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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.374.6553" ID-GBIF-Dataset="6ecd0090-10bd-4484-84a5-85640fd29570" ID-PMC="PMC3909810" ID-Pensoft-Pub="1313-2970-374-23" ID-PubMed="24493963" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-374-23" ModsDocOrigin="ZooKeys 374" ModsDocTitle="Lectotype designations and nomenclatural changes in Xylographus Mellié (Coleoptera, Ciidae)" checkinTime="1451246398404" checkinUser="pensoft" docAuthor="Sandoval-Gomez, Vivian Eliana, Lopes-Andrade, Cristiano &amp; Lawrence, John F." docDate="2014" docId="442EE6114D9C0BF8AF681C3084222B50" docLanguage="en" docName="ZooKeys 374: 23-43" docOrigin="ZooKeys 374" docSource="http://dx.doi.org/10.3897/zookeys.374.6553" docTitle="Xylographus madagascariensis Mellie 1849" docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="FFBB237380611814FFB0FFC82D18CB0D" masterDocTitle="Lectotype designations and nomenclatural changes in Xylographus Mellie (Coleoptera, Ciidae)" masterLastPageNumber="43" masterPageNumber="23" pageNumber="31" updateTime="1668157579771" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Lectotype designations and nomenclatural changes in Xylographus Mellie (Coleoptera, Ciidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sandoval-Gomez, Vivian Eliana</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lopes-Andrade, Cristiano</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lawrence, John F.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>374</mods:number>
</mods:detail>
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<mods:start>23</mods:start>
<mods:end>43</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.374.6553</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.374.6553</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-374-23</mods:identifier>
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<treatment ID-GBIF-Taxon="152050744" LSID="urn:lsid:plazi:treatment:442EE6114D9C0BF8AF681C3084222B50" httpUri="http://treatment.plazi.org/id/442EE6114D9C0BF8AF681C3084222B50" lastPageId="9" lastPageNumber="32" pageId="8" pageNumber="31">
<subSubSection pageId="8" pageNumber="31" type="nomenclature">
<paragraph pageId="8" pageNumber="31">
<taxonomicName LSID="http://species-id.net/wiki/Xylographus_madagascariensis" authority="Mellie, 1849" authorityName="Mellie" authorityYear="1849" class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="madagascariensis">
Xylographus madagascariensis
<normalizedToken originalValue="Mellié">Mellie</normalizedToken>
, 1849
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="31" type="reference_group">
<paragraph pageId="8" pageNumber="31">
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="madagascariensis">Xylographus madagascariensis</taxonomicName>
<bibRefCitation author="Mellie, J" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="15" pageNumber="38" pagination="205 - 274" title="Monographie de l'ancien genre Cis des auteurs." volume="6" year="1849">
<normalizedToken originalValue="Mellié">Mellie</normalizedToken>
1849
</bibRefCitation>
: 224, pl. 9, fig. 18. Type-locality: Madagascar.
</paragraph>
<paragraph pageId="8" pageNumber="31">
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus eichelbaumi" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="eichelbaumi">Xylographus eichelbaumi</taxonomicName>
<bibRefCitation author="Reitter, E" journalOrPublisher="Wiener Entomologische Zeitung" pageId="16" pageNumber="39" pagination="119 - 124" title="Verzeichnis der von Dr. F. Eichelbaum im Jahre 1903 in Deutsch-Ost-Afrika gesammelten Cis-Arten (Coleoptera)." volume="27" year="1908">Reitter 1908</bibRefCitation>
: 119. ssyn. n. Type-locality: Amani, Tanzania.
</paragraph>
<paragraph pageId="8" pageNumber="31">
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus rufipennis" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="rufipennis">Xylographus rufipennis</taxonomicName>
<bibRefCitation author="Pic, M" journalOrPublisher="Moulins" pageId="15" pageNumber="38" pagination="1 - 36" title="Nouveautes diverses." volumeTitle="Melanges Exotico-entomologiques" year="1934">Pic 1934</bibRefCitation>
: 14. syn. n. Type-locality: Gura, Kenya.
</paragraph>
<paragraph pageId="8" pageNumber="31">
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus seychellensis" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="seychellensis">Xylographus seychellensis</taxonomicName>
<bibRefCitation author="Scott, H" journalOrPublisher="Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905. (Ser. 2. Zoology)" pageId="16" pageNumber="39" pagination="1 - 41" title="Coleoptera, Ciidae." volume="Vol. 8" year="1926">Scott 1926</bibRefCitation>
: 10. syn. n. Type-locality: Mahe, Seychelles.
</paragraph>
<paragraph pageId="8" pageNumber="31">
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus tarsalis" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="tarsalis">Xylographus tarsalis</taxonomicName>
<bibRefCitation author="Fahraeus, OI" journalOrPublisher="Vetenskaps-Akademiens, Stockholm" pageId="15" pageNumber="38" pagination="661 - 672" title="Coleoptera Caffrariae, Annis 1838 - 45 a J. A. Wahlberg collecta. Fam. Scolytidae, Paussidae, Bostrichidae et Cioidae. Forhandlingar Kongl." year="1871">
<normalizedToken originalValue="Fåhraeus">Fahraeus</normalizedToken>
1871
</bibRefCitation>
: 670. syn. n. Type-locality: Caffraria (Eastern Cape of South Africa).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="31" type="type series">
<paragraph pageId="8" pageNumber="31">Type series.</paragraph>
<paragraph pageId="8" pageNumber="31">
MADAGASCAR: male lectotype (MNHN), here designated, labeled: &quot;
<taxonomicName genus="Madagascariensis" lsidName="Madagascariensis" pageId="8" pageNumber="31" rank="genus">Madagascariensis</taxonomicName>
Dup. Madagascar.[handwritten] \
<normalizedToken originalValue="Ex-Musæo">Ex-Musaeo</normalizedToken>
Mniszech [printed] \ [red label] LECTOTYPE
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="madagascariensis">Xylographus madagascariensis</taxonomicName>
<normalizedToken originalValue="Mellié">Mellie</normalizedToken>
[handwritten]&quot;.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="32" pageId="8" pageNumber="31" type="type material of junior synonyms">
<paragraph pageId="8" pageNumber="31">Type material of junior synonyms.</paragraph>
<paragraph lastPageId="9" lastPageNumber="32" pageId="8" pageNumber="31">
KENYA: female lectotype (NHM) of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus rufipennis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="rufipennis">
<pageBreakToken pageId="9" pageNumber="32" start="start">Xylographus</pageBreakToken>
rufipennis
</taxonomicName>
Pic, 1934, here designated, labeled: &quot;[red disc]Type [printed] \ R. E, DENT GURA R, 7500 AUG 1929 [printed] \
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus rufipennis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="rufipennis">Xylographus rufipennis</taxonomicName>
n. sp. [handwritten] \ Pres. By Imp. Inst. Ent B. M. 1934-42. [printed] \ [red label] LECTOTYPE
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus rufipennis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="rufipennis">Xylographus rufipennis</taxonomicName>
Pic [handwritten]&quot;; 1 female paralectotype (MNHN), labeled: &quot;R. E, DENT GURA R, 7500 AUG 1929 [printed] \
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus rufipennis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="rufipennis">Xylographus rufipennis</taxonomicName>
n. sp [handwritten] \ ex. British museum [handwritten] \ [yellow label] PARALECTOTYPE
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus rufipennis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="rufipennis">Xylographus rufipennis</taxonomicName>
Pic [handwritten]&quot;. SEYCHELLES: male lectotype (NHM) of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus seychellensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="seychellensis">Xylographus seychellensis</taxonomicName>
<bibRefCitation author="Scott, H" journalOrPublisher="Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905. (Ser. 2. Zoology)" pageId="16" pageNumber="39" pagination="1 - 41" title="Coleoptera, Ciidae." volume="Vol. 8" year="1926">Scott 1926</bibRefCitation>
, here designated, labeled: &quot;[purple disc] LECTOTYPE [printed] \ Mahe, 1908-9 Seychelles Exp. [printed] \ Percy Sladen Trust Exped. Brit. Mus. 1926-246. [printed] \
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus seychellensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="seychellensis">Xylographus seychellensis</taxonomicName>
, Scott TYPE. ♂. [handwritten] \ Figured specimen [printed] (outline whole vis) [handwritten] \ TYPE [printed] \ [red label] LECTOTYPE
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus seychellensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="seychellensis">Xylographus seychellensis</taxonomicName>
Scott [handwritten]&quot;. SOUTH AFRICA: male lectotype (NHRS) of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus tarsalis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="tarsalis">Xylographus tarsalis</taxonomicName>
<bibRefCitation author="Fahraeus, OI" journalOrPublisher="Vetenskaps-Akademiens, Stockholm" pageId="15" pageNumber="38" pagination="661 - 672" title="Coleoptera Caffrariae, Annis 1838 - 45 a J. A. Wahlberg collecta. Fam. Scolytidae, Paussidae, Bostrichidae et Cioidae. Forhandlingar Kongl." year="1871">
<normalizedToken originalValue="Fåhraeus">Fahraeus</normalizedToken>
1871
</bibRefCitation>
, labeled: &quot;Caffraria [printed] \ J. Wahlb. [printed] \ ♂ [printed] \ [red label] Lectotype ♂
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus tarsalis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="tarsalis">Xylographus tarsalis</taxonomicName>
<normalizedToken originalValue="FÅHR">FAHR</normalizedToken>
. Det. Julio Ferrer 1995 [handwritten] \ [green label] Riksmuseum Stockholm [printed]&quot;; 1 male and 1 female paralectotypes (NHRS), labeled: &quot;Caffraria [printed] \ J. Wahlb. [printed] \ [red label] Paralectotype
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus tarsalis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="tarsalis">Xylographus tarsalis</taxonomicName>
<normalizedToken originalValue="FÅHR">FAHR</normalizedToken>
