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<mods:title id="E2CE1BFC11A382B856356C1158664472">A new family for the enigmatic sea pen genus Gyrophyllum Studer, 1891 (Octocorallia, Pennatulacea), a molecular and morphological approach</mods:title>
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<mods:affiliation id="9B2744D10528C4F93858D3932D18A93F">Biodiversidad y Ecología Acuática. Departamento de Zoología, Facultad de Biología, Universidad de Sevilla, Avda. Reina Mercedes 6, 41012 - Sevilla, Spain.</mods:affiliation>
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<mods:namePart id="F44D5A8B8652629CA538318D23327FDA">Drewery, Jim</mods:namePart>
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<emphasis id="FE955D00A8215713837D81C94136FA72" bold="true" box="[564,866,1461,1488]" italics="true" pageId="11" pageNumber="84">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A8215713822381C94654FA6D" author="Studer T." box="[874,1024,1461,1487]" pageId="11" pageNumber="84" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
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<figureCitation id="54DA9D97A821571383A981A44107FA51" box="[736,851,1496,1523]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStartId-0="13.[189,232,1822,1848]" captionStartId-1="14.[189,232,1857,1883]" captionStartId-2="15.[189,232,1787,1813]" captionStartId-3="16.[189,232,1787,1813]" captionStartId-4="17.[189,232,1928,1954]" captionStartId-5="18.[189,232,1576,1602]" captionStartId-6="19.[189,232,1822,1848]" captionStartId-7="20.[189,232,1963,1989]" captionStartId-8="21.[189,232,1961,1987]" captionTargetBox-0="[253,1335,265,1785]" captionTargetBox-1="[244,1343,265,1817]" captionTargetBox-2="[346,1241,265,1749]" captionTargetBox-3="[267,1320,265,1749]" captionTargetBox-4="[429,1159,265,1890]" captionTargetBox-5="[330,1258,265,1538]" captionTargetBox-6="[189,1398,1134,1785]" captionTargetBox-7="[238,1349,265,1925]" captionTargetBox-8="[189,1398,573,1923]" captionTargetId-0="figure-12@13.[253,1335,265,1785]" captionTargetId-1="figure-15@14.[244,1343,265,1817]" captionTargetId-2="figure-11@15.[346,1241,265,1749]" captionTargetId-3="figure-15@16.[267,1320,265,1749]" captionTargetId-4="figure-11@17.[429,1159,265,1890]" captionTargetId-5="figure-14@18.[330,1258,265,1538]" captionTargetId-6="figure-277@19.[189,1398,1134,1785]" captionTargetId-7="figure-14@20.[238,1349,265,1925]" captionTargetId-8="figure-70@21.[189,1398,573,1923]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionTargetPageId-5="18" captionTargetPageId-6="19" captionTargetPageId-7="20" captionTargetPageId-8="21" captionText-0="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." captionText-1="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." captionText-2="Fig. 5. Gyrophyllum hirondellei Studer, 1891. Cross-sections of the axis near the rachis-peduncle limit of G. hirondellei colonies from SCOTIA cruises. A. Colony BECA (OPEN-660), a colony 112 mm in total length, showing an X-shaped cross-section, with an already apparent asymmetry. If growth rings are assumed to be produced annually this specimen is ~5 years old. B. Colony BECA (OPEN-661), a colony 215 mm in total length, showing a highly asymmetric X-shaped cross-section and a symmetrical central core (white dots). This specimen is ~14 years old according to growth rings with the proviso above." captionText-3="Fig. 6. Gyrophyllum hirondellei Studer, 1891. A–B. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). C–D. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. E. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). F. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves." captionText-4="Fig. 7. Gyrophyllum hirondellei Studer, 1891. Details of the ventral edge of a polyp leaf, showing the apertures of an autozooid (black arrows) and a single BPP (white arrows) in differing degrees of development per autozooid. A–B. Colony BECA (OPEN-661). C. Colony MNHM OCT.A.579." captionText-5="Fig. 8. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. B. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’into which each tentacle is partially retracted.C. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. D. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. E. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. F. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. G. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. H. Detail from G, also showing filiform structures." captionText-6="Fig. 9. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. SEM photographs of a tentacle in latero-oral view showing the numerous filiform structures. B. SEM photographs of a tentacle in latero-aboral view showing the naked aboral surface of tentacular axis and filiform structures. C. SEM photographs of a tentacle in lateral view showing digitiform normal pinnulae and, on the left, the numerous and elongate filiform structures. missing collection codes for specimens (as given in all other figs …)" captionText-7="Fig. 10. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Tentacle. B. Polyp leaf, siphonozooids area. C. Calycular pointed processes. D. Rachis exterior." captionText-8="Fig. 11. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Rachis interior. B. Peduncle exterior. C. Peduncle interior." figureDoi-0="http://doi.org/10.5281/zenodo.7374065" figureDoi-1="http://doi.org/10.5281/zenodo.7374067" figureDoi-2="http://doi.org/10.5281/zenodo.7374069" figureDoi-3="http://doi.org/10.5281/zenodo.7374071" figureDoi-4="http://doi.org/10.5281/zenodo.7374073" figureDoi-5="http://doi.org/10.5281/zenodo.7374075" figureDoi-6="http://doi.org/10.5281/zenodo.7374079" figureDoi-7="http://doi.org/10.5281/zenodo.7374082" figureDoi-8="http://doi.org/10.5281/zenodo.7374084" httpUri-0="https://zenodo.org/record/7374065/files/figure.png" httpUri-1="https://zenodo.org/record/7374067/files/figure.png" httpUri-2="https://zenodo.org/record/7374069/files/figure.png" httpUri-3="https://zenodo.org/record/7374071/files/figure.png" httpUri-4="https://zenodo.org/record/7374073/files/figure.png" httpUri-5="https://zenodo.org/record/7374075/files/figure.png" httpUri-6="https://zenodo.org/record/7374079/files/figure.png" httpUri-7="https://zenodo.org/record/7374082/files/figure.png" httpUri-8="https://zenodo.org/record/7374084/files/figure.png" pageId="11" pageNumber="84">Figs 3–11</figureCitation>
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<emphasis id="FE955D00A821571381F4826643B2F996" box="[189,486,1562,1588]" italics="true" pageId="11" pageNumber="84">Gyrophyllum hyrondellei</emphasis>
<bibRefCitation id="A870FCE3A821571380A7826640E1F996" author="Studer T." box="[494,693,1562,1588]" pageId="11" pageNumber="84" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891: 94</bibRefCitation>
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<emphasis id="FE955D00A821571381F4822743B1F9D7" box="[189,485,1627,1653]" italics="true" pageId="11" pageNumber="84">Gyrophyllum hyrondellei</emphasis>
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– 
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<bibRefCitation id="A870FCE3A8215713834B82274096F9D4" author="Studer T." box="[514,706,1627,1654]" pageId="11" pageNumber="84" pagination="1 - 64" refId="ref15703" refString="Studer T. 1901. Alcyonaires provenant des campagnes de l'Hirondelle (1886 - 88). Resultats des Campagnes Scientifiques du Prince Albert Ier de Monaco 20: 1 - 64." type="journal article" year="1901">Studer 1901: 35</bibRefCitation>
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<bibRefCitation id="A870FCE3A821571383B8822741EDF9D4" author="Roule L." box="[753,953,1627,1654]" pageId="11" pageNumber="84" pagination="454 - 458" refId="ref15555" refString="Roule L. 1905. Notice preliminaire sur les pennatulides recueillis par le &quot; Travailleur &quot; et le &quot; Talisman &quot;, dans l'Ocean Atlantique, au large du Maroc. Bulletin du Museum national d'histoire naturelle, Paris 11: 454 - 458." type="journal article" year="1905">Roule 1905: 456</bibRefCitation>
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<bibRefCitation id="A870FCE3A821571382A08227471AF9D4" author="Kukenthal W. &amp; Broch H." box="[1001,1358,1627,1654]" pageId="11" pageNumber="84" pagination="113 - 576" refId="ref14475" refString="Kukenthal W. &amp; Broch H. 1911. Pennatulacea. Wissenschaftiliche Ergebnisse Deutsche Tiefsee- Expedition &quot; Valdivia &quot; 13 (2): 113 - 576." type="journal article" year="1911">Kükenthal &amp; Broch 1911: 394</bibRefCitation>
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<bibRefCitation id="A870FCE3A8215713801C82034047F93B" author="Kukenthal W." box="[341,531,1662,1689]" pageId="11" pageNumber="84" pagination="43" refId="ref14436" refString="Kukenthal W. 1915. Pennatularia. In: Das Tierreich Lfg. 43. Verlag von R. Friedlander und Sohn, Berlin. https: // doi. org / 10.5962 / bhl. title. 1092" type="book chapter" year="1915">Kükenthal 1915</bibRefCitation>
(in text): 120. — 
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<bibRefCitation id="A870FCE3A821571383A58203419AF93B" author="Hickson S. J." box="[748,974,1662,1689]" pageId="11" pageNumber="84" pagination="1 - 265" refId="ref14146" refString="Hickson S. J. 1916. The Pennatulacea of the Siboga expedition, with a general survey of the order. Siboga Expeditie Monographs (14) 77: 1 - 265." type="journal article" year="1916">Hickson 1916: 252</bibRefCitation>
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(in text). — 
<treatmentCitation id="4D40A703A8215713852082034719F93B" author="Thomson J. A." box="[1129,1357,1662,1689]" page="56" pageId="11" pageNumber="84" year="1927">
<bibRefCitation id="A870FCE3A8215713852082034719F93B" author="Thomson J. A." box="[1129,1357,1662,1689]" pageId="11" pageNumber="84" pagination="1 - 77" refId="ref15793" refString="Thomson J. A. 1927. Alcyonaires provenant des campagnes scientifiques du Prince Albert 1 er de Monaco. Resultats des Campagnes Scientifiques Prince Albert Ier de Monaco 73: 1 - 77." type="journal article" year="1927">Thomson 1927: 56</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="4D40A703A821571381A282DE43A2F91E" author="Deichmann E." box="[235,502,1698,1724]" page="286" pageId="11" pageNumber="84" year="1936">
<bibRefCitation id="A870FCE3A821571381A282DE43A2F91E" author="Deichmann E." box="[235,502,1698,1724]" pageId="11" pageNumber="84" pagination="1 - 317" refId="ref13659" refString="Deichmann E. 1936. The Alcyonaria of the western part of the Atlantic Ocean. Memoirs of the Museum of Comparative Zoology, Harvard 53: 1 - 317. https: // doi. org / 10.5962 / bhl. title. 49348" type="journal article" year="1936">Deichmann 1936: 286</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="4D40A703A8215713836F82DE4645F91E" author="Tixier-Durivault A. &amp; d'Hondt M. - J." box="[550,1041,1698,1724]" page="1420" pageId="11" pageNumber="84" year="1974">
<bibRefCitation id="A870FCE3A8215713836F82DE4645F91E" author="Tixier-Durivault A. &amp; d'Hondt M. - J." box="[550,1041,1698,1724]" pageId="11" pageNumber="84" pagination="1361 - 1433" refId="ref15906" refString="Tixier-Durivault A. &amp; d'Hondt M. - J. 1974 b. Les octocoralliaires de la campagne Biacores. Bulletin du Museum national d'histoire naturelle, Paris 174: 1361 - 1433." type="journal article" year="1974">Tixier-Durivault &amp; d’Hondt 1974b: 1420</bibRefCitation>
</treatmentCitation>
. — 
<bibRefCitation id="A870FCE3A8215713850882DE46A8F91F" author="Williams G. C." box="[1089,1276,1698,1725]" pageId="11" pageNumber="84" pagination="93 - 140" refId="ref16020" refString="Williams G. C. 1995 a. Living genera of sea pens (Coelenterata: Octocorallia: Pennatulacea): illustrated key and synopses. Zoological Journal of the Linnean Society 114: 93 - 140. https: // doi. org / 10.1111 / j. 1096 - 3642.1995. tb 00929. x" type="journal article" year="1995">Williams 1995a</bibRefCitation>
(in text).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="84FBD299A821571481F4829F4397FBD2" lastPageId="12" lastPageNumber="85" pageId="11" pageNumber="84" type="materials_examined">
<paragraph id="CC5E8112A821571381F4829F43F0F95C" blockId="11.[189,420,1763,1790]" box="[189,420,1763,1790]" pageId="11" pageNumber="84">
<heading id="9716367EA821571381F4829F43F0F95C" bold="true" box="[189,420,1763,1790]" fontSize="11" level="3" pageId="11" pageNumber="84" reason="3">
<emphasis id="FE955D00A821571381F4829F43F0F95C" bold="true" box="[189,420,1763,1790]" italics="true" pageId="11" pageNumber="84">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="CC5E8112A821571481F4836E4397FBD2" blockId="11.[189,1399,1808,2012]" lastBlockId="12.[189,1400,265,1136]" lastPageId="12" lastPageNumber="85" pageId="11" pageNumber="84">
<materialsCitation id="7C898B4FA821571381F4836E4075F8D1" ID-GBIF-Occurrence="3971158304" collectingDate="1971-11-26" collectionCode="MNHM" elevation="0" latitude="38.608334" location="NORTH EASTERN ATLANTIC" longLatPrecision="117" longitude="-28.291666" municipality="Azores" pageId="11" pageNumber="84" specimenCount="1">
<location id="C93ED7C9A821571381F4836E4015F88E" LSID="urn:lsid:plazi:treatment:44483004A821570E837D81C94653FE19:C93ED7C9A821571381F4836E4015F88E" box="[189,577,1810,1836]" latitude="38.608334" longLatPrecision="117" longitude="-28.291666" municipality="Azores" name="NORTH EASTERN ATLANTIC" pageId="11" pageNumber="84">NORTH EASTERN ATLANTIC</location>
– 
<emphasis id="FE955D00A82157138329836D40E3F889" bold="true" box="[608,695,1809,1835]" pageId="11" pageNumber="84">
<collectingMunicipality id="2C3A1B68A82157138329836D40E3F889" box="[608,695,1809,1835]" name="Azores" pageId="11" pageNumber="84">Azores</collectingMunicipality>
</emphasis>
• 
<specimenCount id="DAE74A9BA82157138399836E4172F88E" box="[720,806,1810,1836]" pageId="11" pageNumber="84" type="generic">1 spec.</specimenCount>
; 
<location id="C93ED7C9A8215713827B836E465EF88E" LSID="urn:lsid:plazi:treatment:44483004A821570E837D81C94653FE19:C93ED7C9A8215713827B836E465EF88E" box="[818,1034,1810,1836]" latitude="38.608334" longLatPrecision="117" longitude="-28.291666" municipality="Azores" name="North Sao Miguel" pageId="11" pageNumber="84">North São Miguel</location>
; 
<geoCoordinate id="A9D5E7D5A8215713855E836E46C3F88E" box="[1047,1175,1808,1837]" degrees="38" direction="north" minutes="36.5" orientation="latitude" pageId="11" pageNumber="84" precision="92" value="38.608334">38º36.5′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A821571385EA836C477DF88E" box="[1187,1321,1808,1836]" degrees="28" direction="west" minutes="17.5" orientation="longitude" pageId="11" pageNumber="84" precision="92" value="-28.291666">28º17.5′ W</geoCoordinate>
; depth 
<elevation id="47CC6621A821571381F483494330F8F2" box="[189,356,1845,1872]" pageId="11" pageNumber="84">
1260– 
<quantity id="0B192CF7A8215713804E83494330F8F2" box="[263,356,1845,1872]" metricMagnitude="3" metricUnit="m" metricValue="1.258" pageId="11" pageNumber="84" unit="m" value="1258.0">1258 m</quantity>
</elevation>
; 
<date id="B85FA7D2A8215713803B83494043F8ED" box="[370,535,1845,1872]" pageId="11" pageNumber="84" value="1971-11-26">
<collectingDate id="A81B5E3AA8215713803B83494043F8ED" box="[370,535,1845,1872]" pageId="11" pageNumber="84" value="1971-11-26">26 Nov. 1971</collectingDate>
</date>
; BIAÇORES 1971 exped.; stn.139; complete colony, 
<quantity id="0B192CF7A821571385D3834946A9F8F2" box="[1178,1277,1845,1872]" metricMagnitude="-1" metricUnit="m" metricValue="1.58" pageId="11" pageNumber="84" unit="mm" value="158.0">158 mm</quantity>
in length; MNHM OCT.A.579; 
<collectionCode id="AAF019D7A821571380F383254048F8D1" box="[442,540,1881,1907]" collectionName="United States, Colorado, Colorado Springs, John May Museum of Natural History" country="USA" httpUri="http://grbio.org/cool/q3kg-qwj4" name="John May Museum of Natural History" pageId="11" pageNumber="84">MNHM</collectionCode>
.
</materialsCitation>
<materialsCitation id="7C898B4FA8215713836E83254665F81B" ID-GBIF-Occurrence="3971158310" accessionNumber="S20321" collectingDate="2020-09-29" collectingDateMax="2022-01-01" collectingDateMin="2020-09-29" collectionCode="NORTH, EASTERN" elevation="0" latitude="38.608334" location="South Rockall Slope" longLatPrecision="117" longitude="-28.291666" municipality="South Rockall Slope" pageId="11" pageNumber="84" specimenCode="SCOTIA 1420S" specimenCount="1">
– 
<emphasis id="FE955D00A8215713837083244162F8D0" bold="true" box="[569,822,1880,1906]" pageId="11" pageNumber="84">
<collectingMunicipality id="2C3A1B68A8215713837083244162F8D0" box="[569,822,1880,1906]" pageId="11" pageNumber="84">South Rockall Slope</collectingMunicipality>
</emphasis>
• 
<specimenCount id="DAE74A9BA82157138207832441F7F8D1" box="[846,931,1880,1907]" pageId="11" pageNumber="84" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A821571382E7832B466FF8D1" box="[942,1083,1879,1907]" degrees="56" direction="north" minutes="08.71" orientation="latitude" pageId="11" pageNumber="84" precision="9" value="56.14517">56º08.71′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A8215713850F832B4682F8D1" box="[1094,1238,1879,1907]" degrees="17" direction="west" minutes="34.64" orientation="longitude" pageId="11" pageNumber="84" precision="9" value="-17.577333">17º34.64′ W</geoCoordinate>
–56º07.25′ N, 17º34.89′ W; depth 
<elevation id="47CC6621A8215713808983074005F834" box="[448,593,1915,1942]" pageId="11" pageNumber="84">
997– 
<quantity id="0B192CF7A821571380B483074005F834" box="[509,593,1915,1942]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="11" pageNumber="84" unit="m" value="101.0">101 m</quantity>
</elevation>
; 
<date id="B85FA7D2A8215713832F83074141F837" box="[614,789,1915,1941]" pageId="11" pageNumber="84" value="2020-09-29">
<collectingDate id="A81B5E3AA8215713832F83074141F837" box="[614,789,1915,1941]" pageId="11" pageNumber="84" value="2020-09-29">29 Sep. 2020</collectingDate>
</date>
; 
<specimenCode id="9C472969A82157138263830741A5F837" box="[810,1009,1915,1941]" pageId="11" pageNumber="84">SCOTIA 1420S</specimenCode>
; stn. 
<accessionNumber id="D3B21CF1A82157138576830746CDF837" box="[1087,1177,1915,1941]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S20321" pageId="11" pageNumber="84" type="EnaNcbi">S20321</accessionNumber>
#8008; complete colony, 
<quantity id="0B192CF7A8215713806C83E343C6F81B" box="[293,402,1951,1977]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="11" pageNumber="84" unit="mm" value="103.0">103 mm</quantity>
in length; NMS.Z. 
<date id="B85FA7D2A821571383C783E340ADF81B" box="[654,761,1951,1977]" pageId="11" pageNumber="84" value="2022-01-01">2022.1.1</date>
- BECA (G-3832); NMS
</materialsCitation>
• 
<materialsCitation id="7C898B4FA8215714852883E34757FE81" ID-GBIF-Occurrence="3971158307" accessionNumber="S20321" collectingDate="2020-09-29" collectingDateMax="2022-01-02" collectingDateMin="2020-09-29" collectionCode="NMS, BECA" elevation="0" lastPageId="12" lastPageNumber="85" latitude="56.120834" longLatPrecision="10" longitude="-17.5815" pageId="11" pageNumber="84" specimenCode="SCOTIA 1420S" specimenCount="1">
<specimenCount id="DAE74A9BA8215713852883E34695F81B" box="[1121,1217,1951,1977]" pageId="11" pageNumber="84" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A8215713859183E14726F81B" box="[1240,1394,1949,1977]" degrees="56" direction="north" minutes="08.71" orientation="latitude" pageId="11" pageNumber="84" precision="9" value="56.14517">56º08.71′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A821571381F483BC4302F87E" box="[189,342,1984,2012]" degrees="17" direction="west" minutes="34.64" orientation="longitude" pageId="11" pageNumber="84" precision="9" value="-17.577333">17º34.64′ W</geoCoordinate>
–56º07.25′ N, 17º34.89′ W; depth 
<elevation id="47CC6621A8215713824783BE41CAF87E" box="[782,926,1986,2012]" pageId="11" pageNumber="84">
997– 
<quantity id="0B192CF7A8215713820583BE41CAF87E" box="[844,926,1986,2012]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="11" pageNumber="84" unit="m" value="101.0">101 m</quantity>
</elevation>
; 
<date id="B85FA7D2A821571382FB83BE460BF87E" box="[946,1119,1986,2012]" pageId="11" pageNumber="84" value="2020-09-29">
<collectingDate id="A81B5E3AA821571382FB83BE460BF87E" box="[946,1119,1986,2012]" pageId="11" pageNumber="84" value="2020-09-29">29 Sep. 2020</collectingDate>
</date>
; 
<specimenCode id="9C472969A8215713853A83BE476DF87E" box="[1139,1337,1986,2012]" pageId="11" pageNumber="84">SCOTIA 1420S</specimenCode>
; stn. 
<accessionNumber id="D3B21CF1A826571481F485754342FE81" box="[189,278,265,291]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S20321" pageId="12" pageNumber="85" type="EnaNcbi">S20321</accessionNumber>
#8009; complete colony, 
<quantity id="0B192CF7A82657148301857540CFFE81" box="[584,667,265,291]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="12" pageNumber="85" unit="mm" value="78.0">78 mm</quantity>
in length; NMS.Z. 
<date id="B85FA7D2A8265714823D857541B4FE81" box="[884,992,265,291]" pageId="12" pageNumber="85" value="2022-01-02">2022.1.2</date>
- BECA (G-3835); NMS
</materialsCitation>
• 
<materialsCitation id="7C898B4FA8265714845585754736FECB" ID-GBIF-Occurrence="3971158301" accessionNumber="S20322" collectingDate="2020-09-29" collectingDateMax="2022-01-03" collectingDateMin="2020-09-29" collectionCode="NMS, BECA" elevation="0" latitude="56.120834" longLatPrecision="10" longitude="-17.5815" pageId="12" pageNumber="85" specimenCode="SCOTIA 1420S" specimenCount="1">
<specimenCount id="DAE74A9BA8265714845585754726FE81" box="[1308,1394,265,291]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A826571481F48557431FFEE5" box="[189,331,299,327]" degrees="56" direction="north" minutes="08.71" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.14517">56º08.71′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A8265714801E855743BCFEE5" box="[343,488,299,327]" degrees="17" direction="west" minutes="34.64" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-17.577333">17º34.64′ W</geoCoordinate>
–56º07.25′ N, 17º34.89′ W; depth 
<elevation id="47CC6621A8265714823E855041ABFEE5" box="[887,1023,300,327]" pageId="12" pageNumber="85">
997– 
<quantity id="0B192CF7A826571482FC855041ABFEE5" box="[949,1023,300,327]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="12" pageNumber="85" unit="m" value="101.0">101 m</quantity>
</elevation>
; 
<date id="B85FA7D2A82657148542855046FCFEE4" box="[1035,1192,300,326]" pageId="12" pageNumber="85" value="2020-09-29">
<collectingDate id="A81B5E3AA82657148542855046FCFEE4" box="[1035,1192,300,326]" pageId="12" pageNumber="85" value="2020-09-29">29 Sep. 2020</collectingDate>
</date>
; 
<specimenCode id="9C472969A826571485FD85504725FEE4" box="[1204,1393,300,326]" pageId="12" pageNumber="85">SCOTIA 1420S</specimenCode>
; stn. 
<accessionNumber id="D3B21CF1A826571481B88533431FFECB" box="[241,331,335,361]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S20322" pageId="12" pageNumber="85" type="EnaNcbi">S20322</accessionNumber>
#8010; complete colony, 
<quantity id="0B192CF7A826571483C1853340BFFEC8" box="[648,747,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="85" unit="mm" value="110.0">110 mm</quantity>
in length; NMS.Z. 
<date id="B85FA7D2A826571482878533466EFECB" box="[974,1082,335,361]" pageId="12" pageNumber="85" value="2022-01-03">2022.1.3</date>
- BECA (G-3831); NMS
</materialsCitation>
• 
<materialsCitation id="7C898B4FA826571481F4850F4609FE12" ID-GBIF-Occurrence="3971158312" accessionNumber="S20322" collectingDate="2020-09-29" collectionCode="BECA" elevation="904" latitude="56.173332" longLatPrecision="10" longitude="-17.492834" pageId="12" pageNumber="85" specimenCode="SCOTIA 1420S" specimenCount="1">
<specimenCount id="DAE74A9BA826571481F4850F4345FE2F" box="[189,273,371,397]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A82657148054850D43FEFE2F" box="[285,426,369,397]" degrees="56" direction="north" minutes="08.83" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.147167">56º08.83′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A826571480FC850D4013FE2F" box="[437,583,369,397]" degrees="17" direction="west" minutes="29.77" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-17.496166">17º29.77′ W</geoCoordinate>
–56º10.40′ N, 17º29.57′ W; depth 
<quantity id="0B192CF7A8265714829D850E460FFE2F" box="[980,1115,370,397]" metricMagnitude="2" metricUnit="m" metricValue="9.035" metricValueMax="9.05" metricValueMin="9.02" pageId="12" pageNumber="85" unit="m" value="903.5" valueMax="905.0" valueMin="902.0">
<elevation id="47CC6621A8265714829D850E460FFE2F" box="[980,1115,370,397]" metricMagnitude="2" metricUnit="m" metricValue="9.035" metricValueMax="9.05" metricValueMin="9.02" pageId="12" pageNumber="85" unit="m" value="903.5" valueMax="905.0" valueMin="902.0">902–905 m</elevation>
</quantity>
; 
<date id="B85FA7D2A8265714852E850F4757FE2F" box="[1127,1283,371,397]" pageId="12" pageNumber="85" value="2020-09-29">
<collectingDate id="A81B5E3AA8265714852E850F4757FE2F" box="[1127,1283,371,397]" pageId="12" pageNumber="85" value="2020-09-29">29 Sep. 2020</collectingDate>
</date>
; 
<specimenCode id="9C472969A82657148447850F435FFE12" pageId="12" pageNumber="85">SCOTIA 1420S</specimenCode>
; stn. 
<accessionNumber id="D3B21CF1A8265714800485EA43F3FE12" box="[333,423,406,432]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S20322" pageId="12" pageNumber="85" type="EnaNcbi">S20322</accessionNumber>
#11835; 1 fragment; BECA OPEN-665 (G-3834); BECA
</materialsCitation>
• 
<materialsCitation id="7C898B4FA8265714853285EA461EFE54" ID-GBIF-Occurrence="3971158314" accessionNumber="S20323" collectingDate="2020-09-30" collectionCode="BECA" elevation="766" latitude="56.38" location="South West Rockall Slope" longLatPrecision="10" longitude="-17.378666" pageId="12" pageNumber="85" specimenCode="OPEN-660" specimenCount="1">
<specimenCount id="DAE74A9BA8265714853285EA4687FE12" box="[1147,1235,406,432]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A826571485A885E84726FE12" box="[1249,1394,404,432]" degrees="56" direction="north" minutes="24.30" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.405">56º24.30′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A826571481F485CB431BFE71" box="[189,335,439,467]" degrees="17" direction="west" minutes="23.04" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-17.384">17º23.04′ W</geoCoordinate>
–56º22.80′ N, 17º22.72′ W; depth 
<quantity id="0B192CF7A826571483A885C5413EFE71" box="[737,874,441,467]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" metricValueMax="7.709999999999999" metricValueMin="7.609999999999999" pageId="12" pageNumber="85" unit="m" value="766.0" valueMax="771.0" valueMin="761.0">
<elevation id="47CC6621A826571483A885C5413EFE71" box="[737,874,441,467]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" metricValueMax="7.709999999999999" metricValueMin="7.609999999999999" pageId="12" pageNumber="85" unit="m" value="766.0" valueMax="771.0" valueMin="761.0">761–771 m</elevation>
</quantity>
; 
<date id="B85FA7D2A8265714823F85C54640FE71" box="[886,1044,441,467]" pageId="12" pageNumber="85" value="2020-09-30">
<collectingDate id="A81B5E3AA8265714823F85C54640FE71" box="[886,1044,441,467]" pageId="12" pageNumber="85" value="2020-09-30">30 Sep. 2020</collectingDate>
</date>
; SCOTIA 1420S; stn. 
<accessionNumber id="D3B21CF1A8265714845485C54723FE71" box="[1309,1399,441,467]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S20323" pageId="12" pageNumber="85" type="EnaNcbi">S20323</accessionNumber>
#8006; complete colony, 
<quantity id="0B192CF7A826571480BE85A04009FE55" box="[503,605,476,503]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="12" pageNumber="85" unit="mm" value="112.0">112 mm</quantity>
in length; 
<collectionCode id="AAF019D7A826571483A585A0416DFE54" box="[748,825,476,502]" pageId="12" pageNumber="85">BECA</collectionCode>
<specimenCode id="9C472969A8265714820F85A0419CFE54" box="[838,968,476,502]" collectionCode="OPEN-" pageId="12" pageNumber="85">OPEN-660</specimenCode>
(G-3832).
</materialsCitation>
<materialsCitation id="7C898B4FA8265714851E85A1435DFDC2" ID-GBIF-Occurrence="3971158302" accessionNumber="S20323" collectingDate="2020-09-30" collectionCode="BECA" elevation="766" latitude="56.38" location="South West Rockall Slope" longLatPrecision="10" longitude="-17.378666" pageId="12" pageNumber="85" specimenCode="OPEN-660" specimenCount="1">
– 
<emphasis id="FE955D00A8265714853A85A04356FDBB" bold="true" pageId="12" pageNumber="85">
<location id="C93ED7C9A8265714853A85A04356FDBB" LSID="urn:lsid:plazi:treatment:44483004A821570E837D81C94653FE19:C93ED7C9A8265714853A85A04356FDBB" latitude="56.38" longLatPrecision="10" longitude="-17.378666" name="South West Rockall Slope" pageId="12" pageNumber="85">South West Rockall Slope</location>
</emphasis>
• 
<specimenCount id="DAE74A9BA8265714806C8583432AFDB8" box="[293,382,511,538]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A826571480C685824075FDB8" box="[399,545,510,538]" degrees="57" direction="north" minutes="07.11" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="57.1185">57º07.11′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A8265714837885824093FDB8" box="[561,711,510,538]" degrees="19" direction="west" minutes="59.63" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-19.993834">19º59.63′ W</geoCoordinate>
–57º04.87′ N, 20º00.47′ W; depth 
<quantity id="0B192CF7A826571485268583474DFDB8" box="[1135,1305,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.0055" metricValueMax="1.009" metricValueMin="1.002" pageId="12" pageNumber="85" unit="m" value="1005.5" valueMax="1009.0" valueMin="1002.0">
<elevation id="47CC6621A826571485268583474DFDB8" box="[1135,1305,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.0055" metricValueMax="1.009" metricValueMin="1.002" pageId="12" pageNumber="85" unit="m" value="1005.5" valueMax="1009.0" valueMin="1002.0">1002–1009 m</elevation>
</quantity>
; 
<date id="B85FA7D2A82657148463858342AFFD9F" pageId="12" pageNumber="85" value="2020-10-03">
<collectingDate id="A81B5E3AA82657148463858342AFFD9F" pageId="12" pageNumber="85" value="2020-10-03">3 Oct. 2020</collectingDate>
</date>
; SCOTIA 1420S; stn. 
<accessionNumber id="D3B21CF1A82657148341865F4035FD9F" box="[520,609,547,573]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S20331" pageId="12" pageNumber="85" type="EnaNcbi">S20331</accessionNumber>
#7706; complete colony, 
<quantity id="0B192CF7A826571482DE865F41BFFD9F" box="[919,1003,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="12" pageNumber="85" unit="mm" value="84.0">84 mm</quantity>
in length; 
<collectionCode id="AAF019D7A82657148524865F46EEFD9F" box="[1133,1210,547,573]" pageId="12" pageNumber="85">BECA</collectionCode>
<specimenCode id="9C472969A8265714858A865E4711FD9F" box="[1219,1349,546,573]" collectionCode="OPEN-" pageId="12" pageNumber="85">OPEN-659</specimenCode>
(G- 3830).
</materialsCitation>
<materialsCitation id="7C898B4FA8265714805A863A40F6FD04" ID-GBIF-Occurrence="3971158303" accessionNumber="S20323" collectingDate="2020-09-30" collectingDateMax="2022-01-04" collectingDateMin="2021-04-14" collectionCode="BECA" elevation="766" latitude="56.38" location="South West Rockall Slope" longLatPrecision="10" longitude="-17.378666" pageId="12" pageNumber="85" specimenCode="OPEN-660" specimenCount="1">
– 
<emphasis id="FE955D00A82657148065863A4072FDFD" bold="true" box="[300,550,581,608]" pageId="12" pageNumber="85">
<location id="C93ED7C9A82657148065863A4072FDFD" LSID="urn:lsid:plazi:treatment:44483004A821570E837D81C94653FE19:C93ED7C9A82657148065863A4072FDFD" box="[300,550,581,608]" latitude="57.081165" longLatPrecision="10" longitude="-20.007833" name="West Rockall Slope" pageId="12" pageNumber="85">West Rockall Slope</location>
</emphasis>
• 
<specimenCount id="DAE74A9BA8265714830C863A40C9FDC2" box="[581,669,582,608]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A826571483E586384169FDC2" box="[684,829,580,608]" degrees="56" direction="north" minutes="42.59" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.709835">56º42.59′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A82657148205863841B5FDC2" box="[844,993,580,608]" degrees="16" direction="west" minutes="30.85" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-16.514166">16º30.85′ W</geoCoordinate>
–56º42.31′ N, 16º37.05′ W; depth 
<quantity id="0B192CF7A826571481F486154312FD21" box="[189,326,617,643]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">
<elevation id="47CC6621A826571481F486154312FD21" box="[189,326,617,643]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">721–792 m</elevation>
</quantity>
; 
<date id="B85FA7D2A8265714801D861543A6FD21" box="[340,498,617,643]" pageId="12" pageNumber="85" value="2021-04-14">
<collectingDate id="A81B5E3AA8265714801D861543A6FD21" box="[340,498,617,643]" pageId="12" pageNumber="85" value="2021-04-14">14 Apr. 2021</collectingDate>
</date>
; SCOTIA 0421S; stn. 
<accessionNumber id="D3B21CF1A826571483B68615410CFD21" box="[767,856,617,643]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S21172" pageId="12" pageNumber="85" type="EnaNcbi">S21172</accessionNumber>
#11670; complete colony, 
<quantity id="0B192CF7A826571485D3861546A9FD21" box="[1178,1277,617,643]" metricMagnitude="-1" metricUnit="m" metricValue="2.03" pageId="12" pageNumber="85" unit="mm" value="203.0">203 mm</quantity>
in length; 
<collectionCode id="AAF019D7A826571481F486F042AAFD04" box="[189,254,652,678]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" country="United Kingdom" httpUri="http://grbio.org/cool/h8hj-4rxx" name="National Museum of Scotland - Natural Sciences" pageId="12" pageNumber="85" type="Museum">NMS</collectionCode>
.Z. 
<date id="B85FA7D2A8265714805186F043D0FD04" box="[280,388,652,678]" pageId="12" pageNumber="85" value="2022-01-04">
<collectingDate id="A81B5E3AA8265714805186F043D0FD04" box="[280,388,652,678]" pageId="12" pageNumber="85" value="2022-01-04">2022.1.4</collectingDate>
</date>
- 
<collectionCode id="AAF019D7A826571480DC86F043B7FD04" box="[405,483,652,678]" pageId="12" pageNumber="85">BECA</collectionCode>
(G-4015); 
<collectionCode id="AAF019D7A8265714832B86F040F6FD04" box="[610,674,652,678]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" country="United Kingdom" httpUri="http://grbio.org/cool/h8hj-4rxx" name="National Museum of Scotland - Natural Sciences" pageId="12" pageNumber="85" type="Museum">NMS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="7C898B4FA826571483FE86F0400AFD4F" ID-GBIF-Occurrence="3971158311" accessionNumber="S21172" collectingDate="2021-04-14" collectionCode="BECA" elevation="757" latitude="56.705166" longLatPrecision="10" longitude="-16.6175" pageId="12" pageNumber="85" specimenCode="SCOTIA 0421S" specimenCount="1">
<specimenCount id="DAE74A9BA826571483FE86F0415EFD05" box="[695,778,652,679]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A8265714825D86F741F5FD05" box="[788,929,651,679]" degrees="56" direction="north" minutes="42.59" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.709835">56º42.59′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A826571482E286F7466FFD05" box="[939,1083,651,679]" degrees="16" direction="west" minutes="30.85" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-16.514166">16º30.85′ W</geoCoordinate>
–56º42.31′ N, 16º37.05′ W; depth 
<quantity id="0B192CF7A8265714804B86D343D3FD68" box="[258,391,687,714]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">
<elevation id="47CC6621A8265714804B86D343D3FD68" box="[258,391,687,714]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">721–792 m</elevation>
</quantity>
; 
<date id="B85FA7D2A826571480D986D34071FD6B" box="[400,549,687,714]" pageId="12" pageNumber="85" value="2021-04-14">
<collectingDate id="A81B5E3AA826571480D986D34071FD6B" box="[400,549,687,714]" pageId="12" pageNumber="85" value="2021-04-14">14Apr.2021</collectingDate>
</date>
; 
<specimenCode id="9C472969A8265714836786D340BDFD6B" box="[558,745,687,713]" pageId="12" pageNumber="85">SCOTIA 0421S</specimenCode>
; stn. 
<accessionNumber id="D3B21CF1A8265714825686D3412CFD6B" box="[799,888,687,713]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S21172" pageId="12" pageNumber="85" type="EnaNcbi">S21172</accessionNumber>
#11671;complete colony, 
<quantity id="0B192CF7A826571485E186D34752FD68" box="[1192,1286,687,714]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" pageId="12" pageNumber="85" unit="mm" value="215.0">215 mm</quantity>
in length; BECA OPEN-661 (G-4016); BECA
</materialsCitation>
• 
<materialsCitation id="7C898B4FA8265714833A86AF40F1FC91" ID-GBIF-Occurrence="3971158313" accessionNumber="S21172" collectingDate="2021-04-14" collectingDateMax="2022-01-05" collectingDateMin="2021-04-14" collectionCode="NMS, BECA" elevation="757" latitude="56.705166" longLatPrecision="10" longitude="-16.6175" pageId="12" pageNumber="85" specimenCode="SCOTIA 0421" specimenCount="1">
<specimenCount id="DAE74A9BA8265714833A86AF4092FD4F" box="[627,710,723,749]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A8265714839986AD4108FD4F" box="[720,860,721,749]" degrees="56" direction="north" minutes="42.59" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.709835">56º42.59′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A8265714822F86AD41A1FD4F" box="[870,1013,721,749]" degrees="16" direction="west" minutes="30.85" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-16.514166">16º30.85′ W</geoCoordinate>
–56º42.31′ N, 16º37.05′ W; depth 
<quantity id="0B192CF7A826571481F4868A4311FCB2" box="[189,325,758,784]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">
<elevation id="47CC6621A826571481F4868A4311FCB2" box="[189,325,758,784]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">721–792 m</elevation>
</quantity>
; 
<date id="B85FA7D2A8265714801B868A43BBFCB2" box="[338,495,758,784]" pageId="12" pageNumber="85" value="2021-04-14">
<collectingDate id="A81B5E3AA8265714801B868A43BBFCB2" box="[338,495,758,784]" pageId="12" pageNumber="85" value="2021-04-14">14 Apr. 2021</collectingDate>
</date>
; 
<specimenCode id="9C472969A826571480B2868A40FFFCB2" box="[507,683,758,784]" pageId="12" pageNumber="85">SCOTIA 0421</specimenCode>
S; stn. 
<accessionNumber id="D3B21CF1A826571483B1868A4105FCB2" box="[760,849,758,784]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S21172" pageId="12" pageNumber="85" type="EnaNcbi">S21172</accessionNumber>
#11672; incomplete colony, lacking peduncle; NMS.Z. 
<date id="B85FA7D2A82657148051876543D0FC96" box="[280,388,793,820]" pageId="12" pageNumber="85" value="2022-01-05">2022.1.5</date>
- BECA (G-4017); NMS
</materialsCitation>
• 
<materialsCitation id="7C898B4FA826571483F087654158FCDB" ID-GBIF-Occurrence="3971158306" accessionNumber="S21172" collectingDate="2021-04-14" collectingDateMax="2022-01-06" collectingDateMin="2021-04-14" collectionCode="NMS, BECA" elevation="757" latitude="56.705166" longLatPrecision="10" longitude="-16.6175" pageId="12" pageNumber="85" specimenCode="SCOTIA 0421" specimenCount="1">
<specimenCount id="DAE74A9BA826571483F087654158FC91" box="[697,780,793,819]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A8265714825F876B41F6FC91" box="[790,930,791,819]" degrees="56" direction="north" minutes="42.59" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.709835">56º42.59′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A826571482E5876B4668FC91" box="[940,1084,791,819]" degrees="16" direction="west" minutes="30.85" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-16.514166">16º30.85′ W</geoCoordinate>
–56º42.31′ N, 16º37.05′ W; depth 
<quantity id="0B192CF7A82657148041874043C6FCF5" box="[264,402,828,855]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">
<elevation id="47CC6621A82657148041874043C6FCF5" box="[264,402,828,855]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">721–792 m</elevation>
</quantity>
; 
<date id="B85FA7D2A826571480E887404016FCF4" box="[417,578,828,854]" pageId="12" pageNumber="85" value="2021-04-14">
<collectingDate id="A81B5E3AA826571480E887404016FCF4" box="[417,578,828,854]" pageId="12" pageNumber="85" value="2021-04-14">14 Apr. 2021</collectingDate>
</date>
; 
<specimenCode id="9C472969A8265714831987404156FCF4" box="[592,770,828,854]" pageId="12" pageNumber="85">SCOTIA 0421</specimenCode>
S; stn. 
<accessionNumber id="D3B21CF1A8265714821A874041F8FCF4" box="[851,940,828,854]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S21172" pageId="12" pageNumber="85" type="EnaNcbi">S21172</accessionNumber>
#11678; complete colony, 
<quantity id="0B192CF7A826571485BB87404702FCF5" box="[1266,1366,828,855]" metricMagnitude="-1" metricUnit="m" metricValue="2.18" pageId="12" pageNumber="85" unit="mm" value="218.0">218 mm</quantity>
in length; NMS.Z. 
<date id="B85FA7D2A8265714803C872343B5FCDB" box="[373,481,863,889]" pageId="12" pageNumber="85" value="2022-01-06">2022.1.6</date>
-BECA (G-4018); NMS
</materialsCitation>
• 
<materialsCitation id="7C898B4FA826571482678723411DFC62" ID-GBIF-Occurrence="3971158309" accessionNumber="S21172" collectingDate="2021-04-14" collectionCode="BECA" elevation="757" latitude="56.705166" location="South East Rosemary Seamount" longLatPrecision="10" longitude="-16.6175" pageId="12" pageNumber="85" specimenCode="OPEN-662" specimenCount="1">
<specimenCount id="DAE74A9BA82657148267872341D3FCD8" box="[814,903,863,890]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A826571482D18722467EFCD8" box="[920,1066,862,890]" degrees="56" direction="north" minutes="42.59" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="56.709835">56º42.59′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A8265714857287224685FCD8" box="[1083,1233,862,890]" degrees="16" direction="west" minutes="30.85" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-16.514166">16º30.85′ W</geoCoordinate>
–56º42.31′ N, 16º37.05′ W; depth 
<quantity id="0B192CF7A826571480EA87FF407EFC3F" box="[419,554,899,925]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">
<elevation id="47CC6621A826571480EA87FF407EFC3F" box="[419,554,899,925]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="7.92" metricValueMin="7.209999999999999" pageId="12" pageNumber="85" unit="m" value="756.5" valueMax="792.0" valueMin="721.0">721–792 m</elevation>
</quantity>
; 
<date id="B85FA7D2A8265714837F87FF4084FC3F" box="[566,720,899,925]" pageId="12" pageNumber="85" value="2021-04-14">
<collectingDate id="A81B5E3AA8265714837F87FF4084FC3F" box="[566,720,899,925]" pageId="12" pageNumber="85" value="2021-04-14">14 Apr. 2021</collectingDate>
</date>
; SCOTIA 0421S; stn. 
<accessionNumber id="D3B21CF1A8265714829D87FF4678FC3F" box="[980,1068,899,925]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S21172" pageId="12" pageNumber="85" type="EnaNcbi">S21172</accessionNumber>
#11801; incomplete colony, lacking peduncle; 
<collectionCode id="AAF019D7A826571480DF87DA43B7FC62" box="[406,483,934,960]" pageId="12" pageNumber="85">BECA</collectionCode>
<specimenCode id="9C472969A826571480A287DA4038FC62" box="[491,620,934,960]" collectionCode="OPEN-" pageId="12" pageNumber="85">OPEN-662</specimenCode>
(G-4019); 
<collectionCode id="AAF019D7A826571483BA87DA4111FC62" box="[755,837,934,960]" pageId="12" pageNumber="85">BECA</collectionCode>
.
</materialsCitation>
<materialsCitation id="7C898B4FA8265714821987DA4320FB88" ID-GBIF-Occurrence="3971158305" accessionNumber="S21172" collectingDate="2021-04-14" collectionCode="BECA" elevation="757" latitude="56.705166" location="South East Rosemary Seamount" longLatPrecision="10" longitude="-16.6175" pageId="12" pageNumber="85" specimenCode="OPEN-662" specimenCount="1">
– 
<emphasis id="FE955D00A8265714822187D94756FC1D" bold="true" box="[872,1282,933,960]" pageId="12" pageNumber="85">
<location id="C93ED7C9A8265714822187D94756FC1D" LSID="urn:lsid:plazi:treatment:44483004A821570E837D81C94653FE19:C93ED7C9A8265714822187D94756FC1D" box="[872,1282,933,960]" latitude="56.705166" longLatPrecision="10" longitude="-16.6175" name="South East Rosemary Seamount" pageId="12" pageNumber="85">South East Rosemary Seamount</location>
</emphasis>
• 
<specimenCount id="DAE74A9BA8265714845487DA4727FC62" box="[1309,1395,934,960]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A826571481F787BB431BFC41" box="[190,335,967,995]" degrees="59" direction="north" minutes="05.85" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="59.0975">59º05.85′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A8265714801687BB43A1FC41" box="[351,501,967,995]" degrees="09" direction="west" minutes="52.94" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-9.882334">09º52.94′ W</geoCoordinate>
–59º04.70′ N, 09º55.40′ W; depth 
<quantity id="0B192CF7A826571482D387B54617FC41" box="[922,1091,969,996]" metricMagnitude="3" metricUnit="m" metricValue="1.0605" metricValueMax="1.0699999999999998" metricValueMin="1.051" pageId="12" pageNumber="85" unit="m" value="1060.5" valueMax="1070.0" valueMin="1051.0">
<elevation id="47CC6621A826571482D387B54617FC41" box="[922,1091,969,996]" metricMagnitude="3" metricUnit="m" metricValue="1.0605" metricValueMax="1.0699999999999998" metricValueMin="1.051" pageId="12" pageNumber="85" unit="m" value="1060.5" valueMax="1070.0" valueMin="1051.0">1051–1070 m</elevation>
</quantity>
; 
<date id="B85FA7D2A8265714851D87B546AAFC41" box="[1108,1278,969,995]" pageId="12" pageNumber="85" value="2021-11-10">
<collectingDate id="A81B5E3AA8265714851D87B546AAFC41" box="[1108,1278,969,995]" pageId="12" pageNumber="85" value="2021-11-10">10 Nov. 2021</collectingDate>
</date>
; SCOTIA 1621S; stn. 
<accessionNumber id="D3B21CF1A8265714800E879043F5FBA5" box="[327,417,1004,1031]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S21553" pageId="12" pageNumber="85" type="EnaNcbi">S21553</accessionNumber>
#11834; complete colony, but peduncle eroded, 
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in length; 
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(G-4095); 
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</materialsCitation>
• 
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<specimenCount id="DAE74A9BA826571480C4807343B5FB88" box="[397,481,1039,1066]" pageId="12" pageNumber="85" type="generic">1 spec.</specimenCount>
; 
<geoCoordinate id="A9D5E7D5A826571480A48072402FFB88" box="[493,635,1038,1066]" degrees="59" direction="north" minutes="05.85" orientation="latitude" pageId="12" pageNumber="85" precision="9" value="59.0975">59º05.85′ N</geoCoordinate>
, 
<geoCoordinate id="A9D5E7D5A826571483CE8072414CFB88" box="[647,792,1038,1066]" degrees="09" direction="west" minutes="52.94" orientation="longitude" pageId="12" pageNumber="85" precision="9" value="-9.882334">09º52.94′ W</geoCoordinate>
–59º04.70′ N, 09º55.40′ W; depth 
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<elevation id="47CC6621A826571485E180734719FB88" box="[1192,1357,1039,1066]" metricMagnitude="3" metricUnit="m" metricValue="1.0605" metricValueMax="1.0699999999999998" metricValueMin="1.051" pageId="12" pageNumber="85" unit="m" value="1060.5" valueMax="1070.0" valueMin="1051.0">1051–1070 m</elevation>
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; 
<date id="B85FA7D2A8265714841080734368FBEF" pageId="12" pageNumber="85" value="2021-11-10">
<collectingDate id="A81B5E3AA8265714841080734368FBEF" pageId="12" pageNumber="85" value="2021-11-10">10 Nov. 2021</collectingDate>
</date>
; 
<specimenCode id="9C472969A82657148000804F43ADFBEE" box="[329,505,1074,1101]" pageId="12" pageNumber="85">SCOTIA 1621</specimenCode>
S; stn. 
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#11833; complete colony, 
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in length; BECA OPEN- 664 (G-4096); BECA
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.
</paragraph>
</subSubSection>
<subSubSection id="84FBD299A826570D81F480EB41A6FE19" lastPageId="21" lastPageNumber="94" pageId="12" pageNumber="85" type="description">
<paragraph id="CC5E8112A826571481F480EB4059FB13" blockId="12.[189,525,1175,1201]" box="[189,525,1175,1201]" pageId="12" pageNumber="85">
<heading id="9716367EA826571481F480EB4059FB13" bold="true" box="[189,525,1175,1201]" fontSize="11" level="3" pageId="12" pageNumber="85" reason="3">
<emphasis id="FE955D00A826571481F480EB4059FB13" bold="true" box="[189,525,1175,1201]" pageId="12" pageNumber="85">Morphological description</emphasis>
</heading>
</paragraph>
<paragraph id="CC5E8112A826570B81F480BA4138FECB" blockId="12.[188,1401,1222,2023]" lastBlockId="19.[188,1398,265,362]" lastPageId="19" lastPageNumber="92" pageId="12" pageNumber="85">
Colonies stout and clavate, pinnate distally (
<figureCitation id="54DA9D97A8265714838480BA416DFB42" box="[717,825,1222,1248]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="13.[189,232,1822,1848]" captionStartId-1="14.[189,232,1857,1883]" captionTargetBox-0="[253,1335,265,1785]" captionTargetBox-1="[244,1343,265,1817]" captionTargetId-0="figure-12@13.[253,1335,265,1785]" captionTargetId-1="figure-15@14.[244,1343,265,1817]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." captionText-1="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi-0="http://doi.org/10.5281/zenodo.7374065" figureDoi-1="http://doi.org/10.5281/zenodo.7374067" httpUri-0="https://zenodo.org/record/7374065/files/figure.png" httpUri-1="https://zenodo.org/record/7374067/files/figure.png" pageId="12" pageNumber="85">Figs 3–4</figureCitation>
), up to 
<quantity id="0B192CF7A826571482DC80BA41ACFB42" box="[917,1016,1222,1248]" metricMagnitude="-1" metricUnit="m" metricValue="2.18" pageId="12" pageNumber="85" unit="mm" value="218.0">218 mm</quantity>
in length in the preserved state. Rachis in two distinct parts: distally a bilaterally symmetrical section bearing polyp leaves, and proximally a stalk of shorter length. Complete rachis is up to 
<quantity id="0B192CF7A8265714823481704188FA85" box="[893,988,1292,1319]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="85" unit="mm" value="110.0">110 mm</quantity>
in length (50.46% of overall length in the whole examined material) and up to 
<quantity id="0B192CF7A826571483E781534154FAE8" box="[686,768,1327,1354]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="85" unit="mm" value="18.0">18 mm</quantity>
in width (measured at mid-length of distal rachis part, not including polyp leaves). Rachis-peduncle limit slightly prominently swollen (
<figureCitation id="54DA9D97A826571485CF812E46A7FACF" box="[1158,1267,1362,1389]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="13.[189,232,1822,1848]" captionStartId-1="14.[189,232,1857,1883]" captionTargetBox-0="[253,1335,265,1785]" captionTargetBox-1="[244,1343,265,1817]" captionTargetId-0="figure-12@13.[253,1335,265,1785]" captionTargetId-1="figure-15@14.[244,1343,265,1817]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." captionText-1="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi-0="http://doi.org/10.5281/zenodo.7374065" figureDoi-1="http://doi.org/10.5281/zenodo.7374067" httpUri-0="https://zenodo.org/record/7374065/files/figure.png" httpUri-1="https://zenodo.org/record/7374067/files/figure.png" pageId="12" pageNumber="85">Figs 3–4</figureCitation>
). Peduncle up to 
<quantity id="0B192CF7A8265714804C810A4332FA32" box="[261,358,1398,1424]" metricMagnitude="-1" metricUnit="m" metricValue="1.08" pageId="12" pageNumber="85" unit="mm" value="108.0">108 mm</quantity>
in length (49.54% of overall length) and up to 
<quantity id="0B192CF7A826571482D4810A41A4FA32" box="[925,1008,1398,1424]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="85" unit="mm" value="11.0">11 mm</quantity>
in width at the widest point (the limit rachis-peduncle). Rachis with up to seven fleshy polyp leaves on each side, projecting somewhat obliquely and extending ventrally upward (
<figureCitation id="54DA9D97A826571483F281C0416CFA74" box="[699,824,1468,1494]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[253,1335,265,1785]" captionTargetId="figure-12@13.[253,1335,265,1785]" captionTargetPageId="13" captionText="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." figureDoi="http://doi.org/10.5281/zenodo.7374065" httpUri="https://zenodo.org/record/7374065/files/figure.png" pageId="12" pageNumber="85">Fig. 3B, F</figureCitation>
). Polyp leaves placed nearly oppositely, difficult to observe in preserved and contracted state (
<figureCitation id="54DA9D97A8265714839E81A3410EFA58" box="[727,858,1503,1530]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[253,1335,265,1785]" captionTargetId="figure-12@13.[253,1335,265,1785]" captionTargetPageId="13" captionText="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." figureDoi="http://doi.org/10.5281/zenodo.7374065" httpUri="https://zenodo.org/record/7374065/files/figure.png" pageId="12" pageNumber="85">Figs 3B, E</figureCitation>
, 
<figureCitation id="54DA9D97A8265714822E81A341DAFA5B" box="[871,910,1503,1529]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[244,1343,265,1817]" captionTargetId="figure-15@14.[244,1343,265,1817]" captionTargetPageId="14" captionText="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi="http://doi.org/10.5281/zenodo.7374067" httpUri="https://zenodo.org/record/7374067/files/figure.png" pageId="12" pageNumber="85">4D</figureCitation>
), increasing in size along the rachis until the mid-zone to last third, then quickly decreasing in size towards the distal part. Rachis with distinctive wide dorsal (
<figureCitation id="54DA9D97A82657148010825A438AF9E2" box="[345,478,1574,1600]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[253,1335,265,1785]" captionTargetId="figure-12@13.[253,1335,265,1785]" captionTargetPageId="13" captionText="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." figureDoi="http://doi.org/10.5281/zenodo.7374065" httpUri="https://zenodo.org/record/7374065/files/figure.png" pageId="12" pageNumber="85">Figs 3B, E</figureCitation>
, 
<figureCitation id="54DA9D97A826571480A4825A4044F9E2" box="[493,528,1574,1600]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[244,1343,265,1817]" captionTargetId="figure-15@14.[244,1343,265,1817]" captionTargetPageId="14" captionText="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi="http://doi.org/10.5281/zenodo.7374067" httpUri="https://zenodo.org/record/7374067/files/figure.png" pageId="12" pageNumber="85">4C</figureCitation>
) and reduced ventral track (
<figureCitation id="54DA9D97A8265714822C825A4196F9E2" box="[869,962,1574,1600]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[253,1335,265,1785]" captionTargetId="figure-12@13.[253,1335,265,1785]" captionTargetPageId="13" captionText="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." figureDoi="http://doi.org/10.5281/zenodo.7374065" httpUri="https://zenodo.org/record/7374065/files/figure.png" pageId="12" pageNumber="85">Figs 3F</figureCitation>
, 
<figureCitation id="54DA9D97A82657148299825A41A0F9E2" box="[976,1012,1574,1600]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[244,1343,265,1817]" captionTargetId="figure-15@14.[244,1343,265,1817]" captionTargetPageId="14" captionText="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi="http://doi.org/10.5281/zenodo.7374067" httpUri="https://zenodo.org/record/7374067/files/figure.png" pageId="12" pageNumber="85">4B</figureCitation>
) due to accumulation of ventral portion of polyp leaves bases. Polyp leaves nearly rectangular, not triangular, maximum length ~ 
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, maximum width ~ 
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. Axis present throughout colony, X-shaped in cross section, up to 
<quantity id="0B192CF7A8265714845982104702F925" box="[1296,1366,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="85" unit="mm" value="2.0">2 mm</quantity>
in maximum diameter at rachis-peduncle limit, becoming progressively asymmetric with age (
<figureCitation id="54DA9D97A8265714844282F34287F96F" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[346,1241,265,1749]" captionTargetId="figure-11@15.[346,1241,265,1749]" captionTargetPageId="15" captionText="Fig. 5. Gyrophyllum hirondellei Studer, 1891. Cross-sections of the axis near the rachis-peduncle limit of G. hirondellei colonies from SCOTIA cruises. A. Colony BECA (OPEN-660), a colony 112 mm in total length, showing an X-shaped cross-section, with an already apparent asymmetry. If growth rings are assumed to be produced annually this specimen is ~5 years old. B. Colony BECA (OPEN-661), a colony 215 mm in total length, showing a highly asymmetric X-shaped cross-section and a symmetrical central core (white dots). This specimen is ~14 years old according to growth rings with the proviso above." figureDoi="http://doi.org/10.5281/zenodo.7374069" httpUri="https://zenodo.org/record/7374069/files/figure.png" pageId="12" pageNumber="85">Fig. 5A– B</figureCitation>
). Autozooids numerous, up to approximately 
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the largest polyp leaves, arranged in one or two (three?) indistinct rows (
<figureCitation id="54DA9D97A8265714835582AA40CAF952" box="[540,670,1750,1776]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[253,1335,265,1785]" captionTargetId="figure-12@13.[253,1335,265,1785]" captionTargetPageId="13" captionText="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." figureDoi="http://doi.org/10.5281/zenodo.7374065" httpUri="https://zenodo.org/record/7374065/files/figure.png" pageId="12" pageNumber="85">Figs 3C, F</figureCitation>
, 
<figureCitation id="54DA9D97A826571483E582AA4084F952" box="[684,720,1750,1776]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[244,1343,265,1817]" captionTargetId="figure-15@14.[244,1343,265,1817]" captionTargetPageId="14" captionText="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi="http://doi.org/10.5281/zenodo.7374067" httpUri="https://zenodo.org/record/7374067/files/figure.png" pageId="12" pageNumber="85">4B</figureCitation>
, 
<figureCitation id="54DA9D97A8265714839682AA417CF952" box="[735,808,1750,1776]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[267,1320,265,1749]" captionTargetId="figure-15@16.[267,1320,265,1749]" captionTargetPageId="16" captionText="Fig. 6. Gyrophyllum hirondellei Studer, 1891. A–B. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). C–D. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. E. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). F. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves." figureDoi="http://doi.org/10.5281/zenodo.7374071" httpUri="https://zenodo.org/record/7374071/files/figure.png" pageId="12" pageNumber="85">6A–D</figureCitation>
) appearing at different levels near the atutozooid apertures along ventral edge of polyp leaf. Anthocodiae up to 
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in length (excluding tentacles) and 
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in width, completely retractile into spiculiferous, and not-always evident ‘calyces’ usually equipped with one prominent lateral blunt to pointed process (BPP hereinafter) up to 
<quantity id="0B192CF7A826571485E48343475CF8F8" box="[1197,1288,1855,1882]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="85" unit="mm" value="3.4">3.4 mm</quantity>
in length (not always present or well developed even in the same polyp leaf, see Discussion) (
<figureCitation id="54DA9D97A826571485F4831E476BF8DF" box="[1213,1343,1890,1917]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[253,1335,265,1785]" captionTargetId="figure-12@13.[253,1335,265,1785]" captionTargetPageId="13" captionText="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." figureDoi="http://doi.org/10.5281/zenodo.7374065" httpUri="https://zenodo.org/record/7374065/files/figure.png" pageId="12" pageNumber="85">Figs 3C, F</figureCitation>
, 
<figureCitation id="54DA9D97A82657148404831F4726F8DF" box="[1357,1394,1891,1917]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[244,1343,265,1817]" captionTargetId="figure-15@14.[244,1343,265,1817]" captionTargetPageId="14" captionText="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi="http://doi.org/10.5281/zenodo.7374067" httpUri="https://zenodo.org/record/7374067/files/figure.png" pageId="12" pageNumber="85">4B</figureCitation>
, 
<figureCitation id="54DA9D97A826571481F483FA42B1F802" box="[189,229,1926,1952]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[429,1159,265,1890]" captionTargetId="figure-11@17.[429,1159,265,1890]" captionTargetPageId="17" captionText="Fig. 7. Gyrophyllum hirondellei Studer, 1891. Details of the ventral edge of a polyp leaf, showing the apertures of an autozooid (black arrows) and a single BPP (white arrows) in differing degrees of development per autozooid. A–B. Colony BECA (OPEN-661). C. Colony MNHM OCT.A.579." figureDoi="http://doi.org/10.5281/zenodo.7374073" httpUri="https://zenodo.org/record/7374073/files/figure.png" pageId="12" pageNumber="85">7A</figureCitation>
). Along ventral edge of polyp leaves, autozooids and spiculiferous BPPs alternate (
<figureCitation id="54DA9D97A826571485E583FA46A7F802" box="[1196,1267,1926,1952]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[429,1159,265,1890]" captionTargetId="figure-11@17.[429,1159,265,1890]" captionTargetPageId="17" captionText="Fig. 7. Gyrophyllum hirondellei Studer, 1891. Details of the ventral edge of a polyp leaf, showing the apertures of an autozooid (black arrows) and a single BPP (white arrows) in differing degrees of development per autozooid. A–B. Colony BECA (OPEN-661). C. Colony MNHM OCT.A.579." figureDoi="http://doi.org/10.5281/zenodo.7374073" httpUri="https://zenodo.org/record/7374073/files/figure.png" pageId="12" pageNumber="85">Fig. 7</figureCitation>
). Tentacles of autozooid up to 
<quantity id="0B192CF7A826571480D483D543ACF861" box="[413,504,1961,1988]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="85" unit="mm" value="3.5">3.5 mm</quantity>
in length in preserved state, with two kinds of processes, standard pinnules arranged in two lateral series (
<figureCitation id="54DA9D97A8265714836783B040DFF844" box="[558,651,1996,2022]" captionStart="Fig" captionStartId="18.[189,232,1576,1602]" captionTargetBox="[330,1258,265,1538]" captionTargetId="figure-14@18.[330,1258,265,1538]" captionTargetPageId="18" captionText="Fig. 8. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. B. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’into which each tentacle is partially retracted.C. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. D. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. E. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. F. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. G. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. H. Detail from G, also showing filiform structures." figureDoi="http://doi.org/10.5281/zenodo.7374075" httpUri="https://zenodo.org/record/7374075/files/figure.png" pageId="12" pageNumber="85">Figs 8F</figureCitation>
, 
<figureCitation id="54DA9D97A826571483D383B040EBF844" box="[666,703,1996,2022]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[189,1398,1134,1785]" captionTargetId="figure-277@19.[189,1398,1134,1785]" captionTargetPageId="19" captionText="Fig. 9. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. SEM photographs of a tentacle in latero-oral view showing the numerous filiform structures. B. SEM photographs of a tentacle in latero-aboral view showing the naked aboral surface of tentacular axis and filiform structures. C. SEM photographs of a tentacle in lateral view showing digitiform normal pinnulae and, on the left, the numerous and elongate filiform structures. missing collection codes for specimens (as given in all other figs …)" figureDoi="http://doi.org/10.5281/zenodo.7374079" httpUri="https://zenodo.org/record/7374079/files/figure.png" pageId="12" pageNumber="85">9C</figureCitation>
), and numerous filiform structures only present along the oral axial surface (
<figureCitation id="54DA9D97A839570B80D585754011FE81" box="[412,581,265,291]" captionStart="Fig" captionStartId="18.[189,232,1576,1602]" captionTargetBox="[330,1258,265,1538]" captionTargetId="figure-14@18.[330,1258,265,1538]" captionTargetPageId="18" captionText="Fig. 8. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. B. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’into which each tentacle is partially retracted.C. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. D. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. E. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. F. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. G. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. H. Detail from G, also showing filiform structures." figureDoi="http://doi.org/10.5281/zenodo.7374075" httpUri="https://zenodo.org/record/7374075/files/figure.png" pageId="19" pageNumber="92">Figs 8C, E–H</figureCitation>
, 
<figureCitation id="54DA9D97A839570B831B857540C9FE81" box="[594,669,265,291]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[189,1398,1134,1785]" captionTargetId="figure-277@19.[189,1398,1134,1785]" captionTargetPageId="19" captionText="Fig. 9. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. SEM photographs of a tentacle in latero-oral view showing the numerous filiform structures. B. SEM photographs of a tentacle in latero-aboral view showing the naked aboral surface of tentacular axis and filiform structures. C. SEM photographs of a tentacle in lateral view showing digitiform normal pinnulae and, on the left, the numerous and elongate filiform structures. missing collection codes for specimens (as given in all other figs …)" figureDoi="http://doi.org/10.5281/zenodo.7374079" httpUri="https://zenodo.org/record/7374079/files/figure.png" pageId="19" pageNumber="92">9A–C</figureCitation>
). Siphonozooids minute, 
<quantity id="0B192CF7A839570B82858575462DFE81" box="[972,1145,265,292]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="5.5" metricValueMin="3.2" pageId="19" pageNumber="92" unit="mm" value="0.435" valueMax="0.55" valueMin="0.32">0.32–0.55 mm</quantity>
in diameter (average 
<quantity id="0B192CF7A839570B81F68550437DFEE5" box="[191,297,300,327]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="92" unit="mm" value="0.46">0.46 mm</quantity>
, N = 20), numerous, scattered on the lateral (actually proximal and distal) sides of polyp leaves (
<figureCitation id="54DA9D97A839570B818C85334374FECB" box="[197,288,335,361]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[267,1320,265,1749]" captionTargetId="figure-15@16.[267,1320,265,1749]" captionTargetPageId="16" captionText="Fig. 6. Gyrophyllum hirondellei Studer, 1891. A–B. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). C–D. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. E. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). F. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves." figureDoi="http://doi.org/10.5281/zenodo.7374071" httpUri="https://zenodo.org/record/7374071/files/figure.png" pageId="19" pageNumber="92">Fig. 6E</figureCitation>
), and rachis dorsal track (more difficult to detect).
</paragraph>
<caption id="989ED19AA827571581F48362434AF84A" ID-DOI="http://doi.org/10.5281/zenodo.7374065" ID-Zenodo-Dep="7374065" httpUri="https://zenodo.org/record/7374065/files/figure.png" pageId="13" pageNumber="86" startId="13.[189,232,1822,1848]" targetBox="[253,1335,265,1785]" targetPageId="13">
<paragraph id="CC5E8112A827571581F48362434AF84A" blockId="13.[189,1400,1822,2024]" pageId="13" pageNumber="86">
<emphasis id="FE955D00A827571581F48362435AF89A" bold="true" box="[189,270,1822,1848]" pageId="13" pageNumber="86">Fig. 3.</emphasis>
<taxonomicName id="0BE1FA91A82757158051836240B7F89A" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[280,739,1822,1848]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="13" pageNumber="86" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A827571580518362406BF89A" box="[280,575,1822,1848]" italics="true" pageId="13" pageNumber="86">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A82757158300836240B7F89A" author="Studer T." box="[585,739,1822,1848]" pageId="13" pageNumber="86" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
, colonies from SCOTIA cruises. 
<emphasis id="FE955D00A8275715853F836246FAF89A" bold="true" box="[1142,1198,1822,1848]" pageId="13" pageNumber="86">A–B</emphasis>
. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. 
<emphasis id="FE955D00A827571583A2833D40ABF8F9" bold="true" box="[747,767,1857,1883]" pageId="13" pageNumber="86">C</emphasis>
. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). 
<emphasis id="FE955D00A8275715806C83F44308F800" bold="true" box="[293,348,1928,1954]" pageId="13" pageNumber="86">D–E</emphasis>
. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. 
<emphasis id="FE955D00A8275715854983F44646F800" bold="true" box="[1024,1042,1928,1954]" pageId="13" pageNumber="86">F</emphasis>
. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows).
</paragraph>
</caption>
<caption id="989ED19AA824571681F4833D4030F84A" ID-DOI="http://doi.org/10.5281/zenodo.7374067" ID-Zenodo-Dep="7374067" httpUri="https://zenodo.org/record/7374067/files/figure.png" pageId="14" pageNumber="87" startId="14.[189,232,1857,1883]" targetBox="[244,1343,265,1817]" targetPageId="14">
<paragraph id="CC5E8112A824571681F4833D4030F84A" blockId="14.[189,1399,1857,2025]" pageId="14" pageNumber="87">
<emphasis id="FE955D00A824571681F4833D4345F8F9" bold="true" box="[189,273,1857,1883]" pageId="14" pageNumber="87">Fig. 4.</emphasis>
<taxonomicName id="0BE1FA91A82457168057833D40A7F8FE" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[286,755,1857,1884]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="14" pageNumber="87" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A82457168057833D401CF8F9" box="[286,584,1857,1883]" italics="true" pageId="14" pageNumber="87">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A8245716831C833D40A7F8FE" author="Studer T." box="[597,755,1857,1884]" pageId="14" pageNumber="87" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
. 
<emphasis id="FE955D00A8245716824D833D414DF8F9" bold="true" box="[772,793,1857,1883]" pageId="14" pageNumber="87">A</emphasis>
. Holotype colony of 
<taxonomicName id="0BE1FA91A82457168563833E4681F8F9" authorityName="Studer" authorityYear="1891" box="[1066,1237,1857,1884]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="14" pageNumber="87" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A82457168563833E4681F8F9" box="[1066,1237,1857,1884]" italics="true" pageId="14" pageNumber="87">G. hirondellei</emphasis>
</taxonomicName>
(MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. 
<emphasis id="FE955D00A8245716857683194607F8DD" bold="true" box="[1087,1107,1893,1919]" pageId="14" pageNumber="87">B</emphasis>
. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. 
<emphasis id="FE955D00A824571681F483D742A2F867" bold="true" box="[189,246,1963,1989]" pageId="14" pageNumber="87">C–D</emphasis>
. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino.
</paragraph>
</caption>
<caption id="989ED19AA825571781F48287435FF84A" ID-DOI="http://doi.org/10.5281/zenodo.7374069" ID-Zenodo-Dep="7374069" httpUri="https://zenodo.org/record/7374069/files/figure.png" pageId="15" pageNumber="88" startId="15.[189,232,1787,1813]" targetBox="[346,1241,265,1749]" targetPageId="15">
<paragraph id="CC5E8112A825571781F48287435FF84A" blockId="15.[188,1400,1787,2024]" pageId="15" pageNumber="88">
<emphasis id="FE955D00A825571781F48287435FF8B7" bold="true" box="[189,267,1787,1813]" pageId="15" pageNumber="88">Fig. 5.</emphasis>
<taxonomicName id="0BE1FA91A8255717805D8287408CF8B7" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[276,728,1787,1813]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="15" pageNumber="88" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A8255717805D82874063F8B7" box="[276,567,1787,1813]" italics="true" pageId="15" pageNumber="88">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A825571783768287408CF8B7" author="Studer T." box="[575,728,1787,1813]" pageId="15" pageNumber="88" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
. Cross-sections of the axis near the rachis-peduncle limit of 
<taxonomicName id="0BE1FA91A82557178197836343D0F89A" authorityName="Studer" authorityYear="1891" box="[222,388,1822,1849]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="15" pageNumber="88" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A82557178197836343D0F89A" box="[222,388,1822,1849]" italics="true" pageId="15" pageNumber="88">G. hirondellei</emphasis>
</taxonomicName>
colonies from SCOTIA cruises. 
<emphasis id="FE955D00A825571782428362414BF89A" bold="true" box="[779,799,1822,1848]" pageId="15" pageNumber="88">A</emphasis>
. Colony BECA (OPEN-660), a colony 112 mm in total length, showing an X-shaped cross-section, with an already apparent asymmetry. If growth rings are assumed to be produced annually this specimen is ~5 years old. 
<emphasis id="FE955D00A825571782A2831941ABF8DD" bold="true" box="[1003,1023,1893,1919]" pageId="15" pageNumber="88">B</emphasis>
. Colony BECA (OPEN-661), a colony 215 mm in total length, showing a highly asymmetric X-shaped cross-section and a symmetrical central core (white dots). This specimen is ~14 years old according to growth rings with the proviso above.
</paragraph>
</caption>
<caption id="989ED19AA83A570881F4828743F3F84B" ID-DOI="http://doi.org/10.5281/zenodo.7374071" ID-Zenodo-Dep="7374071" httpUri="https://zenodo.org/record/7374071/files/figure.png" pageId="16" pageNumber="89" startId="16.[189,232,1787,1813]" targetBox="[267,1320,265,1749]" targetPageId="16">
<paragraph id="CC5E8112A83A570881F4828743F3F84B" blockId="16.[188,1399,1787,2025]" pageId="16" pageNumber="89">
<emphasis id="FE955D00A83A570881F48287435FF8B7" bold="true" box="[189,267,1787,1813]" pageId="16" pageNumber="89">Fig. 6.</emphasis>
<taxonomicName id="0BE1FA91A83A5708805B82874081F8B7" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[274,725,1787,1813]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A83A5708805B82874062F8B7" box="[274,566,1787,1813]" italics="true" pageId="16" pageNumber="89">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A83A5708837782874081F8B7" author="Studer T." box="[574,725,1787,1813]" pageId="16" pageNumber="89" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
. 
<emphasis id="FE955D00A83A570883A98287414CF8B7" bold="true" box="[736,792,1787,1813]" pageId="16" pageNumber="89">A–B</emphasis>
. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). 
<emphasis id="FE955D00A83A57088037833D43E3F8F9" bold="true" box="[382,439,1857,1883]" pageId="16" pageNumber="89">C–D</emphasis>
. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. 
<emphasis id="FE955D00A83A570885AD831946ACF8DD" bold="true" box="[1252,1272,1893,1919]" pageId="16" pageNumber="89">E</emphasis>
. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). 
<emphasis id="FE955D00A83A5708807783D74304F867" bold="true" box="[318,336,1963,1989]" pageId="16" pageNumber="89">F</emphasis>
. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves.
</paragraph>
</caption>
<caption id="989ED19AA83B570981F483F4474BF84B" ID-DOI="http://doi.org/10.5281/zenodo.7374073" ID-Zenodo-Dep="7374073" httpUri="https://zenodo.org/record/7374073/files/figure.png" pageId="17" pageNumber="90" startId="17.[189,232,1928,1954]" targetBox="[429,1159,265,1890]" targetPageId="17">
<paragraph id="CC5E8112A83B570981F483F4474BF84B" blockId="17.[189,1398,1928,2025]" pageId="17" pageNumber="90">
<emphasis id="FE955D00A83B570981F483F4435BF800" bold="true" box="[189,271,1928,1954]" pageId="17" pageNumber="90">Fig. 7.</emphasis>
<taxonomicName id="0BE1FA91A83B5709805283F440BEF800" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[283,746,1928,1954]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="17" pageNumber="90" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A83B5709805283F44017F800" box="[283,579,1928,1954]" italics="true" pageId="17" pageNumber="90">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A83B5709830783F440BEF800" author="Studer T." box="[590,746,1928,1954]" pageId="17" pageNumber="90" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
. Details of the ventral edge of a polyp leaf, showing the apertures of an autozooid (black arrows) and a single BPP (white arrows) in differing degrees of development per autozooid. 
<emphasis id="FE955D00A83B5709834283B24017F84A" bold="true" box="[523,579,1998,2024]" pageId="17" pageNumber="90">A–B</emphasis>
. Colony BECA (OPEN-661). 
<emphasis id="FE955D00A83B570982E183B241E9F84A" bold="true" box="[936,957,1998,2024]" pageId="17" pageNumber="90">C</emphasis>
. Colony MNHM OCT.A.579.
</paragraph>
</caption>
<caption id="989ED19AA838570A81F482544104F84B" ID-DOI="http://doi.org/10.5281/zenodo.7374075" ID-Zenodo-Dep="7374075" httpUri="https://zenodo.org/record/7374075/files/figure.png" pageId="18" pageNumber="91" startId="18.[189,232,1576,1602]" targetBox="[330,1258,265,1538]" targetPageId="18">
<paragraph id="CC5E8112A838570A81F482544104F84B" blockId="18.[187,1400,1576,2025]" pageId="18" pageNumber="91">
<emphasis id="FE955D00A838570A81F482544359F9E0" bold="true" box="[189,269,1576,1602]" pageId="18" pageNumber="91">Fig. 8.</emphasis>
<taxonomicName id="0BE1FA91A838570A8050825440B5F9E0" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[281,737,1576,1602]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="18" pageNumber="91" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A838570A80508254406AF9E0" box="[281,574,1576,1602]" italics="true" pageId="18" pageNumber="91">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A838570A8301825440B5F9E0" author="Studer T." box="[584,737,1576,1602]" pageId="18" pageNumber="91" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
, BECA (OPEN-660). 
<emphasis id="FE955D00A838570A82A2825441AAF9E0" bold="true" box="[1003,1022,1576,1602]" pageId="18" pageNumber="91">A</emphasis>
. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. 
<emphasis id="FE955D00A838570A80848212438BF92A" bold="true" box="[461,479,1646,1672]" pageId="18" pageNumber="91">B</emphasis>
. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’ into which each tentacle is partially retracted. 
<emphasis id="FE955D00A838570A82A482C84656F96C" bold="true" box="[1005,1026,1716,1742]" pageId="18" pageNumber="91">C</emphasis>
. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. 
<emphasis id="FE955D00A838570A834E83624048F89A" bold="true" box="[519,540,1822,1848]" pageId="18" pageNumber="91">D</emphasis>
. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. 
<emphasis id="FE955D00A838570A83C2833E40CBF8FE" bold="true" box="[651,671,1858,1884]" pageId="18" pageNumber="91">E</emphasis>
. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. 
<emphasis id="FE955D00A838570A83EF831940ECF8DD" bold="true" box="[678,696,1893,1919]" pageId="18" pageNumber="91">F</emphasis>
. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. 
<emphasis id="FE955D00A838570A844F83F44749F800" bold="true" box="[1286,1309,1928,1954]" pageId="18" pageNumber="91">G</emphasis>
. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. 
<emphasis id="FE955D00A838570A81B283B24346F84A" bold="true" box="[251,274,1998,2024]" pageId="18" pageNumber="91">H</emphasis>
. Detail from G, also showing filiform structures.
