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<document id="6DF4F04B0C282085ACCBFA2D3095547A" ID-CLB-Dataset="31999" ID-DOI="http://dx.doi.org/10.3897/zookeys.702.14787" ID-GBIF-Dataset="820f03e4-cebc-4d61-8c1c-ddd4c75eb6d9" ID-PMC="PMC5673946" ID-Pensoft-Pub="1313-2970-702-45" ID-PubMed="29118600" ID-ZooBank="C246DC85E2A34ACAB00DF11E56F422E1" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-702-45" ModsDocOrigin="ZooKeys 702" ModsDocTitle="A revision of Meladema diving beetles (Coleoptera, Dytiscidae), with the description of a new species from the central Mediterranean based on molecules and morphology" checkinTime="1506355119895" checkinUser="pensoft" docAuthor="Bilton, David T. &amp; Ribera, Ignacio" docDate="2017" docId="7C910B9C365A6F4520DF4B02A273A89C" docLanguage="en" docName="ZooKeys 702: 45-112" docOrigin="ZooKeys 702" docSource="http://dx.doi.org/10.3897/zookeys.702.14787" docTitle="Meladema lanio Fabricius 1775" docType="treatment" docVersion="6" lastPageNumber="84" masterDocId="FFB6B740184B971E7C751767317DFF8E" masterDocTitle="A revision of Meladema diving beetles (Coleoptera, Dytiscidae), with the description of a new species from the central Mediterranean based on molecules and morphology" masterLastPageNumber="112" masterPageNumber="45" pageNumber="77" updateTime="1701381315638" updateUser="ExternalLinkService">
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<mods:title id="660059457EDEC92408170543622D62EE">A revision of Meladema diving beetles (Coleoptera, Dytiscidae), with the description of a new species from the central Mediterranean based on molecules and morphology</mods:title>
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<mods:namePart id="A50AD663DB099B6B2CE678F1A5CEF68F">Bilton, David T.</mods:namePart>
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<mods:namePart id="C33030AEF4FB8C54F8C22357DC19ADE5">Ribera, Ignacio</mods:namePart>
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<mods:date id="F8C3302CD13CAD2CF3D34B8D9F0F9D74">2017</mods:date>
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<mods:classification id="218525B4A2FEC1821A3E16B4E91822CA">journal article</mods:classification>
<mods:identifier id="5588D19E19D45BC4DDB1CBEBF2588A4F" type="DOI">http://dx.doi.org/10.3897/zookeys.702.14787</mods:identifier>
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<treatment id="7C910B9C365A6F4520DF4B02A273A89C" ID-GBIF-Taxon="134839964" LSID="urn:lsid:plazi:treatment:7C910B9C365A6F4520DF4B02A273A89C" httpUri="http://treatment.plazi.org/id/7C910B9C365A6F4520DF4B02A273A89C" lastPageId="39" lastPageNumber="84" pageId="32" pageNumber="77">
<subSubSection id="E08A4F8C4609E7220329B2A540F65F5B" pageId="32" pageNumber="77" type="nomenclature">
<paragraph id="8655181DAE83394700817370570EA0FA" pageId="32" pageNumber="77">
<taxonomicName id="2823493CF707C91CD24D19C6DC85AB08" ID-CoL="3Z3JX" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Meladema lanio (Fabricius, 1775)</taxonomicName>
Figures 3D, 4D, 5D, 11D, 12D, H, 13D, 14D, 15D, 23, 24B, 26.
</paragraph>
</subSubSection>
<subSubSection id="080623DD8544D8C478D546A73BF53E00" pageId="32" pageNumber="77" type="reference_group">
<paragraph id="21C02343B732D9593783DAB1165257C0" pageId="32" pageNumber="77">
<taxonomicName id="8705F36FFFFD8D9240030B0967979759" class="Insecta" family="Dytiscidae" genus="Dytiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dytiscus lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Dytiscus lanio</taxonomicName>
Fabricius, 1775: 231.
</paragraph>
<paragraph id="CEE917B127B22D5DBA8836C32F47D51A" pageId="32" pageNumber="77">
<taxonomicName id="E12F980DA03706BF45CBE02D08742EA5" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes lowei" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lowei">Colymbetes lowei</taxonomicName>
Gray, 1831: 284.
</paragraph>
<paragraph id="4C3F8AFC0733C75E2C7C2FC0A009DDF8" pageId="32" pageNumber="77">
<taxonomicName id="60A26BC121D41BDDBDB27987B7C53DE6" class="Insecta" family="Dytiscidae" genus="Scutopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutopterus lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Scutopterus lanio</taxonomicName>
(Fabricius, 1775):
<bibRefCitation id="9D024CE2AAB3F00E78F117429FA61D09" author="Dejean, PFMA" journalOrPublisher="Mequignon-Marvis Pere et Fils, Paris" pageId="43" pageNumber="88" title="Catalogue des coleopteres de la collection de M. le comte Dejean. Livraisons 1 &amp; 2." year="1833">Dejean 1833</bibRefCitation>
: 61;
<bibRefCitation id="1A26D1EFE8295B567E3D0CD180FD539F" author="Wollaston, TV" journalOrPublisher="Transactions of the Entomological Society of London" pageId="46" pageNumber="91" pagination="203 - 314" title="On additions to the Atlantic Coleoptera." volume="1871" year="1871">Wollaston 1871</bibRefCitation>
: 221.
