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<mods:title id="48192640AC9AA2D409F8ABD85179499F">New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)</mods:title>
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<mods:namePart id="0C0BD048B6D82C4A36962CFD01B94AE1">Perkins, Philip D.</mods:namePart>
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<taxonomicName id="E2AEADD2FFD5FFC4FF6DFD9CB3F19307" authority="Steffan" authorityName="Steffan" authorityYear="1973" box="[151,395,549,576]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFD5FFC4FF6DFD9CB3F19307" bold="true" box="[151,395,549,576]" pageId="7" pageNumber="64">
<emphasis id="17DA0A43FFD5FFC4FF6DFD9CB3509378" bold="true" box="[151,298,549,576]" italics="true" pageId="7" pageNumber="64">Incoltorrida</emphasis>
Steffan
</emphasis>
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<subSubSection id="6DB485DAFFD5FFC4FF6DFDD0B6359246" pageId="7" pageNumber="64" type="reference_group">
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<taxonomicName id="E2AEADD2FFD5FFC4FF6DFDD0B3A293B9" authority="Steffan, 1973: 634" authorityName="Steffan" authorityPageNumber="634" authorityYear="1973" box="[151,472,616,641]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFD5FFC4FF6DFDD0B36993B9" box="[151,275,617,641]" italics="true" pageId="7" pageNumber="64">Incoltorrida</emphasis>
<treatmentCitation id="A40FF040FFD5FFC4FEE3FDD1B3A293B9" author="Steffan, A. W." box="[281,472,616,641]" page="634" pageId="7" pageNumber="64" year="1973">
<bibRefCitation id="413FABA0FFD5FFC4FEE3FDD1B3A293B9" author="Steffan, A. W." box="[281,472,616,641]" pageId="7" pageNumber="64" pagination="633 - 646" refId="ref13951" refString="Steffan, A. W. (1973) Torridincoidarum tres novae species e regione aethiopica (Coleoptera: Sphaeroidea). Revue de Zoologie et de Botanique Africaines, 87, 633 - 646." type="journal article" year="1973">Steffan, 1973: 634</bibRefCitation>
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.
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species
<taxonomicName id="E2AEADD2FFD5FFC4FD90FDD0B18593B9" authority="Steffan, 1973" authorityName="Steffan" authorityYear="1973" box="[618,1023,616,641]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="madagassica">
<emphasis id="17DA0A43FFD5FFC4FD90FDD0B10A93B9" box="[618,880,617,641]" italics="true" pageId="7" pageNumber="64">Incoltorrida madagassica</emphasis>
<bibRefCitation id="413FABA0FFD5FFC4FC8CFDD1B18593B9" author="Steffan, A. W." box="[886,1023,616,641]" pageId="7" pageNumber="64" pagination="633 - 646" refId="ref13951" refString="Steffan, A. W. (1973) Torridincoidarum tres novae species e regione aethiopica (Coleoptera: Sphaeroidea). Revue de Zoologie et de Botanique Africaines, 87, 633 - 646." type="journal article" year="1973">Steffan, 1973</bibRefCitation>
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(by original designation)
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<paragraph id="2511D651FFD5FFC4FF6DFD16B6359246" blockId="7.[151,1437,687,1074]" pageId="7" pageNumber="64">
The following comments are given by
<bibRefCitation id="413FABA0FFD5FFC4FDABFD16B08E93F2" author="Steffan, A. W." box="[593,756,687,714]" pageId="7" pageNumber="64" pagination="633 - 646" refId="ref13951" refString="Steffan, A. W. (1973) Torridincoidarum tres novae species e regione aethiopica (Coleoptera: Sphaeroidea). Revue de Zoologie et de Botanique Africaines, 87, 633 - 646." type="journal article" year="1973">Steffan (1973)</bibRefCitation>
as the differential diagnosis of
<taxonomicName id="E2AEADD2FFD5FFC4FBA6FD09B69F93F2" authorityName="Steffan" authorityYear="1973" box="[1116,1253,688,714]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFD5FFC4FBA6FD09B69F93F2" box="[1116,1253,688,714]" italics="true" pageId="7" pageNumber="64">Incoltorrida</emphasis>
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: “The members of
<taxonomicName id="E2AEADD2FFD5FFC4FF4CFD6DB34593D6" authority="Perkins &amp; Bergsten, 2019" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[182,319,724,750]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="17DA0A43FFD5FFC4FF4CFD6DB34593D6" box="[182,319,724,750]" italics="true" pageId="7" pageNumber="64">Incoltorrida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CE9B738FFD5FFC4FEBCFD6DB3E893D6" box="[326,402,724,750]" pageId="7" pageNumber="64" rank="genus">n. gen.</taxonomicNameLabel>
differ from the members of
<taxonomicName id="E2AEADD2FFD5FFC4FD28FD6DB18293D5" authority="Steffan 1964" authorityName="Steffan" authorityYear="1964" box="[722,1016,723,750]" class="Insecta" family="Torridincolidae" genus="Torridincola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFD5FFC4FD28FD6DB12593D6" box="[722,863,724,750]" italics="true" pageId="7" pageNumber="64">Torridincola</emphasis>
<bibRefCitation id="413FABA0FFD5FFC4FC9CFD6AB18293D5" author="Steffan, A. W." box="[870,1016,723,749]" pageId="7" pageNumber="64" pagination="193 - 200" refId="ref13922" refString="Steffan, A. W. (1964) Torridincolidae, coleopterorum nova familia e regione aethiopica. Entomologische Zeitschrift, Stuttgart, 74, 193 - 200." type="journal article" year="1964">Steffan 1964</bibRefCitation>
</taxonomicName>
, through the following characters: on average more robust body size and more pronounced sculpturing of the elytra; the proximally narrowed keel on frons and vertex; the length proportions of the tarsomeres (mesotarsi T4 approximately 1.5 times longer than T3, compared to twice as long in
<taxonomicName id="E2AEADD2FFD5FFC4FE1BFCF9B0149262" authorityName="Steffan" authorityYear="1964" box="[481,622,832,858]" class="Insecta" family="Torridincolidae" genus="Torridincola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFD5FFC4FE1BFCF9B0149262" box="[481,622,832,858]" italics="true" pageId="7" pageNumber="64">Torridincola</emphasis>
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); The absence of parameres and the prescence of a thorned saccus membranaceus in the male, and the lateral thorns on the styli of the female genital organ.”
</paragraph>
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<subSubSection id="6DB485DAFFD5FFC4FF3DFC3EB1EF9509" pageId="7" pageNumber="64" type="description">
<paragraph id="2511D651FFD5FFC4FF3DFC3EB1EF9509" blockId="7.[151,1437,687,1074]" pageId="7" pageNumber="64">
All of these diagnostic characters hold true for the new species described herein, except the presence in all species of a single seta on each side near the base of the male genitalia (e.g.,
<figureCitation id="BD95CAD4FFD5FFC4FC34FC12B62C92FE" box="[974,1110,939,966]" captionStart="FIGURES 1920" captionStartId="17.[151,264,1761,1786]" captionTargetBox="[235,1353,486,1744]" captionTargetId="figure@17.[235,1353,486,1744]" captionTargetPageId="17" captionText="FIGURES 1920. (19) Incoltorrida madagassica, male genitalia of non-type; (20) Incoltorrida benesculpta, male genitalia of holotype." httpUri="https://zenodo.org/record/3369829/files/figure.png" pageId="7" pageNumber="64">Figs. 19, 20</figureCitation>
); these setae confirm that the basal lobes they are attached to are the homologues of the parameres present in members of
<taxonomicName id="E2AEADD2FFD5FFC4FB5EFC69B74B92D2" authorityName="Steffan" authorityYear="1964" box="[1188,1329,976,1002]" class="Insecta" family="Torridincolidae" genus="Torridincola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFD5FFC4FB5EFC69B74B92D2" box="[1188,1329,976,1002]" italics="true" pageId="7" pageNumber="64">Torridincola</emphasis>
</taxonomicName>
. Because of distortion, these setae are not visible on the permanent microslide mount of the
<typeStatus id="FA1568F3FFD5FFC4FBC3FC4AB6E19535" box="[1081,1179,1011,1037]" pageId="7" pageNumber="64" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E2AEADD2FFD5FFC4FB3BFC4DB7119536" authorityName="Steffan" authorityYear="1973" box="[1217,1387,1012,1038]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="madagassica">
<emphasis id="17DA0A43FFD5FFC4FB3BFC4DB7119536" box="[1217,1387,1012,1038]" italics="true" pageId="7" pageNumber="64">I. madagassica</emphasis>
</taxonomicName>
, the only male available to Steffan at the time of the original description.
</paragraph>
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</treatment>
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