC2A40795128DFD993F47D2AE1" box="[511,617,727,752]" italics="true" pageId="31" pageNumber="32">bogotana</emphasis>
</taxonomicName>
are translucent. The much larger (body length
<quantity id="4CF49B4B2A4079512DBBD993F2E42AE1" box="[1179,1264,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="31" pageNumber="32" unit="mm" value="35.0">35 mm</quantity>
)
<taxonomicName id="4C0C4D2D2A4079512C23D99CF37F2AE1" authorityName="Distant" authorityYear="1883" box="[1283,1387,728,752]" class="Insecta" family="Cicadidae" genus="Fidicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B978EABC2A4079512C23D99CF37F2AE1" box="[1283,1387,728,752]" italics="true" pageId="31" pageNumber="32">G. fumea</emphasis>
</taxonomicName>
has infuscation on the distal half of the fore wings and distal hind wings easily distinguish this species. The most similar Panamanian species is
<taxonomicName id="4C0C4D2D2A40795128CCD864F49A2B29" authorityName="Davis" authorityYear="1939" box="[492,654,800,824]" class="Insecta" family="Cicadidae" genus="Fidicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B978EABC2A40795128CCD864F49A2B29" box="[492,654,800,824]" italics="true" pageId="31" pageNumber="32">G. panamensis</emphasis>
</taxonomicName>
but it can be distinguished by its tawny body (greenish tawny in fresh specimens), the mesothorax only marked with piceous along the parapsidal suture, the abdomen has silvery and piceous pile that produce a piceous arch across the dorsal tergites from tergites 3–7 and the males and females are of similar size in
<taxonomicName id="4C0C4D2D2A4079512879D8C9F7E82BB5" authorityName="Davis" authorityYear="1939" box="[345,508,909,932]" class="Insecta" family="Cicadidae" genus="Fidicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B978EABC2A4079512879D8C9F7E82BB5" box="[345,508,909,932]" italics="true" pageId="31" pageNumber="32">G. panamensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB336AE2A40795129E7D8EBF5622E19" blockId="31.[151,1437,151,1725]" pageId="31" pageNumber="32">
The new species can be distinguished from the remaining species of
<taxonomicName id="4C0C4D2D2A4079512AE1D8EBF2352BD9" baseAuthorityName="Sanborn" baseAuthorityYear="2016" box="[961,1057,943,968]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B978EABC2A4079512AE1D8EBF2352BD9" box="[961,1057,943,968]" italics="true" pageId="31" pageNumber="32">Guyalna</emphasis>
</taxonomicName>
relatively easily as well. The new species has a body length (
<quantity id="4CF49B4B2A40795128EFD890F4532BFD" box="[463,583,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="31" pageNumber="32" unit="mm" value="20.5" valueMax="23.0" valueMin="18.0">18-23 mm</quantity>
) less than
<taxonomicName id="4C0C4D2D2A4079512BE5D891F5F62BFD" authority="Ruschel, 2017" authorityName="Ruschel" authorityYear="2017" box="[709,994,980,1005]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="aurora">
<emphasis id="B978EABC2A4079512BE5D891F5272BFD" box="[709,819,981,1004]" italics="true" pageId="31" pageNumber="32">G. aurora</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512A1CD890F5F62BFD" author="Ruschel, T. P." box="[828,994,980,1005]" firstAuthor="Ruschel" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="246 - 279" part="4281" refId="ref62806" refString="Ruschel, T. P. (2017) Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246 - 279. https: // doi. org / 10.11646 / zootaxa. 4281.1.25" title="Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species" type="journal article" year="2017">Ruschel, 2017</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512ACFD891F3272BFD" authority="(Berg, 1879)" baseAuthorityName="Berg" baseAuthorityYear="1879" box="[1007,1331,980,1005]" class="Insecta" family="Cicadidae" genus="Fidicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bonaerensis">
<emphasis id="B978EABC2A4079512ACFD891F2822BFC" box="[1007,1174,980,1005]" italics="true" pageId="31" pageNumber="32">G. bonaerensis</emphasis>
(
<bibRefCitation id="EF9D4B5F2A4079512D88D890F3382BFD" author="Berg, C." box="[1192,1324,980,1004]" firstAuthor="Berg" journalOrPublisher="Anales de la Sociedad Cientifica Argentina" pageId="31" pageNumber="32" pagination="135 - 144" part="8" refId="ref59377" refString="Berg, C. (1879) Hemiptera Argentina (Continuacion.) Hemiptera Homoptera Latr. Anales de la Sociedad Cientifica Argentina, 8, 135 - 144." title="Hemiptera Argentina (Continuacion.) Hemiptera Homoptera Latr" type="journal article" year="1879">Berg, 1879</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512C61D891F7502C01" authority="(Walker, 1850)" baseAuthorityName="Walker" baseAuthorityYear="1850" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="brisa">
