543 lines
54 KiB
XML
543 lines
54 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice" docDate="2022" docId="A7EB05D68F17564D90265034DD4E3C08" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) felixi Guéorguiev & Wrase & Assmann & Muilwijk & Machard 2022, sp. nov." docType="treatment" docUuid="17C88C87-FE7B-46B8-8602-F57CC6318FD0" docUuidSource="ZooBank" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
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<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:nameIdentifier type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wrase, David W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
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<mods:affiliation>Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
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<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
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<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Machard, Patrice</mods:namePart>
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<mods:affiliation>29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-13</mods:number>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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<mods:number>1</mods:number>
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<mods:url>http://dx.doi.org/10.3897/dez.69.83840</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.69.83840</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-1324-1-71</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:17C88C87-FE7B-46B8-8602-F57CC6318FD0" httpUri="http://treatment.plazi.org/id/A7EB05D68F17564D90265034DD4E3C08" lastPageNumber="71" pageId="0" pageNumber="71">
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<subSubSection pageId="0" pageNumber="71" type="nomenclature">
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<paragraph pageId="0" pageNumber="71">
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3.
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<taxonomicName LSID="http://zoobank.org/17C88C87-FE7B-46B8-8602-F57CC6318FD0" authority="Guéorguiev & Wrase & Assmann & Muilwijk & Machard, 2022" authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) felixi</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="description">
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<paragraph pageId="0" pageNumber="71">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1C</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche & Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5B</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7C</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8C</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10C</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche & Saulcy, 1855), male paralectotype, " Syria ". Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12C</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13B</figureCitation>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="type locality">
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<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="71">Iran, Chahar Mahal va Bakhtiari Province, 10 km W Naghan Town, 31.9410, 50.6014, 1492-1505 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
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<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
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<paragraph pageId="0" pageNumber="71">The distance from Naghan Town to the type locality is about 10 km in straight line and the direction is to the west of the town. The GPS coordinates and height indicated are more or less correct as they well correspond to the river that flows towards the village of Do Polan. In fact, the type locality is closer to the last village than to Naghan Town.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="type material">
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<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
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♂, 'IRAN, /
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<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
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<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
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<normalizedToken originalValue="vā">va</normalizedToken>
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<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
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/ 10km W Naghan Town /
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<geoCoordinate degrees="31" direction="north" minutes="56" orientation="latitude" precision="15" seconds="27" value="31.940832">31°56'27"N</geoCoordinate>
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,
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<geoCoordinate degrees="050" direction="east" minutes="36" orientation="longitude" precision="1" seconds="05.5" value="50.601524">050°36'05.5"E</geoCoordinate>
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/ 1492-1505m, 02.04.2007 / leg. Jan
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<normalizedToken originalValue="Muilwijk’">Muilwijk'</normalizedToken>
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(HMIM). Paratypes: 4♀♀, labeled as holotype (cMUI); 1♂, 'IRAN, /
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<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
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<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
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<normalizedToken originalValue="vā">va</normalizedToken>
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<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
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/ 10km W Naghan Town /
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<geoCoordinate degrees="31" direction="north" minutes="56" orientation="latitude" precision="15" seconds="27" value="31.940832">31°56'27"N</geoCoordinate>
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,
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<geoCoordinate degrees="050" direction="east" minutes="36" orientation="longitude" precision="1" seconds="05.5" value="50.601524">050°36'05.5"E</geoCoordinate>
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/ 1492-1505m, 02.04.2007 / leg. R.F.F.L.
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<normalizedToken originalValue="Felix’">Felix'</normalizedToken>
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(cFEL).
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</paragraph>
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<paragraph pageId="0" pageNumber="71">TME: 6 specimens. TGE: 2♂♂, 1♀.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="etymology">
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<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="71">Latinized patronym name after Ron Felix (Berkel-Enschot, Netherlands), an enthusiastic coleopterologist working on the taxonomy of carabid beetles, with great collecting skills, who collected part of the type series of the new species.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="diagnosis">
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<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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It is distinct from
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
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</taxonomicName>
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sp. nov. in the pronotum significantly wider than head (PW/HW ≥1.45, vs. PW/HW ≤1.45) and elytra noticeably less wide in relation to length (EL/EW ≤1.55, vs. EL/EW ≥1.55). In addition, median lobe of aedeagus (lateral view) of the new species is larger, with shaft broader and apex barely bent up (
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
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</taxonomicName>
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has significantly narrower shaft and apex more distinctly bent up; Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C, D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="description">
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<paragraph pageId="0" pageNumber="71">Description.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
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.
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</emphasis>
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Specimens of moderate size for
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<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
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</taxonomicName>
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species (BL: 6.00-7.00 mm; BW: 2.20-2.50 mm), with relatively short, moderately convex body (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1C</figureCitation>
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).
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
|
||
.
|
||
</emphasis>
|
||
See Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
|
||
.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
|
||
. Integument uniformly reddish-brown, appendages somewhat lighter than body. Surface moderately shiny.
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
|
||
.
|
||
</emphasis>
|
||
Disc of head with less distinct microsculpture. Pronotum with regular isodiametric sculpticells posterolaterally and slightly stretched isodiametric sculpticells anterolaterally, microsculpture faint to absent in middle. Ventral surface (excl. gula) microsculptured, sculpticells isodiametric (mentum, submentum, pro-, mes-, and metepisternum, prosternum and abdominal ventrites laterally) or slightly stretched isodiametric (genae, medial surface of prosternum, metasternum, abdominal ventrites medially and legs).
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
As wide as a half of pronotum (mean PW/HW= 1.49). Eyes moderately convex. Labrum subrectangular, as long as clypeus, with anterior margin straight. Frontoclypeal suture distinct in middle but very fine, lacking at sides. Frontal furrows shallow, punctiform. Paraorbital sulci moderately deep, ending slightly before or at level of posterior supraorbital pore.
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pronotum distinctly wider than long (mean PW/PL= 1.31), with widest point at second quarter. Anterior and posterior transverse impressions indistinct. Anterior bead present medially, very fine. Sides convex anteriorly, nearly straight posteriorly. Metepisternum slightly longer than wide, MA/MI= 0.90-0.95.
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
|
||
.
|
||
</emphasis>
|
||
Cylindrical, one and a half time as long as wide (mean EL/EW= 1.52), wider and quite longer than pronotum (mean EW/PW= 1.31; mean EL/PL= 2.62). Parascutellar striole and striae moderately impressed, wide; striole short, not joining stria 1; striae 1-6 reaching basal bead, 7 reduced before. Following aberration exists in two specimens, striae 3 and 4 posteriorly, each of them fusing separately with stria 2 and joined stria reaches lateral gutter, otherwise striae posteriorly as in subgeneric description. Interval 3 with three or two minute discal setiferous punctures situated as follows: anterior puncture (in most specimens missing on one or both elytron) adjoining stria 2, in middle of interval 3 or adjoining stria 3, remaining two punctures adjoining stria 2. Umbilicate setiferous series mostly consists of 16 punctures on elytron, rarely with 17 punctures.
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur with four setiferous punctures on anterior side ventrally (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche & Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5B</figureCitation>
|
||
). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
|
||
<emphasis bold="true" pageId="0" pageNumber="71">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Urite IX small, oval, with proximal margin slightly asymmetrical and rounded (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7C</figureCitation>
|
||
). Median lobe of aedeagus curved in lateral view, with short basal bulb, long and broad shaft moderately constricted proximally, and apex hardly reversely turned up; median lobe in ventral view straight, ca. 3.7 times longer than wide; apical lamella more or less asymmetrical. Internal sac in lateral view (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C</figureCitation>
|
||
) with ventral sclerite distinctly broadened and rounded distally; same in ventral view (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10C</figureCitation>
|
||
), with dorsal sclerite forming a concave left protuberance and a convex right curve, and ventral sclerite narrow and completely straight. Right paramere rather short and crooked (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche & Saulcy, 1855), male paralectotype, " Syria ". Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12C</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13B</figureCitation>
|
||
). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
|
||
</paragraph>
|
||
<caption doi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">Figure 3.</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
|
||
<taxonomicName authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">Platyderus (Eremoderus) afghanistanicus</taxonomicName>
|
||
, sp. nov., holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus" subGenus="E.">P. (E.) arabicus</taxonomicName>
|
||
, sp. nov., holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" subGenus="E.">P. (E.) brunki</taxonomicName>
|
||
, sp. nov., holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
|
||
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
|
||
Karsch, 1881, lectotype and its labels;
|
||
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
|
||
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
|
||
Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia;
|
||
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
|
||
Reitter, 1909, lectotype and its labels. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="71" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
The new species lives in sympatry with
|
||
<taxonomicName baseAuthorityName="Menetries" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">Platyderus (s. str.) umbratus</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
|
||
, 1832). However,
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
|
||
</taxonomicName>
|
||
has pronotum more coarsely punctate in basolateral parts with punctures almost reaching the lateral margin, mesotibia with four setiferous punctures on its anterior side ventrally, and short urite with apex more symmetrical (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7C</figureCitation>
|
||
), whereas
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. umbratus</emphasis>
|
||
</taxonomicName>
|
||
possesses pronotum shallowly punctate in base with punctures mostly limited to the basal impressions, mesotibia with two setiferous punctures on its anterior side ventrally, and long urite with apex curved to left (as in Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-Table-UUID="ADBAD32C4643AAB3D1823CD894BA80BA" httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" start="Table 1" startId="T1">
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<emphasis bold="true" pageId="0" pageNumber="71">Table 1.</emphasis>
|
||
Morphometric data for species of the '
|
||
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
|
||
</taxonomicName>
|
||
' and '
|
||
<emphasis italics="true" pageId="0" pageNumber="71">
|
||
<taxonomicName authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">Platyderus Eremoderus iranicus</taxonomicName>
|
||
-
|
||
<taxonomicName authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">Platyderus Eremoderus vanensis</taxonomicName>
|
||
</emphasis>
|
||
' groups of
|
||
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<table pageId="0" pageNumber="71">
|
||
<tr pageId="0" pageNumber="71">
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">Species (number of samples)</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BL/mm</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BW/mm</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/HW</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PL</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
PW/
|
||
<normalizedToken originalValue="PА">PA</normalizedToken>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PB</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PA/PB</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/EW</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EW/PW</th>
|
||
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/PL</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
|
||
(6♂♂, 5♀♀)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.00-8.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.35-2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09-1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.81-0.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.55-2.83</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
|
||
(mean)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.64</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakii">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakii</emphasis>
|
||
</taxonomicName>
|
||
(1♂)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.49</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (2♂♂, 4♀♀)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.00-7.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.20-2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.45-1.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.25-1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.33-1.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.79-0.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.48-1.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60-2.66</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (mean)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.62</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (19♂♂, 16♀♀)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">4.30-6.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.50-2.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.21-1.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30-1.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82-0.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.55-1.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.24-1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50-2.77</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (mean)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">5.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.59</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (11♂♂, 11♀♀)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.85-8.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25-2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30-1.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.14-1.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82-0.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.61-1.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.22-1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.52-2.74</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (mean)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P.</emphasis>
|
||
</taxonomicName>
|
||
sp. (1♀, Karabet Pass)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.67</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (1♂, 1♀)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.30-7.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40-2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38-1.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.21-1.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41-1.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.17-1.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83-0.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.63-2.71</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="71">
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
|
||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
|
||
<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (mean)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.67</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
<tableNote pageId="0" pageNumber="71">For abbreviations in line 1, see "Abbreviations to measurements and ratios" (section "Material and methods").</tableNote>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="71" type="habitat">
|
||
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">The specimens were hand collected by R. Felix and J. Muilwijk close to the village Do Polan along the river course. The vegetation there consisted of (over)-grazed grassland, fields and small orchards; characteristic of a cultivated landscape.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="71" type="distribution">
|
||
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="71">
|
||
Southwest Iran: Central Zagros Mountain Chains (Chahar Mahal va Bakhtiari Province, Kiar County; Fig.
|
||
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |