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<taxonomicName id="C0AA429AFFB0FFCAFF30F38EFDDB9385" authority="(Smith, 1849)" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[151,563,153,180]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergi">
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<emphasis id="35DEE50BFFB0FFCAFF30F38EFE6D9382" bold="true" box="[151,389,153,180]" italics="true" pageId="21" pageNumber="98">Panaspis wahlbergi</emphasis>
(
<bibRefCitation id="633B44E8FFB0FFCAFE33F38EFDC39385" author="Smith, A." box="[404,555,153,179]" pageId="21" pageNumber="98" pagination="1 - 28" refId="ref20825" refString="Smith, A. (1849) Appendix. In: Smith, A. (Ed.), Illustrations of the zoology of South Africa, Reptilia, 1849, pp. 1 - 28 + [i]." type="journal article" year="1849">Smith, 1849</bibRefCitation>
)
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(
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,
<figureCitation id="9F91259CFFB0FFCAFF57F3AAFEDE93EE" box="[240,310,189,216]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStartId-0="21.[151,250,1901,1926]" captionStartId-1="22.[151,250,1681,1706]" captionStartId-2="23.[151,250,1990,2015]" captionTargetBox-0="[151,1436,1051,1882]" captionTargetBox-1="[156,1430,185,1651]" captionTargetBox-2="[237,1358,740,1926]" captionTargetId-0="figure@21.[151,1436,1051,1882]" captionTargetId-1="figure@22.[151,1436,180,1655]" captionTargetId-2="figure@23.[229,1362,733,1929]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURE 13—Preserved specimen of Panaspis whalbergii from Bicuar National Park, Huila Province,Angola (CAS 263401).Photo by Luis M. P. Ceríaco." captionText-1="FIGURE 14—Live photos of P. whalbergii from Bicuar National Park, Huila Province (above, CAS 263401) and near Mumbué, Bié Province (below, AMB 11675). Photos by Luis M. P. Ceríaco." captionText-2="FIGURE 15—Distribution map of Panaspis wahlbergii in Angola." figureDoi-0="http://doi.org/10.5281/zenodo.3693459" figureDoi-1="http://doi.org/10.5281/zenodo.3693461" figureDoi-2="http://doi.org/10.5281/zenodo.3693463" httpUri-0="https://zenodo.org/record/3693459/files/figure.png" httpUri-1="https://zenodo.org/record/3693461/files/figure.png" httpUri-2="https://zenodo.org/record/3693463/files/figure.png" pageId="21" pageNumber="98">1315</figureCitation>
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<taxonomicName id="C0AA429AFFB0FFCAFF30F3E0FDAC9239" ID-CoL="CCQTR" authority="Wahlbergii (Bocage 1895: 52)" authorityName="Wahlbergii (Bocage" authorityYear="1895" box="[151,580,246,271]" class="Reptilia" family="Scincidae" genus="Ablepharus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="35DEE50BFFB0FFCAFF30F3E0FE699239" box="[151,385,247,271]" italics="true" pageId="21" pageNumber="98">Ablepharus Wahlbergii</emphasis>
(
<bibRefCitation id="633B44E8FFB0FFCAFE29F3E0FDD59239" author="Bocage, J. V. B." box="[398,573,246,271]" pageId="21" pageNumber="98" refId="ref16491" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage 1895: 52</bibRefCitation>
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<treatmentCitationGroup id="27BA1E37FFB0FFCAFF30F201FCA49218" box="[151,844,277,302]" pageId="21" pageNumber="98">
<taxonomicName id="C0AA429AFFB0FFCAFF30F201FD8D9218" authority="Boulenger 1905" box="[151,613,277,302]" class="Reptilia" family="Scincidae" genus="Ablepharus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergii">
<emphasis id="35DEE50BFFB0FFCAFF30F201FE979218" box="[151,383,278,302]" italics="true" pageId="21" pageNumber="98">Ablepharus wahlbergii</emphasis>
(
<treatmentCitation id="860B1F08FFB0FFCAFE2BF202FD8D9218" author="Boulenger, G. A." box="[396,613,277,302]" page="111" pageId="21" pageNumber="98" year="1905">
<bibRefCitation id="633B44E8FFB0FFCAFE2BF202FD8D9218" author="Boulenger, G. A." box="[396,613,277,302]" pageId="21" pageNumber="98" pagination="105 - 115" refId="ref17078" refString="Boulenger, G. A. (1905) A list of the batrachians and reptiles collected by Dr. W. J. Ansorge in Angola with descriptions of new species. Annals and Magazine of Natural History, Series 7, 16, 105 - 115, pl. 4. https: // doi. org / 10.1080 / 03745480509443656" type="journal article" year="1905">Boulenger 1905: 111</bibRefCitation>
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; Loveridge 1957: 219)
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<emphasis id="35DEE50BFFB0FFCAFF30F223FE4A927A" box="[151,418,308,332]" italics="true" pageId="21" pageNumber="98">Afroablepharus wahlbergi</emphasis>
(
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<bibRefCitation id="633B44E8FFB0FFCAFE09F223FD6E927B" author="FitzSimons, V. F. M." box="[430,646,308,333]" pageId="21" pageNumber="98" pagination="259 - 274" refId="ref18412" refString="FitzSimons, V. F. M. (1937) Notes on the reptiles and amphibians collected and described from South Africa by Andrew Smith. Annals of the Transvaal Museum, 17, 259 - 274." type="journal article" year="1937">FitzSimons 1937: 269</bibRefCitation>
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;
<treatmentCitation id="860B1F08FFB0FFCAFD28F223FCCA927A" author="Greer, A. E." box="[655,802,308,332]" page="32" pageId="21" pageNumber="98" year="1974">
<bibRefCitation id="633B44E8FFB0FFCAFD28F223FCCA927A" author="Greer, A. E." box="[655,802,308,332]" pageId="21" pageNumber="98" pagination="1 - 67" refId="ref18673" refString="Greer, A. E. (1974) The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology, 31, 1 - 67. https: // doi. org / 10.1071 / ajzs 031" type="journal article" year="1974">Greer 1974: 32</bibRefCitation>
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;
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<bibRefCitation id="633B44E8FFB0FFCAFC8CF222FC5C927A" author="Perret, J. L." box="[811,948,308,333]" pageId="21" pageNumber="98" pagination="5 - 16" refId="ref20205" refString="Perret, J. L. (1975) La differentiation dans le genre Panaspis Cope (Reptilia: Scincidae). Bulletin de la Societe Neuchateloise des Sciences Naturelles, 98, 5 - 16." type="journal article" year="1975">Perret 1975: 8</bibRefCitation>
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;
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<bibRefCitation id="633B44E8FFB0FFCAFC19F223FB75927A" author="Schmitz, A. &amp; Ineich, I. &amp; Chirio, L." box="[958,1181,308,333]" pageId="21" pageNumber="98" pagination="1 - 28" refId="ref20615" refString="Schmitz, A., Ineich, I. &amp; Chirio, L. (2005) Molecular review of the genus Panaspis sensu lato (Reptilia: Scincidae) in Cameroon, with special reference to the status of the proposed subgenera. Zootaxa, 863 (1), 1 - 28. https: // doi. org / 10.11646 / zootaxa. 863.1.1" type="journal article" year="2005">
Schmitz
<emphasis id="35DEE50BFFB0FFCAFBB5F222FBAB927A" box="[1042,1091,308,332]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2005: 19
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;
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Bates
<emphasis id="35DEE50BFFB0FFCAFB45F222FAFB927A" box="[1250,1299,308,332]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2014: 257
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)
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<emphasis id="35DEE50BFFB0FFCAFF30F243FE8A925D" box="[151,354,339,363]" italics="true" pageId="21" pageNumber="98">Panaspis wahlbergi</emphasis>
(
<treatmentCitation id="860B1F08FFB0FFCAFED6F245FDC6925D" author="Branch, W. R." box="[369,558,338,363]" page="159" pageId="21" pageNumber="98" year="1998">
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<bibRefCitation id="633B44E8FFB0FFCAFD9DF245FC94925D" author="Jacobsen, N. H. G. &amp; Broadley, D. G." box="[570,892,338,363]" pageId="21" pageNumber="98" pagination="61 - 71" refId="ref18995" refString="Jacobsen, N. H. G. &amp; Broadley, D. G. (2000) A new species of Panaspis Cope (Reptilia: Scincidae) from southern Africa. African Journal of Herpetology, 49 (1), 61 - 71. https: // doi. org / 10.1080 / 21564574.2000.9650017" type="journal article" year="2000">Jacobsen &amp; Broadley 2000: 63</bibRefCitation>
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;
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<bibRefCitation id="633B44E8FFB0FFCAFC2FF245FB2D925D" author="Branch, W. R. &amp; Conradie, W." box="[904,1221,338,363]" pageId="21" pageNumber="98" pagination="194 - 209" refId="ref16659" refString="Branch, W. R. &amp; Conradie, W. (2015) IV. Herpetofauna da regiao da Lagoa Carumbo. In: Huntley, B. &amp; Francisco, P. (Eds.), Avaliacao Rapida da Biodiversidade da Regiao da Lagoa Carumbo, Lunda Norte-Angola. Ministerio do Ambiente de Angola, Luanda, Angola, pp. 194 - 209." type="book chapter" year="2015">Branch &amp; Conradie 2015: 200</bibRefCitation>
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Medina
<emphasis id="35DEE50BFFB0FFCAFA81F243FAB3925D" box="[1318,1371,339,363]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2016: 410
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</treatmentCitation>
;
<treatmentCitation id="860B1F08FFB0FFCAFF5CF266FE1D92BC" author="Branch, W. R. &amp; Baptista, N. &amp; Conradie, W." box="[251,501,369,394]" page="296" pageId="21" pageNumber="98" year="2019">
<bibRefCitation id="633B44E8FFB0FFCAFF5CF266FE1D92BC" author="Branch, W. R. &amp; Baptista, N. &amp; Conradie, W." box="[251,501,369,394]" pageId="21" pageNumber="98" pagination="283 - 334" refId="ref16727" refString="Branch, W. R., Pinto. P. V., Baptista, N. &amp; Conradie, W. (2019 a) The Reptiles of Angola: History, Diversity, Endemism and Hotspots. In: Huntley, B, Russo, V., Lages &amp; Ferrand, N. (Eds.) Biodiversity of Angola, Science &amp; Conservation: A Modern Synthesis. Springer, Cham, pp. 283 - 334. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 13" type="book chapter" year="2019">
Branch
<emphasis id="35DEE50BFFB0FFCAFEEDF265FE9592BC" box="[330,381,370,394]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2019a: 296
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)
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</paragraph>
<paragraph id="07153919FFB0FFCAFF30F286FC4D929E" blockId="21.[151,1437,246,486]" box="[151,933,400,425]" pageId="21" pageNumber="98">
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<taxonomicName id="C0AA429AFFB0FFCAFF30F286FD70929E" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[151,664,400,425]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB0FFCAFF30F286FF1C929E" box="[151,244,401,424]" italics="true" pageId="21" pageNumber="98">Panaspis</emphasis>
aff.
<emphasis id="35DEE50BFFB0FFCAFE84F287FE61929E" box="[291,393,400,424]" italics="true" pageId="21" pageNumber="98">wahlbergi</emphasis>
(
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<bibRefCitation id="633B44E8FFB0FFCAFE31F287FD70929E" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[406,664,400,424]" pageId="21" pageNumber="98" pagination="1 - 501" refId="ref19496" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="35DEE50BFFB0FFCAFE51F286FDC2929E" box="[502,554,400,424]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2018: 249
</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="860B1F08FFB0FFCAFD04F287FC75929E" author="Branch, W. R. &amp; Baptista, N. &amp; Conradie, W." box="[675,925,400,425]" page="318" pageId="21" pageNumber="98" year="2019">
<bibRefCitation id="633B44E8FFB0FFCAFD04F287FC75929E" author="Branch, W. R. &amp; Baptista, N. &amp; Conradie, W." box="[675,925,400,425]" pageId="21" pageNumber="98" pagination="283 - 334" refId="ref16727" refString="Branch, W. R., Pinto. P. V., Baptista, N. &amp; Conradie, W. (2019 a) The Reptiles of Angola: History, Diversity, Endemism and Hotspots. In: Huntley, B, Russo, V., Lages &amp; Ferrand, N. (Eds.) Biodiversity of Angola, Science &amp; Conservation: A Modern Synthesis. Springer, Cham, pp. 283 - 334. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 13" type="book chapter" year="2019">
Branch
<emphasis id="35DEE50BFFB0FFCAFD55F286FCCD929E" box="[754,805,400,424]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2019a: 318
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)
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</paragraph>
<paragraph id="07153919FFB0FFCAFF30F2A7FDCF92F1" blockId="21.[151,1437,246,486]" box="[151,551,430,455]" pageId="21" pageNumber="98">
<treatmentCitationGroup id="27BA1E37FFB0FFCAFF30F2A7FDCF92F1" box="[151,551,430,455]" pageId="21" pageNumber="98">
<taxonomicName id="C0AA429AFFB0FFCAFF30F2A7FDCF92F1" ID-CoL="CDTZ3" authority="Baptista et al. 2018" box="[151,551,430,455]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="35DEE50BFFB0FFCAFF30F2A7FF1C92F1" box="[151,244,432,455]" italics="true" pageId="21" pageNumber="98">Panaspis</emphasis>
sp. (
<treatmentCitation id="860B1F08FFB0FFCAFE83F2B9FDC892F1" author="Baptista, N. &amp; Antonio, T. &amp; Branch, W. R." box="[292,544,430,455]" page="400" pageId="21" pageNumber="98" year="2018">
<bibRefCitation id="633B44E8FFB0FFCAFE83F2B9FDC892F1" author="Baptista, N. &amp; Antonio, T. &amp; Branch, W. R." box="[292,544,430,455]" pageId="21" pageNumber="98" pagination="397 - 403" refId="ref15989" refString="Baptista, N., Antonio, T. &amp; Branch, W. R. (2018) Amphibians and reptiles of the Tundavala region of the Angolan Escarpment. Biodiversity &amp; Ecology, 6, 397 - 403. https: // doi. org / 10.7809 / b-e. 00351" type="journal article" year="2018">
Baptista
<emphasis id="35DEE50BFFB0FFCAFED8F2A7FE5B92F1" box="[383,435,431,455]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2018: 400
</bibRefCitation>
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)
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="07153919FFB0FFCAFF30F2D9FD6392D0" blockId="21.[151,1437,246,486]" box="[151,651,461,486]" pageId="21" pageNumber="98">
<treatmentCitationGroup id="27BA1E37FFB0FFCAFF30F2D9FD6392D0" box="[151,651,461,486]" pageId="21" pageNumber="98">
<treatmentCitation id="860B1F08FFB0FFCAFF30F2D9FD6392D0" author="Butler, B. O. &amp; Ceriaco, L. M. &amp; Marques, M. P. &amp; Bandeira, S. &amp; Julio, T. &amp; Heinicke, M. P. &amp; Bauer, A. M." box="[151,651,461,486]" page="233" pageId="21" pageNumber="98" year="2019">
<taxonomicName id="C0AA429AFFB0FFCAFF30F2D9FD6392D0" ID-CoL="8H6GW" authority="(Butler et al. 2019: 233)" baseAuthorityName="Butler" baseAuthorityPageNumber="233" baseAuthorityYear="2019" box="[151,651,461,486]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="namibiana">
<emphasis id="35DEE50BFFB0FFCAFF30F2D9FF1C92D3" box="[151,244,462,485]" italics="true" pageId="21" pageNumber="98">Panaspis</emphasis>
aff.