<emphasis id="B978EABC2A4079512C61D891F3882BFC" box="[1345,1436,980,1005]" italics="true" pageId="31" pageNumber="32">G. brisa</emphasis>
(
<bibRefCitation id="EF9D4B5F2A40795129BED8B3F7282C01" author="Walker, F." bookContentInfo="260 pp." box="[158,316,1015,1040]" firstAuthor="Walker" journalOrPublisher="British Museum Trustees, London" pageId="31" pageNumber="32" refId="ref65130" refString="Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I. British Museum Trustees, London, 260 pp." title="List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I" type="book" year="1850">Walker, 1850</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512872D8BCF4BC2C01" authority="(Walker, 1850)" baseAuthorityName="Walker" baseAuthorityYear="1850" box="[338,680,1015,1040]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="chlorogena">
<emphasis id="B978EABC2A4079512872D8BCF7E62C01" box="[338,498,1015,1040]" italics="true" pageId="31" pageNumber="32">G. chlorogena</emphasis>
(
<bibRefCitation id="EF9D4B5F2A4079512B23D8B3F4B52C01" author="Walker, F." bookContentInfo="260 pp." box="[515,673,1015,1040]" firstAuthor="Walker" journalOrPublisher="British Museum Trustees, London" pageId="31" pageNumber="32" refId="ref65130" refString="Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I. British Museum Trustees, London, 260 pp." title="List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I" type="book" year="1850">Walker, 1850</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512B97D8BCF25D2C01" authority="(Sanborn, 2007 b)" baseAuthorityName="Sanborn" baseAuthorityYear="2007" box="[695,1097,1015,1040]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="flavipronotum">
<emphasis id="B978EABC2A4079512B97D8BCF5622C01" box="[695,886,1015,1040]" italics="true" pageId="31" pageNumber="32">G. flavipronotum</emphasis>
(
<bibRefCitation id="EF9D4B5F2A4079512AA8D8B3F2552C01" author="Sanborn, A. F." box="[904,1089,1015,1040]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="21 - 32" part="1503" refId="ref63200" refString="Sanborn, A. F. (2007 b) Additions to the cicada fauna of Venezuela with the description of a new species and checklist of the Venezuelan cicada fauna (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 1503, 21 - 32." title="Additions to the cicada fauna of Venezuela with the description of a new species and checklist of the Venezuelan cicada fauna (Hemiptera: Cicadoidea: Cicadidae)" type="journal article" year="2007" yearSuffix="b">Sanborn, 2007b</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512D77D8BCF3622C01" authority="(Distant, 1883)" baseAuthorityName="Distant" baseAuthorityYear="1883" box="[1111,1398,1015,1040]" class="Insecta" family="Cicadidae" genus="Fidicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="B978EABC2A4079512D77D8BCF2AA2C01" box="[1111,1214,1016,1040]" italics="true" pageId="31" pageNumber="32">G. fumea</emphasis>
(
<bibRefCitation id="EF9D4B5F2A4079512DEFD8B3F3792C01" author="Distant, W. L." box="[1231,1389,1015,1040]" firstAuthor="Distant" pageId="31" pageNumber="32" pagination="17 - 24" part="15" refId="ref59993" refString="Distant, W. L. (1883) Rhynchota: Homoptera. Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America, Part 15 (1), 17 - 24." title="Rhynchota: Homoptera. Biologia Centrali-Americana; contributions to the knowledge of the fauna and flora of Mexico and Central America" type="journal article" year="1883">Distant, 1883</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512CA4D8BCF78F2C24" authority="Sanborn, 2016 b" authorityName="Sanborn" authorityYear="2016" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="jamesi">
<emphasis id="B978EABC2A4079512CA4D8BCF6F42C25" italics="true" pageId="31" pageNumber="32">G. jamesi</emphasis>
<bibRefCitation id="EF9D4B5F2A40795129C7DF58F78F2C24" author="Sanborn, A. F." box="[231,411,1052,1077]" firstAuthor="Sanborn" journalOrPublisher="Zoosystema" pageId="31" pageNumber="32" pagination="177 - 199" part="38" refId="ref63671" refString="Sanborn, A. F. (2016 b) Five new species, a new genus and a new record of cicadas from French Guiana with four new combinations and three new synonymies (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 38, 177 - 199. https: // doi. org / 10.5252 / z 2016 n 2 a 2" title="Five new species, a new genus and a new record of cicadas from French Guiana with four new combinations and three new synonymies (Insecta, Hemiptera, Cicadoidea, Cicadidae)" type="journal article" year="2016" yearSuffix="b">Sanborn, 2016b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512886DF59F4C32C25" authority="Ruschel, 2017" authorityName="Ruschel" authorityYear="2017" box="[422,727,1052,1077]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="principes">
<emphasis id="B978EABC2A4079512886DF59F4392C25" box="[422,557,1053,1076]" italics="true" pageId="31" pageNumber="32">G. principes</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512B14DF58F4C32C25" author="Ruschel, T. P." box="[564,727,1052,1077]" firstAuthor="Ruschel" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="246 - 279" part="4281" refId="ref62806" refString="Ruschel, T. P. (2017) Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246 - 279. https: // doi. org / 10.11646 / zootaxa. 4281.1.25" title="Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species" type="journal article" year="2017">Ruschel, 2017</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512BC3DF59F2302C25" authority="Boulard, 1998" authorityName="Boulard" authorityYear="1998" box="[739,1060,1052,1077]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="rufapicalis">
<emphasis id="B978EABC2A4079512BC3DF59F56F2C24" box="[739,891,1052,1077]" italics="true" pageId="31" pageNumber="32">G. rufapicalis</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512AA2DF58F2302C25" author="Boulard, M." box="[898,1060,1052,1077]" firstAuthor="Boulard" journalOrPublisher="123 e Congres national des societes historiques et scientifiques, Antilles-Guyane, 1998, Histoire naturelle" pageId="31" pageNumber="32" pagination="75 - 94" part="1998" refId="ref59543" refString="Boulard, M. (1998) Apercu de cicadologie antillo-guyanaise. 123 e Congres national des societes historiques et scientifiques, Antilles-Guyane, 1998, Histoire naturelle, 1998, 75 - 94." title="Apercu de cicadologie antillo-guyanaise." type="proceedings" year="1998">Boulard, 1998</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512D10DF59F3792C25" authority="Ruschel, 2017" authorityName="Ruschel" authorityYear="2017" box="[1072,1389,1052,1077]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sakakibari">
<emphasis id="B978EABC2A4079512D10DF59F2D12C24" box="[1072,1221,1052,1077]" italics="true" pageId="31" pageNumber="32">G. sakakibari</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512DECDF58F3792C25" author="Ruschel, T. P." box="[1228,1389,1052,1077]" firstAuthor="Ruschel" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="246 - 279" part="4281" refId="ref62806" refString="Ruschel, T. P. (2017) Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246 - 279. https: // doi. org / 10.11646 / zootaxa. 4281.1.25" title="Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species" type="journal article" year="2017">Ruschel, 2017</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0C4D2D2A40795129B7DF04F7A12C49" authority="Ruschel, 2017" authorityName="Ruschel" authorityYear="2017" box="[151,437,1087,1112]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="spinula">
<emphasis id="B978EABC2A40795129B7DF04F71E2C49" box="[151,266,1087,1112]" italics="true" pageId="31" pageNumber="32">G. spinula</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512833DF7BF7A12C49" author="Ruschel, T. P." box="[275,437,1087,1112]" firstAuthor="Ruschel" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="246 - 279" part="4281" refId="ref62806" refString="Ruschel, T. P. (2017) Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246 - 279. https: // doi. org / 10.11646 / zootaxa. 4281.1.25" title="Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species" type="journal article" year="2017">Ruschel, 2017</bibRefCitation>
</taxonomicName>
all of which have body lengths greater than
<quantity id="4CF49B4B2A4079512A9BDF7BF21B2C46" box="[955,1039,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="31" pageNumber="32" unit="mm" value="25.0">25 mm</quantity>
. The smaller body size (
<quantity id="4CF49B4B2A4079512C0ADF04F3682C46" box="[1322,1404,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="31" pageNumber="32" unit="mm" value="19.0">19 mm</quantity>
or less) of
<taxonomicName id="4C0C4D2D2A40795129D0DF21F7E42C6C" authority="(Walker, 1850)" baseAuthorityName="Walker" baseAuthorityYear="1850" box="[240,496,1124,1149]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="cuta">
<emphasis id="B978EABC2A40795129D0DF21F72A2C6D" box="[240,318,1125,1148]" italics="true" pageId="31" pageNumber="32">G. cuta</emphasis>
(
<bibRefCitation id="EF9D4B5F2A407951286DDF20F7FC2C6C" author="Walker, F." bookContentInfo="260 pp." box="[333,488,1124,1149]" firstAuthor="Walker" journalOrPublisher="British Museum Trustees, London" pageId="31" pageNumber="32" refId="ref65130" refString="Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I. British Museum Trustees, London, 260 pp." title="List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I" type="book" year="1850">Walker, 1850</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A40795128DCDF21F2732C6C" authority="Gogala, Sporar, Sanborn &amp; Maccagnan, 2015" authorityName="Gogala, Sporar, Sanborn &amp; Maccagnan" authorityYear="2015" box="[508,1127,1124,1149]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nadae">
<emphasis id="B978EABC2A40795128DCDF21F44B2C6C" box="[508,607,1124,1149]" italics="true" pageId="31" pageNumber="32">G. nadae</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512B46DF20F2732C6C" author="Gogala, M. &amp; Sporar, K. &amp; Sanborn, A. F. &amp; Maccagnan, D. H. B." box="[614,1127,1124,1149]" firstAuthor="Gogala" journalOrPublisher="Acta Entomologica Slovenica" pageId="31" pageNumber="32" pagination="105 - 116" part="23" refId="ref60942" refString="Gogala, M., Sporar, K., Sanborn, A. F. &amp; Maccagnan, D. H. B. (2015) New cicada species of the genus Guyalna (Hemiptera: Cicadidae) from Brazil. Acta Entomologica Slovenica, 23, 105 - 116." title="New cicada species of the genus Guyalna (Hemiptera: Cicadidae) from Brazil" type="journal article" year="2015">Gogala, Šporar, Sanborn &amp; Maccagnan, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512D53DF21F36C2C6D" authority="Boulard, 1999" authorityName="Boulard" authorityYear="1999" box="[1139,1400,1124,1149]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B978EABC2A4079512D53DF21F2DA2C6D" box="[1139,1230,1125,1148]" italics="true" pageId="31" pageNumber="32">G. nigra</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512DF5DF20F36C2C6D" author="Boulard, M." box="[1237,1400,1124,1149]" firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="31" pageNumber="32" pagination="77 - 116" part="11 / 12" refId="ref59578" refString="Boulard, M. (1999) Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae). Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 11 / 12, 77 - 116." title="Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae)" type="journal article" year="1999">Boulard, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512CA4DF21F78F2CB1" authority="(Jacobi, 1904)" baseAuthorityName="Jacobi" baseAuthorityYear="1904" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B978EABC2A4079512CA4DF21F3812C6D" box="[1412,1429,1125,1148]" italics="true" pageId="31" pageNumber="32">G</emphasis>
.
<emphasis id="B978EABC2A40795129B7DFC3F6FA2CB1" box="[151,238,1159,1184]" italics="true" pageId="31" pageNumber="32">parvula</emphasis>
(
<bibRefCitation id="EF9D4B5F2A40795129DEDFC3F7872CB1" author="Jacobi, A." box="[254,403,1159,1184]" firstAuthor="Jacobi" journalOrPublisher="Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin" pageId="31" pageNumber="32" pagination="155 - 164" part="1904" refId="ref61160" refString="Jacobi, A. (1904) Neue Cicadiden und Fulgoriden Brasiliens. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1904, 155 - 164. https: // doi. org / 10.5962 / bhl. part. 8959" title="Neue Cicadiden und Fulgoriden Brasiliens" type="journal article" year="1904">Jacobi, 1904</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512888DFCCF5402CB1" authority="Sanborn &amp; Heath, 2014" authorityName="Sanborn &amp; Heath" authorityYear="2014" box="[424,852,1159,1184]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="platyrhina">
<emphasis id="B978EABC2A4079512888DFCCF4282CB1" box="[424,572,1159,1184]" italics="true" pageId="31" pageNumber="32">G. platyrhina</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512B65DFC3F5402CB1" author="Sanborn, A. F. &amp; Heath, M. S." box="[581,852,1159,1184]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="1 - 94" part="3883" refId="ref64044" refString="Sanborn, A. F. &amp; Heath, M. S. (2014) The cicadas of Argentina with new records, a new genus and fifteen new species (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 3883 (1), 1 - 94. https: // doi. org / 10.11646 / zootaxa. 3883.1.1" title="The cicadas of Argentina with new records, a new genus and fifteen new species (Hemiptera: Cicadoidea: Cicadidae)" type="journal article" year="2014">Sanborn &amp; Heath, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0C4D2D2A4079512AADDFCCF2CC2CB1" authority="(Uhler, 1903)" baseAuthorityName="Uhler" baseAuthorityYear="1903" box="[909,1240,1159,1184]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sublaqueata">
<emphasis id="B978EABC2A4079512AADDFCCF58A2C8E" box="[909,926,1160,1183]" italics="true" pageId="31" pageNumber="32">G</emphasis>
.
<emphasis id="B978EABC2A4079512A8DDFC3F2222CB1" box="[941,1078,1159,1184]" italics="true" pageId="31" pageNumber="32">sublaqueata</emphasis>
(
<bibRefCitation id="EF9D4B5F2A4079512D67DFC3F2DB2CB1" author="Uhler, P. R." box="[1095,1231,1159,1184]" firstAuthor="Uhler" journalOrPublisher="Transactions of the Maryland Academy of Sciences" pageId="31" pageNumber="32" pagination="1 - 17" part="2" refId="ref65090" refString="Uhler, P. R. (1903) Enumeration of the Cicadidae of Brazil in the collection of Mr. Herbert H. Smith. Transactions of the Maryland Academy of Sciences, 2, 1 - 17." title="Enumeration of the Cicadidae of Brazil in the collection of Mr. Herbert H. Smith" type="journal article" year="1903">Uhler, 1903</bibRefCitation>
)
</taxonomicName>
distinguish them from the new species. The new species can be distinguished from
<taxonomicName id="4C0C4D2D2A4079512AABDFE9F3592CD5" authority="Boulard &amp; Martinelli, 2011" authorityName="Boulard &amp; Martinelli" authorityYear="2011" box="[907,1357,1196,1221]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="atalapae">
<emphasis id="B978EABC2A4079512AABDFE9F2192CD4" box="[907,1037,1196,1221]" italics="true" pageId="31" pageNumber="32">G. atalapae</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512D35DFE8F3592CD5" author="Boulard, M. &amp; Martinelli, N. M." box="[1045,1357,1196,1221]" firstAuthor="Boulard" journalOrPublisher="Lambillionea" pageId="31" pageNumber="32" pagination="219 - 232" part="61" refId="ref59749" refString="Boulard, M. &amp; Martinelli, N. M. (2011) Nouvelles cigales neotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini). Lambillionea, 61, 219 - 232." title="Nouvelles cigales neotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini)" type="journal article" year="2011">Boulard &amp; Martinelli, 2011</bibRefCitation>
</taxonomicName>
by the orange basal area and smoky distal edge of its fore wing, from
<taxonomicName id="4C0C4D2D2A4079512A4ADF94F33E2CF9" authority="Boulard &amp; Martinelli, 2011" authorityName="Boulard &amp; Martinelli" authorityYear="2011" box="[874,1322,1231,1256]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bleuzeni">
<emphasis id="B978EABC2A4079512A4ADF94F5FC2CF9" box="[874,1000,1231,1256]" italics="true" pageId="31" pageNumber="32">G. bleuzeni</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512AD0DF8BF33E2CF9" author="Boulard, M. &amp; Martinelli, N. M." box="[1008,1322,1231,1256]" firstAuthor="Boulard" journalOrPublisher="Lambillionea" pageId="31" pageNumber="32" pagination="219 - 232" part="61" refId="ref59749" refString="Boulard, M. &amp; Martinelli, N. M. (2011) Nouvelles cigales neotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini). Lambillionea, 61, 219 - 232." title="Nouvelles cigales neotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini)" type="journal article" year="2011">Boulard &amp; Martinelli, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512C17DF94F41D2D1D" authority="Sanborn, Moore &amp; Young, 2008" authorityName="Sanborn, Moore &amp; Young" authorityYear="2008" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="coffea">
<emphasis id="B978EABC2A4079512C17DF94F3882CF9" box="[1335,1436,1232,1256]" italics="true" pageId="31" pageNumber="32">G. coffea</emphasis>
<bibRefCitation id="EF9D4B5F2A40795129B7DFB0F41D2D1D" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." box="[151,521,1268,1293]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="1 - 20" part="1846" refId="ref64508" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">Sanborn, Moore &amp; Young, 2008</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0C4D2D2A4079512B64DFB1F5642D1C" authority="(Goding, 1925)" baseAuthorityName="Goding" baseAuthorityYear="1925" box="[580,880,1268,1293]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="distanti">
<emphasis id="B978EABC2A4079512B64DFB1F4AC2D1C" box="[580,696,1268,1293]" italics="true" pageId="31" pageNumber="32">G. distanti</emphasis>
(
<bibRefCitation id="EF9D4B5F2A4079512BE9DFB0F57C2D1C" author="Goding, F. W." box="[713,872,1268,1293]" firstAuthor="Goding" journalOrPublisher="Revista del Colegio Nacional Rocafuerte" pageId="31" pageNumber="32" pagination="1 - 34" part="7" refId="ref60803" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." title="Synopsis of the Cicadidae of Ecuador" type="journal article" year="1925">Goding, 1925</bibRefCitation>
)
</taxonomicName>
where the mesothorax is marked only along the parapsidal suture with piceous, from
<taxonomicName id="4C0C4D2D2A4079512B19DE5CF5FA2D21" authority="Boulard &amp; Martinelli, 2011" authorityName="Boulard &amp; Martinelli" authorityYear="2011" box="[569,1006,1303,1328]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="densusa">
<emphasis id="B978EABC2A4079512B19DE5CF4A62D21" box="[569,690,1303,1328]" italics="true" pageId="31" pageNumber="32">G. densusa</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512B9ADE53F5FA2D21" author="Boulard, M. &amp; Martinelli, N. M." box="[698,1006,1303,1328]" firstAuthor="Boulard" journalOrPublisher="Lambillionea" pageId="31" pageNumber="32" pagination="219 - 232" part="61" refId="ref59749" refString="Boulard, M. &amp; Martinelli, N. M. (2011) Nouvelles cigales neotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini). Lambillionea, 61, 219 - 232." title="Nouvelles cigales neotropicales de la sous-tribu des Guyalinina (Rhynchota, Cicadidae, Cicadinae, Fidicinini)" type="journal article" year="2011">Boulard &amp; Martinelli, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0C4D2D2A4079512D06DE5CF6DB2D44" authority="Boulard &amp; Martinelli, 1996" authorityName="Boulard &amp; Martinelli" authorityYear="1996" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="jauffreti">
<emphasis id="B978EABC2A4079512D06DE5CF28A2D21" box="[1062,1182,1304,1328]" italics="true" pageId="31" pageNumber="32">G. jauffreti</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512D86DE53F6DB2D44" author="Boulard, M. &amp; Martinelli, N. M." firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="31" pageNumber="32" pagination="11 - 81" part="9" refId="ref59682" refString="Boulard, M. &amp; Martinelli, N. M. (1996) Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 9, 11 - 81." title="Revision des Fidicini, nouveau statut de la tribu, especes connues et nouvelles especes (Cicadomorpha, Cicadidae, Cicadinae). Premiere partie: Sous-tribu nouvelle des Fidicinina" type="journal article" year="1996">Boulard &amp; Martinelli, 1996</bibRefCitation>
</taxonomicName>
by the ochraceous ventral surfaces of these species, from
<taxonomicName id="4C0C4D2D2A4079512A7FDE79F28D2D45" authority="Ruschel, 2017" authorityName="Ruschel" authorityYear="2017" box="[863,1177,1340,1365]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="flavantica">
<emphasis id="B978EABC2A4079512A7FDE79F5FA2D44" box="[863,1006,1340,1365]" italics="true" pageId="31" pageNumber="32">G. flavantica</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512AD6DE78F28D2D45" author="Ruschel, T. P." box="[1014,1177,1340,1365]" firstAuthor="Ruschel" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="246 - 279" part="4281" refId="ref62806" refString="Ruschel, T. P. (2017) Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246 - 279. https: // doi. org / 10.11646 / zootaxa. 4281.1.25" title="Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species" type="journal article" year="2017">Ruschel, 2017</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0C4D2D2A4079512D85DE79F6C92D69" authority="Sanborn, 2016 b" authorityName="Sanborn" authorityYear="2016" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="maxineae">
<emphasis id="B978EABC2A4079512D85DE79F3242D45" box="[1189,1328,1341,1364]" italics="true" pageId="31" pageNumber="32">G. maxineae</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512C18DE78F6C92D69" author="Sanborn, A. F." firstAuthor="Sanborn" journalOrPublisher="Zoosystema" pageId="31" pageNumber="32" pagination="177 - 199" part="38" refId="ref63671" refString="Sanborn, A. F. (2016 b) Five new species, a new genus and a new record of cicadas from French Guiana with four new combinations and three new synonymies (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 38, 177 - 199. https: // doi. org / 10.5252 / z 2016 n 2 a 2" title="Five new species, a new genus and a new record of cicadas from French Guiana with four new combinations and three new synonymies (Insecta, Hemiptera, Cicadoidea, Cicadidae)" type="journal article" year="2016" yearSuffix="b">Sanborn, 2016b</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0C4D2D2A4079512832DE24F42C2D69" authority="Ruschel, 2017" authorityName="Ruschel" authorityYear="2017" box="[274,568,1375,1400]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tenebrae">
<emphasis id="B978EABC2A4079512832DE24F7862D69" box="[274,402,1375,1400]" italics="true" pageId="31" pageNumber="32">G. tenebrae</emphasis>
<bibRefCitation id="EF9D4B5F2A40795128B8DE1BF42C2D69" author="Ruschel, T. P." box="[408,568,1375,1400]" firstAuthor="Ruschel" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="246 - 279" part="4281" refId="ref62806" refString="Ruschel, T. P. (2017) Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246 - 279. https: // doi. org / 10.11646 / zootaxa. 4281.1.25" title="Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species" type="journal article" year="2017">Ruschel, 2017</bibRefCitation>
</taxonomicName>
by the infuscation at the base of the hind wing extending about a third of hind wing length and the basal cell of the fore wing is colored completely, from
<taxonomicName id="4C0C4D2D2A4079512D0BDEC1F3512D8D" authority="Ruschel, 2017" authorityName="Ruschel" authorityYear="2017" box="[1067,1349,1412,1437]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B978EABC2A4079512D0BDEC1F2872D8C" box="[1067,1171,1412,1437]" italics="true" pageId="31" pageNumber="32">G. pilosa</emphasis>
<bibRefCitation id="EF9D4B5F2A4079512DBFDEC0F3512D8D" author="Ruschel, T. P." box="[1183,1349,1412,1437]" firstAuthor="Ruschel" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="246 - 279" part="4281" refId="ref62806" refString="Ruschel, T. P. (2017) Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species. Zootaxa, 4281 (1), 246 - 279. https: // doi. org / 10.11646 / zootaxa. 4281.1.25" title="Seven new species of the cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Fidicinini) with a re-description of the type species" type="journal article" year="2017">Ruschel, 2017</bibRefCitation>
</taxonomicName>
by the infuscation at the base of the hind wing extending about a third of hind wing length, the basal cell of the fore wing is colored completely and the dense pile possessed by this species. Finally,
<taxonomicName id="4C0C4D2D2A4079512D03DE89F3452DF4" authority="(Goding, 1925)" baseAuthorityName="Goding" baseAuthorityYear="1925" box="[1059,1361,1484,1509]" class="Insecta" family="Cicadidae" genus="Guyalna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="glauca">
<emphasis id="B978EABC2A4079512D03DE89F2872DF4" box="[1059,1171,1484,1509]" italics="true" pageId="31" pageNumber="32">G. glauca</emphasis>
(
<bibRefCitation id="EF9D4B5F2A4079512D87DE88F35D2DF4" author="Goding, F. W." box="[1191,1353,1484,1509]" firstAuthor="Goding" journalOrPublisher="Revista del Colegio Nacional Rocafuerte" pageId="31" pageNumber="32" pagination="1 - 34" part="7" refId="ref60803" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." title="Synopsis of the Cicadidae of Ecuador" type="journal article" year="1925">Goding, 1925</bibRefCitation>
)
</taxonomicName>
has a monochromatic green head and thorax without piceous markings.
</paragraph>
</subSubSection>
<subSubSection id="C31665252A40795129E7DD50F7842EAC" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BB336AE2A40795129E7DD50F7842EAC" blockId="31.[151,1437,151,1725]" pageId="31" pageNumber="32">
<emphasis id="B978EABC2A40795129E7DD50F7742E3C" bold="true" box="[199,352,1556,1581]" pageId="31" pageNumber="32">Distribution.</emphasis>
The
<typeStatus id="54B7880C2A4079512881DD51F7C52E3C" box="[417,465,1557,1581]" pageId="31" pageNumber="32">type</typeStatus>
series was collected in
<collectingRegion id="49C8F84C2A4079512BCCDD50F5BD2E3C" box="[748,937,1556,1581]" country="Panama" name="Darien" pageId="31" pageNumber="32">Darién Province</collectingRegion>
, on Barro Colorado Island and at Curiche,
<collectingRegion id="49C8F84C2A40795129B7DD73F6F72E41" box="[151,227,1591,1616]" country="Panama" name="Embera-Wounaan" pageId="31" pageNumber="32">Choco</collectingRegion>
,
<collectingCountry id="F31B763E2A40795129CEDD73F7742E41" box="[238,352,1591,1616]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
. Curiche is less than
<quantity id="4CF49B4B2A4079512B6FDD7CF4832E41" box="[591,663,1591,1616]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="km" value="30.0">30 km</quantity>
south of the Panamanian border on the west coast of
<collectingCountry id="F31B763E2A4079512DD4DD73F3722E41" box="[1268,1382,1591,1616]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
. The species was reported from Barro Colorado Island,
<collectingRegion id="49C8F84C2A4079512BFADD19F5272E65" box="[730,819,1629,1652]" country="Panama" name="Panama" pageId="31" pageNumber="32">
<collectingCountry id="F31B763E2A4079512BFADD19F5272E65" box="[730,819,1629,1652]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
</collectingRegion>
Canal Zone,
<quantity id="4CF49B4B2A4079512AF0DD18F20D2E65" box="[976,1049,1628,1652]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="32" unit="m" value="120.0">120 m</quantity>
altitude in lowland forest (
<bibRefCitation id="EF9D4B5F2A4079512C72DD18F6C72E89" author="Wolda, H." firstAuthor="Wolda" journalOrPublisher="Oecologia, Berlin" pageId="31" pageNumber="32" pagination="437 - 442" part="80" refId="ref65636" refString="Wolda, H. (1989) Seasonal cues in tropical organisms: rainfall? Not necessarily. Oecologia, Berlin, 80, 437 - 442. https: // doi. org / 10.1007 / BF 00380064" title="Seasonal cues in tropical organisms: rainfall? Not necessarily" type="journal article" year="1989">Wolda 1989</bibRefCitation>
;
<bibRefCitation id="EF9D4B5F2A40795129FEDD3BF7CE2E89" author="Wolda, H. &amp; Ramos, J. A." box="[222,474,1663,1688]" editor="Quintero, D. &amp; Aiello, A." firstAuthor="Wolda" journalOrPublisher="Oxford University Press, New York" pageId="31" pageNumber="32" pagination="271 - 279" refId="ref65723" refString="Wolda, H. &amp; Ramos, J. A. (1992) Cicadas in Panama, their distribution, seasonality and diversity. In: Quintero, D. &amp; Aiello, A. (Eds.), The insects of Panama and Mesoamerica. Selected studies. Oxford University Press, New York, pp. 271 - 279." title="Cicadas in Panama, their distribution, seasonality and diversity" type="book chapter" volumeTitle="The insects of Panama and Mesoamerica. Selected studies" year="1992">Wolda &amp; Ramos 1992</bibRefCitation>
) and from Las Cumbres,
<quantity id="4CF49B4B2A4079512BDBDD3BF5572E89" box="[763,835,1663,1688]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="km" value="15.0">15 km</quantity>
north of
<collectingRegion id="49C8F84C2A4079512A8BDDC4F22F2E89" box="[939,1083,1663,1688]" country="Panama" name="Panama" pageId="31" pageNumber="32">
<collectingCountry id="F31B763E2A4079512A8BDDC4F2102E86" box="[939,1028,1664,1687]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
City
</collectingRegion>
in
<collectingRegion id="49C8F84C2A4079512D41DDC4F2AE2E86" box="[1121,1210,1664,1687]" country="Panama" name="Panama" pageId="31" pageNumber="32">
<collectingCountry id="F31B763E2A4079512D41DDC4F2AE2E86" box="[1121,1210,1664,1687]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="EF9D4B5F2A4079512DE8DD3BF3472E89" author="Wolda, H." box="[1224,1363,1663,1688]" firstAuthor="Wolda" journalOrPublisher="Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen. Biological, Chemical, Geological, Physical and Medical Sciences" pageId="31" pageNumber="32" pagination="369 - 381" part="96" refId="ref65675" refString="Wolda, H. (1993) Diel and seasonal patterns of mating calls in some neotropical cicadas. Acoustic interference? Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen. Biological, Chemical, Geological, Physical and Medical Sciences, 96, 369 - 381." title="Diel and seasonal patterns of mating calls in some neotropical cicadas. Acoustic interference?" type="journal article" year="1993">Wolda 1993</bibRefCitation>
) as an undetermined species.
</paragraph>
</subSubSection>
</treatment>
</document>
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