</paragraph>
</caption>
<paragraph id="CC5E8112A839570B81F485ED40C2FD03" blockId="19.[187,1399,401,674]" pageId="19" pageNumber="92">
Sclerites differentially distributed in various parts of colony: densely placed in rachis, including polyp leaves, calycular BPP (
<figureCitation id="54DA9D97A839570B808285C8407DFE6C" box="[459,553,436,462]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[267,1320,265,1749]" captionTargetId="figure-15@16.[267,1320,265,1749]" captionTargetPageId="16" captionText="Fig. 6. Gyrophyllum hirondellei Studer, 1891. A–B. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). C–D. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. E. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). F. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves." figureDoi="http://doi.org/10.5281/zenodo.7374071" httpUri="https://zenodo.org/record/7374071/files/figure.png" pageId="19" pageNumber="92">Fig. 6D</figureCitation>
), and along dorsal and ventral tracks, around openings of siphonozooids (
<figureCitation id="54DA9D97A839570B818C85AB434BFE53" box="[197,287,471,497]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[267,1320,265,1749]" captionTargetId="figure-15@16.[267,1320,265,1749]" captionTargetPageId="16" captionText="Fig. 6. Gyrophyllum hirondellei Studer, 1891. A–B. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). C–D. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. E. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). F. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves." figureDoi="http://doi.org/10.5281/zenodo.7374071" httpUri="https://zenodo.org/record/7374071/files/figure.png" pageId="19" pageNumber="92">Fig. 6E</figureCitation>
), along abaxial side of tentacular axis of autozooids (
<figureCitation id="54DA9D97A839570B82C985AB4655FE53" box="[896,1025,471,497]" captionStart="Fig" captionStartId="18.[189,232,1576,1602]" captionTargetBox="[330,1258,265,1538]" captionTargetId="figure-14@18.[330,1258,265,1538]" captionTargetPageId="18" captionText="Fig. 8. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. B. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’into which each tentacle is partially retracted.C. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. D. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. E. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. F. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. G. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. H. Detail from G, also showing filiform structures." figureDoi="http://doi.org/10.5281/zenodo.7374075" httpUri="https://zenodo.org/record/7374075/files/figure.png" pageId="19" pageNumber="92">Fig. 8G–H</figureCitation>
); however, much more scattered on body of autozooids (
<figureCitation id="54DA9D97A839570B809B85864000FDB7" box="[466,596,506,533]" captionStart="Fig" captionStartId="18.[189,232,1576,1602]" captionTargetBox="[330,1258,265,1538]" captionTargetId="figure-14@18.[330,1258,265,1538]" captionTargetPageId="18" captionText="Fig. 8. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. B. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’into which each tentacle is partially retracted.C. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. D. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. E. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. F. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. G. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. H. Detail from G, also showing filiform structures." figureDoi="http://doi.org/10.5281/zenodo.7374075" httpUri="https://zenodo.org/record/7374075/files/figure.png" pageId="19" pageNumber="92">Fig. 8C–D</figureCitation>
). Sclerites present in a reticular manner on polyp leaves, observable not only on surface (
<figureCitation id="54DA9D97A839570B80F386624041FD95" box="[442,533,541,568]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[267,1320,265,1749]" captionTargetId="figure-15@16.[267,1320,265,1749]" captionTargetPageId="16" captionText="Fig. 6. Gyrophyllum hirondellei Studer, 1891. A–B. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). C–D. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. E. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). F. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves." figureDoi="http://doi.org/10.5281/zenodo.7374071" httpUri="https://zenodo.org/record/7374071/files/figure.png" pageId="19" pageNumber="92">Fig. 6E</figureCitation>
) but also internally. Walls between consecutive autozooids thinner than outer surrounding wall (
<figureCitation id="54DA9D97A839570B809D863D4000FDF9" box="[468,596,577,603]" captionStart="Fig" captionStartId="18.[189,232,1576,1602]" captionTargetBox="[330,1258,265,1538]" captionTargetId="figure-14@18.[330,1258,265,1538]" captionTargetPageId="18" captionText="Fig. 8. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. B. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’into which each tentacle is partially retracted.C. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. D. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. E. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. F. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. G. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. H. Detail from G, also showing filiform structures." figureDoi="http://doi.org/10.5281/zenodo.7374075" httpUri="https://zenodo.org/record/7374075/files/figure.png" pageId="19" pageNumber="92">Fig. 8A–B</figureCitation>
). Similar reticular structures visible at rachis-peduncle limit (
<figureCitation id="54DA9D97A839570B8451863D473AFDF9" box="[1304,1390,577,603]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[267,1320,265,1749]" captionTargetId="figure-15@16.[267,1320,265,1749]" captionTargetPageId="16" captionText="Fig. 6. Gyrophyllum hirondellei Studer, 1891. A–B. Detail of the colony MNHM OCT.A.579 in upperventral and lateral view, respectively, showing BPPs (white arrows) and partially retracted autozooids (black arrows). C–D. Detail of polyp leaves and ventral edge of a polyp leaf of the colony BECA (OPEN-660), note trabecular appearance of lateral surfaces and well developed BPPs. E. Detail of lateral surface of a polyp leaf, showing trabecular arrangement of sclerites and siphonozooid openings (arrowed). F. Partial section at rachis-peduncle limit, showing the thick trabecular wall and the axis with longitudinal groves." figureDoi="http://doi.org/10.5281/zenodo.7374071" httpUri="https://zenodo.org/record/7374071/files/figure.png" pageId="19" pageNumber="92">Fig. 6F</figureCitation>
) and on penduncle. Sclerites absent in pinnules (including filiform processes), polyp body, and pharynx. No minute bodies observed in peduncle.
</paragraph>
<paragraph id="CC5E8112A839570B81F486B54194FC14" blockId="19.[187,1400,712,951]" pageId="19" pageNumber="92">
Sclerites from polyp body and tentacular axis as blunt tree-flanged rods up to 
<quantity id="0B192CF7A839570B852C86B44684FD41" box="[1125,1232,712,739]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="92" unit="mm" value="0.25">0.25 mm</quantity>
and 
<quantity id="0B192CF7A839570B844586B54723FD41" box="[1292,1399,713,739]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="19" pageNumber="92" unit="mm" value="0.31">0.31 mm</quantity>
in length, respectively (
<figureCitation id="54DA9D97A839570B809586904030FCA4" box="[476,612,748,774]" captionStart="Fig" captionStartId="18.[189,232,1576,1602]" captionTargetBox="[330,1258,265,1538]" captionTargetId="figure-14@18.[330,1258,265,1538]" captionTargetPageId="18" captionText="Fig. 8. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). A. Transversal section of a polyp leaf, showing its trabecular walls, and the thin barriers between consecutive gastrovascular cavities that are disposed in a line. B. Detail from A (the gastrovascular cavity on the left), showing also pharynx, mesenteria, sclerites of the tentacular axis (visible due to the transparency of oral disc), and (sectioned) one of the oral ‘pouches’into which each tentacle is partially retracted.C. Internal view of the autozooids body wall into which the tentacular crown retracts (this becomes the outer body wall when the autozooid is extended). Note on the bottom right one of the tentacles and several sclerites (arrowed) in the thin body wall of the autozooid. D. Autozooid body wall treated with clove oil to clear tissue with scattered sclerites now easily observed in situ. E. Single tentacle in oral view (after critical point treatment) showing the numerous filiform structures. F. Single tentacle in lateral view (after critical point treatment) showing a series of normal pinnulae, part of the aboral side of tentacle and filiform structures. G. Single tentacle treated with clove oil to observe the presence and disposition of sclerites along the tentacular axis. H. Detail from G, also showing filiform structures." figureDoi="http://doi.org/10.5281/zenodo.7374075" httpUri="https://zenodo.org/record/7374075/files/figure.png" pageId="19" pageNumber="92">Figs 8C–D</figureCitation>
, 
<figureCitation id="54DA9D97A839570B833C869040FFFCA4" box="[629,683,748,774]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[238,1349,265,1925]" captionTargetId="figure-14@20.[238,1349,265,1925]" captionTargetPageId="20" captionText="Fig. 10. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Tentacle. B. Polyp leaf, siphonozooids area. C. Calycular pointed processes. D. Rachis exterior." figureDoi="http://doi.org/10.5281/zenodo.7374082" httpUri="https://zenodo.org/record/7374082/files/figure.png" pageId="19" pageNumber="92">10A</figureCitation>
). Sclerites from polyp leaves (including those surrounding siphonozooids), calycular BPPs, rachis and peduncle as elongated three-flanged rods. Those from polyp leaves up to 
<quantity id="0B192CF7A839570B801F874E4395FCEF" box="[342,449,818,845]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="92" unit="mm" value="0.52">0.52 mm</quantity>
in length (
<figureCitation id="54DA9D97A839570B830F874E40E7FCEE" box="[582,691,818,844]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[238,1349,265,1925]" captionTargetId="figure-14@20.[238,1349,265,1925]" captionTargetPageId="20" captionText="Fig. 10. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Tentacle. B. Polyp leaf, siphonozooids area. C. Calycular pointed processes. D. Rachis exterior." figureDoi="http://doi.org/10.5281/zenodo.7374082" httpUri="https://zenodo.org/record/7374082/files/figure.png" pageId="19" pageNumber="92">Fig. 10B</figureCitation>
). Sclerites from calycular BPPs up to 
<quantity id="0B192CF7A839570B8536874E46BEFCEF" box="[1151,1258,818,845]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="19" pageNumber="92" unit="mm" value="0.48">0.48 mm</quantity>
(
<figureCitation id="54DA9D97A839570B85B5874E473EFCEE" box="[1276,1386,818,844]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[238,1349,265,1925]" captionTargetId="figure-14@20.[238,1349,265,1925]" captionTargetPageId="20" captionText="Fig. 10. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Tentacle. B. Polyp leaf, siphonozooids area. C. Calycular pointed processes. D. Rachis exterior." figureDoi="http://doi.org/10.5281/zenodo.7374082" httpUri="https://zenodo.org/record/7374082/files/figure.png" pageId="19" pageNumber="92">Fig. 10C</figureCitation>
). Sclerites from exterior surface of rachis up to 
<quantity id="0B192CF7A839570B8380872A4164FCD2" box="[713,816,854,880]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="19" pageNumber="92" unit="mm" value="0.44">0.44 mm</quantity>
in length (
<figureCitation id="54DA9D97A839570B82E187294640FCD2" box="[936,1044,853,880]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[238,1349,265,1925]" captionTargetId="figure-14@20.[238,1349,265,1925]" captionTargetPageId="20" captionText="Fig. 10. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Tentacle. B. Polyp leaf, siphonozooids area. C. Calycular pointed processes. D. Rachis exterior." figureDoi="http://doi.org/10.5281/zenodo.7374082" httpUri="https://zenodo.org/record/7374082/files/figure.png" pageId="19" pageNumber="92">Fig. 10D</figureCitation>
). Sclerites from inner rachis up to 
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in length (
<figureCitation id="54DA9D97A839570B80F78705407EFC31" box="[446,554,889,915]" captionStart="Fig" captionStartId="21.[189,232,1961,1987]" captionTargetBox="[189,1398,573,1923]" captionTargetId="figure-70@21.[189,1398,573,1923]" captionTargetPageId="21" captionText="Fig. 11. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Rachis interior. B. Peduncle exterior. C. Peduncle interior." figureDoi="http://doi.org/10.5281/zenodo.7374084" httpUri="https://zenodo.org/record/7374084/files/figure.png" pageId="19" pageNumber="92">Fig. 11A</figureCitation>
). Sclerites from surface of peduncle up to 
<quantity id="0B192CF7A839570B8551870446D4FC31" box="[1048,1152,888,915]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="19" pageNumber="92" unit="mm" value="0.36">0.36 mm</quantity>
in length (
<figureCitation id="54DA9D97A839570B85B487054733FC31" box="[1277,1383,889,915]" captionStart="Fig" captionStartId="21.[189,232,1961,1987]" captionTargetBox="[189,1398,573,1923]" captionTargetId="figure-70@21.[189,1398,573,1923]" captionTargetPageId="21" captionText="Fig. 11. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Rachis interior. B. Peduncle exterior. C. Peduncle interior." figureDoi="http://doi.org/10.5281/zenodo.7374084" httpUri="https://zenodo.org/record/7374084/files/figure.png" pageId="19" pageNumber="92">Fig. 11B</figureCitation>
). Sclerites from inner peduncle up to 
<quantity id="0B192CF7A839570B832887E0409EFC14" box="[609,714,924,951]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="92" unit="mm" value="0.35">0.35 mm</quantity>
in length (
<figureCitation id="54DA9D97A839570B820187E041E6FC14" box="[840,946,924,950]" captionStart="Fig" captionStartId="21.[189,232,1961,1987]" captionTargetBox="[189,1398,573,1923]" captionTargetId="figure-70@21.[189,1398,573,1923]" captionTargetPageId="21" captionText="Fig. 11. Gyrophyllum hirondellei Studer, 1891, BECA (OPEN-660). SEM photographs of sclerites. A. Rachis interior. B. Peduncle exterior. C. Peduncle interior." figureDoi="http://doi.org/10.5281/zenodo.7374084" httpUri="https://zenodo.org/record/7374084/files/figure.png" pageId="19" pageNumber="92">Fig. 11C</figureCitation>
).
</paragraph>
<caption id="989ED19AA839570B81F483624342F84A" ID-DOI="http://doi.org/10.5281/zenodo.7374079" ID-Zenodo-Dep="7374079" httpUri="https://zenodo.org/record/7374079/files/figure.png" pageId="19" pageNumber="92" startId="19.[189,232,1822,1848]" targetBox="[189,1398,1134,1785]" targetPageId="19">
<paragraph id="CC5E8112A839570B81F483624342F84A" blockId="19.[189,1399,1822,2024]" pageId="19" pageNumber="92">
<emphasis id="FE955D00A839570B81F483624358F89A" bold="true" box="[189,268,1822,1848]" pageId="19" pageNumber="92">Fig. 9.</emphasis>
<taxonomicName id="0BE1FA91A839570B805F83624088F89A" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[278,732,1822,1848]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="19" pageNumber="92" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A839570B805F8362406FF89A" box="[278,571,1822,1848]" italics="true" pageId="19" pageNumber="92">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A839570B830A83624088F89A" author="Studer T." box="[579,732,1822,1848]" pageId="19" pageNumber="92" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
, BECA (OPEN-660). 
<emphasis id="FE955D00A839570B82AA836241A3F89A" bold="true" box="[995,1015,1822,1848]" pageId="19" pageNumber="92">A</emphasis>
. SEM photographs of a tentacle in latero-oral view showing the numerous filiform structures. 
<emphasis id="FE955D00A839570B82F8833E4191F8FE" bold="true" box="[945,965,1858,1884]" pageId="19" pageNumber="92">B</emphasis>
. SEM photographs of a tentacle in latero-aboral view showing the naked aboral surface of tentacular axis and filiform structures. 
<emphasis id="FE955D00A839570B845F8318477FF8DC" bold="true" box="[1302,1323,1892,1918]" pageId="19" pageNumber="92">C</emphasis>
. SEM photographs of a tentacle in lateral view showing digitiform normal pinnulae and, on the left, the numerous and elongate filiform structures. missing collection codes for specimens (as given in all other figs …)
</paragraph>
</caption>
<caption id="989ED19AA83E570C81F483D74710F84A" ID-DOI="http://doi.org/10.5281/zenodo.7374082" ID-Zenodo-Dep="7374082" httpUri="https://zenodo.org/record/7374082/files/figure.png" pageId="20" pageNumber="93" startId="20.[189,232,1963,1989]" targetBox="[238,1349,265,1925]" targetPageId="20">
<paragraph id="CC5E8112A83E570C81F483D74710F84A" blockId="20.[189,1399,1963,2025]" pageId="20" pageNumber="93">
<emphasis id="FE955D00A83E570C81F483D7434AF867" bold="true" box="[189,286,1963,1989]" pageId="20" pageNumber="93">Fig. 10.</emphasis>
<taxonomicName id="0BE1FA91A83E570C806083D740A3F867" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[297,759,1963,1989]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="20" pageNumber="93" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A83E570C806083D74005F867" box="[297,593,1963,1989]" italics="true" pageId="20" pageNumber="93">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A83E570C831483D740A3F867" author="Studer T." box="[605,759,1963,1989]" pageId="20" pageNumber="93" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
, BECA (OPEN-660). SEM photographs of sclerites. 
<emphasis id="FE955D00A83E570C81F483B24286F84A" bold="true" box="[189,210,1998,2024]" pageId="20" pageNumber="93">A</emphasis>
. Tentacle. 
<emphasis id="FE955D00A83E570C801883B24331F84A" bold="true" box="[337,357,1998,2024]" pageId="20" pageNumber="93">B</emphasis>
. Polyp leaf, siphonozooids area. 
<emphasis id="FE955D00A83E570C83A283B24154F84A" bold="true" box="[747,768,1998,2024]" pageId="20" pageNumber="93">C</emphasis>
. Calycular pointed processes. 
<emphasis id="FE955D00A83E570C852E83B24628F84A" bold="true" box="[1127,1148,1998,2024]" pageId="20" pageNumber="93">D</emphasis>
. Rachis exterior.
</paragraph>
</caption>
<paragraph id="CC5E8112A83F570D81F485744342FE80" blockId="21.[189,278,264,290]" box="[189,278,264,290]" pageId="21" pageNumber="94">
<heading id="9716367EA83F570D81F485744342FE80" bold="true" box="[189,278,264,290]" fontSize="11" level="3" pageId="21" pageNumber="94" reason="3">
<emphasis id="FE955D00A83F570D81F485744342FE80" bold="true" box="[189,278,264,290]" pageId="21" pageNumber="94">Colour</emphasis>
</heading>
</paragraph>
<paragraph id="CC5E8112A83F570D81F4854B41A6FE19" blockId="21.[188,1400,311,443]" pageId="21" pageNumber="94">
Freshly collected colonies were light brown at peduncle, rachis stalk and the section of rachis bearing polyp leaves dorsally, but darker brown on surfaces of polyps and calycular BPPs with the sclerites visible as whitish trabecular structures. The autozooids themselves were dark brown. Preserved colonies are whitish to light brown (
<figureCitation id="54DA9D97A83F570D80B385DD4037FE19" box="[506,611,417,443]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="13.[189,232,1822,1848]" captionStartId-1="14.[189,232,1857,1883]" captionTargetBox-0="[253,1335,265,1785]" captionTargetBox-1="[244,1343,265,1817]" captionTargetId-0="figure-12@13.[253,1335,265,1785]" captionTargetId-1="figure-15@14.[244,1343,265,1817]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Fig. 3. Gyrophyllum hirondellei Studer, 1891, colonies from SCOTIA cruises. A–B. BECA (OPEN- 660), ventral and dorsal sides, see also Fig. 4. C. Detail from colony NMS.Z.2022.1.6, showing parts of polyp leaves ventral edge with well-developed BPPs (white arrows) and others without BPPs (black arrows). D–E. Ventral and dorsal sides of the colony NMS.Z.2022.1.4. F. Detail from D, showing most of the proximal polyp leaves without BPPs, some can be seen on the distalmost polyp leaves (white arrows)." captionText-1="Fig. 4. Gyrophyllum hirondellei Studer, 1891. A. Holotype colony of G. hirondellei (MOM INV 120036) deposited in the Musée oceanographique de Monaco, ventral view. B. Detail from A, showing the compacted group of polyp leaves and autozooids ‘calyces’ without evident pointed processes. C–D. Dorsal and ventral view of BIAÇORES colony (MNHM OCT.A.579), see also Fig. 6A–B. Photographs A–B: Michel Dagnino." figureDoi-0="http://doi.org/10.5281/zenodo.7374065" figureDoi-1="http://doi.org/10.5281/zenodo.7374067" httpUri-0="https://zenodo.org/record/7374065/files/figure.png" httpUri-1="https://zenodo.org/record/7374067/files/figure.png" pageId="21" pageNumber="94">Figs 3–4</figureCitation>
) while all sclerites are colourless.
</paragraph>
</subSubSection>
<caption id="989ED19AA83F570D81F483D541D9F844" ID-DOI="http://doi.org/10.5281/zenodo.7374084" ID-Zenodo-Dep="7374084" httpUri="https://zenodo.org/record/7374084/files/figure.png" pageId="21" pageNumber="94" startId="21.[189,232,1961,1987]" targetBox="[189,1398,573,1923]" targetPageId="21">
<paragraph id="CC5E8112A83F570D81F483D541D9F844" blockId="21.[189,1399,1961,2023]" pageId="21" pageNumber="94">
<emphasis id="FE955D00A83F570D81F483D54349F861" bold="true" box="[189,285,1961,1987]" pageId="21" pageNumber="94">Fig. 11.</emphasis>
<taxonomicName id="0BE1FA91A83F570D806183D540A3F861" authority="Studer, 1891" authorityName="Studer" authorityYear="1891" box="[296,759,1961,1987]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="21" pageNumber="94" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A83F570D806183D54004F861" box="[296,592,1961,1987]" italics="true" pageId="21" pageNumber="94">Gyrophyllum hirondellei</emphasis>
<bibRefCitation id="A870FCE3A83F570D831283D540A3F861" author="Studer T." box="[603,759,1961,1987]" pageId="21" pageNumber="94" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer, 1891</bibRefCitation>
</taxonomicName>
, BECA (OPEN-660). SEM photographs of sclerites. 
<emphasis id="FE955D00A83F570D81F483B04286F844" bold="true" box="[189,210,1996,2022]" pageId="21" pageNumber="94">A</emphasis>
. Rachis interior. 
<emphasis id="FE955D00A83F570D80D583B143E4F845" bold="true" box="[412,432,1997,2023]" pageId="21" pageNumber="94">B</emphasis>
. Peduncle exterior. 
<emphasis id="FE955D00A83F570D83D083B040FAF844" bold="true" box="[665,686,1996,2022]" pageId="21" pageNumber="94">C</emphasis>
. Peduncle interior.
</paragraph>
</caption>
<subSubSection id="84FBD299A83C570E81F485744653FE19" pageId="22" pageNumber="95" type="distribution">
<paragraph id="CC5E8112A83C570E81F4857440D9FE80" blockId="22.[189,653,264,291]" box="[189,653,264,291]" pageId="22" pageNumber="95">
<heading id="9716367EA83C570E81F4857440D9FE80" bold="true" box="[189,653,264,291]" fontSize="11" level="3" pageId="22" pageNumber="95" reason="3">
<emphasis id="FE955D00A83C570E81F4857440D9FE80" bold="true" box="[189,653,264,291]" pageId="22" pageNumber="95">Geographical and depth distribution</emphasis>
</heading>
</paragraph>
<paragraph id="CC5E8112A83C570E81F4854B4653FE19" blockId="22.[189,1399,311,443]" pageId="22" pageNumber="95">
At present, 
<taxonomicName id="0BE1FA91A83C570E8001854B4038FEF3" authorityName="Studer" authorityYear="1891" box="[328,620,311,337]" class="Anthozoa" family="Pennatulidae" genus="Gyrophyllum" kingdom="Animalia" order="Pennatulacea" pageId="22" pageNumber="95" phylum="Cnidaria" rank="species" species="hirondellei">
<emphasis id="FE955D00A83C570E8001854B4038FEF3" box="[328,620,311,337]" italics="true" pageId="22" pageNumber="95">Gyrophyllum hirondellei</emphasis>
</taxonomicName>
is known from the North Atlantic, from Rosemary Seamount and Rockall and Hatton Banks to Azores and 
<collectingCountry id="B4F6C182A83C570E83EA85264146FED6" box="[675,786,346,372]" name="Bahamas" pageId="22" pageNumber="95">Bahamas</collectingCountry>
, over a bathymetric range of 
<quantity id="0B192CF7A83C570E8520852646A8FED7" box="[1129,1276,346,373]" metricMagnitude="3" metricUnit="m" metricValue="1.4705000000000001" metricValueMax="2.2199999999999998" metricValueMin="0.7209999999999999" pageId="22" pageNumber="95" unit="m" value="1470.5" valueMax="2220.0" valueMin="721.0">721–2220 m</quantity>
depth, the shallower records being those in reported in this paper (see 
<tableCitation id="8163B4A9A83C570E82CC8501418FFE3A" box="[901,987,381,408]" captionStart="Table 1" captionStartId="4.[189,259,265,291]" captionTargetBox="[195,1386,402,1981]" captionTargetPageId="4" captionText="Table 1 (continued on next two pages). Pennatulaceans included in molecular phylogenetic analyses in this paper. Species and GenBank accession numbers in bold are those sequenced for this study. Abbreviation: n.d. = no data." httpUri="http://table.plazi.org/id/989ED19AA82E571C81F485754044FECB" pageId="22" pageNumber="95" tableUuid="989ED19AA82E571C81F485754044FECB">Table 1</tableCitation>
) (
<bibRefCitation id="A870FCE3A83C570E82BE850146D8FE3A" author="Studer T." box="[1015,1164,381,408]" pageId="22" pageNumber="95" pagination="86 - 95" refId="ref15658" refString="Studer T. 1891. Note preliminaire sur les Alcyonaires provenant des campagnes du yacht l'Hirondelle 1886, 1887, 1888. Part 2. Alcyonacea and Pennatulacea. Memoires de la Societe Zoologique de France 4 (2): 86 - 95." type="journal article" year="1891">Studer 1891</bibRefCitation>
; 
<bibRefCitation id="A870FCE3A83C570E85D085024683FE3A" author="Studer T." box="[1177,1239,382,408]" pageId="22" pageNumber="95" pagination="1 - 64" refId="ref15703" refString="Studer T. 1901. Alcyonaires provenant des campagnes de l'Hirondelle (1886 - 88). Resultats des Campagnes Scientifiques du Prince Albert Ier de Monaco 20: 1 - 64." type="journal article" year="1901">1901</bibRefCitation>
; 
<bibRefCitation id="A870FCE3A83C570E85AC85014726FE3A" author="Roule L." box="[1253,1394,381,408]" pageId="22" pageNumber="95" pagination="454 - 458" refId="ref15555" refString="Roule L. 1905. Notice preliminaire sur les pennatulides recueillis par le &quot; Travailleur &quot; et le &quot; Talisman &quot;, dans l'Ocean Atlantique, au large du Maroc. Bulletin du Museum national d'histoire naturelle, Paris 11: 454 - 458." type="journal article" year="1905">Roule 1905</bibRefCitation>
; 
<bibRefCitation id="A870FCE3A83C570E81F785DD43DFFE18" author="Deichmann E." box="[190,395,416,443]" pageId="22" pageNumber="95" pagination="1 - 317" refId="ref13659" refString="Deichmann E. 1936. The Alcyonaria of the western part of the Atlantic Ocean. Memoirs of the Museum of Comparative Zoology, Harvard 53: 1 - 317. https: // doi. org / 10.5962 / bhl. title. 49348" type="journal article" year="1936">Deichmann 1936</bibRefCitation>
; 
<bibRefCitation id="A870FCE3A83C570E80DF85DD4166FE19" author="Tixier-Durivault A. &amp; d'Hondt M. - J." box="[406,818,417,443]" pageId="22" pageNumber="95" pagination="1361 - 1433" refId="ref15906" refString="Tixier-Durivault A. &amp; d'Hondt M. - J. 1974 b. Les octocoralliaires de la campagne Biacores. Bulletin du Museum national d'histoire naturelle, Paris 174: 1361 - 1433." type="journal article" year="1974">Tixier-Durivault &amp; d’Hondt 1974b</bibRefCitation>
; present account).
</paragraph>
</subSubSection>
</treatment>
</document>