<emphasis id="35DEE50BFFB0FFCAFE84F2D9FE6792D0" box="[291,399,462,486]" italics="true" pageId="21" pageNumber="98">namibiana</emphasis>
(
<bibRefCitation id="633B44E8FFB0FFCAFE3BF2DAFD6C92D0" author="Butler, B. O. &amp; Ceriaco, L. M. &amp; Marques, M. P. &amp; Bandeira, S. &amp; Julio, T. &amp; Heinicke, M. P. &amp; Bauer, A. M." box="[412,644,461,486]" pageId="21" pageNumber="98" pagination="225 - 240" refId="ref16898" refString="Butler, B. O., Ceriaco, L. M., Marques, M. P., Bandeira, S., Julio, T., Heinicke, M. P. &amp; Bauer, A. M. (2019) Herpetological Survey of Huila Province, Southwest Angola, Including First Records from Bicuar National Park. Herpetological Review, 50, 225 - 240." type="journal article" year="2019">
Butler
<emphasis id="35DEE50BFFB0FFCAFE43F2D9FDFF92D0" box="[484,535,462,486]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2019: 233
</bibRefCitation>
)
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</paragraph>
<paragraph id="07153919FFB0FFCAFF30F11FFDFA972F" blockId="21.[151,1437,519,1049]" pageId="21" pageNumber="98">
<taxonomicName id="C0AA429AFFB0FFCAFF30F11FFEF29117" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[151,282,519,545]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB0FFCAFF30F11FFEF29117" box="[151,282,519,545]" italics="true" pageId="21" pageNumber="98">P. wahlbergi</emphasis>
</taxonomicName>
was originally described as
<taxonomicName id="C0AA429AFFB0FFCAFDE7F110FC8B9117" authorityName="Smith" authorityYear="1849" box="[576,867,519,545]" class="Reptilia" family="Scincidae" genus="Cryptoblepharus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB0FFCAFDE7F110FC8B9117" box="[576,867,519,545]" italics="true" pageId="21" pageNumber="98">Cryptoblepharus wahlbergi</emphasis>
</taxonomicName>
by
<bibRefCitation id="633B44E8FFB0FFCAFC2FF110FBFC9117" author="Smith, A." box="[904,1044,519,545]" pageId="21" pageNumber="98" pagination="1 - 28" refId="ref20825" refString="Smith, A. (1849) Appendix. In: Smith, A. (Ed.), Illustrations of the zoology of South Africa, Reptilia, 1849, pp. 1 - 28 + [i]." type="journal article" year="1849">Smith (1849)</bibRefCitation>
based on specimens from the “Country to the eastward of the
<collectingRegion id="C56EF7FBFFB0FFCAFE01F13CFE359173" box="[422,477,555,581]" country="South Africa" name="Western Cape" pageId="21" pageNumber="98">Cape</collectingRegion>
Colony,” later restricted to
<collectingRegion id="C56EF7FBFFB0FFCAFCA6F13CFBE79173" box="[769,1039,555,581]" country="South Africa" name="KwaZulu-Natal" pageId="21" pageNumber="98">KwaZulu-Natal Province</collectingRegion>
,
<collectingCountry id="7FBD7989FFB0FFCAFBBDF13CFB4B9173" box="[1050,1187,555,581]" name="South Africa" pageId="21" pageNumber="98">South Africa</collectingCountry>
, by
<bibRefCitation id="633B44E8FFB0FFCAFB68F13CFA719173" author="FitzSimons, V. F. M." box="[1231,1433,555,581]" pageId="21" pageNumber="98" pagination="259 - 274" refId="ref18412" refString="FitzSimons, V. F. M. (1937) Notes on the reptiles and amphibians collected and described from South Africa by Andrew Smith. Annals of the Transvaal Museum, 17, 259 - 274." type="journal article" year="1937">FitzSimons (1937)</bibRefCitation>
. Similar to all other members of the genus, the species has a complex taxonomic history, especially regarding its genetic allocation (for a review of its taxonomic and nomenclatural history see
<bibRefCitation id="633B44E8FFB0FFCAFC28F164FBB691BB" author="Schmitz, A. &amp; Ineich, I. &amp; Chirio, L." box="[911,1118,626,653]" pageId="21" pageNumber="98" pagination="1 - 28" refId="ref20615" refString="Schmitz, A., Ineich, I. &amp; Chirio, L. (2005) Molecular review of the genus Panaspis sensu lato (Reptilia: Scincidae) in Cameroon, with special reference to the status of the proposed subgenera. Zootaxa, 863 (1), 1 - 28. https: // doi. org / 10.11646 / zootaxa. 863.1.1" type="journal article" year="2005">
Schmitz
<emphasis id="35DEE50BFFB0FFCAFC4AF163FBCB91BB" box="[1005,1059,627,653]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2005
</bibRefCitation>
and
<bibRefCitation id="633B44E8FFB0FFCAFB34F164FA8891B8" author="Medina, M. F. &amp; Bauer, A. M. &amp; Branch, W. R. &amp; Schmitz, A. &amp; Conradie, W. &amp; Nagy, Z. T. &amp; Hibbitts, T. J. &amp; Portik, D. M. &amp; Nielsen, S. V. &amp; Colston, T. J. &amp; Kusamba, C. &amp; Behangana, M. &amp; Rodel, M. - O." box="[1171,1376,627,654]" pageId="21" pageNumber="98" pagination="409 - 423" refId="ref19741" refString="Medina, M. F., Bauer, A. M., Branch, W. R., Schmitz, A., Conradie, W., Nagy, Z. T., Hibbitts, T. J., Ernst, R., Portik, D. M., Nielsen, S. V., Colston, T. J., Kusamba, C., Behangana, M. &amp; Rodel, M. - O. (2016) Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution, 100, 409 - 423. https: // doi. org / 10.1016 / j. ympev. 2016.04.026" type="journal article" year="2016">
Medina
<emphasis id="35DEE50BFFB0FFCAFB4CF163FAC991BB" box="[1259,1313,627,653]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
2016
</bibRefCitation>
). The first records of this species for
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were provided by
<bibRefCitation id="633B44E8FFB0FFCAFD48F180FC659187" author="Bocage, J. V. B." box="[751,909,662,689]" pageId="21" pageNumber="98" refId="ref16491" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage (1895)</bibRefCitation>
, who noted specimens collected by J.A.O. de Anchieta from Cahata and Caconda, in the higher elevation areas of
<collectingRegion id="C56EF7FBFFB0FFCAFCF2F1ACFC5191E3" box="[853,953,699,725]" country="Angola" name="Benguela" pageId="21" pageNumber="98">Benguela</collectingRegion>
and
<collectingRegion id="C56EF7FBFFB0FFCAFC48F1ACFBC291E3" box="[1007,1066,699,725]" country="Angola" name="Huila" pageId="21" pageNumber="98">Huíla</collectingRegion>
provinces, respectively. As he had done with
<taxonomicName id="C0AA429AFFB0FFCAFEA0F1F7FDC591CF" authority=", Bocage (1895)" authorityName=", Bocage" authorityYear="1895" box="[263,557,734,761]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="cabindae">
<emphasis id="35DEE50BFFB0FFCAFEA0F1F7FE6A91CF" box="[263,386,735,761]" italics="true" pageId="21" pageNumber="98">P. cabindae</emphasis>
,
<bibRefCitation id="633B44E8FFB0FFCAFE28F1C8FDC591CF" author="Bocage, J. V. B." box="[399,557,734,761]" pageId="21" pageNumber="98" refId="ref16491" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage (1895)</bibRefCitation>
</taxonomicName>
provided a detailed illustration of Angolan
<taxonomicName id="C0AA429AFFB0FFCAFC5EF1F7FB9691CF" authorityName="Smith" authorityYear="1849" box="[1017,1150,735,761]" class="Reptilia" family="Scincidae" genus="Cryptoblepharus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB0FFCAFC5EF1F7FB9691CF" box="[1017,1150,735,761]" italics="true" pageId="21" pageNumber="98">P. wahlbergi</emphasis>
</taxonomicName>
(
<figureCitation id="9F91259CFFB0FFCAFB2BF1C8FB2491CF" box="[1164,1228,735,761]" captionStart="FIGURE 7" captionStartId="15.[151,242,1181,1206]" captionTargetBox="[151,1436,272,1142]" captionTargetId="figure@15.[151,1436,272,1142]" captionTargetPageId="15" captionText="FIGURE 7—Plate with the illustrations of Panaspis cabindae and Panaspis whalbergii from Bocage (1895) Herpetologie dAngola et du Congo.Adapted from Bocage (1895)." figureDoi="http://doi.org/10.5281/zenodo.3693445" httpUri="https://zenodo.org/record/3693445/files/figure.png" pageId="21" pageNumber="98">Fig. 7</figureCitation>
).
<bibRefCitation id="633B44E8FFB0FFCAFB47F1C8FA7491CF" author="Boulenger, G. A." box="[1248,1436,734,761]" pageId="21" pageNumber="98" pagination="105 - 115" refId="ref17078" refString="Boulenger, G. A. (1905) A list of the batrachians and reptiles collected by Dr. W. J. Ansorge in Angola with descriptions of new species. Annals and Magazine of Natural History, Series 7, 16, 105 - 115, pl. 4. https: // doi. org / 10.1080 / 03745480509443656" type="journal article" year="1905">Boulenger (1905)</bibRefCitation>
referred to this species specimens collected “between Benguella and Bihé” [between what it is currently understood as
<collectingRegion id="C56EF7FBFFB0FFCAFF30F030FF139077" box="[151,251,807,833]" country="Angola" name="Benguela" pageId="21" pageNumber="98">Benguela</collectingRegion>
and
<collectingRegion id="C56EF7FBFFB0FFCAFE94F030FEB09077" box="[307,344,807,833]" country="Angola" name="Bie" pageId="21" pageNumber="98">Bié</collectingRegion>
provinces] by the British explorer William John Ansorge (
<date id="73141FD9FFB0FFCAFC72F031FBB99077" box="[981,1105,806,833]" pageId="21" pageNumber="98" value="1850" valueMax="1913">18501913</date>
). These are the only published records of the species for
<collectingCountry id="7FBD7989FFB0FFCAFE0EF05CFE129053" box="[425,506,843,869]" name="Angola" pageId="21" pageNumber="98">Angola</collectingCountry>
, although some other specimens assignable to the species were collected in the country during the twentieth century and deposited in the collections of Portuguese (IICT), North American (CAS, FMNH) and South African (TM) museums. More recently the species has been relocated in the country.
<bibRefCitation id="633B44E8FFB0FFCAFBF8F084FAD5909B" author="Baptista, N. &amp; Antonio, T. &amp; Branch, W. R." box="[1119,1341,915,941]" pageId="21" pageNumber="98" pagination="397 - 403" refId="ref15989" refString="Baptista, N., Antonio, T. &amp; Branch, W. R. (2018) Amphibians and reptiles of the Tundavala region of the Angolan Escarpment. Biodiversity &amp; Ecology, 6, 397 - 403. https: // doi. org / 10.7809 / b-e. 00351" type="journal article" year="2018">
Baptista
<emphasis id="35DEE50BFFB0FFCAFB1BF083FB19909B" box="[1212,1265,915,941]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
(2018)
</bibRefCitation>
reported the presence of
<taxonomicName id="C0AA429AFFB0FFCAFE99F0AFFE4890E7" box="[318,416,952,977]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="35DEE50BFFB0FFCAFE99F0AFFE4890E7" box="[318,416,952,977]" italics="true" pageId="21" pageNumber="98">Panaspis</emphasis>
</taxonomicName>
sp. in theTundavala area.
<bibRefCitation id="633B44E8FFB0FFCAFD13F0A0FC9690E7" author="Butler, B. O. &amp; Ceriaco, L. M. &amp; Marques, M. P. &amp; Bandeira, S. &amp; Julio, T. &amp; Heinicke, M. P. &amp; Bauer, A. M." box="[692,894,951,977]" pageId="21" pageNumber="98" pagination="225 - 240" refId="ref16898" refString="Butler, B. O., Ceriaco, L. M., Marques, M. P., Bandeira, S., Julio, T., Heinicke, M. P. &amp; Bauer, A. M. (2019) Herpetological Survey of Huila Province, Southwest Angola, Including First Records from Bicuar National Park. Herpetological Review, 50, 225 - 240." type="journal article" year="2019">
Butler
<emphasis id="35DEE50BFFB0FFCAFD5AF0AFFCDA90E7" box="[765,818,951,977]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
(2019)
</bibRefCitation>
reported one specimen from Bicuar National Park, which they mistakenly labeled as
<taxonomicName id="C0AA429AFFB0FFCAFDA3F0CBFD5390C3" box="[516,699,987,1013]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="namibiana">
<emphasis id="35DEE50BFFB0FFCAFDA3F0CBFDF090C3" box="[516,536,988,1013]" italics="true" pageId="21" pageNumber="98">P.</emphasis>
aff.
<emphasis id="35DEE50BFFB0FFCAFDEDF0CCFD5390C3" box="[586,699,987,1013]" italics="true" pageId="21" pageNumber="98">namibiana</emphasis>
</taxonomicName>
. Other specimens were recently collected by the authors LMPC and MPM in southeastern
<collectingRegion id="C56EF7FBFFB0FFCAFE26F0E8FDE6972F" box="[385,526,1023,1049]" country="Angola" name="Bie" pageId="21" pageNumber="98">Bié Province</collectingRegion>
.
</paragraph>
<caption id="53D56991FFB0FFCAFF30F47AFEF0949C" ID-DOI="http://doi.org/10.5281/zenodo.3693459" ID-Zenodo-Dep="3693459" httpUri="https://zenodo.org/record/3693459/files/figure.png" pageId="21" pageNumber="98" startId="21.[151,250,1901,1926]" targetBox="[151,1436,1051,1882]" targetPageId="21">
<paragraph id="07153919FFB0FFCAFF30F47AFEF0949C" blockId="21.[151,1437,1901,2035]" pageId="21" pageNumber="98">
<emphasis id="35DEE50BFFB0FFCAFF30F47AFEDB94B0" bold="true" box="[151,307,1901,1926]" pageId="21" pageNumber="98">FIGURE 13—</emphasis>
Preserved specimen of
<taxonomicName id="C0AA429AFFB0FFCAFDA5F479FD5294B0" box="[514,698,1902,1926]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="whalbergii">
<emphasis id="35DEE50BFFB0FFCAFDA5F479FD5294B0" box="[514,698,1902,1926]" italics="true" pageId="21" pageNumber="98">Panaspis whalbergii</emphasis>
</taxonomicName>
from Bicuar National Park, Huila Province,Angola (CAS 263401).Photo by Luis M. P. Ceríaco.
</paragraph>
</caption>
<paragraph id="07153919FFB0FFC9FF60F4A3FA8F94D8" blockId="21.[151,1437,1901,2035]" lastBlockId="22.[151,1437,1788,2031]" lastPageId="22" lastPageNumber="99" pageId="21" pageNumber="98">
Both
<bibRefCitation id="633B44E8FFB0FFCAFEA6F4A2FE0A94F8" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[257,482,1972,1999]" pageId="21" pageNumber="98" pagination="1 - 501" refId="ref19496" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="35DEE50BFFB0FFCAFEC2F4A2FE7F94F8" box="[357,407,1972,1998]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
(2018)
</bibRefCitation>
and Branch
<emphasis id="35DEE50BFFB0FFCAFDCEF4A2FD7794F8" box="[617,671,1972,1998]" italics="true" pageId="21" pageNumber="98">et al.</emphasis>
(2019) referred to the central Angolan populations of
<taxonomicName id="C0AA429AFFB0FFCAFB7EF4A2FA8A94F8" authorityName="Smith" authorityYear="1849" box="[1241,1378,1972,1998]" class="Reptilia" family="Scincidae" genus="Cryptoblepharus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB0FFCAFB7EF4A2FA8A94F8" box="[1241,1378,1972,1998]" italics="true" pageId="21" pageNumber="98">P. wahlbergi</emphasis>
</taxonomicName>
as
<taxonomicName id="C0AA429AFFB0FFCAFA22F4A2FEC194C4" authorityName="Smith" authorityYear="1849" class="Reptilia" family="Scincidae" genus="Cryptoblepharus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="98" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB0FFCAFA22F4A2FA7594F8" box="[1413,1437,1973,1998]" italics="true" pageId="21" pageNumber="98">P.</emphasis>
aff.
<emphasis id="35DEE50BFFB0FFCAFF18F4CFFEC194C4" box="[191,297,2008,2034]" italics="true" pageId="21" pageNumber="98">wahlbergi</emphasis>
</taxonomicName>
, suggesting doubts about its conspecificity with the nominotypical form, and even suggesting that the Angolan population may represent a different taxon (
<bibRefCitation id="633B44E8FFB3FFC9FDE8F5EBFCF19420" author="Branch, W. R. &amp; Baptista, N. &amp; Conradie, W." box="[591,793,1788,1814]" pageId="22" pageNumber="99" pagination="283 - 334" refId="ref16727" refString="Branch, W. R., Pinto. P. V., Baptista, N. &amp; Conradie, W. (2019 a) The Reptiles of Angola: History, Diversity, Endemism and Hotspots. In: Huntley, B, Russo, V., Lages &amp; Ferrand, N. (Eds.) Biodiversity of Angola, Science &amp; Conservation: A Modern Synthesis. Springer, Cham, pp. 283 - 334. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 13" type="book chapter" year="2019">
Branch
<emphasis id="35DEE50BFFB3FFC9FD07F5EAFD3B9420" box="[672,723,1788,1814]" italics="true" pageId="22" pageNumber="99">et al.</emphasis>
2019a
</bibRefCitation>
). However, contrary to what
<bibRefCitation id="633B44E8FFB3FFC9FB99F5EBFAE69421" author="Medina, M. F. &amp; Bauer, A. M. &amp; Branch, W. R. &amp; Schmitz, A. &amp; Conradie, W. &amp; Nagy, Z. T. &amp; Hibbitts, T. J. &amp; Portik, D. M. &amp; Nielsen, S. V. &amp; Colston, T. J. &amp; Kusamba, C. &amp; Behangana, M. &amp; Rodel, M. - O." box="[1086,1294,1788,1815]" pageId="22" pageNumber="99" pagination="409 - 423" refId="ref19741" refString="Medina, M. F., Bauer, A. M., Branch, W. R., Schmitz, A., Conradie, W., Nagy, Z. T., Hibbitts, T. J., Ernst, R., Portik, D. M., Nielsen, S. V., Colston, T. J., Kusamba, C., Behangana, M. &amp; Rodel, M. - O. (2016) Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution, 100, 409 - 423. https: // doi. org / 10.1016 / j. ympev. 2016.04.026" type="journal article" year="2016">
Medina
<emphasis id="35DEE50BFFB3FFC9FB34F5EAFB2E9420" box="[1171,1222,1788,1814]" italics="true" pageId="22" pageNumber="99">et al.</emphasis>
(2016)
</bibRefCitation>
demonstrated for the eastern African populations, where a high level of cryptic diversity exists, and to the suspicions of
<bibRefCitation id="633B44E8FFB3FFC9FB3BF436FA99940C" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[1180,1393,1824,1851]" pageId="22" pageNumber="99" pagination="1 - 501" refId="ref19496" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="35DEE50BFFB3FFC9FB5DF436FAC3940C" box="[1274,1323,1824,1850]" italics="true" pageId="22" pageNumber="99">et al.</emphasis>
(2018)
</bibRefCitation>
and Branch
<emphasis id="35DEE50BFFB3FFC9FF41F452FEF09468" box="[230,280,1860,1886]" italics="true" pageId="22" pageNumber="99">et al.</emphasis>
(2019), our results unambiguously place the central Angolan population as conspecific with nominotypic
<taxonomicName id="C0AA429AFFB3FFC9FA96F453FA7D9468" authorityName="Smith" authorityYear="1849" box="[1329,1429,1860,1886]" class="Reptilia" family="Scincidae" genus="Cryptoblepharus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="99" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB3FFC9FA96F453FA7D9468" box="[1329,1429,1860,1886]" italics="true" pageId="22" pageNumber="99">wahlbergi</emphasis>
</taxonomicName>
. The presence in the country of additional undescribed
<taxonomicName id="C0AA429AFFB3FFC9FD14F47FFCFF94B4" authorityName="Smith" authorityYear="1849" box="[691,791,1896,1922]" class="Reptilia" family="Scincidae" genus="Cryptoblepharus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="99" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB3FFC9FD14F47FFCFF94B4" box="[691,791,1896,1922]" italics="true" pageId="22" pageNumber="99">wahlbergi</emphasis>
</taxonomicName>
-like lineages cannot be excluded, as
<bibRefCitation id="633B44E8FFB3FFC9FB2FF47FFA9294B4" author="Bittencourt-Silva, G. B." box="[1160,1402,1896,1922]" pageId="22" pageNumber="99" pagination="1 - 28" refId="ref16291" refString="Bittencourt-Silva, G. B. (2019) A herpetological survey of western Zambia. Amphibian and Reptile Conservation, 13 (2), 1 - 28." type="journal article" year="2019">Bittencourt-Silva (2019)</bibRefCitation>
recently reported the presence of one of these lineages (“
<taxonomicName id="C0AA429AFFB3FFC9FD09F49AFCE39490" box="[686,779,1933,1958]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="35DEE50BFFB3FFC9FD09F49AFCE39490" box="[686,779,1933,1958]" italics="true" pageId="22" pageNumber="99">Panaspis</emphasis>
</taxonomicName>
sp. Katanga 2” sensu
<bibRefCitation id="633B44E8FFB3FFC9FC45F49BFB759491" author="Medina, M. F. &amp; Bauer, A. M. &amp; Branch, W. R. &amp; Schmitz, A. &amp; Conradie, W. &amp; Nagy, Z. T. &amp; Hibbitts, T. J. &amp; Portik, D. M. &amp; Nielsen, S. V. &amp; Colston, T. J. &amp; Kusamba, C. &amp; Behangana, M. &amp; Rodel, M. - O." box="[994,1181,1932,1959]" pageId="22" pageNumber="99" pagination="409 - 423" refId="ref19741" refString="Medina, M. F., Bauer, A. M., Branch, W. R., Schmitz, A., Conradie, W., Nagy, Z. T., Hibbitts, T. J., Ernst, R., Portik, D. M., Nielsen, S. V., Colston, T. J., Kusamba, C., Behangana, M. &amp; Rodel, M. - O. (2016) Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution, 100, 409 - 423. https: // doi. org / 10.1016 / j. ympev. 2016.04.026" type="journal article" year="2016">
Medina
<emphasis id="35DEE50BFFB3FFC9FB92F49AFB8E9490" box="[1077,1126,1932,1958]" italics="true" pageId="22" pageNumber="99">et al.</emphasis>
2016
</bibRefCitation>
) in northwestern
<collectingCountry id="7FBD7989FFB3FFC9FAEFF49BFA709490" box="[1352,1432,1932,1958]" name="Zambia" pageId="22" pageNumber="99">Zambia</collectingCountry>
, not far from the Angolan border. Specimens from Calombe-Luso, Moxico Province, (IICT 2681959 and 2761959) could potentially belong to this lineage as well. Further sampling in the eastern regions of the country is needed to address this possibility.
</paragraph>
</subSubSection>
<caption id="53D56991FFB3FFC9FF30F586FC8D95F8" ID-DOI="http://doi.org/10.5281/zenodo.3693461" ID-Zenodo-Dep="3693461" httpUri="https://zenodo.org/record/3693461/files/figure.png" pageId="22" pageNumber="99" startId="22.[151,250,1681,1706]" targetBox="[156,1430,185,1651]" targetPageId="22">
<paragraph id="07153919FFB3FFC9FF30F586FC8D95F8" blockId="22.[151,1436,1681,1742]" pageId="22" pageNumber="99">
<emphasis id="35DEE50BFFB3FFC9FF30F586FEDB959C" bold="true" box="[151,307,1681,1706]" pageId="22" pageNumber="99">FIGURE 14—</emphasis>
Live photos of
<taxonomicName id="C0AA429AFFB3FFC9FE68F584FDBE959C" box="[463,598,1682,1706]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="99" phylum="Chordata" rank="species" species="whalbergii">
<emphasis id="35DEE50BFFB3FFC9FE68F584FDBE959C" box="[463,598,1682,1706]" italics="true" pageId="22" pageNumber="99">P. whalbergii</emphasis>
</taxonomicName>
from Bicuar National Park, Huila Province (above, CAS 263401) and near Mumbué, Bié Province (below, AMB 11675). Photos by Luis M. P. Ceríaco.
</paragraph>
</caption>
<subSubSection id="4FB06A92FFB2FFC8FF60F38FFF1B912A" pageId="23" pageNumber="100" type="diagnosis">
<paragraph id="07153919FFB2FFC8FF60F38FFF019275" blockId="23.[151,1437,152,719]" pageId="23" pageNumber="100">
<emphasis id="35DEE50BFFB2FFC8FF60F38FFED09385" bold="true" box="[199,312,152,179]" pageId="23" pageNumber="100">Diagnosis.</emphasis>
<taxonomicName id="C0AA429AFFB2FFC8FEE5F38DFDFE9385" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[322,534,153,179]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="100" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB2FFC8FEE5F38DFDFE9385" box="[322,534,153,179]" italics="true" pageId="23" pageNumber="100">Panaspis wahlbergi</emphasis>
</taxonomicName>
can be distinguished from other members of the genus occurring in
<collectingCountry id="7FBD7989FFB2FFC8FAA6F38EFAA69385" box="[1281,1358,153,179]" name="Angola" pageId="23" pageNumber="100">Angola</collectingCountry>
by the following combination of characteristics: 1) absence of supranasals; 2) ablepharine eye (as defined by
<bibRefCitation id="633B44E8FFB2FFC8FB67F3AAFAD193E1" author="Greer, A. E." box="[1216,1337,189,215]" pageId="23" pageNumber="100" pagination="1 - 67" refId="ref18673" refString="Greer, A. E. (1974) The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology, 31, 1 - 67. https: // doi. org / 10.1071 / ajzs 031" type="journal article" year="1974">Greer 1974</bibRefCitation>
); 3) frontoparietals fused; 4) dorsum coppery-brown, with a broad black lateral band; 5) absence of rows of light spots on the neck; 6) usually a presence of a white ventrolateral stripe, from supralabials to the groin; 7) 24 to 26 midbody scales rows (
<figureCitation id="9F91259CFFB2FFC8FAC0F212FF349275" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="21.[151,250,1901,1926]" captionStartId-1="22.[151,250,1681,1706]" captionTargetBox-0="[151,1436,1051,1882]" captionTargetBox-1="[156,1430,185,1651]" captionTargetId-0="figure@21.[151,1436,1051,1882]" captionTargetId-1="figure@22.[151,1436,180,1655]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 13—Preserved specimen of Panaspis whalbergii from Bicuar National Park, Huila Province,Angola (CAS 263401).Photo by Luis M. P. Ceríaco." captionText-1="FIGURE 14—Live photos of P. whalbergii from Bicuar National Park, Huila Province (above, CAS 263401) and near Mumbué, Bié Province (below, AMB 11675). Photos by Luis M. P. Ceríaco." figureDoi-0="http://doi.org/10.5281/zenodo.3693459" figureDoi-1="http://doi.org/10.5281/zenodo.3693461" httpUri-0="https://zenodo.org/record/3693459/files/figure.png" httpUri-1="https://zenodo.org/record/3693461/files/figure.png" pageId="23" pageNumber="100">Figs. 1314</figureCitation>
).
</paragraph>
<paragraph id="07153919FFB2FFC8FF60F25BFF1B912A" blockId="23.[151,1437,152,719]" pageId="23" pageNumber="100">
<emphasis id="35DEE50BFFB2FFC8FF60F25BFE209251" bold="true" box="[199,456,332,359]" pageId="23" pageNumber="100">Specimens examined.</emphasis>
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<emphasis id="35DEE50BFFB2FFC8FE68F25BFCC39251" bold="true" box="[463,811,332,359]" pageId="23" pageNumber="100">
<collectingCountry id="7FBD7989FFB2FFC8FE68F25BFDA69251" box="[463,590,332,359]" name="Angola" pageId="23" pageNumber="100">ANGOLA</collectingCountry>
:
<collectingRegion id="C56EF7FBFFB2FFC8FDFCF25AFCC09251" box="[603,808,332,359]" country="Angola" name="Benguela" pageId="23" pageNumber="100">Benguela Province</collectingRegion>
:
</emphasis>
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[-13.03333º, 14.25,
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;
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<emphasis id="35DEE50BFFB2FFC8FAB0F25BFF1592BD" bold="true" pageId="23" pageNumber="100">
<collectingRegion id="C56EF7FBFFB2FFC8FAB0F25BFF1192BD" country="Angola" name="Kwanza Sul" pageId="23" pageNumber="100">Kwanza-Sul Province</collectingRegion>
:
</emphasis>
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SE of Quirimbo
</location>
[-10.83998º, 14.308124º,
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85966)
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;
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<emphasis id="35DEE50BFFB2FFC8FC53F267FB6B92BD" bold="true" box="[1012,1155,368,395]" pageId="23" pageNumber="100">
<collectingRegion id="C56EF7FBFFB2FFC8FC53F267FB9792BD" box="[1012,1151,368,395]" country="Angola" name="Bie" pageId="23" pageNumber="100">Bié Province</collectingRegion>
:
</emphasis>
near
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[-13.85417º, 17.47044º,
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</quantity>
] (
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)
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;
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[-11.43333º, 17.15º,
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</quantity>
]
<emphasis id="35DEE50BFFB2FFC8FC0BF283FC5C9299" bold="true" box="[940,948,404,431]" pageId="23" pageNumber="100">(</emphasis>
<specimenCode id="570C9162FFB2FFC8FC13F282FBAE9299" box="[948,1094,404,431]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="23" pageNumber="100" type="Museum">FMNH 74295</specimenCode>
,
<specimenCode id="570C9162FFB2FFC8FBEAF282FB0E9299" box="[1101,1254,404,431]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="100">AMNH 48505</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="B7C23344FFB2FFC8FB53F283FECA92C1" ID-GBIF-Occurrence="2572305921" collectionCode="NHM, CAS" country="Angola" elevation="1675" location="Caconda" pageId="23" pageNumber="100" specimenCount="1" stateProvince="Huila">
<emphasis id="35DEE50BFFB2FFC8FB53F283FA749299" bold="true" box="[1268,1436,404,431]" pageId="23" pageNumber="100">
<collectingRegion id="C56EF7FBFFB2FFC8FB53F283FA709299" box="[1268,1432,404,431]" country="Angola" name="Huila" pageId="23" pageNumber="100">Huíla Province</collectingRegion>
:
</emphasis>
<location id="02756FC2FFB2FFC8FF30F2AEFF0792E5" LSID="urn:lsid:plazi:treatment:8F03880FFFB0FFC7FF30F38EFDA1925E:02756FC2FFB2FFC8FF30F2AEFF0792E5" box="[151,239,441,467]" country="Angola" name="Caconda" pageId="23" pageNumber="100" stateProvince="Huila">Caconda</location>
[-13.73333º, 15.06667º,
<quantity id="C05294FCFFB2FFC8FE48F2AFFDA992E5" box="[495,577,440,467]" metricMagnitude="3" metricUnit="m" metricValue="1.675" pageId="23" pageNumber="100" unit="m" value="1675.0">
<elevation id="8C87DE2AFFB2FFC8FE48F2AFFDA992E5" box="[495,577,440,467]" metricMagnitude="3" metricUnit="m" metricValue="1.675" pageId="23" pageNumber="100" unit="m" value="1675.0">1675 m</elevation>
</quantity>
]
<emphasis id="35DEE50BFFB2FFC8FDE8F2AFFDBF92E5" bold="true" box="[591,599,440,467]" pageId="23" pageNumber="100">(</emphasis>
<collectionCode id="61BBA1DCFFB2FFC8FDF1F2AEFD7B92E5" box="[598,659,441,467]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="23" pageNumber="100" type="Herbarium">NHM</collectionCode>
1906.8.24.72);
<locationDeviation id="16A7203BFFB2FFC8FC92F2AEFBD092E5" box="[821,1080,441,467]" location="Lubango" pageId="23" pageNumber="100">
<quantity id="C05294FCFFB2FFC8FC92F2AEFC6592E5" box="[821,909,441,467]" metricMagnitude="4" metricUnit="m" metricValue="3.0577536" pageId="23" pageNumber="100" unit="mi" value="19.0">19 miles</quantity>
NE of Lubango
</locationDeviation>
[-14.78503º, 13.730408º,
<quantity id="C05294FCFFB2FFC8FAE4F2AEFA7D92E5" box="[1347,1429,440,467]" metricMagnitude="3" metricUnit="m" metricValue="1.691" pageId="23" pageNumber="100" unit="m" value="1691.0">
<elevation id="8C87DE2AFFB2FFC8FAE4F2AEFA7D92E5" box="[1347,1429,440,467]" metricMagnitude="3" metricUnit="m" metricValue="1.691" pageId="23" pageNumber="100" unit="m" value="1691.0">1691 m</elevation>
</quantity>
] (
<collectionCode id="61BBA1DCFFB2FFC8FF39F2CAFF3992C1" box="[158,209,477,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="100" type="Museum">CAS</collectionCode>
85956)
</materialsCitation>
;
<materialsCitation id="B7C23344FFB2FFC8FE8DF2CAFCB892C1" ID-GBIF-Occurrence="2572305913" box="[298,848,476,503]" collectionCode="FMNH" country="Angola" elevation="1760" location="Lubango" pageId="23" pageNumber="100" specimenCode="FMNH 74295" specimenCount="1" stateProvince="Huila">
<location id="02756FC2FFB2FFC8FE8DF2CAFE6C92C1" LSID="urn:lsid:plazi:treatment:8F03880FFFB0FFC7FF30F38EFDA1925E:02756FC2FFB2FFC8FE8DF2CAFE6C92C1" box="[298,388,477,503]" country="Angola" name="Lubango" pageId="23" pageNumber="100" stateProvince="Huila">Lubango</location>
[-14.916667º, 13.5º,
<quantity id="C05294FCFFB2FFC8FDF3F2CAFD4A92C1" box="[596,674,476,503]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="23" pageNumber="100" unit="m" value="1760.0">
<elevation id="8C87DE2AFFB2FFC8FDF3F2CAFD4A92C1" box="[596,674,476,503]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="23" pageNumber="100" unit="m" value="1760.0">1760 m</elevation>
</quantity>
] (
<specimenCode id="570C9162FFB2FFC8FD12F2CAFCA192C1" box="[693,841,476,503]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="23" pageNumber="100" type="Museum">FMNH 74295</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="B7C23344FFB2FFC8FCFDF2CAFF06912A" ID-GBIF-Occurrence="2572305905" collectionCode="CAS" country="Angola" elevation="1243" location="Bicuar National Park" pageId="23" pageNumber="100" specimenCode="CAS 263401" specimenCount="1" stateProvince="Huila">
<location id="02756FC2FFB2FFC8FCFDF2CAFBC392C1" LSID="urn:lsid:plazi:treatment:8F03880FFFB0FFC7FF30F38EFDA1925E:02756FC2FFB2FFC8FCFDF2CAFBC392C1" box="[858,1067,477,503]" country="Angola" name="Bicuar National Park" pageId="23" pageNumber="100" stateProvince="Huila">Bicuar National Park</location>
[-15.1034º, 14.8311º,
<quantity id="C05294FCFFB2FFC8FAA1F2CAFABD92C1" box="[1286,1365,476,503]" metricMagnitude="3" metricUnit="m" metricValue="1.2429999999999999" pageId="23" pageNumber="100" unit="m" value="1243.0">
<elevation id="8C87DE2AFFB2FFC8FAA1F2CAFABD92C1" box="[1286,1365,476,503]" metricMagnitude="3" metricUnit="m" metricValue="1.2429999999999999" pageId="23" pageNumber="100" unit="m" value="1243.0">1243 m</elevation>
</quantity>
] (
<specimenCode id="570C9162FFB2FFC8FACFF2CAFF0F912A" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="100" type="Museum">CAS 263401</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FB06A92FFB2FFC7FF60F133FDA1925E" lastPageId="24" lastPageNumber="101" pageId="23" pageNumber="100" type="distribution">
<paragraph id="07153919FFB2FFC7FF60F133FEBF93CD" blockId="23.[151,1437,152,719]" lastBlockId="24.[151,1437,153,251]" lastPageId="24" lastPageNumber="101" pageId="23" pageNumber="100">
<emphasis id="35DEE50BFFB2FFC8FF60F133FE899109" bold="true" box="[199,353,548,575]" pageId="23" pageNumber="100">Distribution.</emphasis>
<bibRefCitation id="633B44E8FFB2FFC8FECFF132FDA39176" author="Medina, M. F. &amp; Bauer, A. M. &amp; Branch, W. R. &amp; Schmitz, A. &amp; Conradie, W. &amp; Nagy, Z. T. &amp; Hibbitts, T. J. &amp; Portik, D. M. &amp; Nielsen, S. V. &amp; Colston, T. J. &amp; Kusamba, C. &amp; Behangana, M. &amp; Rodel, M. - O." box="[360,587,549,576]" pageId="23" pageNumber="100" pagination="409 - 423" refId="ref19741" refString="Medina, M. F., Bauer, A. M., Branch, W. R., Schmitz, A., Conradie, W., Nagy, Z. T., Hibbitts, T. J., Ernst, R., Portik, D. M., Nielsen, S. V., Colston, T. J., Kusamba, C., Behangana, M. &amp; Rodel, M. - O. (2016) Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution, 100, 409 - 423. https: // doi. org / 10.1016 / j. ympev. 2016.04.026" type="journal article" year="2016">
Medina
<emphasis id="35DEE50BFFB2FFC8FE64F131FE139109" box="[451,507,549,575]" italics="true" pageId="23" pageNumber="100">et al.</emphasis>
(2016)
</bibRefCitation>
restricted to
<collectingCountry id="7FBD7989FFB2FFC8FD7CF132FC849109" box="[731,876,549,575]" name="Mozambique" pageId="23" pageNumber="100">Mozambique</collectingCountry>
and northeastern
<collectingCountry id="7FBD7989FFB2FFC8FB96F132FB289109" box="[1073,1216,549,575]" name="South Africa" pageId="23" pageNumber="100">South Africa</collectingCountry>
, but we here extend that distribution into south-central
<collectingCountry id="7FBD7989FFB2FFC8FDB1F15EFD829155" box="[534,618,585,611]" name="Angola" pageId="23" pageNumber="100">Angola</collectingCountry>
. Historical records from
<collectingCountry id="7FBD7989FFB2FFC8FC27F15EFC1E9155" box="[896,1014,585,611]" name="Zimbabwe" pageId="23" pageNumber="100">Zimbabwe</collectingCountry>
and
<collectingCountry id="7FBD7989FFB2FFC8FB8AF15EFB699155" box="[1069,1153,585,611]" name="Zambia" pageId="23" pageNumber="100">Zambia</collectingCountry>
might also represent true
<taxonomicName id="C0AA429AFFB2FFC8FF30F17AFEED91B1" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[151,261,621,647]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="100" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB2FFC8FF30F17AFEED91B1" box="[151,261,621,647]" italics="true" pageId="23" pageNumber="100">wahlbergi</emphasis>
</taxonomicName>
, potentially connecting the Angolan population to those of
<collectingCountry id="7FBD7989FFB2FFC8FC3AF17AFBC691B1" box="[925,1070,621,647]" name="Mozambique" pageId="23" pageNumber="100">Mozambique</collectingCountry>
and
<collectingCountry id="7FBD7989FFB2FFC8FBC1F17AFB1E91B1" box="[1126,1270,621,647]" name="South Africa" pageId="23" pageNumber="100">South Africa</collectingCountry>
. However, this distribution requires further assessment, as other cryptic lineages may also be present, as evidenced by the recent discovery in northwestern
<collectingCountry id="7FBD7989FFB2FFC8FE1BF1A2FDF891F9" box="[444,528,693,719]" name="Zambia" pageId="23" pageNumber="100">Zambia</collectingCountry>
of one of the Katangan cryptic lineages of
<taxonomicName id="C0AA429AFFB2FFC8FC57F1A2FBB691F9" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[1008,1118,693,719]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="100" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFB2FFC8FC57F1A2FBB691F9" box="[1008,1118,693,719]" italics="true" pageId="23" pageNumber="100">wahlbergi</emphasis>
</taxonomicName>
by
<bibRefCitation id="633B44E8FFB2FFC8FB2EF1A2FA7191F9" author="Bittencourt-Silva, G. B." box="[1161,1433,693,719]" pageId="23" pageNumber="100" pagination="1 - 28" refId="ref16291" refString="Bittencourt-Silva, G. B. (2019) A herpetological survey of western Zambia. Amphibian and Reptile Conservation, 13 (2), 1 - 28." type="journal article" year="2019">Bittencourt-Silva (2019)</bibRefCitation>
. In
<collectingCountry id="7FBD7989FFBDFFC7FF14F38EFEE09385" box="[179,264,153,179]" name="Angola" pageId="24" pageNumber="101">Angola</collectingCountry>
,
<taxonomicName id="C0AA429AFFBDFFC7FEB4F38DFE779385" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[275,415,153,179]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="101" phylum="Chordata" rank="species" species="wahlbergi">
<emphasis id="35DEE50BFFBDFFC7FEB4F38DFE779385" box="[275,415,153,179]" italics="true" pageId="24" pageNumber="101">P. wahlbergi</emphasis>
</taxonomicName>
is limited to the central and southern areas of the Angolan plateau, from
<collectingRegion id="C56EF7FBFFBDFFC7FB6EF38EFA759385" box="[1225,1437,153,179]" country="Angola" name="Benguela" pageId="24" pageNumber="101">Benguela Province</collectingRegion>
to
<collectingRegion id="C56EF7FBFFBDFFC7FF13F3AAFF3393E1" box="[180,219,189,215]" country="Angola" name="Bie" pageId="24" pageNumber="101">Bié</collectingRegion>
(
<figureCitation id="9F91259CFFBDFFC7FF4DF3AAFED593E1" box="[234,317,188,215]" captionStart="FIGURE 15" captionStartId="23.[151,250,1990,2015]" captionTargetBox="[237,1358,740,1926]" captionTargetId="figure@23.[229,1362,733,1929]" captionTargetPageId="23" captionText="FIGURE 15—Distribution map of Panaspis wahlbergii in Angola." figureDoi="http://doi.org/10.5281/zenodo.3693463" httpUri="https://zenodo.org/record/3693463/files/figure.png" pageId="24" pageNumber="101">Fig. 15</figureCitation>
), but is most likely that the species also occurs in the neighboring areas of
<collectingRegion id="C56EF7FBFFBDFFC7FB23F3AAFB3393E1" box="[1156,1243,189,215]" country="Angola" name="Moxico" pageId="24" pageNumber="101">Moxico</collectingRegion>
and Cuando-Cubango provinces.
</paragraph>
<caption id="53D56991FFB2FFC8FF30F4D1FCAA94E9" ID-DOI="http://doi.org/10.5281/zenodo.3693463" ID-Zenodo-Dep="3693463" box="[151,834,1990,2015]" httpUri="https://zenodo.org/record/3693463/files/figure.png" pageId="23" pageNumber="100" startId="23.[151,250,1990,2015]" targetBox="[237,1358,740,1926]" targetPageId="23">
<paragraph id="07153919FFB2FFC8FF30F4D1FCAA94E9" blockId="23.[151,834,1990,2015]" box="[151,834,1990,2015]" pageId="23" pageNumber="100">
<emphasis id="35DEE50BFFB2FFC8FF30F4D1FEDB94E9" bold="true" box="[151,307,1990,2015]" pageId="23" pageNumber="100">FIGURE 15—</emphasis>
Distribution map of
<taxonomicName id="C0AA429AFFB2FFC8FDA6F4D0FD3994E9" box="[513,721,1991,2015]" class="Reptilia" family="Scincidae" genus="Panaspis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="100" phylum="Chordata" rank="species" species="wahlbergii">
<emphasis id="35DEE50BFFB2FFC8FDA6F4D0FD3994E9" box="[513,721,1991,2015]" italics="true" pageId="23" pageNumber="100">Panaspis wahlbergii</emphasis>
</taxonomicName>
in Angola.
</paragraph>
</caption>
<paragraph id="07153919FFBDFFC7FF60F23FFDA1925E" blockId="24.[151,1436,296,360]" pageId="24" pageNumber="101">
<emphasis id="35DEE50BFFBDFFC7FF60F23FFD859275" bold="true" box="[199,621,296,323]" pageId="24" pageNumber="101">Habitat and Natural History notes.</emphasis>
The species is found in Miombo Woodlands, usually under leaf-litter, under rocks, or moving through grass (
<figureCitation id="9F91259CFFBDFFC7FE4AF25AFDD4925E" box="[493,572,333,360]" captionStart="FIGURE 16" captionStartId="24.[151,250,1792,1817]" captionTargetBox="[194,1393,384,1761]" captionTargetId="figure@24.[189,1398,380,1766]" captionTargetPageId="24" captionText="FIGURE 16—Habitat of Panaspis wahlbergi. Above Bicuar National Park, Huíla Province. Below Mumbué, Bié Province. Photos by Luis M. P. Ceríaco" figureDoi="http://doi.org/10.5281/zenodo.3693465" httpUri="https://zenodo.org/record/3693465/files/figure.png" pageId="24" pageNumber="101">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>