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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.29.6004" ID-PMC="PMC3881348" ID-Pensoft-Pub="1314-2003-29-75" ID-Pensoft-UUID="1D50FFB28D558455FFD9FFD4FF96FFFA" ID-PubMed="24399896" ID-Zenodo-Dep="576182" ModsDocID="1314-2003-29-75" checkinTime="1451251807655" checkinUser="pensoft" docAuthor="Levin, Geoffrey A." docDate="2013" docId="C0B569C5EC5F5F6896DDC3C676C0086A" docLanguage="en" docName="PhytoKeys 29: 75-87" docOrigin="PhytoKeys 29" docPubDate="2013-11-22" docSource="http://dx.doi.org/10.3897/phytokeys.29.6004" docTitle="Drypetes alba Griseb., Nachr. Königl. Ges. Wiss. Georg-Augusts-Univ. 1: 165. 1865." docType="treatment" docVersion="3" id="1D50FFB28D558455FFD9FFD4FF96FFFA" lastPageNumber="81" masterDocId="1D50FFB28D558455FFD9FFD4FF96FFFA" masterDocTitle="A synopsis of the New World species of Drypetes section Drypetes (Putranjivaceae) with asymmetrical fruits, including description of a new species" masterLastPageNumber="87" masterPageNumber="75" pageNumber="78" updateTime="1668140962603" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A synopsis of the New World species of Drypetes section Drypetes (Putranjivaceae) with asymmetrical fruits, including description of a new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Levin, Geoffrey A.</mods:namePart>
<mods:affiliation>Illinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 South Oak Street, Champaign, Illinois 61820</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-11-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>29</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>75</mods:start>
<mods:end>87</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.29.6004</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.29.6004</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-29-75</mods:identifier>
<mods:identifier type="Pensoft-UUID">1D50FFB28D558455FFD9FFD4FF96FFFA</mods:identifier>
<mods:identifier type="Zenodo-Dep">576182</mods:identifier>
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<treatment ID-GBIF-Taxon="182228906" LSID="urn:lsid:plazi:treatment:C0B569C5EC5F5F6896DDC3C676C0086A" httpUri="http://treatment.plazi.org/id/C0B569C5EC5F5F6896DDC3C676C0086A" lastPageId="6" lastPageNumber="81" pageId="3" pageNumber="78">
<subSubSection pageId="3" pageNumber="78" type="nomenclature">
<paragraph pageId="3" pageNumber="78">
1b.
<taxonomicName LSID="C0B569C5-EC5F-5F68-96DD-C3C676C0086A" authority="latifolia Griseb., Nachr. Koenigl. Ges. Wiss. Georg-Augusts-Univ. 1: 165. 1865." authorityName="Griseb., Nachr. Königl. Ges. Wiss. Georg-Augusts-Univ. 1: 165. 1865." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Drypetes alba" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="alba">
Drypetes alba latifolia Griseb., Nachr.
<normalizedToken originalValue="Königl">Koenigl</normalizedToken>
. Ges. Wiss. Georg-Augusts-Univ. 1: 165. 1865.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="78" type="reference_group">
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="crocea">Drypetes crocea</taxonomicName>
Poit. var.
<emphasis italics="true" pageId="3" pageNumber="78">latifolia</emphasis>
(Griseb.)
<normalizedToken originalValue="Müll">Muell</normalizedToken>
. Arg., in A. P. de Candolle, Prodr. 15(2): 456. 1866. Type: Based on
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="3" pageNumber="78">Drypetes alba</emphasis>
</taxonomicName>
Poit. var.
<emphasis italics="true" pageId="3" pageNumber="78">latifolia</emphasis>
Griseb.
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName authorityName="C. Wright" authorityYear="1870" baseAuthorityName="Griseb." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="latifolia">Drypetes latifolia</taxonomicName>
(Griseb.) C.Wright, Anales Acad. Ci.
<normalizedToken originalValue="Méd">Med</normalizedToken>
. Habana 7: 151. 1870. Type: Based on
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="3" pageNumber="78">Drypetes alba</emphasis>
</taxonomicName>
Poit. var.
<emphasis italics="true" pageId="3" pageNumber="78">latifolia</emphasis>
Griseb.
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Liliopsida" family="Poaceae" genus="Koelera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="3" pageNumber="78" phylum="Tracheophyta" rank="genus">Koelera</taxonomicName>
?
<emphasis italics="true" pageId="3" pageNumber="78">serrata</emphasis>
Maycock, Fl. Barbad. 38. 1830. Type: No specimens cited or located [according to
<bibRefCitation author="Stafleu, FA" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B14" refString="Stafleu, FA, Cowan, RS, 1981. Taxonomic literature, ed. 2, vol. III: Lh-O. Bohn, Scheltema, &amp; Holkema, Utrecht, Netherlands, 1-980. http://www.biodiversitylibrary.org/item/104137." title="Taxonomic literature, ed. 2, vol. III: Lh-O. Bohn, Scheltema, &amp; Holkema, Utrecht, Netherlands, 1 - 980. http: // www. biodiversitylibrary. org / item / 104137" year="1981">Stafleu and Cowan (1981)</bibRefCitation>
, Maycock probably made no herbarium; no type has ever been cited for this name].
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="serrata">Drypetes serrata</taxonomicName>
(Maycock) Krug &amp; Urb., Bot. Jahrb. Syst. 15: 354. 1892. Type: Based on
<taxonomicName authorityName="Maycock" authorityYear="1830" class="Liliopsida" family="Poaceae" genus="Koelera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="3" pageNumber="78" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis italics="true" pageId="3" pageNumber="78">Koelera serrata</emphasis>
</taxonomicName>
Maycock.
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="serrulata">Drypetes serrulata</taxonomicName>
Pax &amp; K.Hoffm. in H.G.A.Engler, Pflanzenr., IV, 147, XV: 267. 1922, nom. superfl. Type: Based on
<taxonomicName authorityName="Maycock" authorityYear="1830" class="Liliopsida" family="Poaceae" genus="Koelera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="3" pageNumber="78" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis italics="true" pageId="3" pageNumber="78">Koelera serrata</emphasis>
</taxonomicName>
Maycock.
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="glomerata">Drypetes glomerata</taxonomicName>
Griseb., Abh.
<normalizedToken originalValue="Königl">Koenigl</normalizedToken>
. Ges. Wiss.
<normalizedToken originalValue="Göttingen">Goettingen</normalizedToken>
7: 75. 1857. Type: Guadeloupe.n.d.,
<emphasis italics="true" pageId="3" pageNumber="78">E. P. Duchassaing s.n</emphasis>
. (holotype: GOET, photo plants.jstor.org/specimen/goet006390).
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="glomerata">Drypetes glomerata</taxonomicName>
Griseb. var.
<emphasis italics="true" pageId="3" pageNumber="78">genuina</emphasis>
<normalizedToken originalValue="Müll">Muell</normalizedToken>
.Arg. in A.P.de Candolle, Prodr. 15(2): 454. 1866, nom. inval. Type: Based on
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="glomerata">
<emphasis italics="true" pageId="3" pageNumber="78">Drypetes glomerata</emphasis>
</taxonomicName>
Griseb.
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="incurva">Drypetes incurva</taxonomicName>
<normalizedToken originalValue="Müll">Muell</normalizedToken>
. Arg., Linnaea 32: 82. 1863. Type: Cuba, prope Havana, 1833 (fr),
<emphasis italics="true" pageId="3" pageNumber="78">R. de la Sagra 607</emphasis>
(lectotype, designated here: G-DC, duplicate: K).
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="glomerata">Drypetes glomerata</taxonomicName>
Griseb. var.
<emphasis italics="true" pageId="3" pageNumber="78">olivacea</emphasis>
<normalizedToken originalValue="Müll">Muell</normalizedToken>
. Arg. in DC., Prodr. (DC.) 15(2): 454. 1866. Type: Cuba, 1860-1864,
<emphasis italics="true" pageId="3" pageNumber="78">C. Wright 1929</emphasis>
(holotype: G-DC, isotypes: BM, F, GH, K, MO, NY).
</paragraph>
<paragraph pageId="3" pageNumber="78">
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="serrata">Drypetes serrata</taxonomicName>
(Maycock) Krug &amp; Urb.var.
<emphasis italics="true" pageId="3" pageNumber="78">olivacea</emphasis>
(
<normalizedToken originalValue="Müll">Muell</normalizedToken>
. Arg.) Krug &amp; Urb., Bot. Jahrb. Syst. 15: 355. 1892. Type: Based on
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="3" pageNumber="78" phylum="Angiospermae" rank="species" species="glomerata">
<emphasis italics="true" pageId="3" pageNumber="78">Drypetes glomerata</emphasis>
</taxonomicName>
Griseb. var.
<emphasis italics="true" pageId="3" pageNumber="78">olivacea</emphasis>
<normalizedToken originalValue="Müll">Muell</normalizedToken>
. Arg.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="78" type="type">
<paragraph pageId="3" pageNumber="78">Type.</paragraph>
<paragraph pageId="3" pageNumber="78">
Cuba, occ., 1863 (fr),
<emphasis italics="true" pageId="3" pageNumber="78">Wright 1927</emphasis>
(lectotype, designated here: GOET003380, photo plants.jstor.org/specimen/goet003380; duplicates: GH pro parte, K pro parte, MO pro parte, NY pro parte).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="79" type="distribution">
<paragraph pageId="4" pageNumber="79">
<pageBreakToken pageId="4" pageNumber="79" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="79">Cuba, Jamaica, and the Lesser Antilles (Antigua, Guadeloupe, Martinique, Barbados).</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="79" type="ecology">
<paragraph pageId="4" pageNumber="79">Ecology.</paragraph>
<paragraph pageId="4" pageNumber="79">Forests on limestone and schist, from sea level to 1100 m.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="79" type="phenology">
<paragraph pageId="4" pageNumber="79">Phenology.</paragraph>
<paragraph pageId="4" pageNumber="79">Flowering October to March. Fruiting December to July.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="79" type="conservation status">
<paragraph pageId="4" pageNumber="79">Conservation status.</paragraph>
<paragraph pageId="4" pageNumber="79">
Least concern.
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="4" pageNumber="79">latifolia</emphasis>
is widespread and probably secure in Cuba and Jamaica. Its status in the Lesser Antilles is difficult to assess because of a paucity of specimens.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79" type="discussion">
<paragraph pageId="4" pageNumber="79">Discussion.</paragraph>
<paragraph pageId="4" pageNumber="79">
The plants I include in
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
have been segregated into species or varieties in various ways since the middle of the 19th century (
<bibRefCitation author="Grisebach, AHR" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B4" refString="Grisebach, AHR, 1857. Systematische Untersuchungen Ueber die Vegetation der Karaiben, insbesondere Insel Guadeloupe. Abhandlungen der Koeniglichen Gesellschaft der Wissenschaften zu Goettingen 7: 151-286. http://www.biodiversitylibrary.org/item/109580." title="Systematische Untersuchungen Ueber die Vegetation der Karaiben, insbesondere Insel Guadeloupe. Abhandlungen der Koeniglichen Gesellschaft der Wissenschaften zu Goettingen 7: 151 - 286. http: // www. biodiversitylibrary. org / item / 109580" year="1857">Grisebach 1857</bibRefCitation>
;
<bibRefCitation author="Grisebach, AHR" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B5" refString="Grisebach, AHR, 1865. Diagnosen neuer Euphorbiaceen aus Cuba. Nachrichten von der Koeniglichen Gesellschaft der Wissenschaften und von der Georg-Augusts-Universitaet 1: 161-181. http://www.digizeitschriften.de/dms/resolveppn/?PPN=GDZPPN002510227." title="Diagnosen neuer Euphorbiaceen aus Cuba. Nachrichten von der Koeniglichen Gesellschaft der Wissenschaften und von der Georg-Augusts-Universitaet 1: 161 - 181. http: // www. digizeitschriften. de / dms / resolveppn /? PPN = GDZPPN 002510227" year="1865">1865</bibRefCitation>
;
<bibRefCitation author="Howard, RA" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B6" refString="Howard, RA, 1989. Flora of the Lesser Antilles, Dicotyledoneae-part 2. Arnold Arboretum, Harvard Univ., Jamaica Plain, Massachusetts, 1-604." title="Flora of the Lesser Antilles, Dicotyledoneae-part 2. Arnold Arboretum, Harvard Univ., Jamaica Plain, Massachusetts, 1 - 604." year="1989">Howard 1989</bibRefCitation>
;
<bibRefCitation author="Leon, H" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B8" refString="Leon, H, Alain, H, 1953. Flora de Cuba. Fernandez, Havana, 1-556." title="Flora de Cuba. Fernandez, Havana, 1 - 556." year="1953">
<normalizedToken originalValue="León">Leon</normalizedToken>
and Alain 1953
</bibRefCitation>
;
<bibRefCitation author="Mueller, J" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B11" refString="Mueller, J, 1863. Euphorbiaceae. Vorlaeufige Mittheilungen aus dem fuer DeCandolle's Prodromus bestimmten Manuscript ueber diese Familie. Linnaea 32: 1-126. http://www.biodiversitylibrary.org/item/10877." title="Euphorbiaceae. Vorlaeufige Mittheilungen aus dem fuer DeCandolle's Prodromus bestimmten Manuscript ueber diese Familie. Linnaea 32: 1 - 126. http: // www. biodiversitylibrary. org / item / 10877" year="1863">
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
1863
</bibRefCitation>
;
<bibRefCitation author="Mueller, J" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B12" refString="Mueller, J, 1866. Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15(2). Masson, Paris, 1-1286. http://www.biodiversitylibrary.org/item/7166." title="Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15 (2). Masson, Paris, 1 - 1286. http: // www. biodiversitylibrary. org / item / 7166" year="1866">1866</bibRefCitation>
;
<bibRefCitation author="Pax, F" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B13" refString="Pax, F, Hoffmann, K, 1922. Euphorbiaceae-Phyllanthoideae-Phyllantheae. In: Engler A (Ed) Das Pflanzenreich, IV, 147 XV (Heft 81). 1-349. http://www.biodiversitylibrary.org/item/61957." title="Euphorbiaceae-Phyllanthoideae-Phyllantheae. In: Engler A (Ed) Das Pflanzenreich, IV, 147 XV (Heft 81). 1 - 349. http: // www. biodiversitylibrary. org / item / 61957" year="1922">Pax and Hoffmann 1922</bibRefCitation>
). Characters that have been used include those of the leaves (petiole length and blade color, shape, and degree of marginal serration), staminate flowers (size, pedicel length, and stamen exsertion), and pistillate flowers/fruits (pedicel length relative to fruit length, style presence/absence, and fruit size). Most of the distinctions were based on observations of the one or two specimens available to earlier workers and these disappear when more specimens are examined. For example,
<bibRefCitation author="Grisebach, AHR" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B4" refString="Grisebach, AHR, 1857. Systematische Untersuchungen Ueber die Vegetation der Karaiben, insbesondere Insel Guadeloupe. Abhandlungen der Koeniglichen Gesellschaft der Wissenschaften zu Goettingen 7: 151-286. http://www.biodiversitylibrary.org/item/109580." title="Systematische Untersuchungen Ueber die Vegetation der Karaiben, insbesondere Insel Guadeloupe. Abhandlungen der Koeniglichen Gesellschaft der Wissenschaften zu Goettingen 7: 151 - 286. http: // www. biodiversitylibrary. org / item / 109580" year="1857">Grisebach (1857)</bibRefCitation>
,
<bibRefCitation author="Mueller, J" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B12" refString="Mueller, J, 1866. Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15(2). Masson, Paris, 1-1286. http://www.biodiversitylibrary.org/item/7166." title="Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15 (2). Masson, Paris, 1 - 1286. http: // www. biodiversitylibrary. org / item / 7166" year="1866">
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
(1866)
</bibRefCitation>
, and
<bibRefCitation author="Leon, H" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B8" refString="Leon, H, Alain, H, 1953. Flora de Cuba. Fernandez, Havana, 1-556." title="Flora de Cuba. Fernandez, Havana, 1 - 556." year="1953">
<normalizedToken originalValue="León">Leon</normalizedToken>
and Alain (1953)
</bibRefCitation>
distinguished
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
(equivalent to var.
<emphasis italics="true" pageId="4" pageNumber="79">alba</emphasis>
is this treatment) from
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="glomerata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes glomerata</emphasis>
</taxonomicName>
or its synonym
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="serrata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes serrata</emphasis>
</taxonomicName>
(here synonyms of var.
<emphasis italics="true" pageId="4" pageNumber="79">latifolia</emphasis>
) on the basis of the former having staminate flowers that are more than 1 mm long borne on longer pedicels and with exserted stamens in contrast to the latter having staminate flowers that are about 1 mm long borne on short pedicels and with included stamens. These differences appear to be an artifact of flower age: specimens with immature flowers, generally with the anthers indehiscent, were called
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="glomerata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes glomerata</emphasis>
</taxonomicName>
or
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="serrata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes serrata</emphasis>
</taxonomicName>
, whereas those bearing fully mature flowers with dehiscent anthers were called
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
.
<bibRefCitation author="Pax, F" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B13" refString="Pax, F, Hoffmann, K, 1922. Euphorbiaceae-Phyllanthoideae-Phyllantheae. In: Engler A (Ed) Das Pflanzenreich, IV, 147 XV (Heft 81). 1-349. http://www.biodiversitylibrary.org/item/61957." title="Euphorbiaceae-Phyllanthoideae-Phyllantheae. In: Engler A (Ed) Das Pflanzenreich, IV, 147 XV (Heft 81). 1 - 349. http: // www. biodiversitylibrary. org / item / 61957" year="1922">Pax and Hoffmann (1922)</bibRefCitation>
separated the same taxa using petiole length: 5-8 mm for
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
vs. about 1 cm for
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="serrulata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes serrulata</emphasis>
</taxonomicName>
, the superfluous name they used for what Grisebach and
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
called
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="glomerata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes glomerata</emphasis>
</taxonomicName>
. In their concept,
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="serrulata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes serrulata</emphasis>
</taxonomicName>
is restricted to the Lesser Antilles whereas
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
is found throughout the Greater Antilles. Measurement of specimens shows that plants from the Lesser Antilles have slightly longer petioles (7-12 mm vs. 5-10 mm) than those from farther west, but clearly the variation is great and broadly overlapping. As I treat them, the two varieties have completely overlapping petiole lengths (6-10 mm long for var.
<emphasis italics="true" pageId="4" pageNumber="79">alba</emphasis>
vs. 5-12 mm for var.
<emphasis italics="true" pageId="4" pageNumber="79">latifolia</emphasis>
).
<bibRefCitation author="Howard, RA" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B6" refString="Howard, RA, 1989. Flora of the Lesser Antilles, Dicotyledoneae-part 2. Arnold Arboretum, Harvard Univ., Jamaica Plain, Massachusetts, 1-604." title="Flora of the Lesser Antilles, Dicotyledoneae-part 2. Arnold Arboretum, Harvard Univ., Jamaica Plain, Massachusetts, 1 - 604." year="1989">Howard (1989)</bibRefCitation>
reported that
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="serrata">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes serrata</emphasis>
</taxonomicName>
, which he considered to be restricted to the Lesser Antilles, had larger fruits than
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
of the Greater Antilles, but he did not provide comparative measurements. Although mature fruits of
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
are rarely found on herbarium specimens, those I have seen are about 12-13 mm long throughout its range. Differences in leaf color, shape, and degree of marginal serration, alone or in combination, were used by
<bibRefCitation author="Grisebach, AHR" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B5" refString="Grisebach, AHR, 1865. Diagnosen neuer Euphorbiaceen aus Cuba. Nachrichten von der Koeniglichen Gesellschaft der Wissenschaften und von der Georg-Augusts-Universitaet 1: 161-181. http://www.digizeitschriften.de/dms/resolveppn/?PPN=GDZPPN002510227." title="Diagnosen neuer Euphorbiaceen aus Cuba. Nachrichten von der Koeniglichen Gesellschaft der Wissenschaften und von der Georg-Augusts-Universitaet 1: 161 - 181. http: // www. digizeitschriften. de / dms / resolveppn /? PPN = GDZPPN 002510227" year="1865">Grisebach (1865)</bibRefCitation>
and
<bibRefCitation author="Mueller, J" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B12" refString="Mueller, J, 1866. Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15(2). Masson, Paris, 1-1286. http://www.biodiversitylibrary.org/item/7166." title="Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15 (2). Masson, Paris, 1 - 1286. http: // www. biodiversitylibrary. org / item / 7166" year="1866">
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
(1866)
</bibRefCitation>
to describe new varieties based on single specimens, but these characters have been ignored by subsequent authors, presumably because they found, as I have, that these characters vary considerably even within individuals and certainly do not show consistent patterns.
</paragraph>
<paragraph lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79">
The only character that seems consistently to differentiate
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="4" pageNumber="79" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="4" pageNumber="79">Drypetes alba</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="4" pageNumber="79">alba</emphasis>
from var.
<emphasis italics="true" pageId="4" pageNumber="79">latifolia</emphasis>
is the presence of a style about 1 mm long in the former and its ab
<pageBreakToken pageId="5" pageNumber="80" start="start">sence</pageBreakToken>
in the latter, the stigma being sessile. This character was first observed by
<bibRefCitation author="Mueller, J" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B11" refString="Mueller, J, 1863. Euphorbiaceae. Vorlaeufige Mittheilungen aus dem fuer DeCandolle's Prodromus bestimmten Manuscript ueber diese Familie. Linnaea 32: 1-126. http://www.biodiversitylibrary.org/item/10877." title="Euphorbiaceae. Vorlaeufige Mittheilungen aus dem fuer DeCandolle's Prodromus bestimmten Manuscript ueber diese Familie. Linnaea 32: 1 - 126. http: // www. biodiversitylibrary. org / item / 10877" year="1863">
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
(1863)
</bibRefCitation>
when he described
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="incurva">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes incurva</emphasis>
</taxonomicName>
having a sessile stigma; he later noted the same condition in
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="glomerata">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes glomerata</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
, 1866). I have found that all specimens from an individual island show the same condition and I have seen no intermediate specimens. Although this character is consistent geographically, in the absence of additional differences it seems too minor to support more than a varietal distinction. The distribution of the varieties is curious, with var.
<emphasis italics="true" pageId="5" pageNumber="80">latifolia</emphasis>
found both east and west of var.
<emphasis italics="true" pageId="5" pageNumber="80">alba</emphasis>
. DNA sequence data might elucidate this interesting distribution and clarify the evolutionary history of
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes alba</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="5" pageNumber="80">
<bibRefCitation author="Grisebach, AHR" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B5" refString="Grisebach, AHR, 1865. Diagnosen neuer Euphorbiaceen aus Cuba. Nachrichten von der Koeniglichen Gesellschaft der Wissenschaften und von der Georg-Augusts-Universitaet 1: 161-181. http://www.digizeitschriften.de/dms/resolveppn/?PPN=GDZPPN002510227." title="Diagnosen neuer Euphorbiaceen aus Cuba. Nachrichten von der Koeniglichen Gesellschaft der Wissenschaften und von der Georg-Augusts-Universitaet 1: 161 - 181. http: // www. digizeitschriften. de / dms / resolveppn /? PPN = GDZPPN 002510227" year="1865">Grisebach (1865)</bibRefCitation>
based
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes alba</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="5" pageNumber="80">latifolia</emphasis>
on
<emphasis italics="true" pageId="5" pageNumber="80">Wright 1927</emphasis>
. It has long been recognized that this collection, like many of
<normalizedToken originalValue="Wright’s">Wright's</normalizedToken>
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes</emphasis>
</taxonomicName>
collections, is a mixture of two species, in this case
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes alba</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Krug &amp; Urb" authorityYear="1892" baseAuthorityName="Sw." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="lateriflora">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes lateriflora</emphasis>
</taxonomicName>
(
<bibRefCitation author="Krug, CWL" journalOrPublisher="Monographs in Systematic Botany from Missouri Botanical Garden" pageId="9" pageNumber="84" refId="B7" refString="Krug, CWL, Urban, I, 1892. Additamenta ad cognitionem florae Indiae occidentalis. Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie 15: 286-437. http://www.biodiversitylibrary.org/item/674." title="Additamenta ad cognitionem florae Indiae occidentalis. Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie 15: 286 - 437. http: // www. biodiversitylibrary. org / item / 674" year="1892">Krug and Urban 1892</bibRefCitation>
;
<bibRefCitation author="Pax, F" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B13" refString="Pax, F, Hoffmann, K, 1922. Euphorbiaceae-Phyllanthoideae-Phyllantheae. In: Engler A (Ed) Das Pflanzenreich, IV, 147 XV (Heft 81). 1-349. http://www.biodiversitylibrary.org/item/61957." title="Euphorbiaceae-Phyllanthoideae-Phyllantheae. In: Engler A (Ed) Das Pflanzenreich, IV, 147 XV (Heft 81). 1 - 349. http: // www. biodiversitylibrary. org / item / 61957" year="1922">Pax and Hoffmann 1922</bibRefCitation>
). The material at GOET includes two sheets, both from the Grisebach Herbarium, and therefore presumably is the original material studied by Grisebach. These have the additional numbers 46 and 47 on the labels. The sheet labeled 46 (GOET 7917) consists of staminate and pistillate flowering branches of
<taxonomicName authorityName="Krug &amp; Urb" authorityYear="1892" baseAuthorityName="Sw." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="lateriflora">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes lateriflora</emphasis>
</taxonomicName>
and the sheet labeled 47 (GOET 3380) consists of a fruiting specimen of
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes alba</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="5" pageNumber="80">latifolia</emphasis>
. Grisebach briefly described the leaves, staminate flowers, and fruits (
<normalizedToken originalValue="“drupa”">&quot;drupa&quot;</normalizedToken>
), thus he must have considered both sheets to be his new variety. In deciding which material best matches the protologue, the staminate flowers argue for GOET 7917 (
<taxonomicName authorityName="Krug &amp; Urb" authorityYear="1892" baseAuthorityName="Sw." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="lateriflora">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes lateriflora</emphasis>
</taxonomicName>
)and the fruits argue for GOET 3380 (
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes alba</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="5" pageNumber="80">latifolia</emphasis>
). However Grisebach described the leaves as being subentire. The leaves of
<taxonomicName authorityName="Krug &amp; Urb" authorityYear="1892" baseAuthorityName="Sw." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="lateriflora">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes lateriflora</emphasis>
</taxonomicName>
on GOET 7917 are completely entire, whereas the leaves of
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes alba</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="5" pageNumber="80">latifolia</emphasis>
on GOET 3380 are very shallowly crenulate-serrulate. The latter sheet thus better matches the protologue of D.
<emphasis italics="true" pageId="5" pageNumber="80">alba</emphasis>
var.
<emphasis italics="true" pageId="5" pageNumber="80">latifolia</emphasis>
and therefore I designate it as the lectotype.
</paragraph>
<paragraph pageId="5" pageNumber="80">
<bibRefCitation author="Mueller, J" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B11" refString="Mueller, J, 1863. Euphorbiaceae. Vorlaeufige Mittheilungen aus dem fuer DeCandolle's Prodromus bestimmten Manuscript ueber diese Familie. Linnaea 32: 1-126. http://www.biodiversitylibrary.org/item/10877." title="Euphorbiaceae. Vorlaeufige Mittheilungen aus dem fuer DeCandolle's Prodromus bestimmten Manuscript ueber diese Familie. Linnaea 32: 1 - 126. http: // www. biodiversitylibrary. org / item / 10877" year="1863">
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
(1863)
</bibRefCitation>
based
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="incurva">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes incurva</emphasis>
</taxonomicName>
on two collections,
<emphasis italics="true" pageId="5" pageNumber="80">de la Sagra 607</emphasis>
and
<emphasis italics="true" pageId="5" pageNumber="80">Wright 593</emphasis>
, pro parte [this collection number includes material of
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="incurva">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes incurva</emphasis>
</taxonomicName>
(=
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="alba">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes alba</emphasis>
</taxonomicName>
var.
<emphasis italics="true" pageId="5" pageNumber="80">latifolia</emphasis>
) and
<taxonomicName authorityName="Krug &amp; Urb" authorityYear="1892" baseAuthorityName="Sw." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="lateriflora">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes lateriflora</emphasis>
</taxonomicName>
]. Later,
<bibRefCitation author="Mueller, J" journalOrPublisher="Systematic Botany" pageId="9" pageNumber="84" refId="B12" refString="Mueller, J, 1866. Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15(2). Masson, Paris, 1-1286. http://www.biodiversitylibrary.org/item/7166." title="Euphorbiaceae. In: de Candolle AP (Ed) Prodromus systematis naturalis regni vegetabilis, vol. 15 (2). Masson, Paris, 1 - 1286. http: // www. biodiversitylibrary. org / item / 7166" year="1866">
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
(1866)
</bibRefCitation>
cited only
<emphasis italics="true" pageId="5" pageNumber="80">de la Sagra 607</emphasis>
under
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="incurva">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes incurva</emphasis>
</taxonomicName>
, placing
<emphasis italics="true" pageId="5" pageNumber="80">Wright 593</emphasis>
under
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="crocea">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes crocea</emphasis>
</taxonomicName>
Poit., a synonym of
<taxonomicName authorityName="Krug &amp; Urb" authorityYear="1892" baseAuthorityName="Sw." class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="lateriflora">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes lateriflora</emphasis>
</taxonomicName>
. Based on the protologue of
<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Drypetes" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="5" pageNumber="80" phylum="Angiospermae" rank="species" species="incurva">
<emphasis italics="true" pageId="5" pageNumber="80">Drypetes incurva</emphasis>
</taxonomicName>
, either sheet at G-DC could be chosen as the lectotype, but because it is not a mixed collection and therefore minimizes the potential for confusion, I designate
<emphasis italics="true" pageId="5" pageNumber="80">de la Sagra 607</emphasis>
as the lectotype.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80" type="selected specimens examined">
<paragraph pageId="5" pageNumber="80">Selected specimens examined.</paragraph>
<paragraph pageId="5" pageNumber="80">
<emphasis bold="true" pageId="5" pageNumber="80">
CUBA.
<normalizedToken originalValue="Camagüey">Camagueey</normalizedToken>
:
</emphasis>
Banao, 300-500 m, Nov 1975 (♂ fl),
<emphasis italics="true" pageId="5" pageNumber="80">Alvarez et al. 28778</emphasis>
(HAJB).
<emphasis bold="true" pageId="5" pageNumber="80">
<normalizedToken originalValue="Guantánamo">Guantanamo</normalizedToken>
:
</emphasis>
San Antonia del Sur, Puriales de Caujeri, Sierra de Purial cerca de Arroyo, 800 m, 30 May 1982 (fr),
<emphasis italics="true" pageId="5" pageNumber="80">Bisse et al. 47259</emphasis>
(HAJB).
<emphasis bold="true" pageId="5" pageNumber="80">
<normalizedToken originalValue="Holguín">Holguin</normalizedToken>
:
</emphasis>
Sierra de Nipe, prope
<normalizedToken originalValue="Río">Rio</normalizedToken>
Piloto, 350 m, 16 Dec 1915 (♂ fl),
<emphasis italics="true" pageId="5" pageNumber="80">Ekman 6694</emphasis>
(F, K, NY, U, US).
<emphasis bold="true" pageId="5" pageNumber="80">Isla de la Juventud:</emphasis>
Caleta Cocodrilos, 8 Mar 1916 (fr),
<emphasis italics="true" pageId="5" pageNumber="80">Britton et al. 15305</emphasis>
(CM, F, NY, US).
<emphasis bold="true" pageId="5" pageNumber="80">Matanzas:</emphasis>
Cienega, Peninsula de Zapata, montes al norte de Sto.
<normalizedToken originalValue="Tomás">Tomas</normalizedToken>
, 19 Apr 1977 (fr),
<emphasis italics="true" pageId="5" pageNumber="80">Bisse et al. 34469</emphasis>
(HAJB).
<emphasis bold="true" pageId="5" pageNumber="80">
Pinar del
<normalizedToken originalValue="Río">Rio</normalizedToken>
:
</emphasis>
La
<normalizedToken originalValue="Guásima">Guasima</normalizedToken>
, Rangel, Jan 1950 (♀ fl),
<emphasis italics="true" pageId="5" pageNumber="80">Liogier 1261</emphasis>
(GH, US).
<emphasis bold="true" pageId="5" pageNumber="80">
Sancti
<normalizedToken originalValue="Spíritus">Spiritus</normalizedToken>
:
</emphasis>
Trinidad Mountains, Arroyo Grande, 650-750 m, 11-12 Mar 1910 (fr),
<emphasis italics="true" pageId="5" pageNumber="80">Britton &amp;Wilson 5459</emphasis>
(F, NY).
<emphasis bold="true" pageId="5" pageNumber="80">Santiago de Cuba:</emphasis>
Bayate, 20 Feb 1917 (fr),
<emphasis italics="true" pageId="5" pageNumber="80">Ekman 8544</emphasis>
(F, K, LL, NY, U, US).
</paragraph>
<paragraph pageId="6" pageNumber="81">
<emphasis bold="true" pageId="6" pageNumber="81">
<pageBreakToken pageId="6" pageNumber="81" start="start">Jamaica</pageBreakToken>
. Trelawny:
</emphasis>
Boothe district, ca. 3 mi. north of Troy, 1600 m, 14 Mar 1955 (♀ fl, fr),
<emphasis italics="true" pageId="6" pageNumber="81">Proctor 9956</emphasis>
(NY, US).
</paragraph>
<paragraph pageId="6" pageNumber="81">
<emphasis bold="true" pageId="6" pageNumber="81">Antigua and Barbuda. Antigua:</emphasis>
environs de St. Jean, Dec 1902 (♂ fl),
<emphasis italics="true" pageId="6" pageNumber="81">Duss 80</emphasis>
(NY).
</paragraph>
<paragraph pageId="6" pageNumber="81">
<emphasis bold="true" pageId="6" pageNumber="81">Guadeloupe.</emphasis>
Marie-Galante, bois de Folle-Anse, 1896 (fr),
<emphasis italics="true" pageId="6" pageNumber="81">Duss 3628</emphasis>
(F, MO pro parte, NY, US pro parte).
</paragraph>
<paragraph pageId="6" pageNumber="81">
<emphasis bold="true" pageId="6" pageNumber="81">Martinique.</emphasis>
Morne Saint-Martin, pied de la montagne
<normalizedToken originalValue="Pelée">Pelee</normalizedToken>
, 1878 (♂ fl),
<emphasis italics="true" pageId="6" pageNumber="81">Duss 50</emphasis>
(NY).
</paragraph>
</subSubSection>
</treatment>
</document>
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