. Det. Julio Ferrer 1995 [handwritten] \ [green label] Riksmuseum Stockholm [printed]&quot;. TANZANIA: female holotype (NHMW) of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus eichelbaumi" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="eichelbaumi">Xylographus eichelbaumi</taxonomicName>
<bibRefCitation author="Reitter, E" journalOrPublisher="Wiener Entomologische Zeitung" pageId="16" pageNumber="39" pagination="119 - 124" title="Verzeichnis der von Dr. F. Eichelbaum im Jahre 1903 in Deutsch-Ost-Afrika gesammelten Cis-Arten (Coleoptera)." volume="27" year="1908">Reitter 1908</bibRefCitation>
, labeled: &quot;6. [handwritten] \ Amani [printed] \ D. O. Afrika
<normalizedToken originalValue="Eichelbaum’">Eichelbaum'</normalizedToken>
03 [printed] \ 13 April 1903 in
<taxonomicName class="Agaricomycetes" family="Polyporaceae" genus="Fomes" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Fomes nigrolachatus" order="Polyporales" pageId="9" pageNumber="32" phylum="Basidiomycota" rank="species" species="nigrolachatus">Fomes nigrolachatus</taxonomicName>
[handwritten] \ Amani. Deutsch Ostafr. [handwritten] \
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus eichelbaumi" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="eichelbaumi">Xylographus eichelbaumi</taxonomicName>
m. Typ. 1907. [handwritten] \ Eichelbaumi Usambara Reitt. [handwritten] \ [red label] HOLOTYPE
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus eichelbaumi" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="eichelbaumi">Xylographus eichelbaumi</taxonomicName>
Reitt. [handwritten]&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="32" type="remarks">
<paragraph pageId="9" pageNumber="32">Remarks.</paragraph>
<paragraph pageId="9" pageNumber="32">
<bibRefCitation author="Scott, H" journalOrPublisher="Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905. (Ser. 2. Zoology)" pageId="16" pageNumber="39" pagination="1 - 41" title="Coleoptera, Ciidae." volume="Vol. 8" year="1926">Scott (1926)</bibRefCitation>
stated he has not examined the type of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="madagascariensis">Xylographus madagascariensis</taxonomicName>
and that he described
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus seychellensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="seychellensis">Xylographus seychellensis</taxonomicName>
with some hesitation. If he had examined the known male type of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="madagascariensis">Xylographus madagascariensis</taxonomicName>
, he would have observed that it was just slightly more elongate than the specimens he had at hand, with no other differences. Such a small difference in body elongation is expected to occur in
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="genus">Xylographus</taxonomicName>
species with broad geopraphical distribution (see, for instance, the known variation in
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus globipennis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="globipennis">Xylographus globipennis</taxonomicName>
;
<bibRefCitation author="Sandoval-Gomez, VE" journalOrPublisher="Entomological Science" pageId="16" pageNumber="39" pagination="326 - 332" title="Xylographus globipennis Reitter, 1911 (Coleoptera: Ciidae): a barely studied species with broad distribution in the Afrotropical region." url="10.1111/j.1479-8298.2011.00458.x" volume="14" year="2011">
<normalizedToken originalValue="Sandoval-Gómez">Sandoval-Gomez</normalizedToken>
et al. 2011
</bibRefCitation>
). In order to make sure they were all conspecifics, we dissected named male
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus seychellensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="seychellensis">Xylographus seychellensis</taxonomicName>
compared to the type and also the lectotype of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="madagascariensis">Xylographus madagascariensis</taxonomicName>
, and we observed the sclerites of abdominal terminalia to be exactly the same. The lectotype of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus tarsalis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="tarsalis">Xylographus tarsalis</taxonomicName>
is a male
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="madagascariensis">Xylographus madagascariensis</taxonomicName>
with weak secondary sexual characteristics.
<bibRefCitation author="Ferrer, J" journalOrPublisher="Entomofauna, Zeitschrift fuer Entomologie" pageId="15" pageNumber="38" pagination="405 - 416" title="Contribution a la connaissance des Ciidae. Designation des Lectotypes des especes descrites par Fahraeus (1871) et Reitter (1908) preservees aux Musees de Stockholm et de Wien." volume="18" year="1997">Ferrer (1997)</bibRefCitation>
stated that two female paralectotypes of
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus tarsalis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="tarsalis">Xylographus tarsalis</taxonomicName>
were deposited in NHRS. After studying the material, we have seen that they are a male and a female instead. The names
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus eichelbaumi" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="eichelbaumi">Xylographus eichelbaumi</taxonomicName>
and
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus rufipennis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="rufipennis">Xylographus rufipennis</taxonomicName>
were based on females, which clearly correspond to females named
<taxonomicName class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylographus madagascariensis" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="madagascariensis">Xylographus madagascariensis</taxonomicName>
that we examined.
</paragraph>
</subSubSection>
</treatment>
</document>
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