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<document id="1F7CA6EE4503DCDDA9030DB206965274" ID-CLB-Dataset="46730" ID-DOI="10.5281/zenodo.201491" ID-GBIF-Dataset="30aff672-89b2-441c-af59-1a1b9eb6a7c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="201491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365005065" checkinUser="plazi" docAuthor="Leschen, Richard A. B." docDate="2011" docId="C563080BFFC93A65FF216B76F17BF840" docLanguage="en" docName="zt02908p044.pdf" docOrigin="Zootaxa 2908" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laricobius taiwanensis Yu &amp; Montgomery 2007" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="395A7073FFD23A7EFFB66A19F02FFFF6" masterDocTitle="World review of Laricobius (Coleoptera: Derodontidae)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="28" updateTime="1698295152186" updateUser="ExternalLinkService">
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<mods:title id="A77EF2B22499B402EEDB8A21A4D29350">World review of Laricobius (Coleoptera: Derodontidae)</mods:title>
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<mods:namePart id="F9B2FFF63F96FB41E8D8A151CA7D02CD">Leschen, Richard A. B.</mods:namePart>
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<mods:date id="D255EB085AB8CA5BE8765C4D990C9303">2011</mods:date>
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<treatment id="C563080BFFC93A65FF216B76F17BF840" ID-DOI="http://doi.org/10.5281/zenodo.5661250" ID-GBIF-Taxon="119410649" ID-Zenodo-Dep="5661250" LSID="urn:lsid:plazi:treatment:C563080BFFC93A65FF216B76F17BF840" httpUri="http://treatment.plazi.org/id/C563080BFFC93A65FF216B76F17BF840" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="05D0EA96FFC93A65FF216B76F33AFE7F" box="[151,789,367,393]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="4D75B91DFFC93A65FF216B76F33AFE7F" blockId="27.[151,789,367,426]" box="[151,789,367,393]" pageId="27" pageNumber="28">
<heading id="163D0E71FFC93A65FF216B76F33AFE7F" bold="true" box="[151,789,367,393]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="7FBE650FFFC93A65FF216B76F33AFE7F" bold="true" box="[151,789,367,393]" pageId="27" pageNumber="28">
20.
<taxonomicName id="8ACAC29EFFC93A65FF756B76F33AFE7F" authority="Yu &amp; Montgomery, 2007" authorityName="Yu &amp; Montgomery" authorityYear="2007" box="[195,789,367,393]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="7FBE650FFFC93A65FF756B76F1F6FE7F" bold="true" box="[195,473,367,393]" italics="true" pageId="27" pageNumber="28">Laricobius taiwanensis</emphasis>
<bibRefCitation id="295BC4ECFFC93A65FE576B76F33AFE7F" author="Yu" box="[481,789,367,393]" pageId="27" pageNumber="28" refString="Yu, G. &amp; Montgomery, M. E. (2007) A new species of Laricobius (Coleoptera: Derodontidae) from Taiwan. Formosan Entomology, 27, 341 - 347." type="journal article" year="2007">Yu &amp; Montgomery, 2007</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="05D0EA96FFC93A65FF216B8BF10FFE5C" box="[151,288,402,426]" pageId="27" pageNumber="28" type="description">
<paragraph id="4D75B91DFFC93A65FF216B8BF10FFE5C" blockId="27.[151,789,367,426]" box="[151,288,402,426]" pageId="27" pageNumber="28">(figs 20, 48)</paragraph>
</subSubSection>
<subSubSection id="05D0EA96FFC93A65FF216BC3F4DEFDA8" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="4D75B91DFFC93A65FF216BC3F4DEFDA8" blockId="27.[151,1437,474,1975]" pageId="27" pageNumber="28">
<emphasis id="7FBE650FFFC93A65FF216BC3F13EFE05" bold="true" box="[151,273,474,499]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Body bicolored. Head dark; ocelli absent. Prothorax transverse with well-developed lateral carina and sides explanate. Pronotum dark; posterior tooth present. Scutellum dark. Surface of elytra usually not incurvate; elytral punctures not confluent. Epipleura dark. Ventrites dark. Femora dark. Tibiae usually dark. Aedeagus with median carina on phallobase; medium lobe subacute; apices of parameres without an internal ridge.
</paragraph>
</subSubSection>
<subSubSection id="05D0EA96FFC93A65FF736873F165FA48" pageId="27" pageNumber="28" type="description">
<paragraph id="4D75B91DFFC93A65FF736873F165FA48" blockId="27.[151,1437,474,1975]" pageId="27" pageNumber="28">
<emphasis id="7FBE650FFFC93A65FF736873F178FD75" bold="true" box="[197,343,618,643]" pageId="27" pageNumber="28">Description.</emphasis>
Length
<quantity id="8A3214F8FFC93A65FE026873F278FD74" box="[436,599,618,642]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" metricValueMax="2.54" metricValueMin="1.58" pageId="27" pageNumber="28" unit="mm" value="2.06" valueMax="2.54" valueMin="1.58">1.582.54 mm</quantity>
(x = 2.67, n = 15). Body bicolored with head, prothorax, scutellum, elytra along sides to level of abdominal ventrite 5 (usually extending to stria 4 and covering striae 510), along suture (though maybealit reduced or absent), epipleuron, and venter black or chocolate brown; elytra dark red brown; antenna with A1 dark brown, funicle yellow brown, and club red-brown; palpi and legs dark brown, tarsi brown, but lighter in some specimens. Head without ocelli; u-shaped furrow present; interocular distance about
<date id="39749FDDFFC93A65FAE568E3F540FCE4" box="[1363,1391,762,786]" pageId="27" pageNumber="28">4x</date>
the width of the eye; macropunctures rounded and distinct, impressed and sometimes fused into trenches and set into depressions; micropunctures moderately coarse; setae erect, elongate with an average length as long as 2/3 to full width of the eye. Antenna with length of A3 about equal to A4, ratios 2.1:1.4:1.1:1.2:1.3:1.1:1.1:1.0:1.3:1.2:1.9; A11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides weakly convex, converging anteriorly from widest point and weakly convex, subparallel anteriorly and convergent posteriorly; about 0.79 x as long as wide (pronotal length/greatest pronotal width = 0.750.83, x = 0.79); depth =
<quantity id="8A3214F8FFC93A65FE8969EFF1CCFBF8" box="[319,483,1014,1038]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" metricValueMax="3.8" metricValueMin="1.6" pageId="27" pageNumber="28" unit="mm" value="0.27" valueMax="0.38" valueMin="0.16">0.160.38 mm</quantity>
(x = 0.29); pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle 90º to about 80° and subacute, not forming a tooth; posterior angle distinct, rounded, and with a short tooth; foveae visible in dorsal view; macropunctures on disc distinct, well-impressed and well-separated; micropunctures coarse; setae erect, moderately long with length 2/3 to full width of eye. Elytra about 2.91 x as long as wide (elytral length/greatest elytral width = 1.863.71, x = 2.91) and 2.92 x as long as pronotum (elytral length/ pronotal length = 1.733.30, x = 2.12); surface not or weakly-incurvate at basal third; macropunctures not coalescing to form grooves, separated on an average by 1/2 puncture diameter; micropunctures fine; microsculpture not visible; setae suberect to decumbent, short and on average about 2/3 to full length of the eye. Aedeagus with phallobase transverse (wider than long), 2/3 the length of the median lobe, median carina present; median lobe projecting slightly beyond apices of parameres, somewhat narrow with its greatest width equal to width of the parameres at their basal 1/3, apex subacute, median groove or carina absent, ostium subapical; parameres, slender and narrow, apically acute, very short setae present apically and along inner margin, internal subapical ridge absent. Spiculum gastrale narrow.
</paragraph>
</subSubSection>
<subSubSection id="05D0EA96FFC93A65FF736FD3F36EF929" pageId="27" pageNumber="28" type="discussion">
<paragraph id="4D75B91DFFC93A65FF736FD3F2EBF960" blockId="27.[151,1437,474,1975]" pageId="27" pageNumber="28">
<emphasis id="7FBE650FFFC93A65FF736FD3F162FA15" bold="true" box="[197,333,1482,1507]" pageId="27" pageNumber="28">Comments.</emphasis>
There are no discernable ocelli in this species, though the color of the cuticle that surrounds the deeper cephalic macropunctures may be lighter. Among the species that lack ocelli,
<taxonomicName id="8ACAC29EFFC93A65FBF06FE9F4C7F9F1" box="[1094,1256,1520,1543]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="7FBE650FFFC93A65FBF06FE9F4C7F9F1" box="[1094,1256,1520,1543]" italics="true" pageId="27" pageNumber="28">L. taiwanensis</emphasis>
</taxonomicName>
has the sides of the prothorax weakly convex, which will distinguish this species from the others. Having the dark tibiae will distinguish
<taxonomicName id="8ACAC29EFFC93A65FF6C6C21F155F9B9" box="[218,378,1592,1615]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="7FBE650FFFC93A65FF6C6C21F155F9B9" box="[218,378,1592,1615]" italics="true" pageId="27" pageNumber="28">L. taiwanensis</emphasis>
</taxonomicName>
from
<taxonomicName id="8ACAC29EFFC93A65FE086C21F248F9B9" box="[446,615,1592,1615]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="naganoensis">
<emphasis id="7FBE650FFFC93A65FE086C21F248F9B9" box="[446,615,1592,1615]" italics="true" pageId="27" pageNumber="28">L. naganoensis</emphasis>
</taxonomicName>
, the presence of of the posterior pronotal tooth will distinguish
<taxonomicName id="8ACAC29EFFC93A65FA9F6C21F0E0F985" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="7FBE650FFFC93A65FA9F6C21F0E0F985" italics="true" pageId="27" pageNumber="28">L. taiwanensis</emphasis>
</taxonomicName>
from
<taxonomicName id="8ACAC29EFFC93A65FEAA6C45F186F985" box="[284,425,1627,1651]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="osakensis">
<emphasis id="7FBE650FFFC93A65FEAA6C45F186F985" box="[284,425,1627,1651]" italics="true" pageId="27" pageNumber="28">L. osakensis</emphasis>
</taxonomicName>
, and the dark red coloration and presence of a median carina on the basal piece will distinguish
<taxonomicName id="8ACAC29EFFC93A65FEAF6C99F195F961" box="[281,442,1664,1687]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="7FBE650FFFC93A65FEAF6C99F195F961" box="[281,442,1664,1687]" italics="true" pageId="27" pageNumber="28">L. taiwanensis</emphasis>
</taxonomicName>
from
<emphasis id="7FBE650FFFC93A65FE496C99F2EBF960" box="[511,708,1662,1687]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="8ACAC29EFFC93A65FE496C99F2EFF960" box="[511,704,1662,1687]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="kangdingensis">L. kangdingensis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="4D75B91DFFC93A65FF736CBBF50FF94C" blockId="27.[151,1437,474,1975]" box="[197,1312,1698,1723]" pageId="27" pageNumber="28">
The
<typeStatus id="927107BFFFC93A65FF416CBBF176F94C" box="[247,345,1698,1722]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(NMNS) was not examined, but several
<typeStatus id="927107BFFFC93A65FC926CBAF3A0F94D" box="[804,911,1699,1723]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
were, belonging to the same series.
</paragraph>
<paragraph id="4D75B91DFFC93A65FF736CDEF36EF929" blockId="27.[151,1437,474,1975]" box="[197,833,1734,1759]" pageId="27" pageNumber="28">
<emphasis id="7FBE650FFFC93A65FF736CDEF120F929" bold="true" box="[197,271,1735,1759]" pageId="27" pageNumber="28">Hosts.</emphasis>
Plants:
<taxonomicName id="8ACAC29EFFC93A65FEDF6CD1F235F929" box="[361,538,1735,1759]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis id="7FBE650FFFC93A65FEDF6CD1F235F929" box="[361,538,1735,1759]" italics="true" pageId="27" pageNumber="28">Tsuga chinensis</emphasis>
</taxonomicName>
. Adelgid:
<emphasis id="7FBE650FFFC93A65FD276CDEF36EF929" box="[657,833,1735,1759]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="8ACAC29EFFC93A65FD276CDEF312F929" box="[657,829,1735,1759]" class="Insecta" family="Adelgidae" genus="Adelges" kingdom="Animalia" order="Aphidomorpha" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tsugae">Adelges tsugae</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="05D0EA96FFC93A65FF736CF3F316F8F4" box="[197,825,1770,1795]" pageId="27" pageNumber="28" type="biology_ecology">
<paragraph id="4D75B91DFFC93A65FF736CF3F316F8F4" blockId="27.[151,1437,474,1975]" box="[197,825,1770,1795]" pageId="27" pageNumber="28">
<emphasis id="7FBE650FFFC93A65FF736CF3F10BF8F5" bold="true" box="[197,292,1770,1795]" pageId="27" pageNumber="28">Biology.</emphasis>
Habitat and rearing (
<bibRefCitation id="295BC4ECFFC93A65FDA46CF3F302F8F4" author="Yu" box="[530,813,1770,1795]" pageId="27" pageNumber="28" refString="Yu, G. &amp; Montgomery, M. E. (2007) A new species of Laricobius (Coleoptera: Derodontidae) from Taiwan. Formosan Entomology, 27, 341 - 347." type="journal article" year="2007">Yu &amp; Montgomery 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="05D0EA96FFC93A65FF736D17F27CF8D0" box="[197,595,1806,1831]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="4D75B91DFFC93A65FF736D17F27CF8D0" blockId="27.[151,1437,474,1975]" box="[197,595,1806,1831]" pageId="27" pageNumber="28">
<emphasis id="7FBE650FFFC93A65FF736D17F171F8D1" bold="true" box="[197,350,1806,1831]" pageId="27" pageNumber="28">Distribution.</emphasis>
<collectingCountry id="35DDF98DFFC93A65FED36D17F194F8D0" box="[357,443,1806,1830]" name="Taiwan" pageId="27" pageNumber="28">Taiwan</collectingCountry>
: Kao Hsiung.
</paragraph>
</subSubSection>
<subSubSection id="05D0EA96FFC93A65FF736D2AF17BF840" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="4D75B91DFFC93A65FF736D2AF17BF840" blockId="27.[151,1437,474,1975]" pageId="27" pageNumber="28">
<emphasis id="7FBE650FFFC93A65FF736D2AF1C1F8BD" bold="true" box="[197,494,1842,1867]" pageId="27" pageNumber="28">
<typeStatus id="927107BFFFC93A65FF736D2AF12FF8BD" box="[197,256,1843,1867]" pageId="27" pageNumber="28">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="927107BFFFC93A65FE436D2BF2BBF8BC" box="[501,660,1842,1866]" pageId="27" pageNumber="28" type="paratype">PARATYPES</typeStatus>
: 21 (10,
<collectionCode id="2BDB21D8FFC93A65FD416D2AF364F8BC" box="[759,843,1843,1866]" pageId="27" pageNumber="28">BMNH</collectionCode>
[with all original labels hw]; 4,
<collectionCode id="2BDB21D8FFC93A65FB0F6D2AF53DF8BC" box="[1209,1298,1843,1866]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="27" pageNumber="28">NMNH</collectionCode>
; 3,
<collectionCode id="2BDB21D8FFC93A65FA8D6D2AF554F8BC" LSID="urn:lsid:biocol.org:col:34880" box="[1339,1403,1843,1866]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="27" pageNumber="28">YPM</collectionCode>
; 4,
<collectionCode id="2BDB21D8FFC93A65FF216D4FF0C9F898" box="[151,230,1878,1902]" httpUri="http://grbio.org/cool/e3b9-k2i3" name="United States Department of Agriculture" pageId="27" pageNumber="28">USDA</collectionCode>
, one each with Havill Sample # 08-50, Havill Sample # 08-51),
<collectingCountry id="35DDF98DFFC93A65FC0B6D4EF406F899" box="[957,1065,1879,1903]" name="Taiwan" pageId="27" pageNumber="28">TAIWAN</collectingCountry>
Kao Hsiung Co. Road 20, 2 Km west of Ya Kou Hotel
<date id="39749FDDFFC93A65FE2E6D63F21FF864" box="[408,560,1914,1939]" pageId="27" pageNumber="28" value="1994-05-26">26 May 1994</date>
S.T. Murphy/
<typeStatus id="927107BFFFC93A65FD7A6D63F375F864" box="[716,858,1914,1938]" pageId="27" pageNumber="28" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="8ACAC29EFFC93A65FCD46D63F570F865" authority="Yu and Montgomery" authorityName="Yu and Montgomery" box="[866,1375,1914,1939]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="taiwanensis">Laricobius taiwanensis, Yu and Montgomery</taxonomicName>
(yellow green label).
</paragraph>
</subSubSection>
</treatment>
</document>