594 lines
96 KiB
XML
594 lines
96 KiB
XML
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<mods:title id="69ABDB10438ABD76FA17200D787858AD">Contribution to the knowledge of Cloeodes Traver 1938 (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="8088FA5875E507AE2938CF8935047BF6">Kluge, Nikita J.</mods:namePart>
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<mods:title id="8EBC95A0399E5322CC78C4444E40098A">Zootaxa</mods:title>
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<mods:part id="D7D76B8F36A35E3FFEB0D1D34379C42E">
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<mods:date id="3D3D5B97DC24D2047D5716FF4804A45B">2017</mods:date>
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<mods:number id="92FF360F2AFB43BB9787C4A2FB512B91">2017-11-11</mods:number>
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<mods:number id="44F168E41C3C4A0E43281CA5D8B3B64B">4319</mods:number>
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<treatment id="4267878DFFBCFFC82687FC99FDC1FB0B" ID-DOI="http://doi.org/10.5281/zenodo.6042299" ID-GBIF-Taxon="134017032" ID-Zenodo-Dep="6042299" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B" httpUri="http://treatment.plazi.org/id/4267878DFFBCFFC82687FC99FDC1FB0B" lastPageId="30" lastPageNumber="121" pageId="23" pageNumber="114">
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<subSubSection id="82D46510FFBCFFC12687FC99FEB2FCB1" pageId="23" pageNumber="114" type="nomenclature">
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<paragraph id="CA71369BFFBCFFC12687FC99FE7EFC90" blockId="23.[151,505,836,895]" box="[151,505,836,862]" pageId="23" pageNumber="114">
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<heading id="913981F7FFBCFFC12687FC99FE7EFC90" bold="true" box="[151,505,836,862]" fontSize="11" level="1" pageId="23" pageNumber="114" reason="1">
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<emphasis id="F8BAEA89FFBCFFC12687FC99FE7EFC90" bold="true" box="[151,505,836,862]" pageId="23" pageNumber="114">
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<taxonomicName id="0DCE4D18FFBCFFC12687FC99FE28FC90" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[151,431,836,862]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="114" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
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<emphasis id="F8BAEA89FFBCFFC12687FC99FE28FC90" bold="true" box="[151,431,836,862]" italics="true" pageId="23" pageNumber="114">Cloeodes nigrohumeris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E38957F2FFBCFFC127A7FC99FE7EFC90" box="[439,505,836,862]" pageId="23" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="CA71369BFFBCFFC12687FCBBFEB2FCB1" blockId="23.[151,505,836,895]" box="[151,309,870,895]" pageId="23" pageNumber="114">
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(
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<figureCitation id="52F52A1EFFBCFFC1268FFCBBFEAAFCB1" box="[159,301,870,895]" captionStart-0="FIGURES 92 – 97" captionStart-1="FIGURES 98 – 103" captionStart-2="FIGURES 104 – 109" captionStart-3="FIGURES 110 – 115" captionStart-4="FIGURES 116 – 118" captionStart-5="FIGURES 119 – 132" captionStartId-0="25.[151,264,1837,1860]" captionStartId-1="26.[151,264,1937,1960]" captionStartId-2="27.[151,264,1666,1689]" captionStartId-3="28.[151,264,1961,1984]" captionStartId-4="29.[151,264,1368,1391]" captionStartId-5="31.[151,264,1977,2000]" captionTargetBox-0="[151,1434,347,1817]" captionTargetBox-1="[174,1418,193,1924]" captionTargetBox-2="[195,1394,195,1620]" captionTargetBox-3="[165,1429,193,1948]" captionTargetBox-4="[191,1395,214,1346]" captionTargetBox-5="[200,1368,511,1956]" captionTargetId-0="figure@25.[151,1436,347,1817]" captionTargetId-1="figure@26.[174,1418,193,1925]" captionTargetId-2="figure@27.[195,1400,183,1654]" captionTargetId-3="figure@28.[165,1429,193,1948]" captionTargetId-4="figure@29.[170,1417,193,1348]" captionTargetId-5="figure@31.[197,1404,507,1970]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionTargetPageId-4="29" captionTargetPageId-5="31" captionText-0="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." captionText-1="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." captionText-2="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." captionText-3="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." captionText-4="FIGURES 116 – 118. Cloeodes nigrohumeris sp. n. 116, subimaginal gonostyli extracted from mature larva; 117, exuviae of subimaginal genitals; 118, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines) (117 – 118, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." captionText-5="FIGURES 119 – 132. Cloeodes spp., tergalii. 119 – 125, tergalii I – VII of Cloeodes nigrohumeris sp. n.; 126 – 132, tergalii I – VII of Cloeodes auwe. For both individuals vertical line corresponds to length of abdominal segment IV." httpUri-0="https://zenodo.org/record/888653/files/figure.png" httpUri-1="https://zenodo.org/record/888655/files/figure.png" httpUri-2="https://zenodo.org/record/888657/files/figure.png" httpUri-3="https://zenodo.org/record/888659/files/figure.png" httpUri-4="https://zenodo.org/record/888661/files/figure.png" httpUri-5="https://zenodo.org/record/888663/files/figure.png" pageId="23" pageNumber="114">Figs 92–125</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="82D46510FFBCFFC12687FC73FCD5FC0A" box="[151,850,942,964]" pageId="23" pageNumber="114" type="description">
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<paragraph id="CA71369BFFBCFFC12687FC73FCD5FC0A" blockId="23.[151,850,942,964]" box="[151,850,942,964]" pageId="23" pageNumber="114">
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“Genus nr.
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<taxonomicName id="0DCE4D18FFBCFFC12719FC73FEF2FC0A" authority="Roback" authorityName="Traver" authorityPageNumber="133" authorityYear="1932" box="[265,373,942,964]" class="Insecta" family="Baetidae" genus="Neocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="114" phylum="Arthropoda" rank="genus">
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<emphasis id="F8BAEA89FFBCFFC12719FC73FEF2FC0A" box="[265,373,942,964]" italics="true" pageId="23" pageNumber="114">Neocloeon</emphasis>
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</taxonomicName>
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”:
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<bibRefCitation id="AE5F4B6AFFBCFFC1279EFC73FD9FFC0A" author="Roback" box="[398,536,942,964]" pageId="23" pageNumber="114" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI- Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, 14, 129 - 199." type="journal article" year="1966">Roback 1966</bibRefCitation>
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: 133, Pl. 3:
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<figureCitation id="52F52A1EFFBCFFC12480FC73FC86FC0A" box="[656,769,942,964]" captionStart="FIGURES 37 – 57" captionStartId="15.[151,264,1964,1987]" captionTargetBox="[185,1399,538,1939]" captionTargetId="figure@15.[170,1417,517,1951]" captionTargetPageId="15" captionText="FIGURES 37 – 57. Cloeodes spp., tergalii. 37 – 43, tergalii I – VII of Cloeodes vibratorius sp. n.; 44 – 57, tergalii I – VII of two individuals of Cloeodes redactus. For all three individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888639/files/figure.png" pageId="23" pageNumber="114">Figs 37–40</figureCitation>
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(larva).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="82D46510FFBCFFC12687FC2CFEB2FBE0" pageId="23" pageNumber="114" type="etymology">
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<paragraph id="CA71369BFFBCFFC12687FC2CFEB2FBE0" blockId="23.[151,1437,1009,2006]" pageId="23" pageNumber="114">
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<emphasis id="F8BAEA89FFBCFFC12687FC2CFE99FBC4" bold="true" box="[151,286,1009,1034]" pageId="23" pageNumber="114">Etymology.</emphasis>
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The adjective
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<taxonomicName id="0DCE4D18FFBCFFC127D5FC2CFDDCFBC7" authorityName="Kluge" authorityYear="2017" box="[453,603,1009,1033]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="114" phylum="Arthropoda" rank="species" species="nigrohumeris">
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<emphasis id="F8BAEA89FFBCFFC127D5FC2CFDDCFBC7" box="[453,603,1009,1033]" italics="true" pageId="23" pageNumber="114">nigrohumeris</emphasis>
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</taxonomicName>
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refers to pair of black lines on anterolateral parts of imaginal mesonotum (“shoulders”).
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</paragraph>
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</subSubSection>
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<subSubSection id="82D46510FFBCFFC126D7FBE4FDCCF914" pageId="23" pageNumber="114" type="materials_examined">
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||
<paragraph id="CA71369BFFBCFFC126D7FBE4FBCAFAE4" blockId="23.[151,1437,1009,2006]" pageId="23" pageNumber="114">
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<materialsCitation id="7AA63CC6FFBCFFC126D7FBE4FA9AFBB8" ID-GBIF-Occurrence="1632929429" collectingDate="2013-08-18" collectorName="N. Kluge & Sheyko" country="Peru" county="Material" latitude="-7.2" location="Quebrada Huanganaico" longLatPrecision="1303" longitude="-75.36667" municipality="Pampa Hermosa" pageId="23" pageNumber="114" specimenCount="1" specimenCount-male="1" stateProvince="Region Loreto" typeStatus="holotype">
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<emphasis id="F8BAEA89FFBCFFC126D7FBE4FE36FB9C" bold="true" box="[199,433,1081,1106]" pageId="23" pageNumber="114">
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<collectingCounty id="23104E17FFBCFFC126D7FBE4FEA8FB9C" box="[199,303,1081,1106]" pageId="23" pageNumber="114">Material</collectingCounty>
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examined.
