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<document id="C13C862D10305260E8818F3039C84CCC" ID-CLB-Dataset="22256" ID-DOI="10.11646/zootaxa.4772.3.7" ID-GBIF-Dataset="8b277405-a91c-4d0c-b77e-8b1d4820c7cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819764" ID-ZooBank="8DE2F40F-0931-4002-97C4-5603E363B7E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175825495" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel Ángel" docDate="2020" docId="6C5796057B76FFCEFF0939BA9A0B2D88" docLanguage="en" docName="zootaxa.4772.3.7.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrodactylus montanus Arce-Perez &amp; Moron 2000" docType="treatment" docVersion="7" lastPageNumber="578" masterDocId="906EEE7D7B7EFFC5FF9E3F1F990C2A65" masterDocTitle="Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae Melolonthinae) from the Central American Nucleus" masterLastPageNumber="584" masterPageNumber="567" pageNumber="575" updateTime="1698839127964" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="21208A2A9AD360FDD56C82086422EDDB">Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae Melolonthinae) from the Central American Nucleus</mods:title>
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<mods:namePart id="A351D65248DAB0FD86D0F507CC0E5DEC">Arce-Pérez, Roberto</mods:namePart>
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<mods:namePart id="2D2EB703A6932DE8A86E48EA9B1B3777">Morón, Miguel Ángel</mods:namePart>
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<mods:date id="0D35D7CB2B4C8632317538D025FB746A">2020</mods:date>
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<treatment id="6C5796057B76FFCEFF0939BA9A0B2D88" ID-DOI="http://doi.org/10.5281/zenodo.3843997" ID-GBIF-Taxon="164035925" ID-Zenodo-Dep="3843997" LSID="urn:lsid:plazi:treatment:6C5796057B76FFCEFF0939BA9A0B2D88" httpUri="http://treatment.plazi.org/id/6C5796057B76FFCEFF0939BA9A0B2D88" lastPageId="11" lastPageNumber="578" pageId="8" pageNumber="575">
<subSubSection id="ACE474987B76FFCDFF0939BA98632C86" pageId="8" pageNumber="575" type="nomenclature">
<paragraph id="E44127137B76FFCDFF0939BA9A2E2CDA" blockId="8.[151,802,1700,1763]" box="[151,802,1700,1728]" pageId="8" pageNumber="575">
<heading id="BF09907F7B76FFCDFF0939BA9A2E2CDA" bold="true" box="[151,802,1700,1728]" fontSize="11" level="1" pageId="8" pageNumber="575" reason="1">
<taxonomicName id="23FE5C907B76FFCDFF0939BA9A2E2CDA" authority="Arce-Perez &amp; Moron, 2000" authorityName="Arce-Perez &amp; Moron" authorityYear="2000" box="[151,802,1700,1728]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="575" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="D68AFB017B76FFCDFF0939BA9A2E2CDA" bold="true" box="[151,802,1700,1728]" pageId="8" pageNumber="575">
<emphasis id="D68AFB017B76FFCDFF0939BA98C72CDA" bold="true" box="[151,459,1701,1728]" italics="true" pageId="8" pageNumber="575">Macrodactylus montanus</emphasis>
<bibRefCitation id="806F5AE27B76FFCDFE4E39BB9A2E2CDA" author="Arce-Perez, R. &amp; Moron, M. A." box="[464,802,1700,1727]" pageId="8" pageNumber="575" pagination="123 - 239" refId="ref10619" refString="Arce-Perez, R. &amp; Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) en Mexico y Estados Unidos de America. Acta Zoologica Mexicana, New Series, 97, 123 - 239. Available from: http: // www. scielo. org. mx / scielo. php? script = sci _ arttext &amp; pid = S 0065 - 17372000000100009 &amp; lng = es &amp; nrm = iso &amp; tlng = es (accessed 27 February 2020)" type="journal article" year="2000">Arce-Pérez &amp; Morón, 2000</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="E44127137B76FFCDFF0939D698632C86" blockId="8.[151,802,1700,1763]" box="[151,367,1736,1763]" pageId="8" pageNumber="575">Figs. 20, 45–47, 86</paragraph>
</subSubSection>
<subSubSection id="ACE474987B76FFCDFF09380F9BBF2D4E" box="[151,691,1808,1835]" pageId="8" pageNumber="575" type="materials_examined">
<paragraph id="E44127137B76FFCDFF09380F9BBF2D4E" blockId="8.[151,1437,1808,2015]" box="[151,691,1808,1835]" pageId="8" pageNumber="575">
<emphasis id="D68AFB017B76FFCDFF09380F988D2D4E" bold="true" box="[151,385,1808,1835]" pageId="8" pageNumber="575">Material examined.</emphasis>
<specimenCount id="F2F8EC9A7B76FFCDFE19380E9B2F2D4E" box="[391,547,1809,1835]" pageId="8" pageNumber="575" type="generic">37 specimens</specimenCount>
:
<specimenCount id="F2F8EC9A7B76FFCDFDB0380E9B642D4E" box="[558,616,1809,1835]" pageId="8" pageNumber="575" type="male">18 ♂</specimenCount>
,
<specimenCount id="F2F8EC9A7B76FFCDFDEA380E9BA12D4E" box="[628,685,1809,1835]" pageId="8" pageNumber="575" type="female">19 ♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="ACE474987B76FFCEFF59382B9A0B2D88" lastPageId="11" lastPageNumber="578" pageId="8" pageNumber="575" type="diagnosis">
<paragraph id="E44127137B76FFCDFF59382B9ADD2DBA" blockId="8.[151,1437,1808,2015]" pageId="8" pageNumber="575">
<emphasis id="D68AFB017B76FFCDFF59382B984E2D2A" bold="true" box="[199,322,1844,1871]" pageId="8" pageNumber="575">Diagnosis.</emphasis>
Body length
<quantity id="23068AF67B76FFCDFE46382A9B582D2A" box="[472,596,1845,1871]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="8" pageNumber="575" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
; integument of the body black; clypeus red, antennal flagellum, elytra, legs and, pygidium reddish yellow; dorsum and venter with thin, scattered pale yellow vestiture; tarsi with rings of short, white setae (Fig. 20); parameres circular, setose, with a narrow ridge on its internal margins, expanded into a welldeveloped distal shelf-like ledge, apices strongly lanceolate, lateral margins on distal half with sparse, mediumsized setae (Figs. 45–46). Female similar to male; genital plates (Fig. 47).
</paragraph>
<paragraph id="E44127137B77FFCCFF0938869B612D9C" blockId="9.[151,1437,1945,2042]" pageId="9" pageNumber="576">
<emphasis id="D68AFB017B77FFCCFF093886983A2DD7" bold="true" box="[151,310,1945,1970]" pageId="9" pageNumber="576">Figures 48–56.</emphasis>
Dorsal view of
<emphasis id="D68AFB017B77FFCCFE4338859B702DD7" box="[477,636,1946,1970]" italics="true" pageId="9" pageNumber="576">
<taxonomicName id="23FE5C907B77FFCCFE4338859B7B2DD7" authorityName="Dejean" authorityYear="1821" box="[477,631,1946,1970]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="genus">Macrodactylus</taxonomicName>
.
</emphasis>
<emphasis id="D68AFB017B77FFCCFD1C38869B972DD7" bold="true" box="[642,667,1945,1970]" pageId="9" pageNumber="576">48</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFD3738859AD22DD4" authority="Bates" authorityName="Bates" authorityYear="1887" box="[681,990,1946,1970]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="D68AFB017B77FFCCFD3738859A972DD4" box="[681,923,1946,1970]" italics="true" pageId="9" pageNumber="576">Macrodactylus nigripes</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B77FFCCFC7738869D0E2DD7" bold="true" box="[1001,1026,1945,1970]" pageId="9" pageNumber="576">49</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFB8F38859DDA2DD4" authority="Bates" authorityName="Bates" authorityYear="1887" box="[1041,1238,1946,1970]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="D68AFB017B77FFCCFB8F38859D982DD7" box="[1041,1172,1946,1970]" italics="true" pageId="9" pageNumber="576">M. rufescens</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B77FFCCFB7C38869DF72DD7" bold="true" box="[1250,1275,1945,1970]" pageId="9" pageNumber="576">50</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFA97388599DE2DB0" authority="Bates" authorityName="Bates" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="seiceicollis">
<emphasis id="D68AFB017B77FFCCFA9738859C912DD7" box="[1289,1437,1946,1970]" italics="true" pageId="9" pageNumber="576">M. seiceicollis</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B77FFCCFF4338A299FA2DB3" bold="true" box="[221,246,1981,2006]" pageId="9" pageNumber="576">51</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFE9D38A198CD2DB0" authority="Bates" authorityName="Bates" authorityYear="1887" box="[259,449,1982,2005]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="sericinus">
<emphasis id="D68AFB017B77FFCCFE9D38A198732DB0" box="[259,383,1982,2005]" italics="true" pageId="9" pageNumber="576">M. sericinus</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B77FFCCFE5238A298E92DB3" bold="true" box="[460,485,1981,2006]" pageId="9" pageNumber="576">52</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFE6C38A19B8B2DB0" authority="Bate" authorityName="Bates" authorityYear="1887" box="[498,647,1982,2005]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="suavis">
<emphasis id="D68AFB017B77FFCCFE6C38A19B5E2DB0" box="[498,594,1982,2005]" italics="true" pageId="9" pageNumber="576">M. suavis</emphasis>
Bate
</taxonomicName>
<emphasis id="D68AFB017B77FFCCFD1938A19B9D2DB0" box="[647,657,1982,2005]" italics="true" pageId="9" pageNumber="576">s</emphasis>
;
<emphasis id="D68AFB017B77FFCCFD0038A29BBB2DB3" bold="true" box="[670,695,1981,2006]" pageId="9" pageNumber="576">53</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFD5A38A19A622DB0" authority="Bates" authorityName="Bates" authorityYear="1887" box="[708,878,1982,2006]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="sylphis">
<emphasis id="D68AFB017B77FFCCFD5A38A19A202DB3" box="[708,812,1982,2006]" italics="true" pageId="9" pageNumber="576">M. sylphis</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B77FFCCFCE638A29A9D2DB3" bold="true" box="[888,913,1981,2006]" pageId="9" pageNumber="576">54</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFC0038A19DEA2DB3" authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" authorityYear="2005" box="[926,1254,1982,2006]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="D68AFB017B77FFCCFC0038A19D0A2DB3" box="[926,1030,1982,2006]" italics="true" pageId="9" pageNumber="576">M. tibialis</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
;
<emphasis id="D68AFB017B77FFCCFB6F38A19C062DB3" bold="true" box="[1265,1290,1982,2006]" pageId="9" pageNumber="576">55</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFA8938A1987D2D9F" authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" authorityYear="2011" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="zaragozai">
<emphasis id="D68AFB017B77FFCCFA8938A19C902DB0" box="[1303,1436,1982,2005]" italics="true" pageId="9" pageNumber="576">M. zaragozai</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
;
<emphasis id="D68AFB017B77FFCCFEE238FE98992D9F" bold="true" box="[380,405,2017,2042]" pageId="9" pageNumber="576">56</emphasis>
,
<taxonomicName id="23FE5C907B77FFCCFE3C38FD9B612D9C" authority="Bates." authorityName="Bates." box="[418,621,2018,2042]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="D68AFB017B77FFCCFE3C38FD9B242D9F" box="[418,552,2018,2042]" italics="true" pageId="9" pageNumber="576">M. zunilensis</emphasis>
Bates.
</taxonomicName>
</paragraph>
<paragraph id="E44127137B74FFCFFF593F869A0A2AB2" blockId="10.[151,1436,152,287]" pageId="10" pageNumber="577">
<emphasis id="D68AFB017B74FFCFFF593F86988A2AD6" bold="true" box="[199,390,152,179]" pageId="10" pageNumber="577">Natural history.</emphasis>
This species inhabits temperate-cool or humid areas between 1,800 and
<quantity id="23068AF67B74FFCFFB203F869C172AD6" box="[1214,1307,153,179]" metricMagnitude="0" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="577" unit="m" value="2.8">2,800 m</quantity>
(additional information in
<bibRefCitation id="806F5AE27B74FFCFFEA03FA29B622AB2" author="Arce-Perez, R. &amp; Moron, M. A." box="[318,622,189,215]" pageId="10" pageNumber="577" pagination="123 - 239" refId="ref10619" refString="Arce-Perez, R. &amp; Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) en Mexico y Estados Unidos de America. Acta Zoologica Mexicana, New Series, 97, 123 - 239. Available from: http: // www. scielo. org. mx / scielo. php? script = sci _ arttext &amp; pid = S 0065 - 17372000000100009 &amp; lng = es &amp; nrm = iso &amp; tlng = es (accessed 27 February 2020)" type="journal article" year="2000">Arce-Pérez &amp; Morón 2000</bibRefCitation>
,
<bibRefCitation id="806F5AE27B74FFCFFDE73FA39BB82AB2" author="Arce-Perez, R. &amp; Moron, M. A." box="[633,692,188,215]" pageId="10" pageNumber="577" pagination="23 - 27" refId="ref10716" refString="Arce-Perez, R. &amp; Moron, M. A. (2005) New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Honduras and Nicaragua. Zootaxa, 1012 (1), 23 - 27. https: // doi. org / 10.11646 / zootaxa. 1012.1.3" type="journal article" year="2005">2005</bibRefCitation>
,
<bibRefCitation id="806F5AE27B74FFCFFD213FA29BF72AB2" author="Arce-Perez, R. &amp; Moron, M. A." box="[703,763,189,215]" pageId="10" pageNumber="577" refId="ref10871" refString="Arce-Perez, R. &amp; Moron, M. A. (2011) Los Macrodactylus (Coleoptera: Melolonthidae) de America Central. Editorial Academica Espanola, Saarbrucken, 275 pp." type="book" year="2011">2011</bibRefCitation>
).
</paragraph>
<paragraph id="E44127137B74FFCFFF593FFF980C2B7A" blockId="10.[151,1436,152,287]" pageId="10" pageNumber="577">
<emphasis id="D68AFB017B74FFCFFF593FFF9B082A9E" bold="true" box="[199,516,224,251]" pageId="10" pageNumber="577">Geographical distribution.</emphasis>
<materialsCitation id="54962D4E7B74FFCFFD973FFF9BD82A9E" ID-GBIF-Occurrence="2611565326" box="[521,724,224,251]" country="Mexico" location="Chiapas" pageId="10" pageNumber="577" specimenCount="1" stateProvince="Chiapas">
<collectingCountry id="9CE967837B74FFCFFD973FFF9B532A9F" box="[521,607,224,250]" name="Mexico" pageId="10" pageNumber="577">Mexico</collectingCountry>
(
<collectingRegion id="263AE9F17B74FFCFFDF33FFF9BC12A9E" box="[621,717,224,251]" country="Mexico" name="Chiapas" pageId="10" pageNumber="577">Chiapas</collectingRegion>
)
</materialsCitation>
,
<materialsCitation id="54962D4E7B74FFCFFD403FFF9D132A9F" ID-GBIF-Occurrence="2611565324" box="[734,1055,224,251]" country="El Salvador" location="Santa Ana" pageId="10" pageNumber="577" specimenCount="1" stateProvince="Santa Ana">
<collectingCountry id="9CE967837B74FFCFFD403FFF9A6D2A9F" box="[734,865,224,250]" name="El Salvador" pageId="10" pageNumber="577">El Salvador</collectingCountry>
(
<collectingRegion id="263AE9F17B74FFCFFCF03FFE9AE92A9E" box="[878,997,225,251]" country="El Salvador" name="Santa Ana" pageId="10" pageNumber="577">Santa Ana</collectingRegion>
), and
</materialsCitation>
<materialsCitation id="54962D4E7B74FFCFFBBB3FFF99F02B7A" ID-GBIF-Occurrence="2611565316" country="Honduras" location="Ocotepeque" pageId="10" pageNumber="577" specimenCount="1" stateProvince="Ocotepeque">
<collectingCountry id="9CE967837B74FFCFFBBB3FFF9D9E2A9F" box="[1061,1170,224,250]" name="Honduras" pageId="10" pageNumber="577">Honduras</collectingCountry>
(
<collectingRegion id="263AE9F17B74FFCFFB3E3FFE9C262A9E" box="[1184,1322,225,251]" country="Honduras" name="Ocotepeque" pageId="10" pageNumber="577">Ocotepeque</collectingRegion>
,
<collectingRegion id="263AE9F17B74FFCFFAAA3FFF9C992A9E" box="[1332,1429,224,251]" country="Honduras" name="Lempira" pageId="10" pageNumber="577">Lempira</collectingRegion>
) (Fig. 86)
</materialsCitation>
.
</paragraph>
<paragraph id="E44127137B74FFCFFF0938709BF22D96" blockId="10.[151,1437,1903,2036]" pageId="10" pageNumber="577">
<emphasis id="D68AFB017B74FFCFFF093870983A2DED" bold="true" box="[151,310,1903,1928]" pageId="10" pageNumber="577">Figures 57–83.</emphasis>
Diagnostic estructures of male and female of
<taxonomicName id="23FE5C907B74FFCFFC91386F9AAA2DED" authorityName="Dejean" authorityYear="1821" box="[783,934,1904,1928]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="D68AFB017B74FFCFFC91386F9AAA2DED" box="[783,934,1904,1928]" italics="true" pageId="10" pageNumber="577">Macrodactylus</emphasis>
</taxonomicName>
. Parameres, distal view; parameres, lateral view; genital plates, ventral view.
<emphasis id="D68AFB017B74FFCFFE23388B98F02DC9" bold="true" box="[445,508,1939,1964]" pageId="10" pageNumber="577">57–59</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFD95388B9A4D2DCE" authority="Bates" authorityName="Bates" authorityYear="1887" box="[523,833,1940,1964]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="D68AFB017B74FFCFFD95388B9BF22DCE" box="[523,766,1940,1964]" italics="true" pageId="10" pageNumber="577">Macrodactylus nigripes</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B74FFCFFCD0388C9A812DC9" bold="true" box="[846,909,1939,1964]" pageId="10" pageNumber="577">60–62</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFC03388B9D682DCE" authority="Bates" authorityName="Bates" authorityYear="1887" box="[925,1124,1940,1964]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="D68AFB017B74FFCFFC03388B9D2D2DC9" box="[925,1057,1940,1964]" italics="true" pageId="10" pageNumber="577">M. rufescens</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B74FFCFFBEF388C9DBC2DC9" bold="true" box="[1137,1200,1939,1964]" pageId="10" pageNumber="577">63–65</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFB5E388B9C942DCE" authority="Bates" authorityName="Bates" box="[1216,1432,1940,1964]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="seiceicollis">
<emphasis id="D68AFB017B74FFCFFB5E388B9C592DC9" box="[1216,1365,1940,1964]" italics="true" pageId="10" pageNumber="577">M. seiceicollis</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B74FFCFFF0938A899DA2DB5" bold="true" box="[151,214,1975,2000]" pageId="10" pageNumber="577">66–68</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFF7C38A798932DAA" authority="Bates" authorityName="Bates" authorityYear="1887" box="[226,415,1976,1999]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="sericinus">
<emphasis id="D68AFB017B74FFCFFF7C38A798522DAA" box="[226,350,1976,1999]" italics="true" pageId="10" pageNumber="577">M. sericinus</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B74FFCFFE3738A898E42DB5" bold="true" box="[425,488,1975,2000]" pageId="10" pageNumber="577">69–71</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFE6A38A79B842DAA" authority="Bate" authorityName="Bates" authorityYear="1887" box="[500,648,1976,1999]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="suavis">
<emphasis id="D68AFB017B74FFCFFE6A38A79B582DAA" box="[500,596,1976,1999]" italics="true" pageId="10" pageNumber="577">M. suavis</emphasis>
Bate
</taxonomicName>
<emphasis id="D68AFB017B74FFCFFD1638A79B9E2DAA" box="[648,658,1976,1999]" italics="true" pageId="10" pageNumber="577">s</emphasis>
;
<emphasis id="D68AFB017B74FFCFFD0138A89BD22DB5" bold="true" box="[671,734,1975,2000]" pageId="10" pageNumber="577">72–74</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFD7438A79A9E2DAA" authority="Bates" authorityName="Bates" authorityYear="1887" box="[746,914,1976,2000]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="sylphis">
<emphasis id="D68AFB017B74FFCFFD7438A79A5D2DB5" box="[746,849,1976,2000]" italics="true" pageId="10" pageNumber="577">M. sylphis</emphasis>
Bates
</taxonomicName>
;
<emphasis id="D68AFB017B74FFCFFC0238A79AD72DB5" bold="true" box="[924,987,1976,2000]" pageId="10" pageNumber="577">75–77</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFC7938A79C202DB5" authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" authorityYear="2005" box="[999,1324,1976,2000]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="D68AFB017B74FFCFFC7938A79D422DB5" box="[999,1102,1976,2000]" italics="true" pageId="10" pageNumber="577">M. tibialis</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
;
<emphasis id="D68AFB017B74FFCFFAA838A89C792DB5" bold="true" box="[1334,1397,1975,2000]" pageId="10" pageNumber="577">78–80</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFA1F38A798D02D91" authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" authorityYear="2011" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="zaragozai">
<emphasis id="D68AFB017B74FFCFFA1F38A799F72D96" italics="true" pageId="10" pageNumber="577">M. zaragozai</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
;
<emphasis id="D68AFB017B74FFCFFE7838C49B292D91" bold="true" box="[486,549,2011,2036]" pageId="10" pageNumber="577">81–83</emphasis>
,
<taxonomicName id="23FE5C907B74FFCFFDAC38C39BF22D96" authority="Bates." authorityName="Bates." box="[562,766,2012,2036]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="D68AFB017B74FFCFFDAC38C39BB42D91" box="[562,696,2012,2036]" italics="true" pageId="10" pageNumber="577">M. zunilensis</emphasis>
Bates.
</taxonomicName>
</paragraph>
<paragraph id="E44127137B75FFCEFF093CEB98852E30" blockId="11.[151,1436,1012,1109]" pageId="11" pageNumber="578">
<emphasis id="D68AFB017B75FFCEFF093CEB980B2E68" bold="true" box="[151,263,1012,1037]" pageId="11" pageNumber="578">Figure 84.</emphasis>
Distribution of
<taxonomicName id="23FE5C907B75FFCEFE2E3CEA9B4B2E68" authorityName="Dejean" authorityYear="1821" box="[432,583,1013,1037]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="D68AFB017B75FFCEFE2E3CEA9B4B2E68" box="[432,583,1013,1037]" italics="true" pageId="11" pageNumber="578">Macrodactylus</emphasis>
</taxonomicName>
in the Central American Nucleus:
<taxonomicName id="23FE5C907B75FFCEFC2A3CE99D302E68" authorityName="Arce-Perez &amp; Moron" authorityYear="2020" box="[948,1084,1014,1037]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="eniocanoi">
<emphasis id="D68AFB017B75FFCEFC2A3CE99D302E68" box="[948,1084,1014,1037]" italics="true" pageId="11" pageNumber="578">M. eniocanoi</emphasis>
</taxonomicName>
(yellow circle),
<taxonomicName id="23FE5C907B75FFCEFB743CE99C902E68" authorityName="Arce-Perez &amp; Moron" authorityYear="2020" box="[1258,1436,1013,1037]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="escuintlaensis">
<emphasis id="D68AFB017B75FFCEFB743CE99C902E68" box="[1258,1436,1013,1037]" italics="true" pageId="11" pageNumber="578">M. escuintlaensis</emphasis>
</taxonomicName>
(blue circle),
<taxonomicName id="23FE5C907B75FFCEFE833B0598A92E54" authorityName="Bates" authorityYear="1887" box="[285,421,1049,1073]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="fulvescens">
<emphasis id="D68AFB017B75FFCEFE833B0598A92E54" box="[285,421,1049,1073]" italics="true" pageId="11" pageNumber="578">M. fulvescens</emphasis>
</taxonomicName>
(orange circle),
<taxonomicName id="23FE5C907B75FFCEFDD43B059BE92E54" authorityName="Arce-Perez &amp; Moron" authorityYear="2005" box="[586,741,1049,1073]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="hondurensis">
<emphasis id="D68AFB017B75FFCEFDD43B059BE92E54" box="[586,741,1049,1073]" italics="true" pageId="11" pageNumber="578">M. hondurensis</emphasis>
</taxonomicName>
(purple circle),
<taxonomicName id="23FE5C907B75FFCEFC1B3B059D2F2E54" authorityName="Bates" authorityYear="1887" box="[901,1059,1049,1073]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="lineatocollis">
<emphasis id="D68AFB017B75FFCEFC1B3B059D2F2E54" box="[901,1059,1049,1073]" italics="true" pageId="11" pageNumber="578">M. lineatocollis</emphasis>
</taxonomicName>
(red circle),
<taxonomicName id="23FE5C907B75FFCEFB3D3B059C062E54" authorityName="Bates" authorityYear="1887" box="[1187,1290,1049,1073]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="sylphis">
<emphasis id="D68AFB017B75FFCEFB3D3B059C062E54" box="[1187,1290,1049,1073]" italics="true" pageId="11" pageNumber="578">M. sylphis</emphasis>
</taxonomicName>
(green circle),
<taxonomicName id="23FE5C907B75FFCEFF093B2199F32E30" authorityName="Arce-Perez &amp; Moron" authorityYear="2005" box="[151,255,1085,1109]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="D68AFB017B75FFCEFF093B2199F32E30" box="[151,255,1085,1109]" italics="true" pageId="11" pageNumber="578">M. tibialis</emphasis>
</taxonomicName>
(pink circle).
</paragraph>
<paragraph id="E44127137B75FFCEFF0938AF9A0B2D88" blockId="11.[151,1436,1968,2029]" pageId="11" pageNumber="578">
<emphasis id="D68AFB017B75FFCEFF0938AF980B2DAC" bold="true" box="[151,263,1968,1993]" pageId="11" pageNumber="578">Figure 85.</emphasis>
Distribution of
<taxonomicName id="23FE5C907B75FFCEFE3138AE9B4A2DAC" authorityName="Dejean" authorityYear="1821" box="[431,582,1969,1993]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="D68AFB017B75FFCEFE3138AE9B4A2DAC" box="[431,582,1969,1993]" italics="true" pageId="11" pageNumber="578">Macrodactylus</emphasis>
</taxonomicName>
in the Central American Nucleus:
<taxonomicName id="23FE5C907B75FFCEFC2E38AE9D242DAC" authorityName="Arce-Perez &amp; Moron" authorityYear="2000" box="[944,1064,1969,1993]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="carrilloi">
<emphasis id="D68AFB017B75FFCEFC2E38AE9D242DAC" box="[944,1064,1969,1993]" italics="true" pageId="11" pageNumber="578">M. carrilloi</emphasis>
</taxonomicName>
(yellow circle),
<taxonomicName id="23FE5C907B75FFCEFB4838AE9C6C2DAC" authorityName="Bates" authorityYear="1887" box="[1238,1376,1969,1993]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="D68AFB017B75FFCEFB4838AE9C6C2DAC" box="[1238,1376,1969,1993]" italics="true" pageId="11" pageNumber="578">M. costulatus</emphasis>
</taxonomicName>
(blue circle),
<taxonomicName id="23FE5C907B75FFCEFF7B38CA987D2D88" authorityName="Bates" authorityYear="1887" box="[229,369,2005,2029]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="D68AFB017B75FFCEFF7B38CA987D2D88" box="[229,369,2005,2029]" italics="true" pageId="11" pageNumber="578">M. championi</emphasis>
</taxonomicName>
(green circle),
<taxonomicName id="23FE5C907B75FFCEFD9238CA9B852D89" authorityName="Bates" authorityYear="1887" box="[524,649,2005,2028]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="sericinus">
<emphasis id="D68AFB017B75FFCEFD9238CA9B852D89" box="[524,649,2005,2028]" italics="true" pageId="11" pageNumber="578">M. sericinus</emphasis>
</taxonomicName>
(red circle).
</paragraph>
</subSubSection>
</treatment>
</document>
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