295 lines
34 KiB
XML
295 lines
34 KiB
XML
<document id="167D7B51AE33B410AD7A0B000160C898" ID-DOI="10.5281/zenodo.184571" ID-GBIF-Dataset="b623d656-73b0-4edc-a7fe-e86b55d5a10d" ID-ISSN="1175-5326" ID-Zenodo-Dep="184571" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459988450462" checkinUser="plazi" docAuthor="Steenkiste, Niels Van, Volonterio, Odile, Schockaert, Ernest & Artois, Tom" docDate="2008" docId="443987C2BB2D267CFF4D2C886B72FD15" docLanguage="en" docName="zt01914p033.pdf" docOrigin="Zootaxa 1914" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Vauclusia multistriata Steenkiste, Volonterio, Schockaert & Artois, 2008, n.sp." docType="treatment" docVersion="7" lastPageNumber="26" masterDocId="B800FFBABB3A2665FFDA2C6A6E37FFF4" masterDocTitle="Marine Rhabdocoela (Platyhelminthes, Rhabditophora) from Uruguay, with the description of eight new species and two new genera" masterLastPageNumber="33" masterPageNumber="1" pageNumber="24" updateTime="1698232526720" updateUser="plazi">
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<mods:titleInfo id="2BB95AC15171E15FAAFF7B5667B5BA0C">
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<mods:title id="D74BD5096474474D079AFEC9C28E8072">Marine Rhabdocoela (Platyhelminthes, Rhabditophora) from Uruguay, with the description of eight new species and two new genera</mods:title>
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<mods:name id="AD64BAD9F7C6BA551CF409498F2AEA12" type="personal">
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<mods:roleTerm id="0DA6A6AF6D03EB222E4830A83CC6EDD7">Author</mods:roleTerm>
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<mods:namePart id="75EB73EBB3263172200EC95025D86D09">Steenkiste, Niels Van</mods:namePart>
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<mods:name id="0CD033A67B5C68D46E2FB86D1A1F779B" type="personal">
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<mods:roleTerm id="DF25295B815AA3CCD2122106B52B5DB0">Author</mods:roleTerm>
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<mods:namePart id="CB34706AE2EEC685DF4F4EECFB8AE596">Volonterio, Odile</mods:namePart>
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<mods:name id="A400C258AB438A9D4A0A739E66CE9415" type="personal">
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<mods:roleTerm id="2B4B3E8B94F63E4BB4FCBB70351C09BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8DD5B4EBAC804E1154E4DE99417D0B0D">Schockaert, Ernest</mods:namePart>
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</mods:name>
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<mods:name id="46484080593898F34137B216064E9A67" type="personal">
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<mods:roleTerm id="20B22E615379E7D486B867E5B9668642">Author</mods:roleTerm>
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<mods:namePart id="36ADEF7DD4445546EC786FE8BB556371">Artois, Tom</mods:namePart>
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<mods:typeOfResource id="5F2D0FB5190E79B7E7F4F146F98F2000">text</mods:typeOfResource>
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<mods:title id="E8A625A4C6C1841FB95561EFAB8D9F14">Zootaxa</mods:title>
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<mods:part id="3B7B95A0405A16C156C4A007C19F6056">
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<mods:date id="CCFB404B57C7433B05315D0BEEDA37DC">2008</mods:date>
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<mods:detail id="DFCD42B3DD6BBA90250FD511A4BCB9B9" type="volume">
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<mods:number id="4D909931B205AB8606E39B38D98026DC">1914</mods:number>
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<mods:classification id="E7F67E129C128C71AE188ABC6472F386">journal article</mods:classification>
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<mods:identifier id="218DF01D2D69BC444B6F48E7B1CEA157" type="DOI">10.5281/zenodo.184571</mods:identifier>
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<mods:identifier id="0F410F1B642C81DF596CEBD1A1DE78B7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="443987C2BB2D267CFF4D2C886B72FD15" ID-DOI="http://doi.org/10.5281/zenodo.6230427" ID-GBIF-Taxon="119379738" ID-Zenodo-Dep="6230427" LSID="urn:lsid:plazi:treatment:443987C2BB2D267CFF4D2C886B72FD15" httpUri="http://treatment.plazi.org/id/443987C2BB2D267CFF4D2C886B72FD15" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
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<subSubSection id="848A655FBB2D2672FF4D2C886F0DFEEB" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="CC2F36D4BB2D2672FF4D2C886FD4FF08" blockId="23.[151,483,226,287]" box="[151,483,226,252]" pageId="23" pageNumber="24">
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<heading id="976781B8BB2D2672FF4D2C886FD4FF08" bold="true" box="[151,483,226,252]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="FEE4EAC6BB2D2672FF4D2C886FD4FF08" bold="true" box="[151,483,226,252]" pageId="23" pageNumber="24">
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<taxonomicName id="0B904D57BB2D2672FF4D2C886F97FF08" ID-CoL="7FJQH" box="[151,416,226,252]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
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<emphasis id="FEE4EAC6BB2D2672FF4D2C886F97FF08" bold="true" box="[151,416,226,252]" italics="true" pageId="23" pageNumber="24">Vauclusia multistriata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E5D757BDBB2D2672FE722C886FD4FF08" box="[424,483,226,252]" pageId="23" pageNumber="24" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="CC2F36D4BB2D2672FF4D2D6F6F0DFEEB" blockId="23.[151,483,226,287]" box="[151,314,261,287]" pageId="23" pageNumber="24">
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(
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<figureCitation id="54AB2A51BB2D2672FF452D6F6EC6FEEB" box="[159,241,261,287]" captionStart="FIGURE 9" captionStartId="22.[151,255,1532,1556]" captionTargetBox="[227,1360,522,1501]" captionTargetId="figure@22.[227,1361,514,1508]" captionTargetPageId="22" captionText="FIGURE 9. Lurus evelinae Marcus, 1950 (A) Habitus of a live animal. (B) Stylet. Vauclusia multistriata n. sp. (C) Habitus of a live animal." httpUri="https://zenodo.org/record/184580/files/figure.png" pageId="23" pageNumber="24">Figs. 9</figureCitation>
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C, 10)
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</paragraph>
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</subSubSection>
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<subSubSection id="848A655FBB2D2672FF4D2D3F6B65FE4B" pageId="23" pageNumber="24" type="description">
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<paragraph id="CC2F36D4BB2D2672FF4D2D3F6B65FE4B" blockId="23.[151,1437,341,2007]" pageId="23" pageNumber="24">
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<emphasis id="FEE4EAC6BB2D2672FF4D2D3F6F34FE9B" bold="true" box="[151,259,341,367]" pageId="23" pageNumber="24">Locality.</emphasis>
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La Coronilla, Departamento de Rocha,
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<collectingCountry id="B4877644BB2D2672FD242D3F6D52FE9B" box="[766,869,341,367]" name="Uruguay" pageId="23" pageNumber="24">Uruguay</collectingCountry>
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(
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<geoCoordinate id="A9A45013BB2D2672FCA12D3F6A19FE9B" box="[891,1070,341,367]" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="-33.90514">33°54’18.50”S</geoCoordinate>
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,
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<geoCoordinate id="A9A45013BB2D2672FB9B2D3F6B37FE9B" box="[1089,1280,341,367]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-53.510918">53°30’39.30”W</geoCoordinate>
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). Beach and mouth of the canal near hotel Parque Océanico: sand covered by a thin green layer of organic material and sand with organic material near a small pool in open contact with the ocean (
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<date id="B82E1014BB2D2672FBC02DCF6A90FE4B" box="[1050,1191,421,447]" pageId="23" pageNumber="24" value="2004-08-01">01/08/2004</date>
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):
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<typeStatus id="132B8876BB2D2672FB602DCF6ADAFE4B" box="[1210,1261,421,447]" pageId="23" pageNumber="24">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="848A655FBB2D2672FF1C2DA76A1CFDFB" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="CC2F36D4BB2D2672FF1C2DA76A1CFDFB" blockId="23.[151,1437,341,2007]" pageId="23" pageNumber="24">
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<emphasis id="FEE4EAC6BB2D2672FF1C2DA76F0DFE13" bold="true" box="[198,314,461,487]" pageId="23" pageNumber="24">Material.</emphasis>
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Observations on one live animal. One whole mount, designated
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<typeStatus id="132B8876BB2D2672FBE42DA76A92FE13" box="[1086,1189,461,487]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="AA81AE11BB2D2672FB6F2DA76B3BFE13" box="[1205,1292,461,487]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="23" pageNumber="24">SMNH</collectionCode>
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7501). Two serially-sectioned specimens, of which one designated
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<typeStatus id="132B8876BB2D2672FCC02D9F6D48FDFB" box="[794,895,501,527]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
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(HU no. 363).
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</paragraph>
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</subSubSection>
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<subSubSection id="848A655FBB2D2672FF1C2E776F47FDAB" pageId="23" pageNumber="24" type="etymology">
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<paragraph id="CC2F36D4BB2D2672FF1C2E776F47FDAB" blockId="23.[151,1437,341,2007]" pageId="23" pageNumber="24">
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<emphasis id="FEE4EAC6BB2D2672FF1C2E776F64FDC3" bold="true" box="[198,339,541,567]" pageId="23" pageNumber="24">Etymology.</emphasis>
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The species name refers to the many internal ridges of the stylet. Multus (Lat.): many. Striatus (Lat.): striated.
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</paragraph>
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</subSubSection>
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<subSubSection id="848A655FBB2D267DFF1C2E076C24FE56" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
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<paragraph id="CC2F36D4BB2D2672FF1C2E076F03FCD3" blockId="23.[151,1437,341,2007]" pageId="23" pageNumber="24">
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<emphasis id="FEE4EAC6BB2D2672FF1C2E076F69FD73" bold="true" box="[198,350,621,647]" pageId="23" pageNumber="24">Description.</emphasis>
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Slender, 0.5 mm long animal. Eyes absent. The habitus is characterized by division of the body into three zones: a slender, rostral part where rhabdite glands occur (± 1/5 of the body length), a middle part where all organs are situated (± 3/5 of the body length), and a caudal part, which consists of a clearlydelimited, small tail with an epidermis containing many more rhabdites than the rest of the body (± 1/5 of the body length).
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</paragraph>
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<paragraph id="CC2F36D4BB2D2672FF1F2F5F6F79FB7B" blockId="23.[151,1437,341,2007]" pageId="23" pageNumber="24">
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The ciliated epidermis is cellular and has a height of about 3 μm. The basal membrane is very thin. The cilia measure about 3 μm. In the rostral part of the body, two different kinds of rhabdite glands occur. The first
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<typeStatus id="132B8876BB2D2672FF4D2FEF6EFDFC6B" box="[151,202,901,927]" pageId="23" pageNumber="24">type</typeStatus>
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produces long, lancet-shaped, basophilic rhabdites with a length of 8–10 μm, the other
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<typeStatus id="132B8876BB2D2672FB2C2FEF6B1EFC6B" box="[1270,1321,901,927]" pageId="23" pageNumber="24">type</typeStatus>
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produces smaller, oval, eosinophilic rhabdites with a length of about 2–3 μm. Both kinds of glands extend from above the brain to the rostral body end, where they empty. Caudally from the pharynx, dispersed eosinophilic rhabdite glands occur, which empty in the epidermis. The largest concentration of these rhabdite glands is situated behind the genital system in the caudal body part. The length of the oval, eosinophilic rhabdites that are secreted by these glands varies between 2–4 μm. The epidermis of the caudal region is completely filled with these rhabdites.
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</paragraph>
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<paragraph id="CC2F36D4BB2D2672FF1F28F76DBFFB2B" blockId="23.[151,1437,341,2007]" pageId="23" pageNumber="24">
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The mouth is situated at about 40%. The organisation and structure of the pharynx is identical with that of
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<taxonomicName id="0B904D57BB2D2672FF4D28AF6C64FB2B" authority="Willems et al., 2004" authorityName="Willems et al." authorityYear="2004" box="[151,595,1221,1247]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="conica">
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<emphasis id="FEE4EAC6BB2D2672FF4D28AF6F69FB2A" box="[151,350,1221,1246]" italics="true" pageId="23" pageNumber="24">Vauclusia conica</emphasis>
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<bibRefCitation id="A8014B25BB2D2672FEBF28AF6C64FB2B" author="Willems" box="[357,595,1221,1247]" pageId="23" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article" year="2004">
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Willems
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<emphasis id="FEE4EAC6BB2D2672FE1528AF6C3EFB2A" box="[463,521,1221,1246]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
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, 2004
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</bibRefCitation>
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</taxonomicName>
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(see
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<bibRefCitation id="A8014B25BB2D2672FD4B28AF6D4BFB2B" author="Willems" box="[657,892,1221,1247]" pageId="23" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article" year="2004">
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Willems
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<emphasis id="FEE4EAC6BB2D2672FD2028AF6D02FB2A" box="[762,821,1221,1246]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
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2004
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="CC2F36D4BB2D2672FF1F28876B72F8FB" blockId="23.[151,1437,341,2007]" pageId="23" pageNumber="24">The gonads are paired. The two round testes are situated ventrally, just in front of the pharynx and at both sides of the body. The seminal vesicles are rather small and have an anucleated, membranous epithelium. They are not surrounded by muscle layers. When entering the prostate vesicle, both vasa deferentia join to form a ductus ejaculatorius, which runs centrally through the prostate vesicle. Centrally, this prostate vesicle contains coarse-grained, eosinophilic glands, which are surrounded by fine-grained eosinophilic glands. All glands together take up about half of the total volume of the prostate vesicle. Both the fine-grained and coarse-grained eosinophilic glands have extracapsular nuclei-containing parts, and penetrate the prostate vesicle together with the ejaculatory duct. The prostate vesicle is surrounded by thick, inner circular muscles and outer longitudinal muscles. Distally, the prostate vesicle is connected with a conical and thin-walled stylet. This stylet has a length of 45 μm. Proximally, it is 21 μm broad at its broadest, distally up to 4 μm. The inner side of this stylet is ornamented with a large number of ridges that are oriented in three different directions. For the first 17 μm, the ridges run lengthways, the next 7 μm they run transversely and the last 21 μm again lengthways. The male genital atrium is surrounded by circular muscles and is lined with an anucleated, membranous epithelium, as is the common genital atrium. The common gonopore is situated at about 70%.</paragraph>
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<paragraph id="CC2F36D4BB2D267DFF1F2B776C24FE56" blockId="23.[151,1437,341,2007]" lastBlockId="24.[151,1437,152,418]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">The ovaries lie caudally from the gonopore and together with the more rostrally-situated vitellaria, they form long ovovitellaria, which stretch dorsolaterally at both sides beyond the testes. The oviducts are very short, and open in the long, very muscular female duct, which caudally enters the common genital atrium. It is lined with a high, anucleated epithelium and surrounded by a circular muscle layer, which is very well developed in the proximal half. Distally, it is slightly swollen and some nuclei are seen in the somewhat higher epithelium of the transition zone between the female duct and the common genital atrium. Moreover, the distal half of the female duct is surrounded by a peripheral, longitudinal muscle layer. The female bursa is spherical and lined with a nucleated, membranous epithelium. It contains a great deal of sperm in the lumen. A broad bursal stalk connects the bursa with the proximal end of the female duct. Its epithelium is very high and shows many glands. Where it enters the bursa, a sphincter is present. In the proximal part of the female duct, sperm and a glandular secretion were observed. Eosinophilic and basophilic female glands enter this part of the female duct. A uterus is lacking.</paragraph>
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</subSubSection>
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<caption id="98EF665CBB22267DFF4D29FF6CD7FA3B" httpUri="https://zenodo.org/record/184581/files/figure.png" pageId="24" pageNumber="25" targetBox="[156,1321,483,1402]" targetPageId="24">
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<paragraph id="CC2F36D4BB22267DFF4D29FF6CD7FA3B" blockId="24.[151,1436,1429,1487]" pageId="24" pageNumber="25">
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<emphasis id="FEE4EAC6BB22267DFF4D29FF6F1CFA59" bold="true" box="[151,299,1429,1453]" pageId="24" pageNumber="25">FIGURE 10.</emphasis>
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<taxonomicName id="0B904D57BB22267DFEEC29FC6C12FA59" box="[310,549,1430,1453]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
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<emphasis id="FEE4EAC6BB22267DFEEC29FC6C12FA59" box="[310,549,1430,1453]" italics="true" pageId="24" pageNumber="25">Vauclusia multistriata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E5D757BDBB22267DFDEA29FF6C51FA59" box="[560,614,1429,1453]" pageId="24" pageNumber="25" rank="species">
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<emphasis id="FEE4EAC6BB22267DFDEA29FF6C51FA59" bold="true" box="[560,614,1429,1453]" pageId="24" pageNumber="25">n.sp.</emphasis>
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</taxonomicNameLabel>
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(A) Reconstruction of the entire animal. (B) Reconstruction of the atrial organs. (C) Stylet with internal ridges (from holotype).
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</paragraph>
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</caption>
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<subSubSection id="848A655FBB22267CFF1F2A686AA0FD9D" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
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<paragraph id="CC2F36D4BB22267DFF1F2A686F38F8C1" blockId="24.[151,1436,1538,2005]" pageId="24" pageNumber="25">
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<emphasis id="FEE4EAC6BB22267DFF1F2A686F66F9E8" bold="true" box="[197,337,1538,1564]" pageId="24" pageNumber="25">Discussion.</emphasis>
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The species described here can easily be placed in the taxon
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<emphasis id="FEE4EAC6BB22267DFBEA2A696AA9F9E8" box="[1072,1182,1539,1564]" italics="true" pageId="24" pageNumber="25">Vau c lu s ia</emphasis>
|
||
<bibRefCitation id="A8014B25BB22267DFB702A696BACF9E9" author="Willems" box="[1194,1435,1539,1565]" pageId="24" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article" year="2004">
|
||
Willems
|
||
<emphasis id="FEE4EAC6BB22267DFACF2A696B66F9E8" box="[1301,1361,1539,1564]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
because of the combined presence of the following characters (see
|
||
<bibRefCitation id="A8014B25BB22267DFC7A2A416ABEF9B1" author="Willems" box="[928,1161,1579,1605]" pageId="24" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article" year="2004">
|
||
Willems
|
||
<emphasis id="FEE4EAC6BB22267DFBD22A416A75F9B0" box="[1032,1090,1579,1604]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
): pharynx in the middle of the body, paired testes and seminal vesicles, spherical prostate vesicle and two
|
||
<typeStatus id="132B8876BB22267DFBAE2A396A85F999" box="[1140,1202,1619,1645]" pageId="24" pageNumber="25">types</typeStatus>
|
||
of glandular secretions, conical stylet with different internal ridges, a long male atrium, paired ovovitellaria, a long muscular female duct with a swollen distal part, and the presence of a female bursa and female glands. The construction of the pharynx and the position of the testes are comparable with those of
|
||
<taxonomicName id="0B904D57BB22267DFBDF2AA16B5BF911" authority="Willems et al., 2004" authorityName="Willems et al." authorityYear="2004" box="[1029,1388,1739,1765]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="conica">
|
||
<emphasis id="FEE4EAC6BB22267DFBDF2AA16A46F910" box="[1029,1137,1739,1764]" italics="true" pageId="24" pageNumber="25">V. conica</emphasis>
|
||
<bibRefCitation id="A8014B25BB22267DFBA32AA16B5BF911" author="Willems" box="[1145,1388,1739,1765]" pageId="24" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article" year="2004">
|
||
Willems
|
||
<emphasis id="FEE4EAC6BB22267DFB392AA16B28F910" box="[1251,1311,1739,1764]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, the only other representative of the taxon, although in
|
||
<taxonomicName id="0B904D57BB22267DFD3F2A996DBCF8F8" box="[741,907,1779,1804]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
|
||
<emphasis id="FEE4EAC6BB22267DFD3F2A996DBCF8F8" box="[741,907,1779,1804]" italics="true" pageId="24" pageNumber="25">V. multistriata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5D757BDBB22267DFC482A986DFAF8F8" box="[914,973,1778,1804]" pageId="24" pageNumber="25" rank="species">
|
||
<emphasis id="FEE4EAC6BB22267DFC482A986DFAF8F8" bold="true" box="[914,973,1778,1804]" pageId="24" pageNumber="25">n.sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
the pharynx is situated somewhat more anteriorly.
|
||
</paragraph>
|
||
<paragraph id="CC2F36D4BB22267CFF1F2B296D02FE8E" blockId="24.[151,1436,1538,2005]" lastBlockId="25.[151,1436,151,737]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
A first important difference between the two species occurs in the size of the stylet and the pattern of its ridges. Although
|
||
<taxonomicName id="0B904D57BB22267DFEBD2B016C06F870" box="[359,561,1899,1924]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="conica">
|
||
<emphasis id="FEE4EAC6BB22267DFEBD2B016C06F870" box="[359,561,1899,1924]" italics="true" pageId="24" pageNumber="25">Vauclusia conica</emphasis>
|
||
</taxonomicName>
|
||
is almost three times as long as
|
||
<taxonomicName id="0B904D57BB22267DFC642B016A51F870" box="[958,1126,1899,1924]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
|
||
<emphasis id="FEE4EAC6BB22267DFC642B016A51F870" box="[958,1126,1899,1924]" italics="true" pageId="24" pageNumber="25">V. multistriata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5D757BDBB22267DFBB52B006A9DF870" box="[1135,1194,1898,1924]" pageId="24" pageNumber="25" rank="species">
|
||
<emphasis id="FEE4EAC6BB22267DFBB52B006A9DF870" bold="true" box="[1135,1194,1898,1924]" pageId="24" pageNumber="25">n.sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, its stylet is smaller than the stylet of
|
||
<taxonomicName id="0B904D57BB22267DFEB22BF96C38F858" box="[360,527,1939,1964]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
|
||
<emphasis id="FEE4EAC6BB22267DFEB22BF96C38F858" box="[360,527,1939,1964]" italics="true" pageId="24" pageNumber="25">V. multistriata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5D757BDBB22267DFDC22BF86C64F858" box="[536,595,1938,1964]" pageId="24" pageNumber="25" rank="species">
|
||
<emphasis id="FEE4EAC6BB22267DFDC22BF86C64F858" bold="true" box="[536,595,1938,1964]" pageId="24" pageNumber="25">n.sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
The exact structure of the stylet in
|
||
<taxonomicName id="0B904D57BB22267DFC242BF96A5DF858" box="[1022,1130,1939,1964]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="conica">
|
||
<emphasis id="FEE4EAC6BB22267DFC242BF96A5DF858" box="[1022,1130,1939,1964]" italics="true" pageId="24" pageNumber="25">V. conica</emphasis>
|
||
</taxonomicName>
|
||
could not be ascertained, but it apparently has about eight spirally-running and striated ridges (
|
||
<bibRefCitation id="A8014B25BB22267DFC062BD16AE5F821" author="Willems" box="[988,1234,1979,2005]" pageId="24" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article" year="2004">
|
||
Willems
|
||
<emphasis id="FEE4EAC6BB22267DFB902BD16ABFF820" box="[1098,1160,1979,2004]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
). In contrast, the stylet of
|
||
<taxonomicName id="0B904D57BB23267CFF262CF26F96FF45" box="[252,417,152,177]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
|
||
<emphasis id="FEE4EAC6BB23267CFF262CF26F96FF45" box="[252,417,152,177]" italics="true" pageId="25" pageNumber="26">V. multistriata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5D757BDBB23267CFE732CFD6FD3FF45" box="[425,484,151,177]" pageId="25" pageNumber="26" rank="species">
|
||
<emphasis id="FEE4EAC6BB23267CFE732CFD6FD3FF45" bold="true" box="[425,484,151,177]" pageId="25" pageNumber="26">n.sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has many ridges, transverse as well as longitudinal. Secondly, the prostate glands of
|
||
<emphasis id="FEE4EAC6BB23267CFF602CAA6F12FF2D" box="[186,293,192,217]" italics="true" pageId="25" pageNumber="26">V. c o n i c a</emphasis>
|
||
take up the full volume of the prostate vesicle, which is not the case in
|
||
<taxonomicName id="0B904D57BB23267CFB772CAA6B62FF2D" box="[1197,1365,192,217]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
|
||
<emphasis id="FEE4EAC6BB23267CFB772CAA6B62FF2D" box="[1197,1365,192,217]" italics="true" pageId="25" pageNumber="26">V. multistriata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5D757BDBB23267CFABB2CD56BABFF2D" box="[1377,1436,191,217]" pageId="25" pageNumber="26" rank="species">
|
||
<emphasis id="FEE4EAC6BB23267CFABB2CD56BABFF2D" bold="true" box="[1377,1436,191,217]" pageId="25" pageNumber="26">n.sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Another difference is the presence of a bursal stalk with a glandular epithelium between the bursa and the female duct in
|
||
<taxonomicName id="0B904D57BB23267CFE8B2D7A6FCDFEDD" box="[337,506,272,297]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">
|
||
<emphasis id="FEE4EAC6BB23267CFE8B2D7A6FCDFEDD" box="[337,506,272,297]" italics="true" pageId="25" pageNumber="26">V. multistriata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5D757BDBB23267CFDDF2D656C77FEDD" box="[517,576,271,297]" pageId="25" pageNumber="26" rank="species">
|
||
<emphasis id="FEE4EAC6BB23267CFDDF2D656C77FEDD" bold="true" box="[517,576,271,297]" pageId="25" pageNumber="26">n.sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
In
|
||
<emphasis id="FEE4EAC6BB23267CFDB42D7A6CD2FEDD" box="[622,741,272,297]" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="0B904D57BB23267CFDB42D7A6CD7FEDD" box="[622,736,272,297]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="conica">V. conica</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the bursa links up directly with the proximal part of the female duct through a sphincter. Finally,
|
||
<taxonomicName id="0B904D57BB23267CFDA62D526CDFFEA5" box="[636,744,312,337]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="conica">
|
||
<emphasis id="FEE4EAC6BB23267CFDA62D526CDFFEA5" box="[636,744,312,337]" italics="true" pageId="25" pageNumber="26">V. conica</emphasis>
|
||
</taxonomicName>
|
||
has an inversion of the muscle layer in the transition zone between the common genital atrium and the male atrium.
|
||
</paragraph>
|
||
<paragraph id="CC2F36D4BB23267CFF1F2DE26ADBFE3E" blockId="25.[151,1436,151,737]" pageId="25" pageNumber="26">
|
||
Some of the features that are unique to
|
||
<taxonomicName id="0B904D57BB23267CFD422DE26D57FE55" box="[664,864,392,417]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="conica">
|
||
<emphasis id="FEE4EAC6BB23267CFD422DE26D57FE55" box="[664,864,392,417]" italics="true" pageId="25" pageNumber="26">Vauclusia conica</emphasis>
|
||
</taxonomicName>
|
||
were mentioned in the original diagnosis of this genus by
|
||
<bibRefCitation id="A8014B25BB23267CFEDD2DDA6C34FE3E" box="[263,515,432,458]" pageId="25" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article">
|
||
Willems
|
||
<emphasis id="FEE4EAC6BB23267CFEAA2DDA6F9CFE3D" box="[368,427,432,457]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
, but now these features should be removed from this diagnosis.
|
||
</paragraph>
|
||
<paragraph id="CC2F36D4BB23267CFF1F2DB26AA0FD9D" blockId="25.[151,1436,151,737]" pageId="25" pageNumber="26">
|
||
<emphasis id="FEE4EAC6BB23267CFF1F2DB26CDAFE05" box="[197,749,472,497]" italics="true" pageId="25" pageNumber="26">
|
||
Amended diagnosis (after
|
||
<bibRefCitation id="A8014B25BB23267CFE202DB26CD6FE05" author="Willems" box="[506,737,472,497]" pageId="25" pageNumber="33" refString="Willems, W. R., Artois, T. J., Vermin, W., Backeljau, T., Schockaert, E. R. (2004) Reports on the free-living Plathelminthes from Australia: Typhloplanoida, with description of three new taxa. Zoological Science, 21, 333 - 341." type="journal article" year="2004">Willems et al. 2004</bibRefCitation>
|
||
).
|
||
</emphasis>
|
||
Va u cl u si a:
|
||
<emphasis id="FEE4EAC6BB23267CFCAD2DB26AA0FD9D" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="0B904D57BB23267CFCAD2DB26A7FFE05" box="[887,1096,472,497]" class="Rhabditophora" family="Promesostomidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="family">Promesostomidae</taxonomicName>
|
||
with the pharynx in the middle of the body. Paired testes and seminal vesicles. Spherical prostate vesicle with two
|
||
<typeStatus id="132B8876BB23267CFB6D2E6A6AC4FDED" box="[1207,1267,512,537]" pageId="25" pageNumber="26">types</typeStatus>
|
||
of secretions. Conical stylet with several internal ridges. Paired ovovitellaria. Very long, muscular female duct. Distal part of the female duct swollen. Female bursa and female glands present.
|
||
<collectingCountry id="B4877644BB23267CFC182E3A6DC9FD9D" box="[962,1022,592,617]" name="Malta" pageId="25" pageNumber="26">Male</collectingCountry>
|
||
atrium long.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="848A655FBB23267CFF1F2E126B72FD15" pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph id="CC2F36D4BB23267CFF1F2E126B72FD15" blockId="25.[151,1436,151,737]" pageId="25" pageNumber="26">
|
||
<emphasis id="FEE4EAC6BB23267CFF1F2E126F73FD65" box="[197,324,632,657]" italics="true" pageId="25" pageNumber="26">Diagnosis.</emphasis>
|
||
<taxonomicName id="0B904D57BB23267CFE962E126C7DFD66" box="[332,586,632,658]" class="Rhabditophora" family="Promesostomidae" genus="Vauclusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="multistriata" status="sp. nov.">Vauclusia multistriata</taxonomicName>
|
||
<emphasis id="FEE4EAC6BB23267CFD882E1D6D27FD65" box="[594,784,631,657]" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="E5D757BDBB23267CFD882E1D6CBCFD65" box="[594,651,631,657]" pageId="25" pageNumber="26" rank="species">
|
||
<emphasis id="FEE4EAC6BB23267CFD882E1D6CBCFD65" bold="true" box="[594,651,631,657]" italics="true" pageId="25" pageNumber="26">n.sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: species of
|
||
</emphasis>
|
||
Va uc lu s ia
|
||
<emphasis id="FEE4EAC6BB23267CFC432E126A30FD65" box="[921,1031,632,657]" italics="true" pageId="25" pageNumber="26">with a 45</emphasis>
|
||
μ
|
||
<emphasis id="FEE4EAC6BB23267CFBC72E126B72FD15" italics="true" pageId="25" pageNumber="26">m-long stylet with many internal ridges that run lengthways proximally and distally, and transversely in between. Prostate glands only fill a portion of the prostate vesicle. Duct with glandular epithelium between the bursa and the female duct.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |