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<mods:title id="CA74CAF14289326D1167D0BABEE62B1F">Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines</mods:title>
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<mods:namePart id="987B573D00A064D1A1EB42086C56B8F9">Van Hinsbergh, Victor W. M.</mods:namePart>
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<treatment id="C5011D20FFD6FFF7FF01FABFC2BFAB40" ID-DOI="http://doi.org/10.5281/zenodo.5937023" ID-GBIF-Taxon="154474764" ID-Zenodo-Dep="5937023" LSID="urn:lsid:plazi:treatment:C5011D20FFD6FFF7FF01FABFC2BFAB40" httpUri="http://treatment.plazi.org/id/C5011D20FFD6FFF7FF01FABFC2BFAB40" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414">
<subSubSection id="05B2FFBDFFD6FFF4FF01FABFC767A973" pageId="13" pageNumber="414" type="nomenclature">
<paragraph id="4D17AC36FFD6FFF4FF01FABFC7B7A91C" blockId="13.[151,411,1293,1352]" box="[151,411,1293,1319]" pageId="13" pageNumber="414">
<heading id="165F1B5AFFD6FFF4FF01FABFC7B7A91C" bold="true" box="[151,411,1293,1319]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
<taxonomicName id="8AA8D7B5FFD6FFF4FF01FABFC7B7A91C" ID-CoL="35CH7" box="[151,411,1293,1319]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="garmani">
<emphasis id="7FDC7024FFD6FFF4FF01FABFC7B7A91C" bold="true" box="[151,411,1293,1319]" pageId="13" pageNumber="414">
<emphasis id="7FDC7024FFD6FFF4FF01FABFC6D2A91C" bold="true" box="[151,254,1293,1319]" italics="true" pageId="13" pageNumber="414">Diaphus</emphasis>
aff.
<emphasis id="7FDC7024FFD6FFF4FEA3FABFC7B7A91C" bold="true" box="[309,411,1293,1319]" italics="true" pageId="13" pageNumber="414">garmani</emphasis>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4D17AC36FFD6FFF4FF01FA9DC767A973" blockId="13.[151,411,1293,1352]" box="[151,331,1327,1352]" pageId="13" pageNumber="414">(Figures 4850)</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFD6FFF7FF01FACAC2BFAB40" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414" type="description">
<paragraph id="4D17AC36FFD6FFF4FF01FACAC55AA9B5" blockId="13.[151,936,1400,1453]" box="[151,886,1400,1422]" pageId="13" pageNumber="414">
1995
<taxonomicName id="8AA8D7B5FFD6FFF4FF46FACAC7EBA9B5" authority="Smale" authorityName="Smale" box="[208,455,1400,1422]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="garmani">
<emphasis id="7FDC7024FFD6FFF4FF46FACAC7AEA9B5" box="[208,386,1400,1422]" italics="true" pageId="13" pageNumber="414">Diaphus garmani</emphasis>
Smale
</taxonomicName>
MJ, Watson G, Hecht T, p. 43, plate 17 F1
</paragraph>
<paragraph id="4D17AC36FFD6FFF4FF01FA24C584A997" blockId="13.[151,936,1400,1453]" box="[151,936,1430,1453]" pageId="13" pageNumber="414">
2013a
<taxonomicName id="8AA8D7B5FFD6FFF4FF4DFA24C7A1A996" box="[219,397,1430,1453]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="garmani">
<emphasis id="7FDC7024FFD6FFF4FF4DFA24C7A1A996" box="[219,397,1430,1453]" italics="true" pageId="13" pageNumber="414">Diaphus garmani</emphasis>
</taxonomicName>
(group) Schwarzhans pp. 6468, plate 10, fig. 1216
</paragraph>
<paragraph id="4D17AC36FFD6FFF4FF01FA6BC218AA2D" blockId="13.[151,1436,1497,1918]" pageId="13" pageNumber="414">
Material:
<specimenCount id="5BAE67BFFFD6FFF4FE90FA68C7BEA9C9" box="[262,402,1497,1522]" pageId="13" pageNumber="414" type="generic">9 specimens</specimenCount>
in total. Anda2 (3) RGM 962147,
<materialsCitation id="FDC0A66BFFD6FFF4FCB4FA68C5EAA9C9" ID-GBIF-Occurrence="2013952827" box="[802,966,1497,1522]" collectionCode="RGM" pageId="13" pageNumber="414" specimenCode="RGM 962148" specimenCount="1">
<specimenCode id="1D0E044DFFD6FFF4FCB4FA68C5EEA9C9" box="[802,962,1497,1522]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="414" type="Museum">RGM 962148</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFD6FFF4FC59FA68C243A9C9" ID-GBIF-Occurrence="2013952841" box="[975,1135,1497,1522]" collectionCode="RGM" pageId="13" pageNumber="414" specimenCode="RGM 962149" specimenCount="1">
<specimenCode id="1D0E044DFFD6FFF4FC59FA68C243A9C9" box="[975,1135,1497,1522]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="414" type="Museum">RGM 962149</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFD6FFF4FBEDFA6BC3B4A9C9" ID-GBIF-Occurrence="2013952820" box="[1147,1432,1497,1522]" collectionCode="RGM" pageId="13" pageNumber="414" specimenCode="RGM 962150" specimenCount="1">
Anda3 (2)
<specimenCode id="1D0E044DFFD6FFF4FB6EFA68C3B4A9C9" box="[1272,1432,1497,1522]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="414" type="Museum">RGM 962150</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFD6FFF4FF01FA4FC783AA2D" ID-GBIF-Occurrence="2013952822" box="[151,431,1533,1558]" collectionCode="RGM" pageId="13" pageNumber="414" specimenCode="RGM 962151" specimenCount="1">
Anda4 (1)
<specimenCode id="1D0E044DFFD6FFF4FE86FA4CC783AA2D" box="[272,431,1533,1558]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="414" type="Museum">RGM 962151</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFD6FFF4FE2CFA4FC4FEAA2D" ID-GBIF-Occurrence="2013952858" box="[442,722,1533,1558]" collectionCode="RGM" pageId="13" pageNumber="414" specimenCode="RGM 962152" specimenCount="1">
Anda5 (1)
<specimenCode id="1D0E044DFFD6FFF4FDA5FA4CC4FEAA2D" box="[563,722,1533,1558]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="414" type="Museum">RGM 962152</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFD6FFF4FD4BFA4FC21DAA2D" ID-GBIF-Occurrence="2013952835" box="[733,1073,1533,1558]" collectionCode="RGM" pageId="13" pageNumber="414" specimenCode="RGM 962154" specimenCount="1">
AndaDeVos (2)
<specimenCode id="1D0E044DFFD6FFF4FC04FA4CC21DAA2D" box="[914,1073,1533,1558]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="414" type="Museum">RGM 962154</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4D17AC36FFD6FFF4FF51F993C7F1AAB9" blockId="13.[151,1436,1497,1918]" pageId="13" pageNumber="414">
Besides the otoliths that meet perfectly the characteristics of
<taxonomicName id="8AA8D7B5FFD6FFF4FC1BF990C5D6AA02" box="[909,1018,1570,1593]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="regani">
<emphasis id="7FDC7024FFD6FFF4FC1BF990C5D6AA02" box="[909,1018,1570,1593]" italics="true" pageId="13" pageNumber="414">D. regani</emphasis>
</taxonomicName>
, eight flat specimens with a similar outline of
<taxonomicName id="8AA8D7B5FFD6FFF4FE9AF9F4C754AA66" box="[268,376,1606,1629]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="regani">
<emphasis id="7FDC7024FFD6FFF4FE9AF9F4C754AA66" box="[268,376,1606,1629]" italics="true" pageId="13" pageNumber="414">D. regani</emphasis>
</taxonomicName>
differ from this species by a less wide ostium, the height of which hardly exceeds that of the cauda and pseudocolliculum.
</paragraph>
<paragraph id="4D17AC36FFD6FFF4FF51F93FC5BFAB45" blockId="13.[151,1436,1497,1918]" pageId="13" pageNumber="414">
The available otoliths (OL:OH=
<quantity id="8A5001D3FFD6FFF4FDA3F93FC4EBAA9D" box="[565,711,1677,1702]" metricMagnitude="-2" metricUnit="m" metricValue="3.3274" metricValueMax="3.4544" metricValueMin="3.2004" pageId="13" pageNumber="414" unit="in" value="1.31" valueMax="1.36" valueMin="1.26">1.261.36 in</quantity>
the depicted adult specimens) display a range of preservations ranging from good to eroded. As erosion of other
<taxonomicName id="8AA8D7B5FFD6FFF4FD29F903C532AAF1" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1890" box="[703,798,1713,1738]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFD6FFF4FD29F903C532AAF1" box="[703,798,1713,1738]" italics="true" pageId="13" pageNumber="414">Diaphus</emphasis>
</taxonomicName>
species can also result in an outline that is reminiscent of
<taxonomicName id="8AA8D7B5FFD6FFF4FF01F964C72EAAD6" box="[151,258,1750,1773]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="regani">
<emphasis id="7FDC7024FFD6FFF4FF01F964C72EAAD6" box="[151,258,1750,1773]" italics="true" pageId="13" pageNumber="414">D. regani</emphasis>
</taxonomicName>
, we cannot exclude that this is the case in a few of our otoliths. Nevertheless, the specimen depicted in Figure 49 (RGM 962147) is well preserved. It is related to otoliths in the
<taxonomicName id="8AA8D7B5FFD6FFF4FC64F94BC296AB2A" box="[1010,1210,1785,1810]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="garmani">
<emphasis id="7FDC7024FFD6FFF4FC64F94BC296AB2A" box="[1010,1210,1785,1810]" italics="true" pageId="13" pageNumber="414">Diaphus garmani</emphasis>
</taxonomicName>
group. It is closely related to the extant
<taxonomicName id="8AA8D7B5FFD6FFF4FE1AF8AFC4DAAB0D" authority="Gilbert 1906" authorityName="Gilbert" authorityYear="1906" box="[396,758,1821,1846]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="garmani">
<emphasis id="7FDC7024FFD6FFF4FE1AF8AFC479AB0E" box="[396,597,1821,1846]" italics="true" pageId="13" pageNumber="414">Diaphus garmani</emphasis>
Gilbert 1906
</taxonomicName>
, which has a broad (sub)tropical occurrence, but also has resemblance to
<taxonomicName id="8AA8D7B5FFD6FFF4FEDAF8F3C4C8AB61" authority="Ohe &amp; Araki, 1973" authorityName="Ohe &amp; Araki" authorityYear="1973" box="[332,740,1857,1882]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="hataii">
<emphasis id="7FDC7024FFD6FFF4FEDAF8F3C7DAAB61" box="[332,502,1857,1882]" italics="true" pageId="13" pageNumber="414">Diaphus hataii</emphasis>
<bibRefCitation id="2939D1C7FFD6FFF4FD96F8F3C4C8AB61" author="Ohe, F. &amp; Araki, Y." box="[512,740,1857,1882]" pageId="13" pageNumber="414" pagination="407 - 413" refId="ref19564" refString="Ohe, F. &amp; Araki, Y. (1973) Some Miocene fish otoliths from the Yakuoji Formation, Tsu, Mie Prefecture. Science Reports of the Tohoku University, Series 2 (Geology), Special Volume, 6, 407 - 413." type="journal article" year="1973">Ohe &amp; Araki, 1973</bibRefCitation>
</taxonomicName>
, which is known from the Miocene in
<collectingCountry id="35BFECA6FFD6FFF4FB3CF8F0C2C6AB62" box="[1194,1258,1858,1881]" name="Japan" pageId="13" pageNumber="414">Japan</collectingCountry>
(
<bibRefCitation id="2939D1C7FFD6FFF4FB6AF8F3C6FCAB45" author="Ohe, F. &amp; Araki, Y." pageId="13" pageNumber="414" pagination="407 - 413" refId="ref19564" refString="Ohe, F. &amp; Araki, Y. (1973) Some Miocene fish otoliths from the Yakuoji Formation, Tsu, Mie Prefecture. Science Reports of the Tohoku University, Series 2 (Geology), Special Volume, 6, 407 - 413." type="journal article" year="1973">Ohe &amp; Araki, 1973</bibRefCitation>
) and the Caribbean Pliocene (
<bibRefCitation id="2939D1C7FFD6FFF4FDB7F8D7C5ABAB45" author="Schwarzhans, W. &amp; Aguilera, O." box="[545,903,1893,1918]" pageId="13" pageNumber="414" pagination="83 - 150" refId="ref19902" refString="Schwarzhans, W. &amp; Aguilera, O. (2013) Otoliths of the Myctophidae from the Neogene of tropical America. Palaeo Ichthyologica, 13, 83 - 150." type="journal article" year="2013">Schwarzhans &amp; Aguilera, 2013</bibRefCitation>
).
</paragraph>
<caption id="19D7FCBEFFD5FFF7FF01F895C555AB06" box="[151,889,1831,1854]" httpUri="https://zenodo.org/record/2601297/files/figure.png" pageId="14" pageNumber="415" startId="14.[151,234,1831,1853]" targetBox="[170,1428,179,1806]" targetPageId="14">
<paragraph id="4D17AC36FFD5FFF7FF01F895C555AB06" blockId="14.[151,1281,1831,2007]" box="[151,889,1831,1854]" pageId="14" pageNumber="415">
<emphasis id="7FDC7024FFD5FFF7FF01F895C72EAB06" bold="true" box="[151,258,1831,1853]" pageId="14" pageNumber="415">PLATE 4.</emphasis>
39.
<taxonomicName id="8AA8D7B5FFD5FFF7FEB9F895C417AB05" box="[303,571,1831,1854]" class="Actinopterygii" family="Myctophidae" genus="Ceratoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="warmingii">
<emphasis id="7FDC7024FFD5FFF7FEB9F895C417AB05" box="[303,571,1831,1854]" italics="true" pageId="14" pageNumber="415">Ceratoscopelus warmingii</emphasis>
</taxonomicName>
, RGM 962336, locality Roxas.
</paragraph>
</caption>
<paragraph id="4D17AC36FFD5FFF7FF01F8F4C52BAB67" blockId="14.[151,1281,1831,2007]" box="[151,775,1862,1884]" pageId="14" pageNumber="415">
40.
<taxonomicName id="8AA8D7B5FFD5FFF7FF2BF8F4C7E5AB67" box="[189,457,1862,1884]" class="Actinopterygii" family="Myctophidae" genus="Ceratoscopelus" kingdom="Animalia" order="Myctophiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="species" species="warmingii">
<emphasis id="7FDC7024FFD5FFF7FF2BF8F4C7E5AB67" box="[189,457,1862,1884]" italics="true" pageId="14" pageNumber="415">Ceratoscopelus warmingii</emphasis>
</taxonomicName>
, RGM 962335, locality Roxas.
</paragraph>
<paragraph id="4D17AC36FFD5FFF7FF01F8D7C2BFAB40" blockId="14.[151,1281,1831,2007]" box="[151,1171,1893,1915]" pageId="14" pageNumber="415">
41.
<taxonomicName id="8AA8D7B5FFD5FFF7FF2BF8D7C73FAB40" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1890" box="[189,275,1893,1915]" class="Actinopterygii" family="Myctophidae" genus="Diaphus" kingdom="Animalia" order="Myctophiformes" pageId="14" pageNumber="415" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFD5FFF7FF2BF8D7C73FAB40" box="[189,275,1893,1915]" italics="true" pageId="14" pageNumber="415">Diaphus</emphasis>
</taxonomicName>
sp. 1, RGM 962143, locality Roxas. (a) Lateral face, (b) ventral profile, (c) medial face.
</paragraph>
</subSubSection>
</treatment>
</document>