</paragraph>
<paragraph id="784840061C0B5B8A020F6F09EE709577" pageId="32" pageNumber="77">
<taxonomicName id="EF0F80DC07B38C5E48C508D7283CE724" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Colymbetes lanio</taxonomicName>
(Fabricius, 1775):
<bibRefCitation id="C3BB2274823F22BFB78D325D0B812136" author="Aube, C" journalOrPublisher="Mequignon Pere et Fils, Paris" pageId="42" pageNumber="87" title="Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean." year="1838">
<normalizedToken id="6DC36C7B4213A8CEE6DF074C423A36BE" originalValue="Aubé">Aube</normalizedToken>
1838
</bibRefCitation>
: 221;
<bibRefCitation id="F2CC54005D515DD68BA5D877932C043B" author="Wollaston, TV" journalOrPublisher="Taylor &amp; Francis, London" pageId="46" pageNumber="91" title="Insecta maderensia; being an account of the insects of the islands of the Madeiran group." year="1854">Wollaston 1854</bibRefCitation>
: 82;
<bibRefCitation id="102E15C8BD90D1B3714F2F1D57590705" author="Wollaston, TV" journalOrPublisher="Taylor &amp; Francis, London" pageId="46" pageNumber="91" title="Coleoptera Atlantidum, being an enumeration of the coleopterous insects of the Madeiras, Salvages, and Canaries." year="1865">Wollaston 1865</bibRefCitation>
:68.
</paragraph>
<paragraph id="28793FAEB4C1A794068FA30EF91A9021" pageId="32" pageNumber="77">
<taxonomicName id="A1F20EA5DBB83CA9888F117E67ABCA16" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Meladema lanio</taxonomicName>
(Fabricius, 1775):
<bibRefCitation id="6487942E05E0EB7A2A585E2400D89545" author="Sharp, D" journalOrPublisher="Scientific Transactions of the Royal Dublin Society" pageId="46" pageNumber="91" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
: 632;
<bibRefCitation id="00D087ECF1106E7436998BC781A7282B" author="Regimbart, M" journalOrPublisher="Memoires de la Societe Entomologique de Belgique" pageId="45" pageNumber="90" pagination="1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines." volume="4" year="1895">
<normalizedToken id="95BF2B64DBAE6AE7FA266280AD96F558" originalValue="Régimbart">Regimbart</normalizedToken>
1895
</bibRefCitation>
: 184; Gschwedntner, 1936: 42;
<normalizedToken id="A5CECDF9E2B8C6E4A5E0E4FEBF9CA859" originalValue="Falkenstöm">Falkenstoem</normalizedToken>
1938: 15;
<bibRefCitation id="216533CD2C12AA36C12BAC4E28BBE1E5" pageId="32" pageNumber="77">Guignot 1961</bibRefCitation>
: 770;
<bibRefCitation id="9D5FA72A813EC3386849641F46FF3814" author="Balke, M" journalOrPublisher="Boletim do Museo Municipal do Funchal" pageId="42" pageNumber="87" pagination="55 - 83" title="Verbreitung. Lebensweise, Taxonomie und Historie der Dytisciden der Ilha da Madeira (Coleoptera, Dytiscidae)." volume="41" year="1989">Balke and Hendrich 1989</bibRefCitation>
: 71.
</paragraph>
</subSubSection>
<subSubSection id="ABA81B83DBEEB67F2148D9F24EAF9C1A" pageId="32" pageNumber="77" type="type locality">
<paragraph id="613C54CC06953A503ED6A0AC5F06B3F9" pageId="32" pageNumber="77">Type locality.</paragraph>
<paragraph id="84A0FC175D4B6A6EBEE712165EE4BD3C" pageId="32" pageNumber="77">&quot;Maderae aquis&quot;</paragraph>
</subSubSection>
<subSubSection id="6B8F318B79CE4CFB355BF7594B6BC6FD" pageId="32" pageNumber="77" type="type material">
<paragraph id="D7D2D2B91DF17CD8E85A9C5F9F88AFF6" pageId="32" pageNumber="77">Type material</paragraph>
<paragraph id="0859323AE18786B1F358AAFAF97615B0" pageId="32" pageNumber="77">
(BMNH). Lectotype ♀ (herein designated, Figure 24B): &quot;
<taxonomicName id="737D3DA668BEC58685DE6AE86334A1BE" class="Insecta" family="Dytiscidae" genus="Dytiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dytiscus lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Dytiscus lanio</taxonomicName>
// Fab. Entom. p. 231. n. 8.&quot; [Latin name &amp; 231. n. 8. HW] &quot;
<taxonomicName id="70CBC6DB0D5B8592D45A7C17EC00B46F" class="Insecta" family="Dytiscidae" genus="Dytiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dytiscus lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Dytiscus lanio</taxonomicName>
// Fabricius, 1775// LECTOTYPE// DT Bilton &amp; I Ribera des. 2017&quot; (dry pinned, BMNH, Banks Collection). Supposed syntypes comprise the above specimen and one other female, located in the Zoological Museum, University of Copenhagen (Zimsen, 1964; A. Solodovnikov, pers. comm.). Sree Selvantharan and Aslak Hansen have kindly communicated a photo of the Copenhagen specimen labelled as
<taxonomicName id="6045D295999D662AD54250EB0359DCF4" lsidName="lanio" pageId="32" pageNumber="77" rank="species" species="lanio">lanio</taxonomicName>
(both on its pin, and above it in the cabinet). This is in fact a species of
<taxonomicName id="B6007AA9A25B74E16C0EF396C1F70527" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Cybister</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="6D680CB518C3D1198B42F4C9ED5B7B55" pageId="32" pageNumber="77" type="materials_examined">
<paragraph id="E335B444C3F607DADB086CD83BCD1B98" pageId="32" pageNumber="77">Additional material examined</paragraph>
<paragraph id="607E75C8D4B6491F4A1880491D50C4BC" pageId="32" pageNumber="77">
(genotyped specimens). Portugal, Madeira: 1 ♂ &quot;vi/1998 PORTUGAL Madeira// Ribeira dos Cedros// L. C. Kelly leg.&quot; &quot;M3 Hand//
<taxonomicName id="EA6D5FE3F466FF03E4209CDF822F535F" lsidName="" order="Coleoptera" pageId="32" pageNumber="77" rank="order">Coleoptera</taxonomicName>
&quot; [HW] &quot;
<taxonomicName id="6932F80E33E60A108B2CF5878D2AF621" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
//
<taxonomicName id="9BDE061F2CEF1F257A21EDAD736A1847" lsidName="lanio" pageId="32" pageNumber="77" rank="species" species="lanio">lanio</taxonomicName>
&quot; [HW] &quot;DNA voucher// NHM-IRM8A&quot; (IBE); 1 ♀ &quot;vi/1998 PORTUGAL Madeira// Levada das Faias// L. C. Kelly leg.&quot; &quot;M2 handsearch//
<taxonomicName id="4B086F8F09874AE4EF3E0D9B3955106C" lsidName="" order="Coleoptera" pageId="32" pageNumber="77" rank="order">Coleoptera</taxonomicName>
&quot; [HW] &quot;
<taxonomicName id="7C8D770EEAE16923C5503963D3F4CD58" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
//
<taxonomicName id="719952A27E0EC2AB68FE3039E8DFB0F9" lsidName="lanio" pageId="32" pageNumber="77" rank="species" species="lanio">lanio</taxonomicName>
&quot; [HW] &quot;DNA voucher// NHM-IRM9A&quot; (IBE); 1 ♀ &quot;vi/1998 PORTUGAL Madeira// Levada das Faias// L. C. Kelly leg.&quot; &quot;DNA voucher// NHM-IRM9B&quot; (IBE); 1 ♂ &quot;Madeira, Canhas// lado sur de Paul da Serra// 1398 m
<geoCoordinate id="DD07342E47FBB80875DEA3F8F7BB6920" direction="north" orientation="latitude" precision="15" value="32.743332">32°44'36&quot;N</geoCoordinate>
<geoCoordinate id="C980962D957E7C54B08CDF074582CE4C" direction="west" orientation="longitude" precision="15" value="-17.09889">17°05'56&quot;W</geoCoordinate>
// charca sobre arroyo de pietras// 19.7.2011 A. Rudoy leg.&quot; &quot;DNA Voucher// IBE-DV298&quot; (IBE). All with &quot;
<taxonomicName id="068140D99A28D5E5CD6081550A3C455A" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
//
<taxonomicName id="E9AA4F6C3FDC53D7D077DA974C23BF57" lsidName="lanio" pageId="32" pageNumber="77" rank="species" species="lanio">lanio</taxonomicName>
(Fabricius, 1775)// D T Bilton [or I Ribera] det. 2017&quot;
</paragraph>
</subSubSection>
<subSubSection id="063A7096A1AC9DEC5C28DE55C7A361E6" lastPageId="33" lastPageNumber="78" pageId="32" pageNumber="77" type="materials_examined">
<paragraph id="F2A540276A5FB2C1DA001F6515DBB238" pageId="32" pageNumber="77">Additional material examined</paragraph>
<paragraph id="CF9034E59A4BCA134A7871E3CE51C009" lastPageId="33" lastPageNumber="78" pageId="32" pageNumber="77">
(non-genotyped specimens). Portugal, Madeira: 1 ♀
<normalizedToken id="AEAC9D4EC9442D650C8C8B6B71450C6A" originalValue="“pouzo”">&quot;pouzo&quot;</normalizedToken>
[HW] &quot;var. squamata&quot; [HW] &quot;passage
<normalizedToken id="7AC9CF0ADB2966F00A180DA470C8496B" originalValue="à">a</normalizedToken>
la m.//
<taxonomicName id="03970FDFA609341B88AE1FB5E577290C" lsidName="imbricatum" pageId="32" pageNumber="77" rank="species" species="imbricatum">imbricatum</taxonomicName>
Woll.&quot; [HW] (ISNB); 1 ♀ &quot;Madeira// 90.32.&quot; [HW] (BMNH); 1 ♀ &quot;Madeira// 90.32.&quot; [HW] &quot;
<taxonomicName id="B4DA1948E611CBD2D60351BA8DB9E10D" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Colymbetes</taxonomicName>
//
<taxonomicName id="F3E28883711B2198C3BAC46889B0F992" lsidName="lanio" pageId="32" pageNumber="77" rank="species" species="lanio">lanio</taxonomicName>
// Fabr.&quot; [HW] (BMNH); 1 ♀
<normalizedToken id="11744D73F7846680EDDD77722AB91FED" originalValue="“Madeira”">&quot;Madeira&quot;</normalizedToken>
reverse &quot;48// 99&quot; [small, circular label, HW] &quot;
<taxonomicName id="ADC121D7D2FA6BDFE3B4D0A8984A90E7" class="Insecta" family="Dytiscidae" genus="Colymbetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colymbetes" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Colymbetes</taxonomicName>
//
<taxonomicName id="023794C6F6549A92224BF5AEA13BCC4C" lsidName="lanio" pageId="32" pageNumber="77" rank="species" species="lanio">lanio</taxonomicName>
Fabr.// Madeira&quot; [HW] (BMNH); 1 ♀
<normalizedToken id="F4D6D0F4EB4989855908F733A12F5D66" originalValue="“Madeira”">&quot;Madeira&quot;</normalizedToken>
reverse &quot;48// 99&quot; [small, circular label, HW] (BMNH); 1 ♂ &quot;Sharp Coll.// 1905-313.&quot; &quot;
<taxonomicName id="C3B564285B4F66D47D95EA5356E864E9" class="Insecta" family="Dytiscidae" genus="Dytiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dytiscus lanio" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="species" species="lanio">Dytiscus lanio</taxonomicName>
Fab.// (
<taxonomicName id="C95D00DDF6854A95D6729317B0E28707" class="Insecta" family="Dytiscidae" genus="Scutopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutopterus" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Scutopterus</taxonomicName>
Woll.
<taxonomicName id="FFD81756791B0DA163F462970B8B3018" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="32" pageNumber="77" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
D. S.)// Madeira T. V. Wollaston&quot; [HW] (BMNH); 2 ♂♂, 2 ♀♀ &quot;Madeira// in Fauvel&quot; [HW] &quot;Coll. Odier.// B. M. 1921-288&quot; (BMNH); 3 ♂♂, 3 ♀♀ &quot;MADEIRA// Ribeira da St.// Luzia// 24.VIII.1964// E. W. Classey&quot; [HW] &quot;Brit. Mus.// 1968-48.&quot; [8-48 HW] (BMNH); 3 ♂♂, 3 ♀♀ &quot;MADEIRA:// Ribeira de St. Martinho// 13.VIII.1966// E. W. Classey&quot; [HW] &quot;Brit. Mus.// 1966-446&quot; [6-446 HW] (BMNH); 6 ♂♂, 4 ♀♀ &quot;MADEIRA:// Ribeiro de St.// Martinho// 13.VIII.1966// E. W. Classey&quot; [HW] &quot;Brit. Mus.// 1968-48.&quot; [8-48 HW] (BMNH); 1 ♂, 2 ♀♀ &quot;MADEIRA:// Ribeira da Abilheira// 19/VIII/1966// E. W. Classey&quot; [HW] (BMNH); 2 ♂♂ &quot;23/iii/1995 MADEIRA// Ribera on 103 below// Poiso &amp; Monte ca.// 900m D. T. Bilton leg.&quot; (CBP);
<pageBreakToken id="BCC19FAFFF35C2C56C2D7E57D1CE2B71" pageId="33" pageNumber="78" start="start">2</pageBreakToken>
♂♂, 1 ♀ &quot;March 2001 MADEIRA// Levada @ Rabacal// D. T. Bilton leg.&quot; (CBP); 1 ♂ &quot;Madeira 1.-4.8.86// 1kn
<normalizedToken id="E3B67741CC06084A59190BF476BF051F" originalValue="nördl">noerdl</normalizedToken>
. Poiso// Quellarn d. R. Frio// leg. Balke// Hendrich&quot; &quot;COLL.// HENDRICH// BERLIN&quot; &quot;
<taxonomicName id="31B005CE6C0D4CD22CB3ED66027A968F" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="33" pageNumber="78" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
//
<taxonomicName id="C05A003544448ACB2C12171F8E1E1624" lsidName="lanio" pageId="33" pageNumber="78" rank="species" species="lanio">lanio</taxonomicName>
F.&quot; [HW] (CBP); 3 ♂♂, 3 ♀♀ &quot;Madeira 8 Anchadas da Cruz// 932 m
<geoCoordinate id="73B05F080FF2DC1982B9B88DD97C4320" direction="north" orientation="latitude" precision="15" value="32.809723">32°48'35&quot;N</geoCoordinate>
<geoCoordinate id="775D378BFA2D778475C2B5745B5AC2C8" direction="west" orientation="longitude" precision="15" value="-17.296389">17°17'47&quot;W</geoCoordinate>
// charca de arroyo// 4.07.15 A. Rudoy leg.&quot; (IBE); 1 ♂ &quot;Madeira, Canhas// lado sur de Paul da Serra// 1398 m
<geoCoordinate id="7413ACB6B06559AB11A9005F31987D69" direction="north" orientation="latitude" precision="15" value="32.743332">32°44'36&quot;N</geoCoordinate>
<geoCoordinate id="6D9235C13B7B29F0DB81FF3A5854C7AB" direction="west" orientation="longitude" precision="15" value="-17.09889">17°05'56&quot;W</geoCoordinate>
// charca sobre arroyo de pietras// 19.7.2011 A. Rudoy leg.&quot; (IBE); 1 ♂ &quot;
<taxonomicName id="9E1824394D56F18F414F91BB62F2529E" lsidName="lanio" pageId="33" pageNumber="78" rank="species" species="lanio">lanio</taxonomicName>
Fab.&quot; [HW] (BMNH); 1 ♂ &quot;40// 4 2// 2156&quot; [small, circular label, HW] (BMNH); 1 ♂ &quot;40// 4 2// 2157&quot; [small, circular label, HW] &quot;C. Lanio Fabr.&quot; (BMNH); 2 ♂♂, 2 ♀♀ without labels (BMNH, Wollaston Collection); 1 ♂ &quot;
<taxonomicName id="691A7824076FC12D0C02258507D96719" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="33" pageNumber="78" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
//
<taxonomicName id="5A3A1C379A6D7C95E65FE8AD8F24E3A6" lsidName="lanio" pageId="33" pageNumber="78" rank="species" species="lanio">lanio</taxonomicName>
// Fauvel 1897//
<normalizedToken id="5E26DBF0227A328CB2FE8FDED609A5A7" originalValue="Madère”">Madere&quot;</normalizedToken>
[HW]
<normalizedToken id="B456927E8C473229271360C2328399D7" originalValue="“MUSÉUM">&quot;MUSEUM</normalizedToken>
PARIS// Coll. P. de Peyerimhoff// 1950&quot; (MNHN); 1 ♂
<normalizedToken id="0D90129FE29CFB173EF62FAE900E0A37" originalValue="“MUSÉUM">&quot;MUSEUM</normalizedToken>
PARIS// Coll. P. de Peyerimhoff// 1950&quot; (MNHN). All with &quot;
<taxonomicName id="6D173E4A87F265EA0CD284DB4B6F5E51" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema lanio" order="Coleoptera" pageId="33" pageNumber="78" phylum="Arthropoda" rank="species" species="lanio">Meladema lanio</taxonomicName>
// (Fabricius, 1775)// D T Bilton [or I Ribera] det. 2017&quot;.
</paragraph>
<caption id="0F65B6F109A82F8D88826B5007E4EE14" pageId="33" pageNumber="78">
<paragraph id="4438A82917CF17993E989ED28C260E73" pageId="33" pageNumber="78">
Figure 18.
<taxonomicName id="71F5004EF4E16668A0750AEE97DDF341" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="33" pageNumber="78" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
species, Italy, male elytral sculpture; shoulder and middle left and right, respectively (DNA voucher codes, where applicable). A
<taxonomicName id="C45B770B2C094B37903535F424B0F41B" lsidName="M. lepidoptera" pageId="33" pageNumber="78" rank="species" species="lepidoptera">M. lepidoptera</taxonomicName>
sp. n., Liguria, Levante B intermediate specimen, Campania, S. Michele (IBE-AN694) C intermediate specimen, Apulia, Vieste D
<taxonomicName id="51BBA4981C345A91EC20169DE11263CB" lsidName="M. coriacea" pageId="33" pageNumber="78" rank="species" species="coriacea">M. coriacea</taxonomicName>
, Lazio, Grotta di Pastena. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="BFEC7D9C1B7E2D0B4087E45B9A160980" pageId="33" pageNumber="78">
<paragraph id="BBC86EA198E20EB96EFE7C1C34CDD289" pageId="33" pageNumber="78">
Figure 19.
<taxonomicName id="2697BAD0143F90F8BF64982CF678BEE2" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema coriacea" order="Coleoptera" pageId="33" pageNumber="78" phylum="Arthropoda" rank="species" species="coriacea">Meladema coriacea</taxonomicName>
, Italy, Sicily elytral sculpture; shoulder and middle left and right, respectively (DNA voucher codes, where applicable). A male, Bosco Ficuzzo (IBE-AN691) B female, Monte Maganoce. Scale bar = 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4262EAAE68D5014AE0327A0A9C3A7AB8" lastPageId="39" lastPageNumber="84" pageId="33" pageNumber="78" type="description">
<paragraph id="C6598EA2B6837F81684E88CE2313B7FF" pageId="33" pageNumber="78">Description</paragraph>
<paragraph id="6E63A4A118EC7B1BFE3A3D6FFA8C9376" pageId="33" pageNumber="78">(based on all material examined).Size: Lectotype TL = 20.30 mm; EL = 16.30 mm; MW = 10.18 mm. Other material examined TL = 17.40-21.12 mm; EL = 12.67-15.36 mm; MW = 8.70-10.50 mm.</paragraph>
<paragraph id="B31D117E4F055DE431C4856F2E7F6064" lastPageId="34" lastPageNumber="79" pageId="33" pageNumber="78">
Colour. Dorsum (Figure 3D), dark reddish brown to yellow. Labrum yellowish, red anterolaterally in some specimens; clypeus yellowish except central ca. 1/6 red to blackish, connected to dark pigmentation on frons. Frons with transverse pale strip anterolaterally, adjacent to pale parts of clypeus, otherwise dark reddish brown. Me
<pageBreakToken id="6A53804E579566226A94FA4CA626822D" pageId="34" pageNumber="79" start="start">dial</pageBreakToken>
, paired interocular patches on frons yellow; strongly transverse apicolaterally, almost reaching channel around interior margin of compound eye; confluent in centre in some specimens. Pronotum dark reddish brown on disc; narrowly reddish along anterior margin; lateral margins broadly yellowish. Elytra yellowish brown to greenish brown, with black irrorations; pattern visible without being lifted, but more evident when done so (Figures 3D, 4D). Legs yellowish brown to black; posterior tibiae and tarsi darkest. Antennae and maxillary and labial palpi yellowish to reddish. Venter reddish brown; prementum yellow; posterior genae, mentum and submentum reddish. Pronotal hypomeron and shoulder and outer portion of elytral epipleurs yellowish.
</paragraph>
<paragraph id="19C1C1924F9317E1565F49B71B8F2D47" lastPageId="35" lastPageNumber="80" pageId="34" pageNumber="79">
Head. Labrum shining, with medium to fine, sparse punctures. Reticulation absent anteriorly, clearly evident in posterior half, here fine and composed of small, isodiametric to transverse meshes. Clypeus weakly shining, with medium to fine, sparse punctures and traces of very fine, shallow, close punctures. Frons weakly shining, entire surface with, open reticulation; weak in front of interocular patches, becoming stronger and more evident posteriorly. Meshes transverse to isodiametric apically and medially, strongly elongate posteriorly and onto vertex. Paired epicranial foveae on anterior frons, one immediately behind the other. Anterior foveae transverse, posterior foveae elongate oval. Foveae reticulate; anterior and posterior foveae linked by reticulated channel. Internal and posterior borders of compound eyes distinct, raised relative to level of adjacent cuticle. Lateral margins bordered by distinct narrow chan
<pageBreakToken id="B5FE93BCCBE6D8D195D85DA508F59145" pageId="35" pageNumber="80" start="start">nel</pageBreakToken>
; deeper anteriorly than posteriorly and continuing behind posterior margin of eye onto vertex. Channel with dense punctures, bearing long, stiff, yellow recumbent to decumbent setae.
</paragraph>
<paragraph id="42ACD20873A383BABD4EE12DF11FD8F3" lastPageId="36" lastPageNumber="81" pageId="35" pageNumber="80">
Pronotum. Posterior margin weakly sinuate laterally (Figure 3D). Surface somewhat shining, strongly rugose. Reticulation meshes large, open, flat and with varying sizes and orientations. Transverse irregular row of medium punctures bearing long, yellow recumbent to decumbent setae 1/5 behind anterior pronotal margin; interrupted
<pageBreakToken id="08C946A0DFBB0085FE01FF3DA79FAD89" pageId="36" pageNumber="81" start="start">briefly</pageBreakToken>
in centre, obscured by reticulation inside lateral margins but continuing inside lateral third of posterior margin. Reticulation weak and obsolete anterior to transverse row, surface here clearly double punctate, with very fine, dense and medium, sparse to very sparse punctures. Scattered medium punctures visible elsewhere, amongst meshes of microreticulation. Centre of disc with elongate, narrow, slit-like fovea, partly obscured by reticulation in some specimens, but then traceable as an elongate reticulation channel. Lateral margins slightly raised, shining, without rugose sculpture and with fine, scattered punctures.
</paragraph>
<paragraph id="58439D2405212DD60D2A78247E1729F1" pageId="36" pageNumber="81">Elytra. Shining, without crescentic striolae (except in some females - see below). Surface doubly punctate and reticulate (Figure 5D); with very fine, close and fine to medium, sparse punctures (the latter bearing short, peg-like setae). Reticulation fine, somewhat obsolete, meshes isodiametric; sometimes obscuring very fine punctation. Surface weakly tuberculate towards sides and apex, particularly in posterior third (see Figure 3D). Puncture rows well-marked, continuous almost to elytral apices; punctures shallower posteriorly than anteriorly.</paragraph>
<caption id="09BB3276AE5FA93E1DB86AEB7B696B1F" pageId="36" pageNumber="81">
<paragraph id="3576BEE1C9A042FD01BF6F8E01D92A80" pageId="36" pageNumber="81">
Figure 20.
<taxonomicName id="D48C179890352A5BC89EDEF689280F7E" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="36" pageNumber="81" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
species, variation in male median lobes, lateral view.
<normalizedToken id="E05B83CCF0271FF5E31C97E19E90CD59" originalValue="A–N">A-N</normalizedToken>
<taxonomicName id="CBAB675FC364385574993F6D35232071" lsidName="M. coriacea" pageId="36" pageNumber="81" rank="species" species="coriacea">M. coriacea</taxonomicName>
;
<normalizedToken id="8367709AE9BC58110051582547FD6BC6" originalValue="O–U">O-U</normalizedToken>
<taxonomicName id="BD5997B7DFE76D034565E46FE335C0DE" lsidName="M. lepidoptera" pageId="36" pageNumber="81" rank="species" species="lepidoptera">M. lepidoptera</taxonomicName>
sp. n. (DNA voucher codes, where applicable). A Spain, Huesca,
<normalizedToken id="C0C6D64BE2D4DB546892AB33340BF6E4" originalValue="Bernués">Bernues</normalizedToken>
(IBE-DV292) B Spain, Girona, Port Bou (IBE-DV-293) C Spain, Tarragona, Esblada D Spain,
<normalizedToken id="9F431F54E9434DA63EEAB6C4DA9D70E6" originalValue="Cáceres">Caceres</normalizedToken>
, nr. Plasencia E Spain, Murcia, Fte. Caputa (NHM-IRM13) F Spain,
<normalizedToken id="980B3F330C6B8B75173C0130071545A3" originalValue="Córdoba">Cordoba</normalizedToken>
, Baena (NHM-IRM14C) G Tenerife, Chamorga (MNCN-AI1095) H Morocco, Oued Massa (NHM-IRM2A) I Morocco, Tazekka (NHM-IRM1C) J Morocco, Sidi-Ibrahim (NHM-IR7) K Algeria,
<normalizedToken id="E04496914B9A7B39E1CA891A4E55BE40" originalValue="Aïn">Ain</normalizedToken>
Damous (MNCN-HI6) L Italy, Sicily, Bosco Ficuzzo (IBE-AN691) M Greece, Poros, Kampos N Greece, Ios
<normalizedToken id="7BCD65BA2CEF7E80127D61F547B4F2F1" originalValue="O–P">O-P</normalizedToken>
Corsica, Francardo Q holotype, Corsica, Cap Corse (NHM-IRM12E) R Italy, Montecristo (IBE-DV289) S Sardinia, Villagrande
<normalizedToken id="CE3D0A6501A18FA158D3B2522DE7E7C5" originalValue="Strìsaili">Strisaili</normalizedToken>
(IBE-RA5) T Italy, Elba, Pomonte (IBE-AN692) U Italy, Tuscany, S. Luce (IBE-AN693). Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="5EC342D42AF670BF8AE04D35FF6E23B1" pageId="36" pageNumber="81">
<paragraph id="BF8396EA73B18F030BED5076E31949E2" pageId="36" pageNumber="81">
Figure 21.
<taxonomicName id="F527F7C56B91C9256064190F5561CB1C" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema lepidoptera" order="Coleoptera" pageId="36" pageNumber="81" phylum="Arthropoda" rank="species" species="lepidoptera">Meladema lepidoptera</taxonomicName>
sp. n. elytral sculpture (males); shoulder and middle left and right, respectively (DNA voucher codes). A holotype male, Corsica, Cap Corse (NHM-IRM12E) B Italy, Montecristo (IBE-DV289) C Italy, Elba, Pomonte (IBE-AN692) D Italy, Tuscany, S. Luce (IBE-AN693). Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="667405A9AB9B0EA7F7DCDCD4C228764D" pageId="36" pageNumber="81">
<paragraph id="4755C3DBAED61F6014939684D4251CF3" pageId="36" pageNumber="81">
Figure 22.
<taxonomicName id="8020DA4235C75CC43977B5586DED1DD8" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema lepidoptera" order="Coleoptera" pageId="36" pageNumber="81" phylum="Arthropoda" rank="species" species="lepidoptera">Meladema lepidoptera</taxonomicName>
sp. n., Sardinia, elytral sculpture SEMs; shoulder and middle left and right, respectively (DNA voucher codes). A male, Villagrande
<normalizedToken id="9F1285D61CC658EC7D8B3110CCE1C191" originalValue="Strìsaili">Strisaili</normalizedToken>
(IBE-RA5) B female, Tortoli (IBE-RA18).
</paragraph>
</caption>
<paragraph id="FF772DD94F63EBC6C8CF425D652D61C1" lastPageId="38" lastPageNumber="83" pageId="36" pageNumber="81">
Venter. Prementum shining, tumid in centre, with fine and medium, sparse punctures. Mentum shining; central projection with shallow median emargination. Lateral lobes with medium, sparse punctures, and scattered, whitish recumbent to decumbent setae. Submentum shining, with transverse wrinkles centrally, and elongate wrinkles laterally. Central 1/4 with medium, sparse punctures bearing long, white-yellowish, erect setae. Gula shining, with sparse, shallow, transverse wrinkles; patch of medium-coarse punctures posterolaterally. Genae shining, with obsolete, open, elongate reticu
<pageBreakToken id="CB6D915E74599CCF3C53D5D2243F32E0" pageId="37" pageNumber="82" start="start">lation</pageBreakToken>
. Prosternum shining, with weak, low, irregular transverse ridges laterally. Arched in centre and with fine, moderate to close punctures laterally, bearing long, white-yellowish, recumbent to erect setae; punctures and setae extending in an irregular row onto process, just below arch. Process lanceolate, arched; apex acuminately rounded. Centre of prosternum and process with double punctation of very fine, close to very
<pageBreakToken id="23AAD6E3CE21F0C97D866276B55DD354" pageId="38" pageNumber="83" start="start">close</pageBreakToken>
and medium, sparse to moderate punctures. Pronotal hypomeron shining, impunctate. Elytral epipleurs shining, with fine wrinkles; irregular puncture row close to internal margin, from centre to close to apex, punctures bearing fine, whitish, erect setae. Metaventrite shining, central portion with reticulation reduced to sparse, transverse scratches and very fine, close and fine to medium, sparse punctures. Metaventral process strongly reticulate, with transverse to elongate, rugose meshes and traces of fine, sparse punctures. Metaventral process relatively broad; apex acuminate and upturned slightly anterolaterally. Metacoxal lines not reaching anterior border of metacoxae, disappearing approx 1/10 from margin. Internal laminae of metacoxae shining, sculpture as on centre of metaventrite. Metacoxal lobes sculptured as internal laminae, strongly rounded, with irregular, elongate field of medium to coarse punctures close to lateral margins, bearing fine, white, recumbent to erect setae. External laminae of metacoxae shining, smooth close to process, and with obsolete reticulation elsewhere; without distinct meshes, wrinkled, elongate around anterior and posterior margins; doubly ounctate, with very fine, close and fine, very sparse punctures. Abdominal ventrites shining. Ventrites 3-5 with cluster of golden, erect setae anteromedially. Ventrite 1 with weak reticulation of elongate scratches throughout. Ventrite 2 with similar reticulation; absent close to centre. Ventrite 3 and 4 with scratches restricted to lateral 1/3. Ventrites 2-5 doubly punctate; very fine, moderate and fine, sparse to very sparse punctures; punctures most evident in areas without reticulation. Ventrites 3-5 with transverse irregular row of long, yellowish, recumbent to decumbent setae laterally. Ventrite 6 (Figure 11D) with very fine, moderate punctures and medium to coarse, sparse to moderate punctures; punctures coarser close to apex. Elongate, semicircular wrinkles and channels apicolaterally and centrally; apicolateral sculpture extending basally around central portion of ventrite. Some punctures in channels bearing elongate, whitish, erect setae.
</paragraph>
<paragraph id="7F61A461C336E7B4B0DED50F9363DD9A" pageId="38" pageNumber="83">Male. Foretarsi (Figure 12D) with articulo-setal counts as follows (base to apex): row 1, 8-9; row 2, 10; row 3, 10; row 4, 8. Number of setae in rows may differ from right to left tarsus in same beetle. Curved, golden setae bordering articulo-setal field relatively sparse, particularly laterally. Foretarsal claws (Figures 12D, 13D) elongate, curved. Mesotarsi (Figure 12H) with articulo-setal counts as follows (base to apex): row 1, 7-8; row 2, 8; row 3, 8; row 4, 6 (2 clusters of 3, situated laterally). Curved, golden setae bordering articulo-setal field relatively sparse, especially laterally. Mesotarsal claws (Figure 12H) elongate, curved. Abdominal ventrite 6 (Figure 11D) with apex rounded, with well-marked median emargination. Median lobe (Figure 14D) asymmetrical, sinuation strong, approximately 1/3-0.35 from apex; ventral margin of apical portion relatively straight in lateral view. Parameres (Figure 14D) with inner margin almost right-angled at base, with distinct small projection; outer margin undulated slightly; inner straight.</paragraph>
<paragraph id="0D9B334659C86264951B1E450CFF1217" pageId="38" pageNumber="83">Female. As male, except for simple fore and mesotarsi, and differently shaped abdominal ventrite 6 (with bluntly pointed apex - Figure 15D).</paragraph>
<paragraph id="8511EE814CACE82BF99BDCADADB09F61" lastPageId="39" lastPageNumber="84" pageId="38" pageNumber="83">
Variation. Males and females generally have identical sculpture on the elytra. Two females studied, one from Ribeira da St. Luzia (Figure 26B, BMNH) and one from
<normalizedToken id="EDCAB55F8ED6E4B2057BE19933F67AA2" originalValue="“pouzo”">&quot;pouzo&quot;</normalizedToken>
(ISNB) have distinct crescentic striolae, as have been observed in female
<taxonomicName id="D634DFD771B65D0EA960ECB732C059A9" lsidName="M. imbricata" pageId="38" pageNumber="83" rank="species" species="imbricata">M. imbricata</taxonomicName>
from La Palma (see above). Females sculptured in this manner appear to be relatively rare, the
<pageBreakToken id="B67C22A33FB003C067CA87664756D948" pageId="39" pageNumber="84" start="start">only</pageBreakToken>
other occurrences of this form we can find being mentioned by
<bibRefCitation id="8C0008EF183529346837C5A0E65A23D7" author="Aube, C" journalOrPublisher="Mequignon Pere et Fils, Paris" pageId="42" pageNumber="87" title="Species general des hydrocanthares et gyriniens; pour faire suite au species general des coleopteres de la collection de M. le comte Dejean." year="1838">
<normalizedToken id="A32D49AD9FDF1B3C51BF480011605807" originalValue="Aubé">Aube</normalizedToken>
(1838)
</bibRefCitation>
, who had seen a female with this sculpture (see also
<bibRefCitation id="16E2CF571FE6B86FCA310D6D800B0BF5" author="Sharp, D" journalOrPublisher="Scientific Transactions of the Royal Dublin Society" pageId="46" pageNumber="91" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
) and
<bibRefCitation id="CE2091897BD3BA99E33C8862E85644BC" pageId="39" pageNumber="84">
<normalizedToken id="4FFDF6FDF480605092D319EC79080A19" originalValue="Falkenström">Falkenstroem</normalizedToken>
(1938)
</bibRefCitation>
. Such females may be mistaken for
<taxonomicName id="E26C38BE455AA548994CB460CA7514BC" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
, as was the case with the ISNB animal listed above.
</paragraph>
<caption id="565A0A27D9D716266E1E472921EBD310" pageId="39" pageNumber="84">
<paragraph id="0630190CA1E16A94A5C98E28E9BBE364" pageId="39" pageNumber="84">
Figure 23.
<taxonomicName id="015201910BA0495915E02FE5806BDB72" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="39" pageNumber="84" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
distribution, material examined. Symbols with black border show locations of genotyped specimens. Symbol colours as follows:
<taxonomicName id="82C8044478EF9E5595AD8DA098E9C372" lsidName="M. coriacea" pageId="39" pageNumber="84" rank="species" species="coriacea">M. coriacea</taxonomicName>
- red;
<taxonomicName id="E68B4D5FD9A805B2980684DF87181B94" lsidName="M. lepidoptera" pageId="39" pageNumber="84" rank="species" species="lepidoptera">M. lepidoptera</taxonomicName>
sp. n. - blue;
<taxonomicName id="8BB9E0F667017C0A1B4E77771ED62E13" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
- green;
<taxonomicName id="6E1FC48E61029CB24A1BD48BAED3FD9E" lsidName="M. lanio" pageId="39" pageNumber="84" rank="species" species="lanio">M. lanio</taxonomicName>
- yellow. Bicoloured symbols, hybrid or morphologically intermediate individuals.
</paragraph>
</caption>
<caption id="177261417A84221992404BF3F959A0CB" pageId="39" pageNumber="84">
<paragraph id="152D9AD1E02A0A9BFB13E80D95A8DD14" pageId="39" pageNumber="84">
Figure 24.
<taxonomicName id="29453060CA2B526F4DC56A1DB095AC7B" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="39" pageNumber="84" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
type labels and specimens in situ. A
<taxonomicName id="0FD4ACFDAC42E35496A24474D945F167" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
, holotype, BMNH, Wollaston Collection B
<taxonomicName id="3328F74441A1EEDF921DB0A5B12270E0" lsidName="M. lanio" pageId="39" pageNumber="84" rank="species" species="lanio">M. lanio</taxonomicName>
lectotpe, BMNH, Banks Collection.
</paragraph>
</caption>
<caption id="F5F94A255E27AC04EAFB353320582C3D" pageId="39" pageNumber="84">
<paragraph id="552277BA1E339A26CCDC2E8A6472B391" pageId="39" pageNumber="84">
Figure 25.
<taxonomicName id="B3C4559EBA6EEAE59ED897775B1287E8" class="Insecta" family="Dytiscidae" genus="Meladema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meladema" order="Coleoptera" pageId="39" pageNumber="84" phylum="Arthropoda" rank="genus">Meladema</taxonomicName>
species, Canary Islands, elytral shoulder sculpture SEMs
<normalizedToken id="307527454A3AB74EAD276A840CEC2640" originalValue="A–E">A-E</normalizedToken>
males F female (DNA voucher codes where applicable).
<normalizedToken id="83637327BC18EB5E97EF38627C4AF621" originalValue="A–B">A-B</normalizedToken>
<taxonomicName id="0ACCFDE94821845AFC4D70CE3FFD1449" lsidName="M. coriacea" pageId="39" pageNumber="84" rank="species" species="coriacea">M. coriacea</taxonomicName>
x
<taxonomicName id="9536AA78C3EBAF9E421E96F75D7882F4" lsidName="imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">imbricata</taxonomicName>
hybrid, Tenerife, Bco. del
<normalizedToken id="411A7655E70C1F7588A9CE0BAB1CDBB6" originalValue="Río">Rio</normalizedToken>
600 m (NHM-IRM16A) C
<taxonomicName id="3A36C8AFFF1E21E21124D45D5C0532EF" lsidName="M. coriacea" pageId="39" pageNumber="84" rank="species" species="coriacea">M. coriacea</taxonomicName>
, Tenerife, Bco. de Masca (NHM-IRM19A) D
<taxonomicName id="6AB7FDE49BD6C407C643189AB6D79E0A" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
, La Gomera, El Cedro (NHM-IRM3A) E
<taxonomicName id="ADCC7F8776B773A76941F3AAB9DC7FA7" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
x
<taxonomicName id="4EFBC114CBB59262395EA01129BABD47" lsidName="coriacea" pageId="39" pageNumber="84" rank="species" species="coriacea">coriacea</taxonomicName>
hybrid, Tenerife, Bco. del
<normalizedToken id="73CEF5B7D2C9CAC2563A6E2BBC2A92E3" originalValue="Río">Rio</normalizedToken>
1,600 m (NHM-IRM5B) F
<taxonomicName id="C95C0749EDB27EC7F4BB06E2021CAEBB" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
, La Palma, Bco. Hoyo Verde.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="325E9D0CA51E21953817F79B61DE3547" pageId="39" pageNumber="84" type="differential diagnosis">
<paragraph id="986DA522AB5625A212FA904E56161BE3" pageId="39" pageNumber="84">Differential diagnosis.</paragraph>
<paragraph id="472CB2BAF0D6569431F736AA9C703950" pageId="39" pageNumber="84">
Closest to
<taxonomicName id="69872103C1FD32921C19F88DB2D0E45F" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
; for diagnostic characters see under that species.
</paragraph>
</subSubSection>
<subSubSection id="3D4B97B3A343545D3599ABF9EB9E8044" pageId="39" pageNumber="84" type="distribution">
<paragraph id="F852B4C1545827978A297AF8C26F9865" pageId="39" pageNumber="84">Distribution.</paragraph>
<paragraph id="FCD00DDBC2A4E3B9F4E602D4D054396F" pageId="39" pageNumber="84">
Restricted to the main island of Madeira (Figure 23), where it is relatively widely distributed in permanent streams, particularly in remnant laurel forests in the mountains. Can also be found in man-made levada systems. Much more abundant on Madeira than
<taxonomicName id="08EEFD404CF11D5B655632430ACB1172" lsidName="M. imbricata" pageId="39" pageNumber="84" rank="species" species="imbricata">M. imbricata</taxonomicName>
on the Canaries, but still listed as Vulnerable (B1 + 2b) on the IUCN Red List (
<bibRefCitation id="DFFF3116C3C89CAED7CC359A7D8BCB73" author="Foster, G" journalOrPublisher="Linzer biologische Beitraeg" pageId="44" pageNumber="89" url="http://dx.doi.org/10.2305/IUCN.UK.1996.RLTS.T13015A3406416.en" year="1996 b">Foster 1996b</bibRefCitation>
), reflecting its very small global range.
</paragraph>
</subSubSection>
</treatment>
</document>
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