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<document id="5AAE202004D28E2E31AEE1D22742DAA7" ID-CLB-Dataset="54548" ID-DOI="10.11646/zootaxa.5222.5.1" ID-GBIF-Dataset="a2aef7c8-09e1-48f1-983e-00132055a5a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7471736" ID-ZooBank="F4983B58-7301-4A3F-AC08-3E5120450EA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671694447742" checkinUser="plazi" docAuthor="Evenhuis, Neal L. &amp; Gharali, Babak" docDate="2022" docId="A1541B072616FFA8FF60FAFD1C4FFD71" docLanguage="en" docName="zootaxa.5222.5.1.pdf" docOrigin="Zootaxa 5222 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empidideicus (Anomaloptilus) irwini Evenhuis &amp; Gharali 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="412" masterDocId="5D6D637F261FFFA3FFF7FFEB1E24FFF4" masterDocTitle="The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar" masterLastPageNumber="426" masterPageNumber="401" pageNumber="410" updateTime="1699413895303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DE1E33FF3A0AC08D7D2939C52D9CD5F4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14783FECCC5F36575690E6996C659E19">
<mods:title id="602D18407E348FB441E32AF2015E828B">The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="FAF9296DC2C462A24D7B1C66A80D5894" type="personal">
<mods:role id="7E899BA9AFDC05BCC3179F3AFB24A827">
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<mods:namePart id="57013310735356CBB3FF0BD1430D02E1">Evenhuis, Neal L.</mods:namePart>
<mods:nameIdentifier id="3671EF4B81F89140F4E3C616DED2420A" type="ORCID">0000-0002-1314-755X</mods:nameIdentifier>
<mods:affiliation id="94EAC9301A77AE1C96DC361CFE1CF202">J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817 - 2704, USA neale @ bishopmuseum. org; https: // orcid. org / 0000 - 0002 - 1314 - 755 X</mods:affiliation>
<mods:nameIdentifier id="5F2B27F4AD85E71191F47A789EF071F7" type="email">neale@bishopmuseum.org</mods:nameIdentifier>
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<mods:name id="5693DDC82049D2774F567B5F22ADE6DC" type="personal">
<mods:role id="2B46705902D0B237073B73856AB34DD3">
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<mods:namePart id="B3A881BA8D2FD2430F70A6181F19F9F1">Gharali, Babak</mods:namePart>
<mods:nameIdentifier id="33C9092F63971D43BEC1CE85F9263C86" type="ORCID">0000-0003-2490-156X</mods:nameIdentifier>
<mods:affiliation id="6798B18EBE5CEDD579CA7424D0EC5B6B">Plant Protection Research Department, Qazvin Agricultural and Natural Resources Research and Education Center, AREEO, Qazvin, Iran. bgaharaei @ yahoo. com; https: // orcid. org / 0000 - 0003 - 2490 - 156 X</mods:affiliation>
<mods:nameIdentifier id="CBAE1CE3D9CF2DBD40E741C4569897FD" type="email">bgaharaei@yahoo.com</mods:nameIdentifier>
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<mods:title id="65FD230B8B9FE80B99BC3610644DA16A">Zootaxa</mods:title>
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<mods:part id="53B38A7C1C6465B8FD29975D344EF021">
<mods:date id="73319F13EFA1DD24E75AB33C8B189527">2022</mods:date>
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<mods:number id="C59E3E3B9D828FF9954F294F3736B2AF">2022-12-22</mods:number>
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<mods:number id="F9EE20DA48726D7F18FF9942554761AD">5222</mods:number>
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<treatment id="A1541B072616FFA8FF60FAFD1C4FFD71" ID-DOI="http://doi.org/10.5281/zenodo.7471782" ID-GBIF-Taxon="207927029" ID-Zenodo-Dep="7471782" LSID="urn:lsid:plazi:treatment:A1541B072616FFA8FF60FAFD1C4FFD71" httpUri="http://treatment.plazi.org/id/A1541B072616FFA8FF60FAFD1C4FFD71" lastPageId="11" lastPageNumber="412" pageId="9" pageNumber="410">
<subSubSection id="61E7F99A2616FFAAFF60FAFD1D89FAC5" box="[151,941,1302,1329]" pageId="9" pageNumber="410" type="nomenclature">
<paragraph id="2942AA112616FFAAFF60FAFD1D89FAC5" blockId="9.[151,941,1302,1365]" box="[151,941,1302,1329]" pageId="9" pageNumber="410">
<heading id="720A1D7D2616FFAAFF60FAFD1D89FAC5" bold="true" box="[151,941,1302,1329]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
<emphasis id="1B8976032616FFAAFF60FAFD1D89FAC5" bold="true" box="[151,941,1302,1329]" pageId="9" pageNumber="410">
<taxonomicName id="EEFDD1922616FFAAFF60FAFD1D7AFAC4" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[151,862,1302,1329]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B8976032616FFAAFF60FAFD1C7CFAC5" bold="true" box="[151,600,1302,1329]" italics="true" pageId="9" pageNumber="410">Empidideicus (Anomaloptilus) irwini</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<taxonomicNameLabel id="00BACB782616FFAAFC9DFAFC1D89FAC5" box="[874,941,1303,1329]" pageId="9" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2616FFAAFF60FAD01FCEFA11" pageId="9" pageNumber="410" type="description">
<paragraph id="2942AA112616FFAAFF60FAD01C4DFAA1" blockId="9.[151,941,1302,1365]" box="[151,617,1338,1365]" pageId="9" pageNumber="410">
(
<figureCitation id="B1C6B6942616FFAAFF68FAD01ECCFAA1" box="[159,232,1339,1365]" captionStart="FIGURES 5–10" captionStartId="4.[151,264,1768,1792]" captionTargetBox="[151,1435,181,1745]" captionTargetId="figure-17@4.[151,1435,181,1745]" captionTargetPageId="4" captionText="FIGURES 5–10. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 5, E. (A.) andriambahomananii, n. sp., male; 6, E. (A.) andriambahomananii, n. sp., female; 7, E. (A.) contrasticus, n. sp., male; 8, E. (A.) hauseri, n. sp., male; 9, E. (A.) irwini, n. sp., male; 10, E. (A.) nanus, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471746" httpUri="https://zenodo.org/record/7471746/files/figure.png" pageId="9" pageNumber="410">Figs 9</figureCitation>
,
<figureCitation id="B1C6B6942616FFAAFF03FAD01F36FAA1" box="[244,274,1339,1365]" captionStart="FIGURES 17–19" captionStartId="8.[151,264,1390,1414]" captionTargetBox="[151,1435,741,1367]" captionTargetId="figure-245@8.[151,1435,741,1367]" captionTargetPageId="8" captionText="FIGURES 17–19. Malagasy Empidideicus (Anomaloptilus), heads. 17, E. (A.) andriambahomananii, n. sp., male, oblique view; 18, E. (A.) irwini, n. sp., female, dorsal view; 19, E. (A.) rinhai, n. sp., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471750" httpUri="https://zenodo.org/record/7471750/files/figure.png" pageId="9" pageNumber="410">18</figureCitation>
,
<figureCitation id="B1C6B6942616FFAAFEE9FAD01F18FAA1" box="[286,316,1339,1365]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1965,1989]" captionTargetBox="[151,1363,1475,1942]" captionTargetId="figure-302@8.[151,1435,1475,1942]" captionTargetPageId="8" captionText="FIGURES 20–22. Malagasy Empidideicus (Anomaloptilus), heads. 20, E. (A.) contrasticus, n. sp., male, frontal view; 21, E. (A.) irwini, n. sp., male, oblique view; 22, E. (A.) rhinoclypeatus Evenhuis, male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471752" httpUri="https://zenodo.org/record/7471752/files/figure.png" pageId="9" pageNumber="410">21</figureCitation>
,
<figureCitation id="B1C6B6942616FFAAFEBFFAD01FB4FAA1" box="[328,400,1339,1365]" captionStart="FIGURES 25–33" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 25–33. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="9" pageNumber="410">30, 31</figureCitation>
,
<figureCitation id="B1C6B6942616FFAAFE6BFAD01F9EFAA1" box="[412,442,1339,1365]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="9" pageNumber="410">43</figureCitation>
,
<figureCitation id="B1C6B6942616FFAAFE31FAD11C2AFAA1" box="[454,526,1338,1365]" captionStart="FIGURES 50–58" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 50–58. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="9" pageNumber="410">55, 56</figureCitation>
,
<figureCitation id="B1C6B6942616FFAAFDEDFAD01C45FAA1" box="[538,609,1339,1365]" captionStart="FIGURES 69–70" captionStartId="21.[152,270,1016,1040]" captionTargetBox="[156,1429,422,990]" captionTargetId="figure-129@21.[151,1436,417,992]" captionTargetPageId="21" captionText="FIGURES 69–70. Empidideicus (Anomaloptilus) irwini, n. sp., genitalia. 69, male, A, gonocoxae, ventral view, B, epandrium, dorsal view, C, phallic complex, dorsal view; 70, female." figureDoi="http://doi.org/10.5281/zenodo.7471786" httpUri="https://zenodo.org/record/7471786/files/figure.png" pageId="9" pageNumber="410">69, 70</figureCitation>
)
</paragraph>
<paragraph id="2942AA112616FFAAFF60FA681FCEFA11" blockId="9.[151,1437,1411,1941]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF60FA681EF8FA69" bold="true" box="[151,220,1411,1437]" pageId="9" pageNumber="410">Types</emphasis>
.
<materialsCitation id="9995A04C2616FFAAFF07FA681AE7FA35" ID-GBIF-Occurrence="4003029356" collectingDate="1998-04-18" collectionCode="CAS" collectorName="M. E. Irwin &amp; E. I. Schlinger" country="Madagascar" latitude="-23.191668" location="Ifaty" longLatPrecision="855" longitude="43.616665" pageId="9" pageNumber="410" specimenCount="3" specimenCount-female="2" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="1B8976032616FFAAFF07FA681F73FA69" box="[240,343,1411,1437]" italics="true" pageId="9" pageNumber="410">
<typeStatus id="F64614B32616FFAAFF07FA681F73FA69" box="[240,343,1411,1437]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
</emphasis>
♂ and 2♂,
<specimenCount id="3FFB61982616FFAAFE18FA681C35FA69" box="[495,529,1411,1437]" pageId="9" pageNumber="410" type="female">2♀</specimenCount>
<emphasis id="1B8976032616FFAAFDE9FA681CA8FA68" box="[542,652,1411,1436]" italics="true" pageId="9" pageNumber="410">
<typeStatus id="F64614B32616FFAAFDE9FA681CA8FA68" box="[542,652,1411,1436]" pageId="9" pageNumber="410" type="paratype">paratypes</typeStatus>
</emphasis>
from
<collectingCountry id="51EAEA812616FFAAFD2CFA681D83FA69" box="[731,935,1411,1437]" name="Madagascar" pageId="9" pageNumber="410">MADAGASCAR</collectingCountry>
:
<collectingRegion id="EB3964F32616FFAAFC41FA681A5DFA69" box="[950,1145,1411,1437]" country="Madagascar" name="Toliara" pageId="9" pageNumber="410">Toliara Province</collectingRegion>
:
<location id="2C22FCCA2616FFAAFB7FFA681A9AFA69" LSID="urn:lsid:plazi:treatment:A1541B072616FFA8FF60FAFD1C4FFD71:2C22FCCA2616FFAAFB7FFA681A9AFA69" box="[1160,1214,1411,1437]" country="Madagascar" latitude="-23.191668" longLatPrecision="855" longitude="43.616665" name="Ifaty" pageId="9" pageNumber="410" stateProvince="Toliara">Ifaty</location>
,
<quantity id="EE0507F42616FFAAFB39FA681B3EFA69" box="[1230,1306,1411,1437]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="410" unit="km" value="22.0">22 km</quantity>
N
<collectingRegion id="EB3964F32616FFAAFAB1FA681BBCFA69" box="[1350,1432,1411,1437]" country="Madagascar" name="Toliara" pageId="9" pageNumber="410">Toliara</collectingRegion>
,
<geoCoordinate id="4CC9CCD62616FFAAFF60FA431F23FA37" box="[151,263,1448,1475]" degrees="23" direction="south" minutes="11.5" orientation="latitude" pageId="9" pageNumber="410" precision="92" value="-23.191668">23°11.5′S</geoCoordinate>
,
<geoCoordinate id="4CC9CCD62616FFAAFEE3FA431F55FA37" box="[276,369,1448,1475]" degrees="43" direction="east" minutes="37" orientation="longitude" pageId="9" pageNumber="410" precision="925" value="43.616665">43°37′E</geoCoordinate>
,
<date id="5D438CD12616FFAAFE89FA4C1C34FA35" box="[382,528,1447,1473]" pageId="9" pageNumber="410" value="1998-04-18">
<collectingDate id="4D0775392616FFAAFE89FA4C1C34FA35" box="[382,528,1447,1473]" pageId="9" pageNumber="410" value="1998-04-18">18 Apr 1998</collectingDate>
</date>
, hand netted, spiny forest,
<collectorName id="8408CFC72616FFAAFCB2FA4C1DE0FA35" box="[837,964,1447,1473]" pageId="9" pageNumber="410">M.E. Irwin</collectorName>
,
<collectorName id="8408CFC72616FFAAFC25FA4C1A4AFA35" box="[978,1134,1447,1473]" pageId="9" pageNumber="410">E.I. Schlinger</collectorName>
(
<collectionCode id="4FEC32D42616FFAAFB88FA4C1A9FFA35" box="[1151,1211,1447,1473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="410" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="9995A04C2616FFAAFB38FA4C1BBDFA35" ID-GBIF-Occurrence="4003029365" box="[1231,1433,1447,1473]" collectionCode="CAS" country="Madagascar" pageId="9" pageNumber="410" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="F64614B32616FFAAFB38FA4C1B13FA35" box="[1231,1335,1447,1473]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
in
<collectionCode id="4FEC32D42616FFAAFAA8FA4C1BBDFA35" box="[1375,1433,1447,1473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="410" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="9995A04C2616FFAAFF60FA201FC2FA11" ID-GBIF-Occurrence="4003029367" box="[151,486,1483,1509]" collectionCode="CAS, BPBM" country="Madagascar" pageId="9" pageNumber="410" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="F64614B32616FFAAFF60FA201F20FA11" box="[151,260,1483,1509]" pageId="9" pageNumber="410" type="paratype">Paratypes</typeStatus>
in
<collectionCode id="4FEC32D42616FFAAFEDFFA201F7AFA11" box="[296,350,1483,1509]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="410" type="Museum">CAS</collectionCode>
and
<collectionCode id="4FEC32D42616FFAAFE62FA201FC2FA11" box="[405,486,1483,1509]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="9" pageNumber="410" type="Museum">BPBM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2616FFAAFF30FA051A92F9A5" pageId="9" pageNumber="410" type="diagnosis">
<paragraph id="2942AA112616FFAAFF30FA051A92F9A5" blockId="9.[151,1437,1411,1941]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30FA051F1EF9FC" bold="true" box="[199,314,1518,1544]" pageId="9" pageNumber="410">Diagnosis</emphasis>
. Most similar to
<emphasis id="1B8976032616FFAAFE0CFA041C30F9FC" box="[507,532,1519,1544]" italics="true" pageId="9" pageNumber="410">E.</emphasis>
<taxonomicName id="EEFDD1922616FFAAFDEDFA041CF1F9FC" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[538,725,1519,1545]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032616FFAAFDD3FA041C11F9FD" box="[548,565,1519,1545]" italics="true" pageId="9" pageNumber="410">A</emphasis>
.)
<emphasis id="1B8976032616FFAAFDBBFA041CF1F9FC" box="[588,725,1519,1544]" italics="true" pageId="9" pageNumber="410">contrasticus</emphasis>
</taxonomicName>
<emphasis id="1B8976032616FFAAFD2BFA041D3FF9FD" bold="true" box="[732,795,1519,1545]" pageId="9" pageNumber="410">
<taxonomicNameLabel id="00BACB782616FFAAFD2BFA041D3FF9FD" box="[732,795,1519,1545]" pageId="9" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the short mouthparts, the proboscis not exceeding the oral margin, but can be easily separated from it by yellow scutellum (black in
<emphasis id="1B8976032616FFAAFBE4F9F81A0FF9D8" box="[1043,1067,1555,1580]" italics="true" pageId="9" pageNumber="410">E.</emphasis>
<taxonomicName id="EEFDD1922616FFAAFBC4F9F81ACBF9D8" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1075,1263,1555,1581]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032616FFAAFBCAF9F81A6AF9D9" box="[1085,1102,1555,1581]" italics="true" pageId="9" pageNumber="410">A</emphasis>
.)
<emphasis id="1B8976032616FFAAFB91F9F81ACBF9D8" box="[1126,1263,1555,1580]" italics="true" pageId="9" pageNumber="410">contrasticus</emphasis>
</taxonomicName>
<emphasis id="1B8976032616FFAAFB00F9F81B13F9D9" bold="true" box="[1271,1335,1555,1581]" pageId="9" pageNumber="410">
<taxonomicNameLabel id="00BACB782616FFAAFB00F9F81B13F9D9" box="[1271,1335,1555,1581]" pageId="9" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) and the eyes holoptic or nearly so below the antennae (clearly dichoptic in
<emphasis id="1B8976032616FFAAFC72F9DC1DBAF9A4" box="[901,926,1591,1616]" italics="true" pageId="9" pageNumber="410">E.</emphasis>
<taxonomicName id="EEFDD1922616FFAAFC52F9DC1A7BF9A4" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[933,1119,1591,1617]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032616FFAAFC59F9DC1D9BF9A5" box="[942,959,1591,1617]" italics="true" pageId="9" pageNumber="410">A</emphasis>
.)
<emphasis id="1B8976032616FFAAFC21F9DC1A7BF9A4" box="[982,1119,1591,1616]" italics="true" pageId="9" pageNumber="410">contrasticus</emphasis>
</taxonomicName>
<emphasis id="1B8976032616FFAAFB91F9DC1A81F9A5" bold="true" box="[1126,1189,1591,1617]" pageId="9" pageNumber="410">
<taxonomicNameLabel id="00BACB782616FFAAFB91F9DC1A81F9A5" box="[1126,1189,1591,1617]" pageId="9" pageNumber="410" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2616FFA8FF30F9B11B37FDC9" lastPageId="11" lastPageNumber="412" pageId="9" pageNumber="410" type="description">
<paragraph id="2942AA112616FFAAFF30F9B11F7EF981" blockId="9.[151,1437,1411,1941]" box="[199,346,1626,1653]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30F9B11F75F980" bold="true" box="[199,337,1626,1652]" pageId="9" pageNumber="410">Description</emphasis>
:
</paragraph>
<paragraph id="2942AA112616FFAAFF30F9941D3BF861" blockId="9.[151,1437,1411,1941]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30F9941F20F96D" bold="true" box="[199,260,1663,1689]" pageId="9" pageNumber="410">Male</emphasis>
(
<figureCitation id="B1C6B6942616FFAAFEE3F9941F7DF96D" box="[276,345,1663,1689]" captionStart="FIGURES 5–10" captionStartId="4.[151,264,1768,1792]" captionTargetBox="[151,1435,181,1745]" captionTargetId="figure-17@4.[151,1435,181,1745]" captionTargetPageId="4" captionText="FIGURES 5–10. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 5, E. (A.) andriambahomananii, n. sp., male; 6, E. (A.) andriambahomananii, n. sp., female; 7, E. (A.) contrasticus, n. sp., male; 8, E. (A.) hauseri, n. sp., male; 9, E. (A.) irwini, n. sp., male; 10, E. (A.) nanus, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471746" httpUri="https://zenodo.org/record/7471746/files/figure.png" pageId="9" pageNumber="410">Fig. 9</figureCitation>
). Lengths:
<quantity id="EE0507F42616FFAAFE2EF9941C5BF96D" box="[473,639,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.0899999999999999" metricValueMax="1.2" metricValueMin="0.98" pageId="9" pageNumber="410" unit="mm" value="1.0899999999999999" valueMax="1.2" valueMin="0.98">0.98–1.20 mm</quantity>
. Wing: 1.00–
<quantity id="EE0507F42616FFAAFCE1F9941D59F96D" box="[790,893,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="410" unit="mm" value="1.1">1.10 mm</quantity>
.
<emphasis id="1B8976032616FFAAFC7FF9941DE1F96D" box="[904,965,1663,1689]" italics="true" pageId="9" pageNumber="410">Head</emphasis>
(
<figureCitation id="B1C6B6942616FFAAFC22F9941A0EF96D" box="[981,1066,1663,1689]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1965,1989]" captionTargetBox="[151,1363,1475,1942]" captionTargetId="figure-302@8.[151,1435,1475,1942]" captionTargetPageId="8" captionText="FIGURES 20–22. Malagasy Empidideicus (Anomaloptilus), heads. 20, E. (A.) contrasticus, n. sp., male, frontal view; 21, E. (A.) irwini, n. sp., male, oblique view; 22, E. (A.) rhinoclypeatus Evenhuis, male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471752" httpUri="https://zenodo.org/record/7471752/files/figure.png" pageId="9" pageNumber="410">Fig. 21</figureCitation>
): globular, wider than long; eyes dichoptic above antennae separated at vertex by 1.5× distance between lateral ocelli, but holoptic or nearly so below level of antennae, separated by width of one ommatidium; occiput and postgena black; frons slightly depressed medially, with small black spot medially; face white, tip of oral margin white; antennae with scape yellow, short, subtrapezoidal; pedicel yellow, subconical, wider than long; first flagellomere black, lanceolate, length 2.5× width; second flagellomere black, three-fourths length of first flagellomere, with minute transparent apical style; mentum white; clypeus white, minute; proboscis dark slightly exerted beyond oral margin; labrum sclerotized, stiff, pointed apically, length subequal to proboscis; palpus not evident.
</paragraph>
<caption id="7D82FA992615FFA9FF60FAF61F3EFAAD" ID-DOI="http://doi.org/10.5281/zenodo.7471754" ID-Zenodo-Dep="7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="10" pageNumber="411" startId="10.[151,264,1309,1333]" targetBox="[151,1435,181,1284]" targetPageId="10">
<paragraph id="2942AA112615FFA9FF60FAF61F3EFAAD" blockId="10.[151,1436,1309,1369]" pageId="10" pageNumber="411">
<emphasis id="1B8976032615FFA9FF60FAF61F74FAC1" bold="true" box="[151,336,1309,1333]" pageId="10" pageNumber="411">FIGURES 23–24</emphasis>
. Malagasy
<taxonomicName id="EEFDD1922615FFA9FE3BFAF61D22FAC1" box="[460,774,1309,1333]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="subGenus" subGenus="Anomaloptilus">
<emphasis id="1B8976032615FFA9FE3BFAF61C72FAC1" box="[460,598,1309,1333]" italics="true" pageId="10" pageNumber="411">Empidideicus</emphasis>
(
<emphasis id="1B8976032615FFA9FD9FFAF61CD8FAC1" box="[616,764,1309,1333]" italics="true" pageId="10" pageNumber="411">Anomaloptilus</emphasis>
)
</taxonomicName>
, antennae.
<emphasis id="1B8976032615FFA9FC8AFAF61DB2FAC1" bold="true" box="[893,918,1309,1333]" pageId="10" pageNumber="411">23</emphasis>
,
<emphasis id="1B8976032615FFA9FC51FAF51D91FAC1" box="[934,949,1310,1333]" italics="true" pageId="10" pageNumber="411">E</emphasis>
.
<taxonomicName id="EEFDD1922615FFA9FC31FAF61AF6FAC1" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[966,1234,1309,1333]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032615FFA9FC39FAF61DF9FAC0" box="[974,989,1309,1332]" italics="true" pageId="10" pageNumber="411">A</emphasis>
.)
<emphasis id="1B8976032615FFA9FC01FAF61AF6FAC1" box="[1014,1234,1309,1333]" italics="true" pageId="10" pageNumber="411">andriambahomananii</emphasis>
</taxonomicName>
,
<emphasis id="1B8976032615FFA9FB16FAF61B39FAC1" bold="true" box="[1249,1309,1309,1333]" pageId="10" pageNumber="411">
<taxonomicNameLabel id="00BACB782615FFA9FB16FAF61B39FAC1" box="[1249,1309,1309,1333]" pageId="10" pageNumber="411" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="1B8976032615FFA9FADAFAF61B62FAC1" bold="true" box="[1325,1350,1309,1333]" pageId="10" pageNumber="411">24</emphasis>
,
<emphasis id="1B8976032615FFA9FAA1FAF51B41FAC1" box="[1366,1381,1310,1333]" italics="true" pageId="10" pageNumber="411">E</emphasis>
.
<taxonomicName id="EEFDD1922615FFA9FA81FAF61EF0FAAD" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032615FFA9FA89FAF61BA9FAC0" box="[1406,1421,1309,1332]" italics="true" pageId="10" pageNumber="411">A</emphasis>
.)
<emphasis id="1B8976032615FFA9FF60FAA91EF0FAAD" box="[151,212,1346,1369]" italics="true" pageId="10" pageNumber="411">nanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="00BACB782615FFA9FF17FAAA1F3EFAAD" box="[224,282,1345,1369]" pageId="10" pageNumber="411" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="2942AA112615FFA9FF30FA6C1D27F9C5" blockId="10.[151,1437,1415,2018]" pageId="10" pageNumber="411">
<emphasis id="1B8976032615FFA9FF30FA6C1F3CFA55" box="[199,280,1415,1441]" italics="true" pageId="10" pageNumber="411">Thorax</emphasis>
(
<figureCitation id="B1C6B6942615FFA9FED0FA6C1F5FFA55" box="[295,379,1415,1441]" captionStart="FIGURES 25–33" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 25–33. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="10" pageNumber="411">Fig. 30</figureCitation>
). Mesonotum matte black with white markings in following areas: dorsolaterally from humeral callus to postalar callus, interrupted above wing base, interhumeral mark not coalesced with mark on humeral callus; scutellum yellow; mesonotal disc and dorsum of scutellum with short fine brown hairs; pleura bare; pleura black with white on following: posterodorsal corner of anepisternum and anepimeron, upper half of meron and metepisternum; halter stem grayish brown, knob white.
</paragraph>
<paragraph id="2942AA112615FFA9FF30F9D71FA3F98D" blockId="10.[151,1437,1415,2018]" pageId="10" pageNumber="411">
<emphasis id="1B8976032615FFA9FF30F9D71ED8F9A1" box="[199,252,1596,1621]" italics="true" pageId="10" pageNumber="411">Legs</emphasis>
. Coxae and femora yellow, mid femur with some brown on basodorsal half; rest of legs white except tarsal segments 3–5 brown.
</paragraph>
<paragraph id="2942AA112615FFA9FF30F96F1AFCF8D9" blockId="10.[151,1437,1415,2018]" pageId="10" pageNumber="411">
<emphasis id="1B8976032615FFA9FF30F96F1F25F969" box="[199,257,1668,1693]" italics="true" pageId="10" pageNumber="411">Wing</emphasis>
(
<figureCitation id="B1C6B6942615FFA9FEE6F9681F42F969" box="[273,358,1667,1693]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="10" pageNumber="411">Fig. 43</figureCitation>
). Elongate, length 2.5× width; subhyaline; veins brown; costa ends slightly beyond end of R
<subScript id="B579A8542615FFA9FA8CF97D1BB1F950" attach="left" box="[1403,1429,1686,1700]" fontSize="6" pageId="10" pageNumber="411">4+5</subScript>
; vein Sc incomplete; Rs faded at connection with R
<subScript id="B579A8542615FFA9FD2CF9511CC7F93C" attach="left" box="[731,739,1722,1736]" fontSize="6" pageId="10" pageNumber="411">1</subScript>
; R
<subScript id="B579A8542615FFA9FCF1F9511D04F93C" attach="left" box="[774,800,1722,1736]" fontSize="6" pageId="10" pageNumber="411">4+5</subScript>
slightly curved, ending in wing margin beyond level of end of M
<subScript id="B579A8542615FFA9FEF4F9351F2FF918" attach="left" box="[259,267,1758,1772]" fontSize="6" pageId="10" pageNumber="411">2</subScript>
; vein M
<subScript id="B579A8542615FFA9FE98F9351F53F918" attach="left" box="[367,375,1758,1772]" fontSize="6" pageId="10" pageNumber="411">1</subScript>
curved to wing margin; M
<subScript id="B579A8542615FFA9FD58F9351C93F918" attach="left" box="[687,695,1758,1772]" fontSize="6" pageId="10" pageNumber="411">2</subScript>
straight; cell dm closed by apical crossvein, width one-half that of crossvein r-m; A
<subScript id="B579A8542615FFA9FE81F8E91F5AF8E4" attach="left" box="[374,382,1794,1808]" fontSize="6" pageId="10" pageNumber="411">1</subScript>
very thin, straight to wing margin; anal cell broadly open at wing margin; fringe of hair on posterior margin of wing well shortest at wing tip becoming longer and more sparse toward base.
</paragraph>
<paragraph id="2942AA112615FFA9FF30F8DC1CCCF881" blockId="10.[151,1437,1415,2018]" pageId="10" pageNumber="411">
<emphasis id="1B8976032615FFA9FF30F8DC1F15F8A5" box="[199,305,1847,1873]" italics="true" pageId="10" pageNumber="411">Abdomen</emphasis>
(
<figureCitation id="B1C6B6942615FFA9FEB4F8DC1FBEF8A6" box="[323,410,1847,1874]" captionStart="FIGURES 50–58" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 50–58. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="10" pageNumber="411">Fig. 55</figureCitation>
). Dorsum black, with thin white band posteriorly; tergites with sparsely scattered minute black hairs; sternites brown medially, white laterally.
</paragraph>
<paragraph id="2942AA112615FFA8FF30F8941EF2FEB5" blockId="10.[151,1437,1415,2018]" lastBlockId="11.[151,1437,151,645]" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
<emphasis id="1B8976032615FFA9FF30F8941F15F86D" box="[199,305,1919,1945]" italics="true" pageId="10" pageNumber="411">Genitalia</emphasis>
(
<figureCitation id="B1C6B6942615FFA9FEB5F8941FBCF86D" box="[322,408,1919,1945]" captionStart="FIGURES 69–70" captionStartId="21.[152,270,1016,1040]" captionTargetBox="[156,1429,422,990]" captionTargetId="figure-129@21.[151,1436,417,992]" captionTargetPageId="21" captionText="FIGURES 69–70. Empidideicus (Anomaloptilus) irwini, n. sp., genitalia. 69, male, A, gonocoxae, ventral view, B, epandrium, dorsal view, C, phallic complex, dorsal view; 70, female." figureDoi="http://doi.org/10.5281/zenodo.7471786" httpUri="https://zenodo.org/record/7471786/files/figure.png" pageId="10" pageNumber="411">Fig. 69</figureCitation>
). Hypopygium shiny brown. In ventral view with gonocoxites rounded, rectangular, small cleft medially bearing paired trident-shaped process, paired pegs medially on basal portion of gonocoxite lobes, with thin dark sclerotization basally and large seta medially on each lobe (
<figureCitation id="B1C6B6942615FFA9FC24F82C1A18F815" box="[979,1084,1991,2017]" captionStart="FIGURES 69–70" captionStartId="21.[152,270,1016,1040]" captionTargetBox="[156,1429,422,990]" captionTargetId="figure-129@21.[151,1436,417,992]" captionTargetPageId="21" captionText="FIGURES 69–70. Empidideicus (Anomaloptilus) irwini, n. sp., genitalia. 69, male, A, gonocoxae, ventral view, B, epandrium, dorsal view, C, phallic complex, dorsal view; 70, female." figureDoi="http://doi.org/10.5281/zenodo.7471786" httpUri="https://zenodo.org/record/7471786/files/figure.png" pageId="10" pageNumber="411">Fig. 69A</figureCitation>
); gonostylus subtriangular with long medial process; lateral arms of parameral sheath apodemes extending well below gonocoxa, foliate apically; aedeagal bulb large, rounded; aedeagal apodeme extending slightly beyond lateral arms of parameral sheath, with broad lanceolate lateral rami; upper portion of parameral sheath with bulbous apex in ventral view (
<figureCitation id="B1C6B6942614FFA8FAD2FF341BABFF0D" box="[1317,1423,223,249]" captionStart="FIGURES 69–70" captionStartId="21.[152,270,1016,1040]" captionTargetBox="[156,1429,422,990]" captionTargetId="figure-129@21.[151,1436,417,992]" captionTargetPageId="21" captionText="FIGURES 69–70. Empidideicus (Anomaloptilus) irwini, n. sp., genitalia. 69, male, A, gonocoxae, ventral view, B, epandrium, dorsal view, C, phallic complex, dorsal view; 70, female." figureDoi="http://doi.org/10.5281/zenodo.7471786" httpUri="https://zenodo.org/record/7471786/files/figure.png" pageId="11" pageNumber="412">Fig. 69C</figureCitation>
); epandrium subrectangular with large rounded, sclerotized apicoventral processes; cerci large, not exerted (
<figureCitation id="B1C6B6942614FFA8FA98FEE81EEDFEB5" captionStart="FIGURES 69–70" captionStartId="21.[152,270,1016,1040]" captionTargetBox="[156,1429,422,990]" captionTargetId="figure-129@21.[151,1436,417,992]" captionTargetPageId="21" captionText="FIGURES 69–70. Empidideicus (Anomaloptilus) irwini, n. sp., genitalia. 69, male, A, gonocoxae, ventral view, B, epandrium, dorsal view, C, phallic complex, dorsal view; 70, female." figureDoi="http://doi.org/10.5281/zenodo.7471786" httpUri="https://zenodo.org/record/7471786/files/figure.png" pageId="11" pageNumber="412">Fig. 69B</figureCitation>
).
</paragraph>
<paragraph id="2942AA112614FFA8FF30FEA01B37FDC9" blockId="11.[151,1437,151,645]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30FEA01F3AFE91" bold="true" box="[199,286,331,357]" pageId="11" pageNumber="412">Female</emphasis>
: As in male except eyes more dichoptic (
<figureCitation id="B1C6B6942614FFA8FD1CFEA01D1BFE91" box="[747,831,331,357]" captionStart="FIGURES 17–19" captionStartId="8.[151,264,1390,1414]" captionTargetBox="[151,1435,741,1367]" captionTargetId="figure-245@8.[151,1435,741,1367]" captionTargetPageId="8" captionText="FIGURES 17–19. Malagasy Empidideicus (Anomaloptilus), heads. 17, E. (A.) andriambahomananii, n. sp., male, oblique view; 18, E. (A.) irwini, n. sp., female, dorsal view; 19, E. (A.) rinhai, n. sp., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471750" httpUri="https://zenodo.org/record/7471750/files/figure.png" pageId="11" pageNumber="412">Fig. 18</figureCitation>
); thorax subshiny (
<figureCitation id="B1C6B6942614FFA8FBE2FEA01A4CFE91" box="[1045,1128,331,357]" captionStart="FIGURES 25–33" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 25–33. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="11" pageNumber="412">Fig. 31</figureCitation>
); tergal pattern of abdomen (
<figureCitation id="B1C6B6942614FFA8FF68FE841ECBFE7D" box="[159,239,366,393]" captionStart="FIGURES 50–58" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 50–58. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="11" pageNumber="412">Fig. 56</figureCitation>
) similar to that in
<emphasis id="1B8976032614FFA8FE41FE841FEBFE7C" box="[438,463,367,392]" italics="true" pageId="11" pageNumber="412">E.</emphasis>
<taxonomicName id="EEFDD1922614FFA8FE23FE841C7CFE7D" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[468,600,367,393]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032614FFA8FE2AFE841FCAFE7D" box="[477,494,367,393]" italics="true" pageId="11" pageNumber="412">A</emphasis>
.)
<emphasis id="1B8976032614FFA8FDF3FE841C7CFE7D" box="[516,600,367,393]" italics="true" pageId="11" pageNumber="412">hauseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00BACB782614FFA8FDA9FE841CBFFE7D" box="[606,667,367,393]" pageId="11" pageNumber="412" rank="species">
<emphasis id="1B8976032614FFA8FDA9FE841CB0FE7D" bold="true" box="[606,660,367,393]" pageId="11" pageNumber="412">n. sp</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="B1C6B6942614FFA8FD5EFE841CDDFE7D" box="[681,761,366,393]" captionStart="FIGURES 50–58" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 50–58. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="11" pageNumber="412">Fig. 54</figureCitation>
) but dark areas more pronounced. Genitalia (
<figureCitation id="B1C6B6942614FFA8FB1DFE841B1AFE7D" box="[1258,1342,367,393]" captionStart="FIGURES 69–70" captionStartId="21.[152,270,1016,1040]" captionTargetBox="[156,1429,422,990]" captionTargetId="figure-129@21.[151,1436,417,992]" captionTargetPageId="21" captionText="FIGURES 69–70. Empidideicus (Anomaloptilus) irwini, n. sp., genitalia. 69, male, A, gonocoxae, ventral view, B, epandrium, dorsal view, C, phallic complex, dorsal view; 70, female." figureDoi="http://doi.org/10.5281/zenodo.7471786" httpUri="https://zenodo.org/record/7471786/files/figure.png" pageId="11" pageNumber="412">Fig. 70</figureCitation>
): genital fork U-shaped, with two inwardly-directed process on lateral arms, median portion of “U” membranous, evanescent laterally, spermatheca spheroid-ellipsoid, sclerotized brown, longer than wide, small invagination apically with numerous glandular trichomes, with striations on basal three fourths; apical spermathecal duct subequal to length of spermatheca, thin, transparent; sperm pump minute, slightly sclerotized, without evident apical or basal valves; basal duct 1.5× length of apical duct, corrugated, slightly wider than apical duct; common duct absent.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2614FFA8FF30FDAC1C4FFD71" pageId="11" pageNumber="412" type="etymology">
<paragraph id="2942AA112614FFA8FF30FDAC1C4FFD71" blockId="11.[151,1437,151,645]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30FDAC1F6CFD95" bold="true" box="[199,328,583,609]" pageId="11" pageNumber="412">Etymology</emphasis>
. The specific epithet honors the collector, Michael E. Irwin and for his continued support of taxonomic research on Malagasy
<taxonomicName id="EEFDD1922614FFA8FDE7FD801C4CFD71" box="[528,616,619,645]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="11" pageNumber="412" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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