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</emphasis>
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<typeStatus id="15758839FFBCFFC127A9FBE4FDA1FB9C" box="[441,550,1081,1106]" pageId="23" pageNumber="114" type="holotype">Holotype</typeStatus>
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: L-S-I
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<specimenCount id="DCC8FD12FFBCFFC12460FBE7FD04FB9C" box="[624,643,1082,1106]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
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{specimen [XIX](3)2013},
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<collectingCountry id="B2D9760BFFBCFFC125D2FBE7FB8BFB9F" box="[962,1036,1082,1105]" name="Peru" pageId="23" pageNumber="114">PERU</collectingCountry>
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,
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<collectingRegion id="080AF879FFBCFFC12209FBE4FB39FB9F" box="[1049,1214,1081,1106]" country="Peru" name="Loreto" pageId="23" pageNumber="114">Region Loreto</collectingRegion>
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, Provincia
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<collectingRegion id="080AF879FFBCFFC1232EFBE4FA1FFB9C" box="[1342,1432,1081,1106]" country="Peru" name="Ucayali" pageId="23" pageNumber="114">Ucayali</collectingRegion>
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,
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<collectingMunicipality id="2A15ACE1FFBCFFC12687FB83FED4FBBB" box="[151,339,1118,1141]" pageId="23" pageNumber="114">Pampa Hermosa</collectingMunicipality>
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,
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<location id="CF116040FFBCFFC1274FFB80FDECFBB8" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B:CF116040FFBCFFC1274FFB80FDECFBB8" box="[351,619,1117,1142]" country="Peru" county="Material" latitude="-7.2" longLatPrecision="1303" longitude="-75.36667" municipality="Pampa Hermosa" name="Quebrada Huanganaico" pageId="23" pageNumber="114" stateProvince="Region Loreto">Quebrada Huanganaico</location>
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,
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<geoCoordinate id="AFFA505CFFBCFFC12466FB83FD45FBB8" box="[630,706,1118,1142]" direction="south" orientation="latitude" pageId="23" pageNumber="114" precision="925" value="-7.2">7°12'S</geoCoordinate>
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,
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<geoCoordinate id="AFFA505CFFBCFFC124DFFB80FCB5FBB8" box="[719,818,1117,1142]" direction="west" orientation="longitude" pageId="23" pageNumber="114" precision="925" value="-75.36667">75°22'W</geoCoordinate>
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,
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<date id="BE70105BFFBCFFC1252EFB83FC54FBB8" box="[830,979,1118,1142]" pageId="23" pageNumber="114" value="2013-08-18">
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||
<collectingDate id="AE34E9B3FFBCFFC1252EFB83FC54FBB8" box="[830,979,1118,1142]" pageId="23" pageNumber="114" value="2013-08-18">18.VIII.2013</collectingDate>
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</date>
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, coll.
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<collectorName id="673B534DFFBCFFC12207FB83FBF8FBB8" box="[1047,1151,1117,1142]" pageId="23" pageNumber="114">N. Kluge</collectorName>
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& L.
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<collectorName id="673B534DFFBCFFC122D5FB80FA9AFBB8" box="[1221,1309,1117,1142]" pageId="23" pageNumber="114">Sheyko</collectorName>
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="7AA63CC6FFBCFFC12337FB83FBA0FB2C" ID-GBIF-Occurrence="1632929434" collectingDate="2013-07-26" collectingDateMax="2013-08-20" collectingDateMin="2013-07-26" collectorName="N. Kluge & Sheyko" country="Peru" county="Material" latitude="-7.1833334" location="Aguas Termales" longLatPrecision="1303" longitude="-74.95" municipality="Pampa Hermosa" pageId="23" pageNumber="114" specimenCount="19" specimenCount-female="6" specimenCount-larva="7" specimenCount-male="6" stateProvince="Ucayali" typeStatus="paratype">
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||
<typeStatus id="15758839FFBCFFC12337FB83FA1FFBB8" box="[1319,1432,1118,1142]" pageId="23" pageNumber="114" type="paratype">Paratypes</typeStatus>
|
||
: the same locality and collectors,
|
||
<collectingDate id="AE34E9B3FFBCFFC12418FB5FFD4EFB54" box="[520,713,1154,1178]" pageId="23" pageNumber="114" value="2013-08-18" valueMax="2013-08-20" valueMin="2013-08-18">
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||
<date id="BE70105BFFBCFFC12418FB5FFD4EFB54" box="[520,713,1154,1178]" pageId="23" pageNumber="114" value="2013-08-18" valueMax="2013-08-20" valueMin="2013-08-18">18–20.VIII.2013</date>
|
||
</collectingDate>
|
||
: 5 L-S-I
|
||
<specimenCount id="DCC8FD12FFBCFFC1253AFB5FFCBAFB54" box="[810,829,1154,1178]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
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||
, 1 L-S
|
||
<specimenCount id="DCC8FD12FFBCFFC1259CFB5FFC27FB54" box="[908,928,1154,1178]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
|
||
, 9 L-S-I
|
||
<specimenCount id="DCC8FD12FFBCFFC12217FB5FFB92FB54" box="[1031,1045,1154,1178]" pageId="23" pageNumber="114" type="female">♀</specimenCount>
|
||
, 3 L-S
|
||
<specimenCount id="DCC8FD12FFBCFFC12279FB5FFBFFFB54" box="[1129,1144,1154,1178]" pageId="23" pageNumber="114" type="female">♀</specimenCount>
|
||
, 3 L/S
|
||
<specimenCount id="DCC8FD12FFBCFFC122D6FB5FFB5DFB54" box="[1222,1242,1154,1178]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
|
||
, 2 L/S
|
||
<specimenCount id="DCC8FD12FFBCFFC1233CFB5FFABBFB54" box="[1324,1340,1154,1178]" pageId="23" pageNumber="114" type="female">♀</specimenCount>
|
||
, 1 larva. Provincia
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||
<collectingRegion id="080AF879FFBCFFC12719FB78FEE4FB70" box="[265,355,1189,1214]" country="Peru" name="Ucayali" pageId="23" pageNumber="114">Ucayali</collectingRegion>
|
||
,
|
||
<locationDeviation id="DBC32FB9FFBCFFC1277EFB7BFDF8FB70" box="[366,639,1189,1214]" pageId="23" pageNumber="114">
|
||
<quantity id="0D369B7EFFBCFFC1277EFB7BFE31FB70" box="[366,438,1189,1214]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="114" unit="km" value="17.0">17 km</quantity>
|
||
NNE Contamana
|
||
</locationDeviation>
|
||
,
|
||
<location id="CF116040FFBCFFC1249BFB7BFCC5FB70" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B:CF116040FFBCFFC1249BFB7BFCC5FB70" box="[651,834,1189,1214]" latitude="-7.1833334" longLatPrecision="1303" longitude="-74.95" name="Aguas Termales" pageId="23" pageNumber="114" stateProvince="Ucayali">Aguas Termales</location>
|
||
,
|
||
<geoCoordinate id="AFFA505CFFBCFFC1255EFB7BFC1FFB70" box="[846,920,1190,1214]" direction="south" orientation="latitude" pageId="23" pageNumber="114" precision="925" value="-7.1833334">7°11'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="AFFA505CFFBCFFC125B5FB78FB8FFB70" box="[933,1032,1189,1214]" direction="west" orientation="longitude" pageId="23" pageNumber="114" precision="925" value="-74.95">74°57'W</geoCoordinate>
|
||
,
|
||
<collectingDate id="AE34E9B3FFBCFFC12203FB78FB76FB70" box="[1043,1265,1189,1214]" pageId="23" pageNumber="114" value="2013-07-26" valueMax="2013-08-04" valueMin="2013-07-26">
|
||
<date id="BE70105BFFBCFFC12203FB78FB76FB70" box="[1043,1265,1189,1214]" pageId="23" pageNumber="114" value="2013-07-26" valueMax="2013-08-04" valueMin="2013-07-26">26.VII–4.VIII.2013</date>
|
||
</collectingDate>
|
||
, coll.
|
||
<collectorName id="673B534DFFBCFFC12325FB7BFA1BFB70" box="[1333,1436,1189,1214]" pageId="23" pageNumber="114">N. Kluge</collectorName>
|
||
& L.
|
||
<collectorName id="673B534DFFBCFFC126C5FB14FEAAFB2C" box="[213,301,1225,1250]" pageId="23" pageNumber="114">Sheyko</collectorName>
|
||
: 3 L-S-I
|
||
<specimenCount id="DCC8FD12FFBCFFC12781FB17FE23FB2C" box="[401,420,1226,1250]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
|
||
, 1 L-S/I
|
||
<specimenCount id="DCC8FD12FFBCFFC12416FB17FD9DFB2C" box="[518,538,1226,1250]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
|
||
, 2 L-S-I
|
||
<specimenCount id="DCC8FD12FFBCFFC12492FB17FD17FB2C" box="[642,656,1226,1250]" pageId="23" pageNumber="114" type="female">♀</specimenCount>
|
||
, 1 L-S
|
||
<specimenCount id="DCC8FD12FFBCFFC124F6FB17FD72FB2C" box="[742,757,1226,1250]" pageId="23" pageNumber="114" type="female">♀</specimenCount>
|
||
, 3 L/S
|
||
<specimenCount id="DCC8FD12FFBCFFC12554FB17FCDFFB2C" box="[836,856,1226,1250]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
|
||
, 1 L/S
|
||
<specimenCount id="DCC8FD12FFBCFFC125BCFB17FC3BFB2C" box="[940,956,1226,1250]" pageId="23" pageNumber="114" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="DCC8FD12FFBCFFC125D8FB17FBA0FB2C" box="[968,1063,1225,1250]" pageId="23" pageNumber="114" type="larva">7 larvae</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="7AA63CC6FFBCFFC12222FB14FBCEFAE4" ID-GBIF-Occurrence="1632929439" collectingDate="2006-02-01" collectingDateMax="2006-02-16" collectingDateMin="2006-02-01" collectorName="Quebrada El Sabalo & N. Kluge" country="Peru" county="Provincia Maynas" latitude="-4.266667" location="Rio Itaya" longLatPrecision="1306" longitude="-73.45" municipality="Quebrada El Sabalo" pageId="23" pageNumber="114" specimenCount="36" specimenCount-female="1" specimenCount-larva="33" specimenCount-male="2" stateProvince="Ucayali" typeStatus="paratype">
|
||
<collectingCounty id="23104E17FFBCFFC12222FB14FA85FB2C" box="[1074,1282,1225,1250]" pageId="23" pageNumber="114">Provincia Maynas</collectingCounty>
|
||
,
|
||
<collectorName id="673B534DFFBCFFC1231EFB14FF64FAC8" pageId="23" pageNumber="114">
|
||
<collectingMunicipality id="2A15ACE1FFBCFFC1231EFB14FF64FAC8" pageId="23" pageNumber="114">Quebrada El Sabalo</collectingMunicipality>
|
||
</collectorName>
|
||
(right tributary of
|
||
<location id="CF116040FFBCFFC127A6FB30FDA7FAC8" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B:CF116040FFBCFFC127A6FB30FDA7FAC8" box="[438,544,1261,1286]" country="Peru" county="Provincia Maynas" latitude="-4.266667" longLatPrecision="1306" longitude="-73.45" municipality="Quebrada El Sabalo" name="Rio Itaya" pageId="23" pageNumber="114" stateProvince="Ucayali">Rio Itaya</location>
|
||
) between
|
||
<location id="CF116040FFBCFFC12483FB33FC99FAC8" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B:CF116040FFBCFFC12483FB33FC99FAC8" box="[659,798,1262,1286]" country="Peru" county="Provincia Maynas" latitude="-4.266667" longLatPrecision="1306" longitude="-73.45" municipality="Quebrada El Sabalo" name="Puente Itaya" pageId="23" pageNumber="114" stateProvince="Ucayali">Puente Itaya</location>
|
||
(
|
||
<quantity id="0D369B7EFFBCFFC1253EFB30FCF1FAC8" box="[814,886,1261,1286]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="23" pageNumber="114" unit="km" value="57.0">57 km</quantity>
|
||
from
|
||
<location id="CF116040FFBCFFC125ABFB30FB8DFAC8" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B:CF116040FFBCFFC125ABFB30FB8DFAC8" box="[955,1034,1261,1286]" country="Peru" county="Provincia Maynas" latitude="-4.266667" longLatPrecision="1306" longitude="-73.45" municipality="Quebrada El Sabalo" name="Iquitos" pageId="23" pageNumber="114" stateProvince="Ucayali">Iquitos</location>
|
||
) and
|
||
<location id="CF116040FFBCFFC12259FB33FB56FAC8" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B:CF116040FFBCFFC12259FB33FB56FAC8" box="[1097,1233,1261,1286]" country="Peru" county="Provincia Maynas" latitude="-4.266667" longLatPrecision="1306" longitude="-73.45" municipality="Quebrada El Sabalo" name="San Joaquin" pageId="23" pageNumber="114" stateProvince="Ucayali">San Joaquin</location>
|
||
(at
|
||
<location id="CF116040FFBCFFC122EDFB30FA17FAC8" LSID="urn:lsid:plazi:treatment:4267878DFFBCFFC82687FC99FDC1FB0B:CF116040FFBCFFC122EDFB30FA17FAC8" box="[1277,1424,1261,1286]" country="Peru" county="Provincia Maynas" latitude="-4.266667" longLatPrecision="1306" longitude="-73.45" municipality="Quebrada El Sabalo" name="Rio Amazon" pageId="23" pageNumber="114" stateProvince="Ucayali">Rio Amazon</location>
|
||
),
|
||
<geoCoordinate id="AFFA505CFFBCFFC12687FACCFF64FAE4" box="[151,227,1297,1322]" direction="south" orientation="latitude" pageId="23" pageNumber="114" precision="925" value="-4.266667">4°16'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="AFFA505CFFBCFFC126FFFACFFED6FAE4" box="[239,337,1298,1322]" direction="west" orientation="longitude" pageId="23" pageNumber="114" precision="925" value="-73.45">73°27'W</geoCoordinate>
|
||
,
|
||
<collectingDate id="AE34E9B3FFBCFFC1274DFACCFE68FAE4" box="[349,495,1297,1322]" pageId="23" pageNumber="114" value="2006-02-01" valueMax="2006-02-16" valueMin="2006-02-01">
|
||
<date id="BE70105BFFBCFFC1274DFACCFE68FAE4" box="[349,495,1297,1322]" pageId="23" pageNumber="114" value="2006-02-01" valueMax="2006-02-16" valueMin="2006-02-01">1–16.II.2006</date>
|
||
</collectingDate>
|
||
, coll.
|
||
<collectorName id="673B534DFFBCFFC12423FACFFD18FAE4" box="[563,671,1297,1322]" pageId="23" pageNumber="114">N. Kluge</collectorName>
|
||
: 1 L-S-I
|
||
<specimenCount id="DCC8FD12FFBCFFC124EFFACFFC95FAE4" box="[767,786,1298,1322]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
|
||
, 7 L/S
|
||
<specimenCount id="DCC8FD12FFBCFFC1254EFACFFCF5FAE4" box="[862,882,1298,1322]" pageId="23" pageNumber="114" type="male">♂</specimenCount>
|
||
, 5 L/S
|
||
<specimenCount id="DCC8FD12FFBCFFC125D3FACFFC54FAE4" box="[963,979,1298,1322]" pageId="23" pageNumber="114" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="DCC8FD12FFBCFFC125CEFACFFBCEFAE4" box="[990,1097,1297,1322]" pageId="23" pageNumber="114" type="larva">33 larvae</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFBCFFC126D7FAE8FDCCF914" blockId="23.[151,1437,1009,2006]" pageId="23" pageNumber="114">
|
||
<emphasis id="F8BAEA89FFBCFFC126D7FAE8FE32FA80" bold="true" box="[199,437,1333,1358]" pageId="23" pageNumber="114">
|
||
Descriptions.
|
||
<emphasis id="F8BAEA89FFBCFFC1277AFAEBFE29FA83" bold="true" box="[362,430,1334,1357]" italics="true" pageId="23" pageNumber="114">Larva</emphasis>
|
||
.
|
||
</emphasis>
|
||
CUTICULAR COLORATION: Cuticle partly colorless, partly light brownish. Most part of head colorless. Pronotum and/or mesonotum either with colorless, light brownish and darker brow areas (
|
||
<figureCitation id="52F52A1EFFBCFFC1237AFA84FF31FA58" captionStart="FIGURES 92 – 97" captionStartId="25.[151,264,1837,1860]" captionTargetBox="[151,1434,347,1817]" captionTargetId="figure@25.[151,1436,347,1817]" captionTargetPageId="25" captionText="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="23" pageNumber="114">Figs 94</figureCitation>
|
||
,
|
||
<figureCitation id="52F52A1EFFBCFFC126D3FAA3FF77FA58" box="[195,240,1406,1430]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="23" pageNumber="114">103</figureCitation>
|
||
), or at most part colorless with few small brown markings. Mesothorax and metathorax with contrasting brown macula adjacent to dorsal coxal articulation; in some individuals this macula occupies most part of episternum on metathorax and smaller part of episternum on mesothorax; remainder cuticle of thoracic pleura and sterna colorless (
|
||
<figureCitation id="52F52A1EFFBCFFC12742FA34FE33F9CC" box="[338,436,1513,1538]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="23" pageNumber="114">Fig. 102</figureCitation>
|
||
). Legs light with or without diffusive darker macula at middle of femur, knee articulation, tibia just distad of patella-tibial suture and/or base of tarsus (
|
||
<figureCitation id="52F52A1EFFBCFFC12544F9D0FC3EF9E8" box="[852,953,1549,1574]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="23" pageNumber="114">Fig. 102</figureCitation>
|
||
). Abdominal terga at most light, with few contrasting brown maculae (
|
||
<figureCitation id="52F52A1EFFBCFFC127CAF9ECFDB5F984" box="[474,562,1585,1610]" captionStart="FIGURES 92 – 97" captionStartId="25.[151,264,1837,1860]" captionTargetBox="[151,1434,347,1817]" captionTargetId="figure@25.[151,1436,347,1817]" captionTargetPageId="25" captionText="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="23" pageNumber="114">Figs 96</figureCitation>
|
||
,
|
||
<figureCitation id="52F52A1EFFBCFFC1242FF9EFFD21F984" box="[575,678,1586,1610]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="23" pageNumber="114">100–101</figureCitation>
|
||
): tergum I at most brown; tergum II with anterior-median part and pair of submedian myo-sigilla brown; terga III–V at most colorless, tergum VI at most brown, tergum VII with unpaired medio-anterior brown macula, tergum VIII colorless, tergum IX often with lateral parts brown, tergum X light with brownish maculae. Abdominal sterna and paraprocts at most colorless. Caudalii light, often with brown band near apex (
|
||
<figureCitation id="52F52A1EFFBCFFC12740F91CFE4AF914" box="[336,461,1729,1754]" captionStart="FIGURES 92 – 97" captionStartId="25.[151,264,1837,1860]" captionTargetBox="[151,1434,347,1817]" captionTargetId="figure@25.[151,1436,347,1817]" captionTargetPageId="25" captionText="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="23" pageNumber="114">Fig. 92–93</figureCitation>
|
||
,
|
||
<figureCitation id="52F52A1EFFBCFFC127C8F91FFDB8F914" box="[472,575,1730,1754]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="23" pageNumber="114">100–101</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82D46510FFBCFFC826D7F93BFC21FC92" lastPageId="30" lastPageNumber="121" pageId="23" pageNumber="114" type="description">
|
||
<paragraph id="CA71369BFFBCFFC126D7F93BFEDFF8A4" blockId="23.[151,1437,1009,2006]" pageId="23" pageNumber="114">
|
||
HYPODERMAL COLORATION: Abdominal terga of mature and immature larvae of both sexes with contrasting reddish or brown maculae well visible through cuticle: terga II–VII with median macula and pair of transverse stripes on posterior margin; terga II, V and VI, beside this, with pair of lateral maculae (as in imago) (
|
||
<figureCitation id="52F52A1EFFBCFFC1268FF88CFECCF8A4" box="[159,331,1873,1898]" captionStart="FIGURES 92 – 97" captionStartId="25.[151,264,1837,1860]" captionTargetBox="[151,1434,347,1817]" captionTargetId="figure@25.[151,1436,347,1817]" captionTargetPageId="25" captionText="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="23" pageNumber="114">Figs 92, 93, 95</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFBCFFCE26D7F8ABFACBFDD6" blockId="23.[151,1437,1009,2006]" lastBlockId="24.[151,1437,151,933]" lastPageId="24" lastPageNumber="115" pageId="23" pageNumber="114">
|
||
SHAPE AND SETATION: Frontal suture pointed, rectangular; face relatively long; in female larva eyes brought together (as in
|
||
<figureCitation id="52F52A1EFFBCFFC1278EF844FE64F87C" box="[414,483,1945,1970]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1969,1992]" captionTargetBox="[191,1397,203,1957]" captionTargetId="figure@7.[191,1407,193,1957]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Cloeodes vibratorius sp. n., larva. 6, head of female larva; 7, maxilla; 8, apex of maxilla; 9 – 11, left and right mandibles; 12, labrum (apical setae not shown) (10, 12, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888629/files/figure.png" pageId="23" pageNumber="114">Fig. 6</figureCitation>
