170 lines
18 KiB
XML
170 lines
18 KiB
XML
<document id="D7A5DCB640317518A25C58F22F6FFCC8" ENCODING="UTF-8" ID-GBIF-Dataset="a3834a3c-393d-4fc1-9857-4e014cb4a765" ModsDocID="z01117p001" checkinTime="1247232184893" checkinUser="thomas" docAuthor="H. H. Tan & Z. H. Sulaiman" docDate="2006" docId="8FEED6CD6AE7FFFBE261A499A3BACCAC" docLanguage="en" docName="2006_Tan_Sulaiman_gg1.xml" docOrigin="Zootaxa 1117" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:EFF4FFDA-A1B8-44EB-ACB4-F4A9623FB4B9" docTitle="Gastromyzon venustus Tan & Sulaiman, 2006, new species" docType="treatment" docVersion="12" lastPageNumber="18" masterDocId="157B8C3E6EA404C176DDA767AF6D510E" masterDocTitle="Three new species of Gastromyzon (Teleostei: Balitoridae) from the Temburong River basin, Brunei Darussalam, Borneo." masterLastPageNumber="19" masterPageNumber="1" pageNumber="13" updateTime="1701314283152" updateUser="plazi">
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<mods:titleInfo id="243D15E72D8342AE59566069E40664E9">
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<mods:title id="A8FB2B7452D544390B065CD8176A9C30">Three new species of Gastromyzon (Teleostei: Balitoridae) from the Temburong River basin, Brunei Darussalam, Borneo.</mods:title>
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<mods:name id="C501FDC2DBEB6D39875161E46E6DEC72" type="personal">
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<mods:role id="8DD11894AEC7B4E055D605E851372593">
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<mods:roleTerm id="DF269850761484FA33423021A8D5E57C">Author</mods:roleTerm>
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<mods:namePart id="3B2C68A1590E83D9E77EA92D5137E224">H. H. Tan</mods:namePart>
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</mods:name>
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<mods:name id="C5A0CFF3F6E2C36C3F7FB1CD494141F7" type="personal">
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<mods:roleTerm id="53A525EF5831DB1A40CCAE5E7E7C545E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E63F24BE2F3BB1D1AD3552DDFAA53ECB">Z. H. Sulaiman</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2AB0DA2C3ABEB3D26D24D16CF46F6E08">text</mods:typeOfResource>
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<mods:titleInfo id="BD0ABF229837D0291B506972E0E2E801">
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<mods:title id="FD58E0C518A30C8D7D7367E00369ADDF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4DDC2F065067594E8C3CA70FF9986384">
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<mods:date id="26D1D2DE36BB22C08FBB0D6E0872025E">2006</mods:date>
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<mods:detail id="4FBCD720A28C0E34F71161B274DB4EFF" type="volume">
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<mods:number id="75198E52FECDF2748E9A68275D6C16B6">1117</mods:number>
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<mods:start id="69A634909664BD5EF75AE0F8AB8E84D5">1</mods:start>
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<mods:end id="B330AC0403DBDE4C8A5098238290D7D7">19</mods:end>
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<mods:location id="E9D61EF2EFFE8B4523703F82A164F145">
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<mods:url id="597093C8B0AABF6237E95F66EE468575">http://www.zoobank.org/urn:lsid:zoobank.org:pub:EFF4FFDA-A1B8-44EB-ACB4-F4A9623FB4B9</mods:url>
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</mods:location>
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<mods:classification id="DF35F52A00BDC12F4322953D5793F2A3">journal article</mods:classification>
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<mods:identifier id="5CD66BD72A59BDA4E55A3A2448D9FDA8" type="Plazi-Custom">z01117p001</mods:identifier>
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</mods:mods>
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<treatment id="8FEED6CD6AE7FFFBE261A499A3BACCAC" ID-DOI="http://doi.org/10.5281/zenodo.6256477" ID-GBIF-Taxon="100121951" ID-Zenodo-Dep="6256477" LSID="urn:lsid:plazi:treatment:8FEED6CD6AE7FFFBE261A499A3BACCAC" httpUri="http://treatment.plazi.org/id/8FEED6CD6AE7FFFBE261A499A3BACCAC" lastPageNumber="18" pageNumber="13">
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<subSubSection id="769C3E98B06F925D77A51AE86C22447B" type="nomenclature">
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<paragraph id="003B6F2AF361F36CF7D8B9418F358243" pageNumber="13">
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<taxonomicName id="C0C4F8F318EF953FB47014A83C885D07" ID-CoL="3FDS7" LSID-ZBK="urn:lsid:zoobank.org:act:CEC6BB92-06C1-4BB1-8E1E-1939B5547CF5" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon venustus Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="13" phylum="Chordata" rank="species" species="venustus">Gastromyzon venustus</taxonomicName>
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,
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<taxonomicNameLabel id="267A30C22E91C3B28A07764C421249EF" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="605CE2E949CE93FACAE24158D7BD3F22" type="description">
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<paragraph id="F5FE56EF3CCA1A091566A7D4A21C42F4" pageNumber="13">Figs. 5-6</paragraph>
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</subSubSection>
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<subSubSection id="55E0F0E5B47400C2FB8C9B982165646A" type="reference_group">
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<paragraph id="DF35FD3B957BE20EA03C285A01911A12" pageNumber="13">
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<taxonomicName id="2C1C743FACE8FBB7AFA4A3CA00BC442B" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="13" phylum="Chordata" rank="species" species="monticola">Gastromyzon monticola</taxonomicName>
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(non Vaillant) - Choy & Chin, 1994: 762.
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</paragraph>
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</subSubSection>
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<subSubSection id="3457E954AE051776C080E3C589B2AE62" type="nomenclature">
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<paragraph id="8857E076929BF1C1246BD28D18A43379" pageNumber="13">
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Material examined:
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<materialsCitation id="E2477B3A117CB91DF124A7F730610936" ID-GBIF-Occurrence="923901492" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectingCountry id="9346C5248FAD8272B594E39ED207D5C7">BRUNEI DARUSSALAM</collectingCountry>
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(
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<collectingRegion id="C2AA6369B230EF27B7412C1046484366">Temburong district</collectingRegion>
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: Temburong River basin):
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<typeStatus id="7ED65899760C94A94BCA77441FCED739">HOLOTYPE</typeStatus>
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:
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<collectionCode id="195F8A97573E81D029326E6AC0381DC4" collectionName="UBD">UBD</collectionCode>
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uncatalogued, 38.8 mm SL; Sungai Belalong, in front and near to the Kuala Belalong Field Studies Centre (
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<geoCoordinate id="192E20EF473D5C9C2215B4EF604C51F5" direction="north" orientation="latitude" value="4.5473332">04°32'50.4"N</geoCoordinate>
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<geoCoordinate id="2A0C5FBC842BB1CC99D52A88DB6E1C84" direction="east" orientation="longitude" value="115.15767">115°09'27.6"E</geoCoordinate>
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[80 m asl]); H. H. Tan & K. K. P. Lim, 4-
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<date id="4EA44C25E5BF934995B3155E733C28A4" day="07" month="10" value="2001-10-07" year="2001">7 Oct 2001</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="38FB4EA5B23EF0A2E8E14D463C3C4FC3" pageNumber="13">
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<materialsCitation id="D9FFA76D74E2E25F8FE2614200698B0A" ID-GBIF-Occurrence="923901495" country="Brunei Darussalam" stateProvince="Temburong District">
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<typeStatus id="F5690D9562FEE2CE718F610045C23849">PARATYPES</typeStatus>
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:
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<collectionCode id="B7D64496B600F3B158D5BEBC5A72077E" collectionName="UBD">UBD</collectionCode>
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uncatalogued, 3 ex., 37.4-39.6 mm SL
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</materialsCitation>
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;
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<materialsCitation id="13B3092B3E178B380FB0886C6070FE56" ID-GBIF-Occurrence="923901493" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="202CE633AA1AD23058D88561FC27E366" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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47104, 4 ex., 36.6-49.4 mm SL; same locality as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="53E86E108D734F2FBE0BDD14F116B922" type="materials_examined">
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<paragraph id="1779B56C2631B06C1397D03B4CAA5A48" pageNumber="13">
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NON-TYPE MATERIAL:
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<materialsCitation id="FF401A0A7C0BBAE197AA7DDC1107D946" ID-GBIF-Occurrence="923901498" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectingCountry id="6FD077B962A264E1F9CAEBB508FABD5E">BRUNEI DARUSSALAM</collectingCountry>
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(
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<collectingRegion id="AFA14465FE3CB4C20A906C159BF79735">Temburong district</collectingRegion>
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: Temburong basin):
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<collectionCode id="26139EC1E5C3883B36CDE89D3E14A4EF">UBD-B-</collectionCode>
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1b, 1 ex., 39.0 mm SL
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</materialsCitation>
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;
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<materialsCitation id="37AB307379F935AFC80433DDE586D06B" ID-GBIF-Occurrence="923901494" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="75D6FCCEF6EFBF201B38892742D09E75" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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38777, 1 ex., 33.7 mm SL; Sungai Temburong, about 1 km downstream of Kuala Belalong; K. K. P. Lim et al.,
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<date id="226E084FC3889B380AD7045BDEAC2D3C" day="16" month="06" value="1995-06-16" year="1995">16 Jun 1995</date>
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</materialsCitation>
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.
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<materialsCitation id="19094FC4598734F1EBA6825CA6E9F815" ID-GBIF-Occurrence="923901496" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="F620272FA4562D52C0EFF2969BE21698">UBD-SC</collectionCode>
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/92/7/F5b, 2 ex., 28.1-41.5 mm SL; Sungai Belalong, Kuala Belalong Field Study Centre; S. Choy,
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<date id="79E6AD7F37D0EB19F3FA34C05FC58E0B" day="27" month="07" value="1992-07-27" year="1992">27 Jul 1992</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="6BE943534E4C5A7E197A877A6912E6C4" type="diagnosis">
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<paragraph id="583B0E9B8C4C6EC1D8C1F15D93173D88" pageNumber="14">
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Diagnosis.
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<taxonomicName id="719B09B48A5C2E5819F0D53D7CA680F2" LSID-ZBK="urn:lsid:zoobank.org:act:CEC6BB92-06C1-4BB1-8E1E-1939B5547CF5" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon venustus Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="14" phylum="Chordata" rank="species" species="venustus">Gastromyzon venustus</taxonomicName>
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differs from its congeners in having the following unique combination of characters: body plain brown, head dorsum plain brown; dorsal, caudal and anal fins red in life; absence of secondary rostrum; absence of postoral pouch; gill slit angular, subopercular groove present and continuous with pectoral-fin origin; a broadly rounded snout when viewed dorsally; presence of sublacrymal groove; absence of scales on abdomen; 58-63 scales in lateral line; pelvic fin just reaching level of anal fin origin, adpressed dorsal fin not overlapping level of anal fin origin. Maximum size: 49.4 mm SL (ZRC 47104).
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</paragraph>
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</subSubSection>
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<subSubSection id="C24E57567D554B1E93CD782993F15E3C" type="description">
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<paragraph id="38FF0B28B7B6B8FD1745BCBA4298C696" lastPageNumber="16" pageNumber="15">Description. General body shape and appearance as in Figs. 5-6. Meristic and morphometric data appear in Table 1. Head broadly rounded when viewed dorsally, relatively short (27.5-28.7 % SL) and wide (27.1-28.5 % SL, 95.6-102.8 % HL), head relatively flattened (head depth 12.4-14.5 % SL, 43.7-52.8 % HL). Snout elongated (snout length 63.7-71.7 % HL). Tubercles present over entire snout. Sublacrymal groove present, not visible from side. Gill slit strongly angular. Subopercular groove pronounced and continuous to origin of pectoral fin base. Postoral pouch absent. Scales absent from belly. Posterior part of pectoral fin not overlapping anterior part of pelvic fin in adults, but overlapping in juveniles. Pectoral and pelvic fins with serrae on anteriormost rays. Dorsal fin situated at about mid body (predorsal length 53.9-58.1 % SL). Adpressed dorsal fin not overlapping level of anal fin origin. Deepest part of body at dorsal fin origin (body depth at dorsal fin origin 16.4-18.2 % SL). Anus situated just behind posterior base of fused pelvic fins. Caudal peduncle relatively deep (11.0-11.8 % SL) and long (6.8-10.7 % SL).</paragraph>
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</subSubSection>
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<subSubSection id="CA84A3FD2248BE13C8521AF9950E6D99" type="description">
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<paragraph id="CB17CC9F6F7B9EB588C29D5BBF340A13" lastPageNumber="18" pageNumber="17">Body pigmentation and life coloration. See Fig. 5. Body plain yellowish brown, ventrum cream, a tiny golden spot on posterior edge of every body scale. Head dorsum plain yellowish brown. Eye with golden iris. Dorsal fin red with 3 black bars, the subdistal bar most distinct, with interradial membranes and margin red; and with antero-basal black spot. Caudal fin red with 3-4 black bars, anterior central part gold, subdistal band most distinct, with interradial membranes and margin red. Anal fin reddish, with 2 faint black bars and reddish margin. Pectoral and pelvic fins yellowish brown, with reddish margins. Pelvic axillary flap light yellowish brown. Choy and Chin (1994: 762) noted that small individuals have a blackish-blue spotted appearance, particularly if seen under bright sunlight. The fins have red edges and the caudal fin has 4 dark vertical bands.</paragraph>
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<paragraph id="43FD303A1E79823AFA064FCB3319961C" pageNumber="18">Colour in alcohol. See Fig. 6. Body dark brown or black, the ventrum cream. Head dorsum black. Dorsal fin red (when freshly preserved) or brown with 3 black bars, the subdistal bar most distinct, with an antero-basal black spot. Caudal fin red (when freshly preserved) or brown with 3-4 black bars, subdistal bar most distinct, hyaline interradial membrane and margin. Anal fin brown with 2 faint black bars, and a hyaline margin. Pectoral and pelvic fins grey, with reddish to hyaline margin. Pelvic axillary flap brown.</paragraph>
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</subSubSection>
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<subSubSection id="4AA86F4CA2E96DA2BAE5CDCB3F040B86" type="discussion">
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<paragraph id="9E4D823A46657110C62ADCB2679C59A3" pageNumber="18">
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Remarks. In addition to the characters mentioned in the diagnosis,
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<taxonomicName id="10E3166598AA651617483EEAF3569158" LSID-ZBK="urn:lsid:zoobank.org:act:CEC6BB92-06C1-4BB1-8E1E-1939B5547CF5" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon venustus Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="venustus">G. venustus</taxonomicName>
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can be further differentiated from other congeners of the
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<taxonomicName id="80A9B2E78377D4D5F9EE2D575C0B848C" LSID-ZBK="urn:lsid:zoobank.org:act:F931BE01-35E3-4B97-AA7B-8376875F99E1" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon pariclavis Tan & Martin-Smith 1998:362" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="pariclavis">G. pariclavis group</taxonomicName>
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by the following characters: plain body pattern (vs. barred body in
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<taxonomicName id="670DB1D8571EF1F0A98F23C79BC855E0" LSID-ZBK="urn:lsid:zoobank.org:act:F931BE01-35E3-4B97-AA7B-8376875F99E1" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon pariclavis Tan & Martin-Smith 1998:362" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="pariclavis">G. pariclavis</taxonomicName>
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and reticulated body pattern in
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<taxonomicName id="42125B2564D3AF5C25E2ED7389CFB5CA" LSID-ZBK="urn:lsid:zoobank.org:act:FD3C88F0-F897-426B-9FB7-3B70788422CD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon embalohensis Rachmatika 1998:652" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="embalohensis">G. embalohensis</taxonomicName>
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); fewer transverse scales than
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<taxonomicName id="5D29AE08FBA15FF3614F1E9EBCCFC9F7" LSID-ZBK="urn:lsid:zoobank.org:act:F931BE01-35E3-4B97-AA7B-8376875F99E1" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon pariclavis Tan & Martin-Smith 1998:362" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="pariclavis">G. pariclavis</taxonomicName>
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(11-13.1.10-12 vs. 15-18.1.14-15); greater dorsal depth than
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<taxonomicName id="BE7BA606DD2175EE2E5B6842A74EFC04" LSID-ZBK="urn:lsid:zoobank.org:act:FD3C88F0-F897-426B-9FB7-3B70788422CD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon embalohensis Rachmatika 1998:652" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="embalohensis">G. embalohensis</taxonomicName>
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(19.5-21.7 vs. 18.2-18.9 % SL); greater body depth than
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<taxonomicName id="3DA608887ED9402390F872D0E3462C93" LSID-ZBK="urn:lsid:zoobank.org:act:FD3C88F0-F897-426B-9FB7-3B70788422CD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon embalohensis Rachmatika 1998:652" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="embalohensis">G. embalohensis</taxonomicName>
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(16.4-18.2 vs. 14.8-15.7 % SL); dorsal fin base length less than
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<taxonomicName id="08F50808AD56C22058DE2D07E1F707A8" LSID-ZBK="urn:lsid:zoobank.org:act:F931BE01-35E3-4B97-AA7B-8376875F99E1" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon pariclavis Tan & Martin-Smith 1998:362" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="pariclavis">G. pariclavis</taxonomicName>
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(17.1-20.2 vs. 20.1-22.4 % SL).
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</paragraph>
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</subSubSection>
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<subSubSection id="1E0449D165FC041C29C1BBDC74F17764" type="distribution">
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<paragraph id="7ADC1B50FC8077AE568D787CE54A720C" pageNumber="18">
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Distribution.
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<taxonomicName id="0C81E400E1214E6E348EA95A8E0EA06B" LSID-ZBK="urn:lsid:zoobank.org:act:CEC6BB92-06C1-4BB1-8E1E-1939B5547CF5" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon venustus Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="venustus">Gastromyzon venustus</taxonomicName>
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is known only from Sungai Belalong, Temburong River basin, Brunei Darussalam (Fig. 7).
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</paragraph>
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</subSubSection>
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<subSubSection id="9FFA20E19C8DDF54CB8DE3552EE29984" type="etymology">
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<paragraph id="5AC3768DD68177F907E03FF46BDAB096" pageNumber="18">Etymology. From the Latin venustus, meaning charming, beautiful. This is in reference to the distinctive life coloration. An adjective.</paragraph>
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</subSubSection>
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<subSubSection id="7FBAA6D77C1636A8D372D5B32F83927D" type="biology_ecology">
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<paragraph id="857F77075AC7CA41BABA1F63E41F2F59" pageNumber="18">
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Field notes. See under
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<taxonomicName id="C349D3A3C52B34F502A93FA2B91EAD98" LSID-ZBK="urn:lsid:zoobank.org:act:D41EEA6C-4212-4E99-B70E-54D86FB848BD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon cranbrooki Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="18" phylum="Chordata" rank="species" species="cranbrooki">G. cranbrooki</taxonomicName>
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for list of syntopic species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |