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<document id="9179E0F469AC5CCBA48BA3A7228822F4" ID-DOI="10.11646/zootaxa.4061.1.9" ID-GBIF-Dataset="260194f3-503b-4d7b-b425-95305d9f8f6d" ID-ISSN="1175-5326" ID-Zenodo-Dep="270382" ID-ZooBank="1C0F8AB3-7382-4E1A-954D-BA74179D6239" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458391305552" checkinUser="plazi" docAuthor="Nihei, Silvio S., Andrade, Marcos R., Pape, Thomas &amp; Cerretti, Pierfilippo" docDate="2016" docId="AB553714FFB51878FF42FF406918FE18" docLanguage="en" docName="zootaxa.4061.1.9.pdf.imf" docOrigin="Zootaxa 4061 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Shannoniella setinervis Nihei, Andrade, Pape &amp; Cerretti, 2016, sp. nov." docType="treatment" docVersion="6" lastPageNumber="91" masterDocId="576C4F6CFFB1187EFFD5FFD76B28FFC8" masterDocTitle="The Shannoniella sisters (Diptera: Rhinophoridae)" masterLastPageNumber="92" masterPageNumber="85" pageNumber="89" updateTime="1698185520000" updateUser="plazi">
<mods:mods id="9F125326F0B5F7FB41A78B93B6F30C01" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8E08B37BE902C11AC7FEB84C11E1B9D7">
<mods:title id="B87EED008FF73877EEFB5E9FE8F9D3B6">The Shannoniella sisters (Diptera: Rhinophoridae)</mods:title>
</mods:titleInfo>
<mods:name id="B87BF66E398E208CEB4AF07BB41FBCB8" type="personal">
<mods:role id="59B8EF63C275677E73EEF72105FCB1F0">
<mods:roleTerm id="215F09C92DC291B64372F545E74275D3">Author</mods:roleTerm>
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<mods:namePart id="555811FB425AB773BE737ACEE17896E5">Nihei, Silvio S.</mods:namePart>
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<mods:name id="460D687A9783C18595F2582675535DF1" type="personal">
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<mods:namePart id="051B45D2F54E76ABFD1301F2FEB4D8EA">Andrade, Marcos R.</mods:namePart>
</mods:name>
<mods:name id="C86D5C4CA0D082550E78D295277AB042" type="personal">
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<mods:namePart id="A19D56F35B4C8A32F9309D870630C946">Pape, Thomas</mods:namePart>
</mods:name>
<mods:name id="923B3C634954998C3FAE65D59660FAD0" type="personal">
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<mods:namePart id="6FB1FE4C6630D40246CED836C5A44C88">Cerretti, Pierfilippo</mods:namePart>
</mods:name>
<mods:typeOfResource id="0491A19FC4327E0E26FE4E49A19492C4">text</mods:typeOfResource>
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<mods:title id="5D01112329A1A71AC11E5ACD720E6F8A">Zootaxa</mods:title>
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<mods:part id="EE1E2D4F89790C79CB2527D9B2B2B4CB">
<mods:date id="F8BD99FB454BD8EAB79C291D805B1F21">2016</mods:date>
<mods:detail id="191BF863C00167DD13018490247CE162" type="volume">
<mods:number id="32E803E33A7AA3A3D0F68A2A53B4A3B5">4061</mods:number>
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<mods:identifier id="A25279C8AEABFBA9FD475426EFC5B8F0" type="DOI">10.11646/zootaxa.4061.1.9</mods:identifier>
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<treatment id="AB553714FFB51878FF42FF406918FE18" ID-DOI="http://doi.org/10.5281/zenodo.6058383" ID-GBIF-Taxon="127638553" ID-Zenodo-Dep="6058383" LSID="urn:lsid:plazi:treatment:AB553714FFB51878FF42FF406918FE18" httpUri="http://treatment.plazi.org/id/AB553714FFB51878FF42FF406918FE18" lastPageId="6" lastPageNumber="91" pageId="4" pageNumber="89">
<subSubSection id="6BE6D589FFB5187AFF42FF406A3FFF1B" pageId="4" pageNumber="89" type="nomenclature">
<paragraph id="23438602FFB5187AFF42FF406930FF7A" blockId="4.[151,536,151,211]" box="[151,536,151,178]" pageId="4" pageNumber="89">
<heading id="780B316EFFB5187AFF42FF406930FF7A" bold="true" box="[151,536,151,178]" fontSize="11" level="1" pageId="4" pageNumber="89" reason="1">
<emphasis id="11885A10FFB5187AFF42FF406930FF7A" bold="true" box="[151,536,151,178]" pageId="4" pageNumber="89">
<taxonomicName id="E4FCFD81FFB5187AFF42FF406A9BFF79" ID-CoL="8DWV6" box="[151,435,151,177]" class="Insecta" family="Rhinophoridae" genus="Shannoniella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="setinervis" status="sp. nov.">
<emphasis id="11885A10FFB5187AFF42FF406A9BFF79" bold="true" box="[151,435,151,177]" italics="true" pageId="4" pageNumber="89">Shannoniella setinervis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0ABBE76BFFB5187AFE6FFF4F6930FF7A" box="[442,536,152,178]" pageId="4" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="23438602FFB5187AFF42FF6D6A3FFF1B" blockId="4.[151,536,151,211]" box="[151,279,186,211]" pageId="4" pageNumber="89">
(
<figureCitation id="BBC79A87FFB5187AFF4AFF6D6BCDFF1B" box="[159,229,186,211]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="4" pageNumber="89">Figs 2</figureCitation>
ag)
</paragraph>
</subSubSection>
<subSubSection id="6BE6D589FFB5187AFF42FED56844FE63" pageId="4" pageNumber="89" type="reference_group">
<paragraph id="23438602FFB5187AFF42FED56F40FED3" blockId="4.[151,1437,258,2025]" box="[151,1128,258,283]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF42FED56A08FED3" bold="true" box="[151,288,258,283]" pageId="4" pageNumber="89">References.</emphasis>
Cerretti &amp; Pape (2012); Cerretti
<emphasis id="11885A10FFB5187AFD41FED369EDFED3" box="[660,709,258,283]" italics="true" pageId="4" pageNumber="89">et al</emphasis>
. (2014) [both as “
<taxonomicName id="E4FCFD81FFB5187AFC44FED56F0DFED3" ID-CoL="644Q4" box="[913,1061,258,283]" class="Insecta" family="Rhinophoridae" genus="Shannoniella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="11885A10FFB5187AFC44FED56F0DFED3" box="[913,1061,258,283]" italics="true" pageId="4" pageNumber="89">Shannoniella</emphasis>
</taxonomicName>
sp.”].
</paragraph>
<paragraph id="23438602FFB5187AFF12FEF16844FE63" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12FEF16AD0FEF7" bold="true" box="[199,504,294,319]" pageId="4" pageNumber="89">Type-material examined:</emphasis>
<typeStatus id="FC4738A0FFB5187AFDD6FEF16943FEF7" box="[515,619,294,319]" pageId="4" pageNumber="89" type="holotype">Holotype</typeStatus>
♂ (MZSP), “Therezopolis [Teresópolis]/ Est. do Rio [Rio de Janeiro State]
<date id="5742A0C2FFB5187AFF0BFE9C6A09FEAB" box="[222,289,331,355]" pageId="4" pageNumber="89" value="1939-11">XI-39</date>
/ J. F. T. de Freitas [collector]” (white label with black border), “10C” (white label with black border), “Holotipo” (green label), “
<taxonomicName id="E4FCFD81FFB5187AFE1DFEB96974FE4F" ID-CoL="644Q4" box="[456,604,366,391]" class="Insecta" family="Rhinophoridae" genus="Shannoniella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">Shannoniella</taxonomicName>
/
<taxonomicName id="E4FCFD81FFB5187AFDA2FEB969CAFE4F" ID-CoL="8DWV6" box="[631,738,366,391]" class="Insecta" family="Rhinophoridae" genus="Shannoniella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="setinervis">setinervis</taxonomicName>
/ S.S. Nihei des. 2015”.
<typeStatus id="FC4738A0FFB5187AFBD3FEB86F5FFE4F" box="[1030,1143,367,391]" pageId="4" pageNumber="89" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="35FA4D8BFFB5187AFB51FEB86F9AFE4E" box="[1156,1202,367,391]" pageId="4" pageNumber="89" type="generic">2 ♂</specimenCount>
,
<collectingCountry id="5BEBC692FFB5187AFB14FEB96E20FE4F" box="[1217,1288,366,391]" name="Brazil" pageId="4" pageNumber="89">Brazil</collectingCountry>
, Paraná, São José dos Pinhais, Serra do Mar,
<date id="5742A0C2FFB5187AFE28FE4469A1FE63" box="[509,649,403,427]" pageId="4" pageNumber="89" value="1984-07-20">20.VII.1984</date>
, J.A. Rafael (CNC).
</paragraph>
</subSubSection>
<subSubSection id="6BE6D589FFB5187AFF12FE616A23FDD1" pageId="4" pageNumber="89" type="etymology">
<paragraph id="23438602FFB5187AFF12FE616A23FDD1" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12FE616A60FE07" bold="true" box="[199,328,438,463]" pageId="4" pageNumber="89">Etymology</emphasis>
: The species epithet is derived from the Latin
<emphasis id="11885A10FFB5187AFCA4FE6068B6FE06" box="[881,926,439,462]" italics="true" pageId="4" pageNumber="89">seta</emphasis>
= bristle and
<emphasis id="11885A10FFB5187AFB93FE606FB8FE06" box="[1094,1168,439,462]" italics="true" pageId="4" pageNumber="89">nervus</emphasis>
= vein. The species is named after the more extensive row of setulae dorsally on wing vein R4+5 as compared with its sister species
<taxonomicName id="E4FCFD81FFB5187AFA52FE0C6A2CFDD2" class="Insecta" family="Rhinophoridae" genus="Shannoniella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="11885A10FFB5187AFA52FE0C6A2CFDD2" italics="true" pageId="4" pageNumber="89">S. cuspidata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="6BE6D589FFB51878FF12FDF26A8AFE40" lastPageId="6" lastPageNumber="91" pageId="4" pageNumber="89" type="description">
<paragraph id="23438602FFB5187AFF12FDF26FA7FDF6" blockId="4.[151,1437,258,2025]" box="[199,1167,549,574]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12FDF26AB4FDF6" bold="true" box="[199,412,549,574]" pageId="4" pageNumber="89">Description. Male</emphasis>
. Body length:
<quantity id="E4042BE7FFB5187AFD97FDF169B0FDF5" box="[578,664,550,574]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="89" unit="mm" value="4.3">4.3 mm</quantity>
(
<typeStatus id="FC4738A0FFB5187AFD72FDF26827FDF6" box="[679,783,549,574]" pageId="4" pageNumber="89" type="holotype">holotype</typeStatus>
); wing length:
<quantity id="E4042BE7FFB5187AFC63FDF26F24FDF5" box="[950,1036,549,574]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="89" unit="mm" value="3.5">3.5 mm</quantity>
(
<typeStatus id="FC4738A0FFB5187AFBCEFDF26FABFDF6" box="[1051,1155,549,574]" pageId="4" pageNumber="89" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="23438602FFB5187AFF12FD9E6AD0FCF5" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12FD9E6A3EFDAA" box="[199,278,585,610]" italics="true" pageId="4" pageNumber="89">Colour</emphasis>
(
<figureCitation id="BBC79A87FFB5187AFEF0FD9E6A42FDAA" box="[293,362,585,610]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="4" pageNumber="89">Fig. 2</figureCitation>
a). Body mostly light brown. Head light brown with grey pruinosity on gena, postgena and face; scape brown; pedicel yellow; postpedicel orangish brown, the base near arista yellowish; arista orangish brown with yellow base; palpus yellow. Thorax light brown with grey pruinosity on notopleuron, anepisternum, katepisternum, anatergite and katatergite. Coxae light brown with grey pruinosity; trochanters yellowish; femora yellowish with the apical third light brown; tibiae and tarsi light brown. Wing infuscated with light brown, but with 3 clear (non-infuscated), hyaline spots on cells r4+5, dm and m. Upper and lower calypters slightly infuscated on borders. Abdomen light brown.
</paragraph>
<paragraph id="23438602FFB5187AFF12FC9F689DFB49" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12FC9F6A2CFCA9" box="[199,260,840,865]" italics="true" pageId="4" pageNumber="89">Head</emphasis>
(
<figureCitation id="BBC79A87FFB5187AFEC1FC9F6A72FCA9" box="[276,346,840,865]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="4" pageNumber="89">Fig. 2</figureCitation>
b). Eye bare. Arista thickened almost to the tip; arista thickly micropubescent. First and second aristomere elongated, the second as long as the pedicel; third aristomere very long, about 2.9 times as long as the pedicel. Postpedicel elongate, slightly surpassing lower facial margin, about 4.5 times as long as the pedicel. Frontal stripe with subparallel margins, width at vertex about 1.8 times the eye width in dorsal view, and about 0.5 times the head width. No ocellar setae. Inner vertical seta strong and convergent; outer vertical seta not developed. Two lateroclinate orbital setae, the uppermost stronger. Seven frontal setae. Parafacial bare, very narrow. Face very broad, deeply sunken. Vibrissal angle below ventral margin of gena, distinctly projected forward and turned inwards. Vibrissa not easily distinguished from the several perivibrissal setae. Facial ridge straight and setulose along its full length. Postocular setae short. Proboscis and palpus short.
</paragraph>
<paragraph id="23438602FFB5187AFF12FB5B69D9FA69" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12FB5B6A30FB6D" box="[199,280,1164,1189]" italics="true" pageId="4" pageNumber="89">Thorax</emphasis>
. Scutum with long, fine clothing setulae; acrostichals not developed; dorsocentrals 2+3, the presutural ones interspersed by moderately long setulae; no intra-alars; supra-alars 1+1, prealar (first postsutural supra-alar before the wing insertion) not developed. Postpronotum with 2 strong and long setae, with fine clothing setulae. Posthumeral 1; notopleurals 2, the anterior nearly twice as long as the posterior. Prosternum and proepisternum bare. One upcurved proepisternal and one upcurved proepimeral setae. Two katepisternals. Three long anepimeral setulae, one more developed. Katepimeron bare. Anatergite with fine setulae. Scutellum with one pair of long, divergent subapical setae; one pair of median discal setae close to the margin and nearly as long and strong as the subapicals; some irregularly developed discal setulae.
</paragraph>
<paragraph id="23438602FFB5187AFF12FA7A6F94F9F9" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12FA7A6BD4FA0C" box="[199,252,1453,1476]" italics="true" pageId="4" pageNumber="89">Legs</emphasis>
. Fore femur with a row of long posteroventral setae, and the posterior surface with several developed setae. Fore tibia with one weak submedian posterodorsal seta, and one weak submedian posteroventral seta. Mid tibia with one weak submedian posterodorsal seta.
<collectingCountry id="5BEBC692FFB5187AFD18FA23682DF9C5" box="[717,773,1524,1549]" name="India" pageId="4" pageNumber="89">Hind</collectingCountry>
tibia with one submedian anterodorsal seta, one submedian posterodorsal seta, and one submedian anteroventral seta (erect in right leg, but not in left leg).
</paragraph>
<paragraph id="23438602FFB5187AFF12F9EA6A92F8D9" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12F9EA6A29F99C" box="[199,257,1597,1620]" italics="true" pageId="4" pageNumber="89">Wing</emphasis>
(
<figureCitation id="BBC79A87FFB5187AFEC4F9EB6A7FF99D" box="[273,343,1596,1621]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="4" pageNumber="89">Fig. 2</figureCitation>
c). Costal spine not differentiated. Vein R1 entirely setulose dorsally; R4+5 setulose dorsally from the base and reaching the level of crossvein dm-cu. Section of M between crossvein dm-cu and the apical bend about 0.9 times as long as the preceding section (between dm-cu and r-m crossveins). M reaching costa at wing margin; cell r4+5 open, the distance between M and R4+5 at wing margin about three times as long as crossvein r-m. Vein R 4+5 diverging from R2+3, the distance between both at wing margin longer than crossvein dm-cu (this about 3/ 4 times the said distance).
</paragraph>
<paragraph id="23438602FFB5187AFF12F8CB6EB8F890" blockId="4.[151,1437,258,2025]" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12F8CB6A19F8FD" box="[199,305,1820,1845]" italics="true" pageId="4" pageNumber="89">Abdomen</emphasis>
(
<figureCitation id="BBC79A87FFB5187AFEEAF8CB6AABF8FD" box="[319,387,1820,1845]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="4" pageNumber="89">Fig. 2</figureCitation>
d). Tergites covered by long fine setulae. Syntergite 1+2 without a mid-dorsal depression, with several lateral setae, and one pair of median marginal setae. Tergites 3, 4 and 5 each with a row of marginal setae.
</paragraph>
<paragraph id="23438602FFB51878FF12F8B36899FEAD" blockId="4.[151,1437,258,2025]" lastBlockId="6.[151,1437,151,464]" lastPageId="6" lastPageNumber="91" pageId="4" pageNumber="89">
<emphasis id="11885A10FFB5187AFF12F8B36AA9F8B5" box="[199,385,1892,1917]" italics="true" pageId="4" pageNumber="89">Male terminalia</emphasis>
(
<figureCitation id="BBC79A87FFB5187AFE40F8B36AC9F8B5" box="[405,481,1892,1917]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="4" pageNumber="89">Figs 2</figureCitation>
eg). Sternite 5 (
<figureCitation id="BBC79A87FFB5187AFD7EF8B369DBF8B5" box="[683,755,1892,1917]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="4" pageNumber="89">Fig. 2</figureCitation>
e) upright laterally, with a deep median cleft; lateral lobe characterized by a posterior three-dimensional cap-like apophysis, but slightly bilobed laterally; median basal window absent. Tergite 6 divided medially into two hemitergites, each with a row of setae along posterior margin and separated from segment 7+8 by a very narrow membrane. Sternite 6 almost symmetrical, articulated with segment 7+8 on left side, and attached to it by a short membrane on its right side. Epandrium very short and convex. Surstylus well developed, broad in lateral view; surstylus widely fused to epandrium (
<figureCitation id="BBC79A87FFB4187BFB26F89A6E13F8AE" box="[1267,1339,1869,1894]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="5" pageNumber="90">Fig. 2</figureCitation>
f). Cerci slightly shorter than surstylus, not fused medially (
<figureCitation id="BBC79A87FFB4187BFD2DF8A66869F842" box="[760,833,1905,1930]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="5" pageNumber="90">Fig. 2</figureCitation>
f), pointed apically (sub-triangular in lateral view). Processi longi strongly widened at mid-length and medially almost touching each other; processus longus firmly and widely fused with surstylus. Basal plate of hypandrium very short, not concave, hypandrial arms very long and converging. Aedeagus (
<figureCitation id="BBC79A87FFB71878FE70FF406AC3FF78" box="[421,491,151,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="6" pageNumber="91">
Fig.
<quantity id="E4042BE7FFB71878FE09FF4F6AC3FF78" box="[476,491,152,176]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="6" pageNumber="91" unit="g" value="2.0">2</quantity>
</figureCitation>
g) with pregonite well developed, sub-triangular and with 2 setae postero-apically. Postgonite with a long (fine) seta at about mid length of anterior margin. Epiphallus well developed, strongly widened and sub circular distally in lateral view. Dorsolateral processes well fused medially in a single narrow sclerotization. Median process of ventral sclerite of distiphallus firmly fused to the base of ventral sclerite and divided longitudinally into two hemisclerites. Lateroventral lobe of distiphallus in sub-distal position and covered with slightly sclerotized scale-like spinulae. Acrophallus membranous.
</paragraph>
<caption id="7783D68AFFB4187BFF42F9706F05F935" pageId="5" pageNumber="90">
<paragraph id="23438602FFB4187BFF42F9706F05F935" blockId="5.[151,1436,1703,1789]" pageId="5" pageNumber="90">
<emphasis id="11885A10FFB4187BFF42F9706A3EF975" bold="true" box="[151,278,1703,1725]" pageId="5" pageNumber="90">
<figureCitation id="BBC79A87FFB4187BFF42F9706A39F975" box="[151,273,1703,1725]" captionStart="FIGURE 2" captionStartId="5.[151,250,1703,1725]" captionText="FIGURE 2. Shannoniella setinervis sp. nov., paratype male, details of morphology. a. Habitus, left lateral view. b. Head profile. c. Right wing, dorsal view, extension of setulae on vein R 4 + 5 indicated with red arrows. d. Abdomen, dorsal view. e. Sternite 5, ventral view. f. Cerci and surstyli, posterior view. g. Aedeagus, left lateral view." pageId="5" pageNumber="90">FIGURE 2</figureCitation>
.
</emphasis>
<taxonomicName id="E4FCFD81FFB4187BFEF5F9706939F975" box="[288,529,1703,1725]" class="Insecta" family="Rhinophoridae" genus="Shannoniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="setinervis" status="sp. nov.">
<emphasis id="11885A10FFB4187BFEF5F9706939F975" box="[288,529,1703,1725]" italics="true" pageId="5" pageNumber="90">Shannoniella setinervis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0ABBE76BFFB4187BFDCEF9706947F975" box="[539,623,1703,1725]" pageId="5" pageNumber="90" rank="species">
<emphasis id="11885A10FFB4187BFDCEF9706947F975" bold="true" box="[539,623,1703,1725]" pageId="5" pageNumber="90">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="FC4738A0FFB4187BFDAAF97F69FEF975" box="[639,726,1704,1725]" pageId="5" pageNumber="90" type="paratype">paratype</typeStatus>
male, details of morphology.
<emphasis id="11885A10FFB4187BFBC2F9706F02F975" bold="true" box="[1047,1066,1703,1725]" pageId="5" pageNumber="90">a.</emphasis>
Habitus, left lateral view.
<emphasis id="11885A10FFB4187BFA9DF9706E74F975" bold="true" box="[1352,1372,1703,1725]" pageId="5" pageNumber="90">b.</emphasis>
Head profile.
<emphasis id="11885A10FFB4187BFF3CF9116BD3F914" bold="true" box="[233,251,1734,1756]" pageId="5" pageNumber="90">c.</emphasis>
Right wing, dorsal view, extension of setulae on vein R4+5 indicated with red arrows.
<emphasis id="11885A10FFB4187BFBA2F9116FA4F914" bold="true" box="[1143,1164,1734,1756]" pageId="5" pageNumber="90">d.</emphasis>
Abdomen, dorsal view.
<emphasis id="11885A10FFB4187BFA5FF9116EB4F914" bold="true" box="[1418,1436,1734,1756]" pageId="5" pageNumber="90">e.</emphasis>
Sternite 5, ventral view.
<emphasis id="11885A10FFB4187BFE58F9306AB5F935" bold="true" box="[397,413,1767,1789]" pageId="5" pageNumber="90">f.</emphasis>
Cerci and surstyli, posterior view.
<emphasis id="11885A10FFB4187BFD2BF9306839F935" bold="true" box="[766,785,1767,1789]" pageId="5" pageNumber="90">g.</emphasis>
Aedeagus, left lateral view.
</paragraph>
</caption>
<paragraph id="23438602FFB71878FF12FEB86A8AFE40" blockId="6.[151,1437,151,464]" box="[199,418,367,392]" pageId="6" pageNumber="91">
<emphasis id="11885A10FFB71878FF12FEB86A0DFE40" bold="true" box="[199,293,367,392]" pageId="6" pageNumber="91">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="6BE6D589FFB71878FF12FE436918FE18" pageId="6" pageNumber="91" type="distribution">
<paragraph id="23438602FFB71878FF12FE436918FE18" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="91">
<emphasis id="11885A10FFB71878FF12FE436A48FE65" bold="true" box="[199,352,404,429]" pageId="6" pageNumber="91">Distribution.</emphasis>
The species appears to be restricted to the Brazilian Atlantic Forest (here recorded from the states of Rio de Janeiro and Paraná).
</paragraph>
</subSubSection>
</treatment>
</document>