|
||
). Labrum equally wide at base and at middle, with pair of submedian long setae and one pair of sublateral long setae (
|
||
<figureCitation id="52F52A1EFFBCFFC12437F860FD09F818" box="[551,654,1981,2006]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="23" pageNumber="114">Fig. 106</figureCitation>
|
||
). Prostheca of left mandible with 3 blunt processes and 3 pointed processes; median margin of left mandible proximad of prostheca straight or slightly convex, smooth (
|
||
<figureCitation id="52F52A1EFFB3FFCE233CFF4AFA17FF7E" box="[1324,1424,151,176]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="24" pageNumber="115">Fig. 104</figureCitation>
|
||
). Prostheca of right mandible directed medially-proximally, bifurcate, with longest branch directed proximally; median margin of right mandible proximad of prostheca convex or nearly straight, smooth or with serration, without seta-like processes (
|
||
<figureCitation id="52F52A1EFFB3FFCE27FAFED9FDD3FED3" box="[490,596,260,285]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="24" pageNumber="115">Fig. 105</figureCitation>
|
||
). Stipes of maxilla proximally with 4–5 setae on ventro-medial side. Maxillary canines and distal dentiseta similar, all hooked medially (
|
||
<figureCitation id="52F52A1EFFB3FFCE2588FEFAFC7CFE8E" box="[920,1019,295,320]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="24" pageNumber="115">Fig. 109</figureCitation>
|
||
). Maxillary palp 2-segmented, either reaching apex of lacinia, or slightly shorter, or slightly longer (
|
||
<bibRefCitation id="AE5F4B6AFFB3FFCE2579FE91FBE0FEAA" author="Roback" box="[873,1127,332,357]" pageId="24" pageNumber="115" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI- Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, 14, 129 - 199." type="journal article" year="1966">
|
||
Roback 1966:
|
||
<figureCitation id="52F52A1EFFB3FFCE2202FE91FBE0FEAA" box="[1042,1127,332,357]" captionStart="FIGURES 37 – 57" captionStartId="15.[151,264,1964,1987]" captionTargetBox="[185,1399,538,1939]" captionTargetId="figure@15.[170,1417,517,1951]" captionTargetPageId="15" captionText="FIGURES 37 – 57. Cloeodes spp., tergalii. 37 – 43, tergalii I – VII of Cloeodes vibratorius sp. n.; 44 – 57, tergalii I – VII of two individuals of Cloeodes redactus. For all three individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888639/files/figure.png" pageId="24" pageNumber="115">Fig. 38</figureCitation>
|
||
</bibRefCitation>
|
||
). Labium with glossae and paraglossae subequal, both narrowed apically (
|
||
<figureCitation id="52F52A1EFFB3FFCE24BFFEB2FC94FE46" box="[687,787,367,392]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="24" pageNumber="115">Fig. 108</figureCitation>
|
||
). Glossa dorsally with 1–2 stout pointed setae near apex and with 10–12 setae in dorso-lateral row; ventrally with irregularly arranged setae in proximal part and 5–7 setae forming ventro-median row. Paraglossa with latero-apical setae forming one regular row and few (2–4) setae just dorsad of it; with 4–5 setae in ventro-median row; with 4 setae in dorso-median row. Distal segment of labial palp widened apically, with median margin slightly shorter than lateral margin (
|
||
<figureCitation id="52F52A1EFFB3FFCE25CCFE22FBBAFDD6" box="[988,1085,511,536]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="24" pageNumber="115">Fig. 108</figureCitation>
|
||
;
|
||
<bibRefCitation id="AE5F4B6AFFB3FFCE2259FE22FB66FDD6" author="Roback" box="[1097,1249,511,536]" pageId="24" pageNumber="115" refString="Roback, S. S. (1966) The Catherwood Foundation Peruvia-Amazon expedition. VI- Ephemeroptera nymphs. Monographs of the Academy of Natural Sciences of Philadelphia, 14, 129 - 199." type="journal article" year="1966">Roback 1966</bibRefCitation>
|
||
:
|
||
<figureCitation id="52F52A1EFFB3FFCE22FCFE22FAC7FDD6" box="[1260,1344,511,536]" captionStart="FIGURES 37 – 57" captionStartId="15.[151,264,1964,1987]" captionTargetBox="[185,1399,538,1939]" captionTargetId="figure@15.[170,1417,517,1951]" captionTargetPageId="15" captionText="FIGURES 37 – 57. Cloeodes spp., tergalii. 37 – 43, tergalii I – VII of Cloeodes vibratorius sp. n.; 44 – 57, tergalii I – VII of two individuals of Cloeodes redactus. For all three individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888639/files/figure.png" pageId="24" pageNumber="115">Fig. 40</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB3FFCE26D7FDF9FC63FC6A" blockId="24.[151,1437,151,933]" pageId="24" pageNumber="115">
|
||
All thoracic terga without protuberances. Metanotum without vestiges of hind protoptera (
|
||
<figureCitation id="52F52A1EFFB3FFCE22AEFDF9FAA7FDF2" box="[1214,1312,548,573]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="24" pageNumber="115">Fig. 102</figureCitation>
|
||
). Fore legs largest, middle legs smaller and hind legs smallest; on fore leg tarsus much longer than tibia, on middle and hind legs tarsus subequal to tibia; claws elongated, much longer than half of tarsal length (
|
||
<tableCitation id="874C0320FFB3FFCE2262FDB1FB4DFD4B" box="[1138,1226,620,645]" captionStart="TABLE 5" captionStartId="24.[151,239,1272,1295]" captionText="TABLE 5. Leg proportions of male C. nigrohumeris sp. n. measured on outer side of leg of paratype No. [XXI] (4) 2013." pageId="24" pageNumber="115">Table 5</tableCitation>
|
||
;
|
||
<figureCitation id="52F52A1EFFB3FFCE22C9FDB1FABAFD4A" box="[1241,1341,620,645]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="24" pageNumber="115">Fig. 102</figureCitation>
|
||
). Femur parallel-sided, with outer margin straight and inner margin slightly convex; apical projection equally short on femora of all leg pairs. Outer side of femur with 5–7 minute stout setae; apical projection with 2 larger spatulate setae. Inner and dorsal side of femur with few stout setae, which are subequal or smaller than setae on dorsal side. Patella-tibial suture and proximal arc of long setae with contiguous sockets equally developed on tibiae of all leg pairs; inner margin of tibia with irregular small stout pointed setae, apically with 2 larger setae (as in
|
||
<figureCitation id="52F52A1EFFB3FFCE2329FCC2FA08FCF6" box="[1337,1423,799,824]" captionStart="FIGURES 13 – 17" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[191,1395,198,1904]" captionTargetId="figure@8.[191,1395,193,1908]" captionTargetPageId="8" captionText="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri="https://zenodo.org/record/888631/files/figure.png" pageId="24" pageNumber="115">Fig. 16</figureCitation>
|
||
). Outer-apical seta blunt, reaching apex of tibia. Dorsal side of each tarsus in proximal half with long fine setae forming 2 longitudinal rows. Claw longer than 1/2 of tarsus length (
|
||
<figureCitation id="52F52A1EFFB3FFCE259AFCBAFC6BFC4E" box="[906,1004,871,896]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="24" pageNumber="115">Fig. 102</figureCitation>
|
||
). Scales on abdominal terga and sterna numerous, short, triangular or semicircular, colorless and delicate (
|
||
<figureCitation id="52F52A1EFFB3FFCE2593FC51FC50FC6A" box="[899,983,908,933]" captionStart="FIGURES 92 – 97" captionStartId="25.[151,264,1837,1860]" captionTargetBox="[151,1434,347,1817]" captionTargetId="figure@25.[151,1436,347,1817]" captionTargetPageId="25" captionText="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="24" pageNumber="115">Fig. 97</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="9EB16613FFB3FFCE2687FC09FBA1FC25" box="[151,1062,979,1003]" pageId="24" pageNumber="115">
|
||
<paragraph id="CA71369BFFB3FFCE2687FC09FBA1FC25" blockId="24.[151,1062,979,1003]" box="[151,1062,979,1003]" pageId="24" pageNumber="115">
|
||
<emphasis id="F8BAEA89FFB3FFCE2687FC09FE8EFC25" bold="true" box="[151,265,980,1003]" pageId="24" pageNumber="115">TABLE 4.</emphasis>
|
||
Leg proportions of male
|
||
<taxonomicName id="0DCE4D18FFB3FFCE240AFC0EFD59FC25" box="[538,734,979,1003]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="115" phylum="Arthropoda" rank="species" species="redactus">
|
||
<emphasis id="F8BAEA89FFB3FFCE240AFC0EFD59FC25" box="[538,734,979,1003]" italics="true" pageId="24" pageNumber="115">Cloeodes redactus</emphasis>
|
||
</taxonomicName>
|
||
measured on outer side of leg.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CA71369BFFB3FFCE268FFBDAFB56FB79" pageId="24" pageNumber="115">
|
||
<table id="B8CEC43BFFB30029268FFBDBFA8BFB79" box="[159,1292,1030,1207]" gridcols="4" gridrows="4" pageId="24" pageNumber="115" rowsContinueIn="24.[159,1293,1322,1499]">
|
||
<tr id="74FE34D9FFB30029268FFBDBFA8BFBF5" box="[159,1292,1030,1083]" gridrow="0" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFBDBFF62FBF5" box="[159,229,1030,1083]" gridcol="0" gridrow="0" pageId="24" pageNumber="115">Leg:</th>
|
||
<th id="372F5DA5FFB300292722FBDBFD9CFBF5" box="[306,539,1030,1083]" gridcol="1" gridrow="0" pageId="24" pageNumber="115">Larva ♂ femur/tibia/tarsus/claw</th>
|
||
<th id="372F5DA5FFB30029248BFBDBFC07FBF5" box="[667,896,1030,1083]" gridcol="2" gridrow="0" pageId="24" pageNumber="115">Subimago ♂ femur/tibia/tarsomeres</th>
|
||
<th id="372F5DA5FFB300292207FBDBFA8BFBF5" box="[1047,1292,1030,1083]" gridcol="3" gridrow="0" pageId="24" pageNumber="115">Imago ♂ femur/tibia/tarsomeres</th>
|
||
</tr>
|
||
<tr id="74FE34D9FFB30029268FFB93FA8BFBAA" box="[159,1292,1102,1124]" gridrow="1" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFB93FF62FBAA" box="[159,229,1102,1124]" gridcol="0" gridrow="1" pageId="24" pageNumber="115">fore</th>
|
||
<td id="372F5DA5FFB300292722FB93FD9CFBAA" box="[306,539,1102,1124]" gridcol="1" gridrow="1" pageId="24" pageNumber="115">64: 40: 50: 13</td>
|
||
<td id="372F5DA5FFB30029248BFB93FC07FBAA" box="[667,896,1102,1124]" gridcol="2" gridrow="1" pageId="24" pageNumber="115">64: 63: 4:24:18:9:13</td>
|
||
<td id="372F5DA5FFB300292207FB93FA8BFBAA" box="[1047,1292,1102,1124]" gridcol="3" gridrow="1" pageId="24" pageNumber="115">83: 120: 5:57:36:17:13</td>
|
||
</tr>
|
||
<tr id="74FE34D9FFB30029268FFBA5FA8BFB40" box="[159,1292,1144,1166]" gridrow="2" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFBA5FF62FB40" box="[159,229,1144,1166]" gridcol="0" gridrow="2" pageId="24" pageNumber="115">middle</th>
|
||
<td id="372F5DA5FFB300292722FBA5FD9CFB40" box="[306,539,1144,1166]" gridcol="1" gridrow="2" pageId="24" pageNumber="115">61: 36: 35: 13</td>
|
||
<td id="372F5DA5FFB30029248BFBA5FC07FB40" box="[667,896,1144,1166]" gridcol="2" gridrow="2" pageId="24" pageNumber="115">60: 60: 18:7:2:11</td>
|
||
<td id="372F5DA5FFB300292207FBA5FA8BFB40" box="[1047,1292,1144,1166]" gridcol="3" gridrow="2" pageId="24" pageNumber="115">60: 60: 18:7:2:11</td>
|
||
</tr>
|
||
<tr id="74FE34D9FFB30029268FFB7CFA8BFB79" box="[159,1292,1185,1207]" gridrow="3" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFB7CFF62FB79" box="[159,229,1185,1207]" gridcol="0" gridrow="3" pageId="24" pageNumber="115">hind</th>
|
||
<td id="372F5DA5FFB300292722FB7CFD9CFB79" box="[306,539,1185,1207]" gridcol="1" gridrow="3" pageId="24" pageNumber="115">58: 30: 33: 13</td>
|
||
<td id="372F5DA5FFB30029248BFB7CFC07FB79" box="[667,896,1185,1207]" gridcol="2" gridrow="3" pageId="24" pageNumber="115">60: 60: 18:7:2:11</td>
|
||
<td id="372F5DA5FFB300292207FB7CFA8BFB79" box="[1047,1292,1185,1207]" gridcol="3" gridrow="3" pageId="24" pageNumber="115">60: 60: 18:7:2:11</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="9EB16613FFB3FFCE2687FB25FA1BFAC1" box="[151,1436,1271,1295]" pageId="24" pageNumber="115">
|
||
<paragraph id="CA71369BFFB3FFCE2687FB25FA1BFAC1" blockId="24.[151,1436,1271,1295]" box="[151,1436,1271,1295]" pageId="24" pageNumber="115">
|
||
<emphasis id="F8BAEA89FFB3FFCE2687FB25FE8DFAC1" bold="true" box="[151,266,1272,1295]" pageId="24" pageNumber="115">TABLE 5.</emphasis>
|
||
Leg proportions of male
|
||
<taxonomicName id="0DCE4D18FFB3FFCE2406FB25FD45FAC1" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[534,706,1271,1295]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="115" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB3FFCE2406FB25FD45FAC1" box="[534,706,1271,1295]" italics="true" pageId="24" pageNumber="115">C. nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB3FFCE24D9FB25FC83FAC1" box="[713,772,1272,1295]" pageId="24" pageNumber="115" rank="species">
|
||
<emphasis id="F8BAEA89FFB3FFCE24D9FB25FC83FAC1" bold="true" box="[713,772,1272,1295]" pageId="24" pageNumber="115">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
measured on outer side of leg of paratype No. [XXI] (4) 2013.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CA71369BFFB3FFCE268FFAF6FB53FA15" pageId="24" pageNumber="115">
|
||
<table id="B8CEC43BFFB30029268FFAF7FA8AFA15" box="[159,1293,1322,1499]" gridcols="4" gridrows="4" pageId="24" pageNumber="115" rowsContinueFrom="24.[159,1292,1030,1207]">
|
||
<tr id="74FE34D9FFB30029268FFAF7FA8AFA91" box="[159,1293,1322,1375]" gridrow="0" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFAF7FF62FA91" box="[159,229,1322,1375]" gridcol="0" gridrow="0" pageId="24" pageNumber="115">Leg:</th>
|
||
<th id="372F5DA5FFB300292723FAF7FD9BFA91" box="[307,540,1322,1375]" gridcol="1" gridrow="0" pageId="24" pageNumber="115">Larva ♂ femur/tibia/tarsus/claw</th>
|
||
<th id="372F5DA5FFB30029248CFAF7FC06FA91" box="[668,897,1322,1375]" gridcol="2" gridrow="0" pageId="24" pageNumber="115">Subimago ♂ femur/tibia/tarsomeres</th>
|
||
<th id="372F5DA5FFB300292208FAF7FA8AFA91" box="[1048,1293,1322,1375]" gridcol="3" gridrow="0" pageId="24" pageNumber="115">Imago ♂ femur/tibia/tarsomeres</th>
|
||
</tr>
|
||
<tr id="74FE34D9FFB30029268FFAAFFA8AFA46" box="[159,1293,1394,1416]" gridrow="1" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFAAFFF62FA46" box="[159,229,1394,1416]" gridcol="0" gridrow="1" pageId="24" pageNumber="115">fore</th>
|
||
<td id="372F5DA5FFB300292723FAAFFD9BFA46" box="[307,540,1394,1416]" gridcol="1" gridrow="1" pageId="24" pageNumber="115">78: 41: 49: 27</td>
|
||
<td id="372F5DA5FFB30029248CFAAFFC06FA46" box="[668,897,1394,1416]" gridcol="2" gridrow="1" pageId="24" pageNumber="115">75: 62: 4:24:17:9:13</td>
|
||
<td id="372F5DA5FFB300292208FAAFFA8AFA46" box="[1048,1293,1394,1416]" gridcol="3" gridrow="1" pageId="24" pageNumber="115">90: 107: 5:55:33:15:13</td>
|
||
</tr>
|
||
<tr id="74FE34D9FFB30029268FFA41FA8AFA7C" box="[159,1293,1436,1458]" gridrow="2" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFA41FF62FA7C" box="[159,229,1436,1458]" gridcol="0" gridrow="2" pageId="24" pageNumber="115">middle</th>
|
||
<td id="372F5DA5FFB300292723FA41FD9BFA7C" box="[307,540,1436,1458]" gridcol="1" gridrow="2" pageId="24" pageNumber="115">73: 37: 41: 27</td>
|
||
<td id="372F5DA5FFB30029248CFA41FC06FA7C" box="[668,897,1436,1458]" gridcol="2" gridrow="2" pageId="24" pageNumber="115">73: 64: 23:9:3:12</td>
|
||
<td id="372F5DA5FFB300292208FA41FA8AFA7C" box="[1048,1293,1436,1458]" gridcol="3" gridrow="2" pageId="24" pageNumber="115">73: 64: 23:9:3:12</td>
|
||
</tr>
|
||
<tr id="74FE34D9FFB30029268FFA18FA8AFA15" box="[159,1293,1477,1499]" gridrow="3" pageId="24" pageNumber="115">
|
||
<th id="372F5DA5FFB30029268FFA18FF62FA15" box="[159,229,1477,1499]" gridcol="0" gridrow="3" pageId="24" pageNumber="115">hind</th>
|
||
<td id="372F5DA5FFB300292723FA18FD9BFA15" box="[307,540,1477,1499]" gridcol="1" gridrow="3" pageId="24" pageNumber="115">68: 34: 38: 27</td>
|
||
<td id="372F5DA5FFB30029248CFA18FC06FA15" box="[668,897,1477,1499]" gridcol="2" gridrow="3" pageId="24" pageNumber="115">68: 59: 20:8:3:12</td>
|
||
<td id="372F5DA5FFB300292208FA18FA8AFA15" box="[1048,1293,1477,1499]" gridcol="3" gridrow="3" pageId="24" pageNumber="115">68: 59: 20:8:3:12</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB3FFCE26D7F9C1FE72F82B" blockId="24.[151,1437,1564,2021]" pageId="24" pageNumber="115">
|
||
Posterior margin of abdominal tergum I either entirely smooth, or with few irregular short denticles; posterior margins of terga II–X with regular long pointed denticles; on tergum IX, portion of posterior margin behind pair of submedian setae not markedly projected backward, either with diminished denticles (
|
||
<figureCitation id="52F52A1EFFB3FFCE2240F9B9FB24F9B2" box="[1104,1187,1636,1661]" captionStart="FIGURES 92 – 97" captionStartId="25.[151,264,1837,1860]" captionTargetBox="[151,1434,347,1817]" captionTargetId="figure@25.[151,1436,347,1817]" captionTargetPageId="25" captionText="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="24" pageNumber="115">Fig. 97</figureCitation>
|
||
), or without denticles. Posterior margins of abdominal sterna I–IV smooth or with minute denticles on sternum IV; posterior margins of sterna V–VIII with regular pointed denticles, shorter and lighter than denticles on terga. Each sternum II–VI with pair of regular transverse arched rows of long, fine, simple (not furcate) setae with closely contiguous sockets (as in
|
||
<figureCitation id="52F52A1EFFB3FFCE2687F929FF6DF8C2" box="[151,234,1780,1805]" captionStart="FIGURES 18 – 25" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@9.[151,1436,193,1885]" captionTargetPageId="9" captionText="FIGURES 18 – 25. Cloeodes vibratorius sp. n. 18 – 23, larval exuviae: 18, tergum IX; 19, paraprocts; 20, portion of sternum IV with setal row; 21, fore leg; 22, middle leg and half of metanotum; 23, hind leg; 24 – 25, subimaginal exuviae: 24, left half of mesonotum; 25, postsubalar sclerite and lateropostnotal crest (21 – 23, holotype)." httpUri="https://zenodo.org/record/888633/files/figure.png" pageId="24" pageNumber="115">Fig. 20</figureCitation>
|
||
). Tergalii I–VII long, lanceolate and pointed (
|
||
<figureCitation id="52F52A1EFFB3FFCE24F8F929FC02F8C3" box="[744,901,1780,1805]" captionStart="FIGURES 119 – 132" captionStartId="31.[151,264,1977,2000]" captionTargetBox="[200,1368,511,1956]" captionTargetId="figure@31.[197,1404,507,1970]" captionTargetPageId="31" captionText="FIGURES 119 – 132. Cloeodes spp., tergalii. 119 – 125, tergalii I – VII of Cloeodes nigrohumeris sp. n.; 126 – 132, tergalii I – VII of Cloeodes auwe. For both individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888663/files/figure.png" pageId="24" pageNumber="115">Figs 119–125</figureCitation>
|
||
). All tergalii with ribs only on proximal parts of costal and anal margins, without middle rib. Margins bearing costal and anal ribs smooth (not serrated); margin free of ribs slightly notched, with small seta in each notch. Paraprocts with anterior median apodeme poorly developed; each paraproct with unequal pointed denticles on posterior margin, with dense scales as on sterna and terga (
|
||
<figureCitation id="52F52A1EFFB3FFCE237FF882FF42F852" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="24" pageNumber="115">Fig. 107</figureCitation>
|
||
). Each swimming seta of caudalii divided into dark proximal part and colorless delicate distal part (
|
||
<figureCitation id="52F52A1EFFB3FFCE233DF859FA08F852" box="[1325,1423,1924,1949]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="24" pageNumber="115">Fig. 101</figureCitation>
|
||
). Ventral and dorsal side of paracercus and lateral sides of cerci in middle part with one or several long pointed denticles on each 2nd segment.
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB2FFCF26D7FF45FEF4FF36" blockId="25.[151,1436,151,285]" pageId="25" pageNumber="116">
|
||
DEVELOPING MALE GENITALS: In last larval instar, developing subimaginal gonostyli folded under larval cuticle in «
|
||
<taxonomicName id="0DCE4D18FFB2FFCF270EFF61FE26FF1A" authorityName="Novikova & Kluge" authorityYear="1987" box="[286,417,188,212]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8BAEA89FFB2FFCF270EFF61FE26FF1A" box="[286,417,188,212]" italics="true" pageId="25" pageNumber="116">Nigrobaetis</emphasis>
|
||
</taxonomicName>
|
||
-type» pose, with 2nd segment bent by convexity medially, and 3rd segment directed caudally (
|
||
<figureCitation id="52F52A1EFFB2FFCF2715FF02FEE1FF36" box="[261,358,223,248]" captionStart="FIGURES 116 – 118" captionStartId="29.[151,264,1368,1391]" captionTargetBox="[191,1395,214,1346]" captionTargetId="figure@29.[170,1417,193,1348]" captionTargetPageId="29" captionText="FIGURES 116 – 118. Cloeodes nigrohumeris sp. n. 116, subimaginal gonostyli extracted from mature larva; 117, exuviae of subimaginal genitals; 118, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines) (117 – 118, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888661/files/figure.png" pageId="25" pageNumber="116">Fig. 116</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB2FFCF26D7FED9FB36FED3" blockId="25.[151,1436,151,285]" box="[199,1201,260,285]" pageId="25" pageNumber="116">RESPIRATORY MOVEMENTS: Larva is able to make respiratory vibrations by tergalii.</paragraph>
|
||
<caption id="9EB16613FFB2FFCF2687F8F0FDEBF86E" httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="25" pageNumber="116" targetBox="[151,1434,347,1817]" targetPageId="25">
|
||
<paragraph id="CA71369BFFB2FFCF2687F8F0FDEBF86E" blockId="25.[151,1436,1837,1952]" pageId="25" pageNumber="116">
|
||
<emphasis id="F8BAEA89FFB2FFCF2687F8F0FED4F88D" bold="true" box="[151,339,1837,1860]" pageId="25" pageNumber="116">FIGURES 92–97.</emphasis>
|
||
<taxonomicName id="0DCE4D18FFB2FFCF2749F8F3FDC2F88A" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[345,581,1838,1860]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="116" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB2FFCF2749F8F3FDC2F88A" box="[345,581,1838,1860]" italics="true" pageId="25" pageNumber="116">Cloeodes nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB2FFCF245BF8F3FD04F88D" box="[587,643,1838,1859]" pageId="25" pageNumber="116" rank="species">
|
||
<emphasis id="F8BAEA89FFB2FFCF245BF8F3FD04F88D" bold="true" box="[587,643,1838,1859]" pageId="25" pageNumber="116">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
92, immature last instar male larva; 93–96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I–X, abdominal segments I–X.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9EB16613FFB1FFCC2687F84CFB20F82B" httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="26" pageNumber="117" targetBox="[174,1418,193,1924]" targetPageId="26">
|
||
<paragraph id="CA71369BFFB1FFCC2687F84CFB20F82B" blockId="26.[151,1436,1937,2021]" pageId="26" pageNumber="117">
|
||
<emphasis id="F8BAEA89FFB1FFCC2687F84CFEE6F869" bold="true" box="[151,353,1937,1960]" pageId="26" pageNumber="117">FIGURES 98–103.</emphasis>
|
||
<taxonomicName id="0DCE4D18FFB1FFCC2778F84FFDD2F866" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[360,597,1938,1960]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="117" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB1FFCC2778F84FFDD2F866" box="[360,597,1938,1960]" italics="true" pageId="26" pageNumber="117">Cloeodes nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB1FFCC244CF84FFD11F869" box="[604,662,1938,1959]" pageId="26" pageNumber="117" rank="species">
|
||
<emphasis id="F8BAEA89FFB1FFCC244CF84FFD11F869" bold="true" box="[604,662,1938,1959]" pageId="26" pageNumber="117">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
98–99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100–103, larval exuviae: 100–101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9EB16613FFB0FFCD2687F95FFD57F918" httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="27" pageNumber="118" targetBox="[195,1394,195,1620]" targetPageId="27">
|
||
<paragraph id="CA71369BFFB0FFCD2687F95FFD57F918" blockId="27.[151,1436,1666,1750]" pageId="27" pageNumber="118">
|
||
<emphasis id="F8BAEA89FFB0FFCD2687F95FFEE8F956" bold="true" box="[151,367,1666,1689]" pageId="27" pageNumber="118">FIGURES 104–109.</emphasis>
|
||
<taxonomicName id="0DCE4D18FFB0FFCD2768F95EFDE0F957" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[376,615,1667,1689]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="118" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB0FFCD2768F95EFDE0F957" box="[376,615,1667,1689]" italics="true" pageId="27" pageNumber="118">Cloeodes nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F8BAEA89FFB0FFCD247FF95EFD2CF956" bold="true" box="[623,683,1667,1688]" pageId="27" pageNumber="118">
|
||
<taxonomicNameLabel id="E38957F2FFB0FFCD247FF95EFD2CF956" box="[623,683,1667,1688]" pageId="27" pageNumber="118" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larva. 104–105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104–106, 108– 109, holotype). Abbreviations: ds1, ds2, ds3, dentisetae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CA71369BFFB0FFCD26D7F8DEFDB0F83A" blockId="27.[151,1436,1795,2036]" pageId="27" pageNumber="118">
|
||
<emphasis id="F8BAEA89FFB0FFCD26D7F8DEFEC5F8D2" bold="true" box="[199,322,1795,1820]" italics="true" pageId="27" pageNumber="118">Subimago.</emphasis>
|
||
CUTICULAR COLORATION: Cuticle with light brown and colorless areas. Head colorless, only antennae slightly tinged with pale brownish. Pronotum with light brown and colorless areas. Mesonotum at most light brownish, medioparapsidal suture colorless, other sutures darkened (
|
||
<figureCitation id="52F52A1EFFB0FFCD25E8F896FBD6F8AA" box="[1016,1105,1867,1892]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="27" pageNumber="118">Fig. 98</figureCitation>
|
||
). Thoracic pleura with light brown and colorless areas; lateropostnotal crest narrowly colored by dark brown, diffusively bordered by light brown (
|
||
<figureCitation id="52F52A1EFFB0FFCD26FEF84EFEC5F862" box="[238,322,1939,1964]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="27" pageNumber="118">Fig. 99</figureCitation>
|
||
). Base of wing light brown, other wing cuticle lighter. Legs at most colorless; distal margin of each tarsal segment narrowly bordered by brown. Abdominal terga and sterna slightly colored with light brownish, microtrichiae brown. Cerci colorless.
|
||
</paragraph>
|
||
<caption id="9EB16613FFB7FFCA2687F874FADFF811" httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="28" pageNumber="119" targetBox="[165,1429,193,1948]" targetPageId="28">
|
||
<paragraph id="CA71369BFFB7FFCA2687F874FADFF811" blockId="28.[151,1436,1961,2015]" pageId="28" pageNumber="119">
|
||
<emphasis id="F8BAEA89FFB7FFCA2687F874FEEFF871" bold="true" box="[151,360,1961,1984]" pageId="28" pageNumber="119">FIGURES 110–115.</emphasis>
|
||
<taxonomicName id="0DCE4D18FFB7FFCA2761F877FDDAF80E" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[369,605,1962,1984]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="119" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB7FFCA2761F877FDDAF80E" box="[369,605,1962,1984]" italics="true" pageId="28" pageNumber="119">Cloeodes nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB7FFCA2474F877FD19F871" box="[612,670,1962,1983]" pageId="28" pageNumber="119" rank="species">
|
||
<emphasis id="F8BAEA89FFB7FFCA2474F877FD19F871" bold="true" box="[612,670,1962,1983]" pageId="28" pageNumber="119">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9EB16613FFB6FFCB2687FA85FC74F9C6" httpUri="https://zenodo.org/record/888661/files/figure.png" pageId="29" pageNumber="120" targetBox="[191,1395,214,1346]" targetPageId="29">
|
||
<paragraph id="CA71369BFFB6FFCB2687FA85FC74F9C6" blockId="29.[151,1436,1368,1544]" pageId="29" pageNumber="120">
|
||
<emphasis id="F8BAEA89FFB6FFCB2687FA85FEEDFAA0" bold="true" box="[151,362,1368,1391]" pageId="29" pageNumber="120">FIGURES 116–118.</emphasis>
|
||
<taxonomicName id="0DCE4D18FFB6FFCB2764FA84FDE5FAA1" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[372,610,1369,1391]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="120" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB6FFCB2764FA84FDE5FAA1" box="[372,610,1369,1391]" italics="true" pageId="29" pageNumber="120">Cloeodes nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB6FFCB247BFA84FD21FAA0" box="[619,678,1369,1390]" pageId="29" pageNumber="120" rank="species">
|
||
<emphasis id="F8BAEA89FFB6FFCB247BFA84FD21FAA0" bold="true" box="[619,678,1369,1390]" pageId="29" pageNumber="120">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
116, subimaginal gonostyli extracted from mature larva; 117, exuviae of subimaginal genitals; 118, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines) (117–118, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m.IX-X, areas of anterior attachment of median paraproctal muscles (i.e., muscles going from sternum IX to common base of paraprocts); m.sg, median sterno-styligeral muscle; usg, unistyliger.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CA71369BFFB6FFCB26D7F9EBFD61F980" blockId="29.[151,1436,1589,2010]" box="[199,742,1589,1614]" pageId="29" pageNumber="120">HYPODERMAL COLORATION: As in imago.</paragraph>
|
||
<paragraph id="CA71369BFFB6FFCB26D7F987FB52F958" blockId="29.[151,1436,1589,2010]" pageId="29" pageNumber="120">
|
||
TEXTURE: On all legs of both sexes last tarsal segment entirely covered by pointed microlepides; other tarsal segments at most part covered by blunt microlepides, apically by pointed microlepides (
|
||
<tableCitation id="874C0320FFB6FFCB2260F9A0FB43F958" box="[1136,1220,1661,1686]" captionStart="TABLE 1" captionStartId="12.[151,239,152,175]" captionText="TABLE 1. Texture of subimaginal legs and apical spines of imaginal and subimaginal legs." pageId="29" pageNumber="120">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB6FFCB26D7F97CFF32F8CF" blockId="29.[151,1436,1589,2010]" pageId="29" pageNumber="120">
|
||
<emphasis id="F8BAEA89FFB6FFCB26D7F97CFDF8F974" bold="true" box="[199,639,1697,1722]" pageId="29" pageNumber="120">
|
||
<emphasis id="F8BAEA89FFB6FFCB26D7F97CFDFFF974" bold="true" box="[199,632,1697,1722]" italics="true" pageId="29" pageNumber="120">Tarsal spines of subimago and imago</emphasis>
|
||
.
|
||
</emphasis>
|
||
Tarsus of fore leg in male and female without apical spines. Tarsus of middle and hind leg of both sexes with 1 apical spine on initial 3rd tarsomere (next after 1st+2nd tarsomere) (
|
||
<tableCitation id="874C0320FFB6FFCB234CF918FF23F8CF" captionStart="TABLE 1" captionStartId="12.[151,239,152,175]" captionText="TABLE 1. Texture of subimaginal legs and apical spines of imaginal and subimaginal legs." pageId="29" pageNumber="120">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB6FFC826D7F8D3FAEBFEAB" blockId="29.[151,1436,1589,2010]" lastBlockId="30.[151,1437,151,1221]" lastPageId="30" lastPageNumber="121" pageId="29" pageNumber="120">
|
||
<emphasis id="F8BAEA89FFB6FFCB26D7F8D3FEE5F8E8" bold="true" box="[199,354,1805,1830]" italics="true" pageId="29" pageNumber="120">Imago, male.</emphasis>
|
||
COLORATION (
|
||
<figureCitation id="52F52A1EFFB6FFCB2427F8D0FD53F8E8" box="[567,724,1805,1830]" captionStart="FIGURES 110 – 115" captionStartId="28.[151,264,1961,1984]" captionTargetBox="[165,1429,193,1948]" captionTargetId="figure@28.[165,1429,193,1948]" captionTargetPageId="28" captionText="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="29" pageNumber="120">Figs 110–113</figureCitation>
|
||
). Head ocher. Turbinate eyes either unicolor dull-orange or ocher-yellowish, or with proximal half darker and distal half lighter. Thorax ocher; mesonotum with pair of anterolateral contrasting dark brown or black stripes bordering mesonotum anteriad of wing bases and including antelateroparapsidal scutal sutures. Wing membrane colorless; veins at most ocher or light brownish, basal parts of veins C, Sc, MA, MP and CuA darkened with brown. Femora of all legs ocher, with brown or reddish subapical band (hypodermal coloration); fore femur also with brown macula apically (
|
||
<figureCitation id="52F52A1EFFB6FFCB25E0F81CFB09F817" box="[1008,1166,1985,2010]" captionStart="FIGURES 110 – 115" captionStartId="28.[151,264,1961,1984]" captionTargetBox="[165,1429,193,1948]" captionTargetId="figure@28.[165,1429,193,1948]" captionTargetPageId="28" captionText="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="29" pageNumber="120">Figs 112–113</figureCitation>
|
||
). Tibiae, tarsi and claws of all legs pale ocher. Abdominal segments I–VI colorless, segments VII–X pale yellowish, tergites I–VII with following larger or smaller contrasting reddish or brown markings: tergite I with transverse line bordering posterior margin; tergites II–VII with unpaired median spot near anterior margin and pair of transverse stripes near posterior margin; tergites III, V and VI, besides it, with pair of short oblique stripes on lateral parts. Lateral tracheal trunks and bases of sternal tracheae either colorless, or partly colored by black or brown. In proximal part of cercus, middle part of each segment ocher, area around each segment colorless; distal part of cercus entirely colorless.
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB5FFC826D7FEADFB39FDF3" blockId="30.[151,1437,151,1221]" pageId="30" pageNumber="121">
|
||
STRUCTURE. Turbinate eyes with facetted surface wider than stem (
|
||
<figureCitation id="52F52A1EFFB5FFC825FFFEB2FB16FE46" box="[1007,1169,367,392]" captionStart="FIGURES 110 – 115" captionStartId="28.[151,264,1961,1984]" captionTargetBox="[165,1429,193,1948]" captionTargetId="figure@28.[165,1429,193,1948]" captionTargetPageId="28" captionText="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="30" pageNumber="121">Figs 110, 113</figureCitation>
|
||
). Pterostigma with 3–5 oblique veins. Two marginal intercalaries nearly in each space (except few posterior ones), length of most intercalaries less than distance between adjacent longitudinal veins (as in
|
||
<figureCitation id="52F52A1EFFB5FFC825CBFE6AFBA8FE1E" box="[987,1071,439,464]" captionStart="FIGURES 32 – 36" captionStartId="11.[151,264,1813,1836]" captionTargetBox="[173,1416,207,1773]" captionTargetId="figure@11.[170,1417,193,1792]" captionTargetPageId="11" captionText="FIGURES 32 – 36. Cloeodes vibratorius sp. n. 32, fore wing of male; 33, subimaginal gonostyli developing under larval cuticle; 34, exuviae of subimaginal genitals; 35, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 36, gonostylus (32, 34 – 35, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888637/files/figure.png" pageId="30" pageNumber="121">Fig. 32</figureCitation>
|
||
). Hind wing absent. Middle and hind tarsus long (about 0.7 of tibia length), its 1st+2nd segment subequal to 3rd, 4th and 5th segments together (
|
||
<tableCitation id="874C0320FFB5FFC8268EFE22FF70FDD6" box="[158,247,511,536]" captionStart="TABLE 5" captionStartId="24.[151,239,1272,1295]" captionText="TABLE 5. Leg proportions of male C. nigrohumeris sp. n. measured on outer side of leg of paratype No. [XXI] (4) 2013." pageId="30" pageNumber="121">Table 5</tableCitation>
|
||
). Genitals as in
|
||
<figureCitation id="52F52A1EFFB5FFC827A3FE22FD92FDD6" box="[435,533,511,536]" captionStart="FIGURES 116 – 118" captionStartId="29.[151,264,1368,1391]" captionTargetBox="[191,1395,214,1346]" captionTargetId="figure@29.[170,1417,193,1348]" captionTargetPageId="29" captionText="FIGURES 116 – 118. Cloeodes nigrohumeris sp. n. 116, subimaginal gonostyli extracted from mature larva; 117, exuviae of subimaginal genitals; 118, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines) (117 – 118, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888661/files/figure.png" pageId="30" pageNumber="121">Fig. 118</figureCitation>
|
||
: surface between unistyligers striated; gonovectes sharply bent; median sternostyligeral muscle slender, consists of pair of separate bunches, which are convergent V-shaped.
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB5FFC826D7FD95FAFBFD66" blockId="30.[151,1437,151,1221]" pageId="30" pageNumber="121">
|
||
<emphasis id="F8BAEA89FFB5FFC826D7FD95FEFCFDAE" bold="true" box="[199,379,583,608]" italics="true" pageId="30" pageNumber="121">Imago, female.</emphasis>
|
||
COLORATION (
|
||
<figureCitation id="52F52A1EFFB5FFC82443FD9AFD71FDAE" box="[595,758,583,608]" captionStart="FIGURES 110 – 115" captionStartId="28.[151,264,1961,1984]" captionTargetBox="[165,1429,193,1948]" captionTargetId="figure@28.[165,1429,193,1948]" captionTargetPageId="28" captionText="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="30" pageNumber="121">Figs 114–115</figureCitation>
|
||
). Head ocher. Thorax ocher; mesonotum without pair of anterolateral dark stripes characteristic for male. Coloration of wing veins as in male, but somewhat darker. Coloration of legs as in male. Abdomen ocher, with contrasting reddish or brown markings arranged as in male.
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB5FFC826D7FD69FF52FCDB" blockId="30.[151,1437,151,1221]" pageId="30" pageNumber="121">
|
||
STRUCTURE. Eyes relatively large, elevated above head surface (
|
||
<figureCitation id="52F52A1EFFB5FFC825DEFD69FBE9FD03" box="[974,1134,692,717]" captionStart="FIGURES 110 – 115" captionStartId="28.[151,264,1961,1984]" captionTargetBox="[165,1429,193,1948]" captionTargetId="figure@28.[165,1429,193,1948]" captionTargetPageId="28" captionText="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="30" pageNumber="121">Figs 114–115</figureCitation>
|
||
). Wing with one marginal intercalary in some spaces and no one marginal intercalary in other spaces; in other respects wing venation as in male.
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB5FFC826D7FCFDFB5AFCF6" blockId="30.[151,1437,151,1221]" box="[199,1245,799,824]" pageId="30" pageNumber="121">
|
||
<emphasis id="F8BAEA89FFB5FFC826D7FCFDFF71FCF9" bold="true" box="[199,246,800,823]" italics="true" pageId="30" pageNumber="121">Egg</emphasis>
|
||
. Oval,
|
||
<quantity id="0D369B7EFFB5FFC82758FCFDFE19FCF9" box="[328,414,800,824]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="121" unit="mm" value="0.1">0.1 mm</quantity>
|
||
length. Chorion irregularly crumpled, without relief (as in
|
||
<figureCitation id="52F52A1EFFB5FFC82223FCC2FB57FCF6" box="[1075,1232,799,824]" captionStart="FIGURES 145 – 146" captionStartId="35.[151,264,1200,1223]" captionTargetBox="[151,1436,700,1179]" captionTargetId="figure@35.[151,1436,700,1179]" captionTargetPageId="35" captionText="FIGURES 145 – 146. Eggs. 145, Cloeodes vibratorius sp. n.; 146, Cloeodes redactus." httpUri="https://zenodo.org/record/888671/files/figure.png" pageId="30" pageNumber="121">Figs 145, 146</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB5FFC826D7FC99FC21FC92" blockId="30.[151,1437,151,1221]" box="[199,934,836,861]" pageId="30" pageNumber="121">
|
||
<emphasis id="F8BAEA89FFB5FFC826D7FC99FEC9FC93" bold="true" box="[199,334,836,861]" pageId="30" pageNumber="121">Dimension.</emphasis>
|
||
Fore wing length: male 4.0–
|
||
<quantity id="0D369B7EFFB5FFC82482FC99FD6DFC92" box="[658,746,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="30" pageNumber="121" unit="mm" value="4.1">4.1 mm</quantity>
|
||
; female
|
||
<quantity id="0D369B7EFFB5FFC82559FC99FC26FC92" box="[841,929,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="30" pageNumber="121" unit="mm" value="4.2">4.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82D46510FFB5FFC826D7FCBAFDC1FB0B" pageId="30" pageNumber="121" type="distribution">
|
||
<paragraph id="CA71369BFFB5FFC826D7FCBAFDCBFC4E" blockId="30.[151,1437,151,1221]" box="[199,588,871,896]" pageId="30" pageNumber="121">
|
||
<emphasis id="F8BAEA89FFB5FFC826D7FCBAFEE7FC4E" bold="true" box="[199,352,871,896]" pageId="30" pageNumber="121">Distribution.</emphasis>
|
||
Peruvian Amazonia.
|
||
</paragraph>
|
||
<paragraph id="CA71369BFFB5FFC826D7FC51FDC1FB0B" blockId="30.[151,1437,151,1221]" pageId="30" pageNumber="121">
|
||
<emphasis id="F8BAEA89FFB5FFC826D7FC51FEE5FC6B" bold="true" box="[199,354,908,933]" pageId="30" pageNumber="121">Comparison.</emphasis>
|
||
Larva of
|
||
<taxonomicName id="0DCE4D18FFB5FFC827C6FC51FD59FC6A" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[470,734,908,932]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB5FFC827C6FC51FD59FC6A" box="[470,734,908,932]" italics="true" pageId="30" pageNumber="121">Cloeodes nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB5FFC824F7FC51FCAEFC6B" box="[743,809,908,933]" pageId="30" pageNumber="121" rank="species">
|
||
<emphasis id="F8BAEA89FFB5FFC824F7FC51FCAEFC6B" bold="true" box="[743,809,908,933]" pageId="30" pageNumber="121">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="0DCE4D18FFB5FFC825D4FC51FBBCFC6A" box="[964,1083,908,932]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="spaceki">
|
||
<emphasis id="F8BAEA89FFB5FFC825D4FC51FBBCFC6A" box="[964,1083,908,932]" italics="true" pageId="30" pageNumber="121">C. spaceki</emphasis>
|
||
</taxonomicName>
|
||
by long claws, shape of distal dentiseta of maxilla, widened distal segment of labial palp and cuticular coloration of abdomen; judging by the photo of mature larva ready to molt to subimago (
|
||
<bibRefCitation id="AE5F4B6AFFB5FFC824C1FC09FC40FC22" author="Massariol" box="[721,967,980,1005]" pageId="30" pageNumber="121" refString="Massariol, F. C., Lima, L. R. C., Pinheiro, U. D. S., Quieroz, L. L., Oliveira, L. G. & Salles, F. F. (2013) Two-winged Cloeodes in Brazil: New species, stage description, and key to South American species. Journal of Insect Science (Tucson), 13 (Article 60), 1 - 20." type="journal article" year="2013">
|
||
Massariol
|
||
<emphasis id="F8BAEA89FFB5FFC8255AFC08FCFBFC22" box="[842,892,980,1004]" italics="true" pageId="30" pageNumber="121">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
: Fig, 7), male subimago and imago of
|
||
<taxonomicName id="0DCE4D18FFB5FFC82392FC09FF6CFBDE" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="spaceki">
|
||
<emphasis id="F8BAEA89FFB5FFC82392FC09FF6CFBDE" italics="true" pageId="30" pageNumber="121">C. spaceki</emphasis>
|
||
</taxonomicName>
|
||
has a pair of contrasting dark stripes on mesonotum as in
|
||
<taxonomicName id="0DCE4D18FFB5FFC8256DFC25FBB2FBDE" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[893,1077,1016,1040]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB5FFC8256DFC25FBB2FBDE" box="[893,1077,1016,1040]" italics="true" pageId="30" pageNumber="121">C. nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB5FFC8222CFC2AFBFBFBDE" box="[1084,1148,1015,1040]" pageId="30" pageNumber="121" rank="species">
|
||
<emphasis id="F8BAEA89FFB5FFC8222CFC2AFBFBFBDE" bold="true" box="[1084,1148,1015,1040]" pageId="30" pageNumber="121">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Larva of the new species
|
||
<taxonomicName id="0DCE4D18FFB5FFC82687FBC1FECAFBFA" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[151,333,1052,1076]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB5FFC82687FBC1FECAFBFA" box="[151,333,1052,1076]" italics="true" pageId="30" pageNumber="121">C. nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB5FFC82746FBC1FE11FBFB" box="[342,406,1052,1077]" pageId="30" pageNumber="121" rank="species">
|
||
<emphasis id="F8BAEA89FFB5FFC82746FBC1FE11FBFB" bold="true" box="[342,406,1052,1077]" pageId="30" pageNumber="121">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="0DCE4D18FFB5FFC8243CFBC1FD26FBFA" box="[556,673,1052,1076]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="spaceki">
|
||
<emphasis id="F8BAEA89FFB5FFC8243CFBC1FD26FBFA" box="[556,673,1052,1076]" italics="true" pageId="30" pageNumber="121">C. spaceki</emphasis>
|
||
</taxonomicName>
|
||
by constant presence of reddish hypodermal markings on abdomen (
|
||
<figureCitation id="52F52A1EFFB5FFC8268FFBE2FE99FB96" box="[159,286,1087,1112]" captionStart="FIGURES 92 – 97" captionStartId="25.[151,264,1837,1860]" captionTargetBox="[151,1434,347,1817]" captionTargetId="figure@25.[151,1436,347,1817]" captionTargetPageId="25" captionText="FIGURES 92 – 97. Cloeodes nigrohumeris sp. n. 92, immature last instar male larva; 93 – 96, mature last instar male larva ready to molt to subimago: 93, intact condition; 94, its detached cuticle of pronotum and mesonotum; 95, its extracted abdomen with hypodermal coloration; 96, its detached cuticle of abdomen. 97, middle of posterior margin of abdominal tergum IX. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888653/files/figure.png" pageId="30" pageNumber="121">Figs 92–93</figureCitation>
|
||
) and by peculiar subapical seta on dorsal side of glossa (
|
||
<figureCitation id="52F52A1EFFB5FFC82584FBE2FC72FB96" box="[916,1013,1087,1112]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="30" pageNumber="121">Fig. 108</figureCitation>
|
||
). Tergalius IV of
|
||
<taxonomicName id="0DCE4D18FFB5FFC822A7FB9DFAABFB96" box="[1207,1324,1088,1112]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="spaceki">
|
||
<emphasis id="F8BAEA89FFB5FFC822A7FB9DFAABFB96" box="[1207,1324,1088,1112]" italics="true" pageId="30" pageNumber="121">C. spaceki</emphasis>
|
||
</taxonomicName>
|
||
is figured as rounded apically (
|
||
<bibRefCitation id="AE5F4B6AFFB5FFC8276DFBB9FDEAFBB2" author="Massariol" box="[381,621,1124,1149]" pageId="30" pageNumber="121" refString="Massariol, F. C., Lima, L. R. C., Pinheiro, U. D. S., Quieroz, L. L., Oliveira, L. G. & Salles, F. F. (2013) Two-winged Cloeodes in Brazil: New species, stage description, and key to South American species. Journal of Insect Science (Tucson), 13 (Article 60), 1 - 20." type="journal article" year="2013">
|
||
Massariol
|
||
<emphasis id="F8BAEA89FFB5FFC827E4FBB8FDA3FBB2" box="[500,548,1124,1148]" italics="true" pageId="30" pageNumber="121">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
: Fig, 42b), while tergalii of
|
||
<taxonomicName id="0DCE4D18FFB5FFC825B2FBB9FBDFFBB2" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[930,1112,1124,1148]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
|
||
<emphasis id="F8BAEA89FFB5FFC825B2FBB9FBDFFBB2" box="[930,1112,1124,1148]" italics="true" pageId="30" pageNumber="121">C. nigrohumeris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E38957F2FFB5FFC8224EFBB9FB1AFBB3" box="[1118,1181,1124,1149]" pageId="30" pageNumber="121" rank="species">
|
||
<emphasis id="F8BAEA89FFB5FFC8224EFBB9FB1AFBB3" bold="true" box="[1118,1181,1124,1149]" pageId="30" pageNumber="121">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are pointed (
|
||
<figureCitation id="52F52A1EFFB5FFC8233EFBB9FA17FBB2" box="[1326,1424,1124,1149]" captionStart="FIGURES 119 – 132" captionStartId="31.[151,264,1977,2000]" captionTargetBox="[200,1368,511,1956]" captionTargetId="figure@31.[197,1404,507,1970]" captionTargetPageId="31" captionText="FIGURES 119 – 132. Cloeodes spp., tergalii. 119 – 125, tergalii I – VII of Cloeodes nigrohumeris sp. n.; 126 – 132, tergalii I – VII of Cloeodes auwe. For both individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888663/files/figure.png" pageId="30" pageNumber="121">Fig. 122</figureCitation>
|
||
). This difference can be either species-specific, or caused by poor condition of tergalii in the
|
||
<typeStatus id="15758839FFB5FFC8229EFB54FB39FB6F" box="[1166,1214,1161,1185]" pageId="30" pageNumber="121">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="0DCE4D18FFB5FFC82338FB55FA1BFB6E" box="[1320,1436,1160,1184]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="121" phylum="Arthropoda" rank="species" species="spaceki">
|
||
<emphasis id="F8BAEA89FFB5FFC82338FB55FA1BFB6E" box="[1320,1436,1160,1184]" italics="true" pageId="30" pageNumber="121">C. spaceki</emphasis>
|
||
</taxonomicName>
|
||
(F.F. Salles, personal communication).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |