180 lines
21 KiB
XML
180 lines
21 KiB
XML
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<mods:title id="BF8D27A25E492B124E3C635439A4C9B1">A review of Elocomosta Hansen with a description of a new species with reduced eyes from China (Coleoptera, Hydrophilidae, Sphaeridiinae)</mods:title>
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<mods:namePart id="182E91798466F08564DA924F301AC287">Lin, Renchao</mods:namePart>
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<mods:namePart id="032812A747E01AD8585C0E1791ADBA32">Jia, Fenglong</mods:namePart>
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<mods:namePart id="EBAD5278347E9A952D939962AAB1A786">Fikacek, Martin</mods:namePart>
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<mods:date id="2B517C8BC28A9BEFDB7C3373B40B1808">2016</mods:date>
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<treatment id="5E60B8315AA8D611868086880D61D69F" ID-GBIF-Taxon="127854870" LSID="urn:lsid:plazi:treatment:5E60B8315AA8D611868086880D61D69F" httpUri="http://treatment.plazi.org/id/5E60B8315AA8D611868086880D61D69F" lastPageId="9" lastPageNumber="90" pageId="5" pageNumber="86">
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<subSubSection id="B22B1F419D87C697272CE397FDD19E51" pageId="5" pageNumber="86" type="multiple">
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<paragraph id="558305D7CE5FD3BADD3A86340E087088" pageId="5" pageNumber="86">Taxon classification Animalia Coleoptera Hydrophilidae</paragraph>
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</subSubSection>
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<subSubSection id="4BCDF5E7E5747214A0183C5E491B59CF" pageId="5" pageNumber="86" type="nomenclature">
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<paragraph id="C21B829FF18A017DC2111926F3963ABA" pageId="5" pageNumber="86">
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<taxonomicName id="C92257795A52153E1D1EC8F30377584A" ID-CoL="39C7J" authority="Hansen, 1989" authorityName="Hansen" authorityYear="1989" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra Hansen, 1989</taxonomicName>
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Figs 1-3, 8, 9, 11, 13, 15-16, 18, 21-22, 24, 26, 28, 32
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</paragraph>
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</subSubSection>
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<subSubSection id="8F27866B22BD35E65DDFBB2176552E6F" pageId="6" pageNumber="87" type="reference_group">
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<paragraph id="4FAAD10F4E9F8BF2218D427B174BC98D" pageId="6" pageNumber="87">
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<taxonomicName id="02432C8C499C4545842CE99CC2EA36B4" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="nigra">
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<pageBreakToken id="783A9863079BC5BCCB55461F27864FC8" pageId="6" pageNumber="87" start="start">Elocomosta</pageBreakToken>
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nigra
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</taxonomicName>
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Hansen, 1989: 255.
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</paragraph>
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</subSubSection>
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<subSubSection id="3454363418B5C0208A1B28FA195D2438" pageId="6" pageNumber="87" type="type material">
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<paragraph id="67F9142440D9990B9350C78F2178258A" pageId="6" pageNumber="87">Type material examined.</paragraph>
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<paragraph id="2BB8818F01A285DF5ABF4CE90A13BF8C" pageId="6" pageNumber="87">
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Paratype. 1 female (NMPC): SARAWAK / Semengoh For. / Reserve. 11 mi. / SW Kuching / 1-4.vi.1968 / R. W. Taylor // PARATYPE /
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<taxonomicName id="C423AC79AA9C2044691ADFC74E0715FE" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="genus">Elocomosta</taxonomicName>
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/
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<taxonomicName id="69673F74AF8FEFE6BC480D8F18C83CD4" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="nigra">nigra</taxonomicName>
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/ M. Hansen.
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</paragraph>
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</subSubSection>
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<subSubSection id="340A11EEEB8DE39388BAF6C9A793485A" pageId="7" pageNumber="88" type="materials_examined">
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<paragraph id="562E2C48479772754FBE1A26496C263B" pageId="7" pageNumber="88">
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<pageBreakToken id="A8978E4B5906FAB5CA1DA2DFF665B9EC" pageId="7" pageNumber="88" start="start">Additional</pageBreakToken>
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material examined.
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</paragraph>
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<paragraph id="FEB2CF33679F210DE2EA5A69A7229229" pageId="7" pageNumber="88">1 female (NMPC): BORNEO: Sarawak / Kuching, Santubong / 26.3.1990 / leg. A. RIEDEL.</paragraph>
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</subSubSection>
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<subSubSection id="ED5CAECE20A3D5EE63AA1D553944DFA4" lastPageId="9" lastPageNumber="90" pageId="7" pageNumber="88" type="redescription">
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<paragraph id="A16E78B8D1EAA23EAC15900D2CD66960" pageId="7" pageNumber="88">Redescription.</paragraph>
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<paragraph id="ACEEF8E8E1E4D59010EDC95DD71E8F0D" lastPageId="9" lastPageNumber="90" pageId="7" pageNumber="88">
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Habitus. Body widely oval in dorsal view, moderately convex in lateral view. Length 2.5-2.9 mm, width 1.7-1.9 mm. Coloration. Dorsum of head,
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<pageBreakToken id="BBDCDA708576FEEAF30AEBBF2BD2E274" pageId="8" pageNumber="89" start="start">pronotum</pageBreakToken>
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and elytron uniformly black, margins only very indistinctly and narrowly paler. Maxillary palpi and antennae yellow-brown. Venter black to dark brown, tarsi and mouthparts reddish to reddish yellow. Head. Clypeus with sparse and very fine punctures, interstices smooth; anterior and lateral marginal portions of clypeus without microsculpture. Frons with punctation slightly coarser than on clypeus. Eyes well developed, strongly constricted laterally by anterior and posterior projections of frons; dorsal portion large, dorsally eyes divided by distance of 4.4
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<normalizedToken id="AFE4BA3F73059E0A36A4074A36E3AB0D" originalValue="×">x</normalizedToken>
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the dorsal width of one eye, ventral portion small. Labrum nearly completely concealed by clypeus. Surface of mentum without microsculpture. Thorax. Pronotum with very fine and sparse punctation, punctures slightly smaller than on frons; surface between punctures smooth, without
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<pageBreakToken id="61AD898EF4C85908DF80E6677DB020D4" pageId="9" pageNumber="90" start="start">microsculpture</pageBreakToken>
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; posterolateral corners rectangular, not projecting posteriad. Elytra not explanate laterally, interval punctation consisting of punctures coarser than on pronotum; epipleura moderately wide. Mesoventral plate circular, ca. as long as wide, with distinct but narrow marginal bead. Metaventrite short, metaventral process wide at base, narrowing anteriad. Metanepisternum with anterolateral tooth. Profemora with hydrofuge dense pubescence in basal 0.8. Male genitalia. Parameres ca. 1.4
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<normalizedToken id="A4D7ED918F1F5FC4AEA1410AFC08EA1B" originalValue="×">x</normalizedToken>
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as long as phallobase, narrowly parallel-sided subapically, pointed at apex. Phallobase wide and long, posteriorly without manubrium. Median lobe a little longer than parameres, but with basal portion deeply inserted into phallobase, nearly parallel-sided below gonopore, apically abruptly narrowed into pointed apex; gonopore situated subapically.
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</paragraph>
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</subSubSection>
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<subSubSection id="7061F1FE9FEB132849E8D717D77D9558" pageId="9" pageNumber="90" type="biology_ecology">
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<paragraph id="8A43B28924699F55B1D3C5B3CE632D7C" pageId="9" pageNumber="90">Biology.</paragraph>
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<paragraph id="DF4D9EE404BD6F3A4460B2A2EEBBC88B" pageId="9" pageNumber="90">
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Terrestrial species, part of type series was collected from rainforest leaf litter (
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<bibRefCitation id="4729A8C862A3C8784AED30E424318B0E" author="Hansen, M" journalOrPublisher="Entomologica scandinavica" pageId="11" pageNumber="92" pagination="251 - 262" title="New genera of Sphaeridiinae (Coleoptera: Hydrophilidae)." url="10.1163/187631289X00320" volume="20" year="1989">Hansen 1989</bibRefCitation>
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). No biology data are available on locality labels in the specimens examined by us.
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</paragraph>
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</subSubSection>
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<subSubSection id="28E089533D56997DF7FC04FCBE543F4D" pageId="9" pageNumber="90" type="distribution">
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<paragraph id="E126C3B0F4B077E5632B8191F2860BCC" pageId="9" pageNumber="90">Distribution.</paragraph>
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<paragraph id="984BD9FC122E315AC55E9ADDFC5108CF" pageId="9" pageNumber="90">Endemic to Borneo (Sarawak State, Malaysia).</paragraph>
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<caption id="B8D3CD7398D297F84604E9D83D227DE2" pageId="9" pageNumber="90">
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<paragraph id="77AF42E35CF4733AEA84182DFC62D065" pageId="9" pageNumber="90">
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Figures 9-17. External morphology of
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<taxonomicName id="1F1CE45D24EEBE6578CC6DF0A8FC6C1A" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">Elocomosta</taxonomicName>
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species. 9-10 ventral morphology, general view (9
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<taxonomicName id="96C36D281E30B1A9EA740EB9A6211F8C" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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Hansen, 1989 10
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<taxonomicName id="6B69A44DFC8D32DCC82C420A1295A324" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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sp. n.) 11-12 head, ventral view (11
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<taxonomicName id="48A47F48BE22839F25454DFC77DC1A68" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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12
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<taxonomicName id="464419E597AF5DA5358983FEFC6368F7" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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) 13-14 detail of the ocular region of the head, dorsal view (13
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<taxonomicName id="ECB6E318F28F8C363E3E10CF51B92345" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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14
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<taxonomicName id="88E57CECD6F7D8BC39F64454B823BB74" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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) 15-16 head appendages of
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<taxonomicName id="0311812685750237DBE12E9755EAB18C" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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(15 antenna 16 maxillary palpus and galea) 17 anterior margin of head of
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<taxonomicName id="53B43D760AB702996DA404ED0D7E6A16" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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in dorsal view, showing largely exposed labrum.
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</paragraph>
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</caption>
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<caption id="F163BA62D88E24FB42B5023A3D518E22" pageId="9" pageNumber="90">
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<paragraph id="FBE7FA8CF37F0BA96199867D0D305C78" pageId="9" pageNumber="90">
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Figures 18-25. External morphology of
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<taxonomicName id="1C1EB2730BE827E0A05C8CFD770FB89A" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">Elocomosta</taxonomicName>
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species. 18-19 profemur (18
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<taxonomicName id="67B3A11977A48AD3C4388E0DBA2BB74C" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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Hansen, 1989 19
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<taxonomicName id="C6A223300BC654A9EFB33B8E49928BEE" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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sp. n.). 20-21 punctation of pronotum and elytral base (20
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<taxonomicName id="93D8ADC1A3EAE7DBA7D0D8AA48B7461F" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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21
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<taxonomicName id="E6C03EA89C555196E3FE376FA04C11CD" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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) 22-23 meso- and metaventral morphology (22
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<taxonomicName id="83B5EC486152D0777D6B2B537B0A9FD7" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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23
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<taxonomicName id="10AB1AD7F73F1B6EFDA98060510BA72C" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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) 24-25 details of mesoventral plate and metaventral process (24
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<taxonomicName id="48D8723EEDDE4F84C8CBA38202F8855B" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
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25
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<taxonomicName id="4A6BA7AF89EAC77F1E126EF82FEAA17D" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
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).
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</paragraph>
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</caption>
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<caption id="593E6DCE2FEB606FAA13CDEE31227488" pageId="9" pageNumber="90">
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<paragraph id="57A8AE3F45DFD38F0C062698B3550991" pageId="9" pageNumber="90">
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Figures 26-33. External morphology of
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<taxonomicName id="BC1CB65291CE6F06832561B9469C2E27" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">Elocomosta</taxonomicName>
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species. 26-27 lateral portion of pronotum (26
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<taxonomicName id="00AAAD125BA0C5F8579A9DB444778889" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
|
||
Hansen, 1989 27
|
||
<taxonomicName id="81C99CED5274EFB020EB72F74A222EAA" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
|
||
sp. n.). 28-29 details of elytral punctation and trichobothria (28
|
||
<taxonomicName id="3D93A481D547BE458B99970136359C91" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
|
||
29
|
||
<taxonomicName id="549EFBB79F20878BE2A191DF77478F0F" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
|
||
) 30 prosternum of
|
||
<taxonomicName id="4AFB5CAA34B49AECA545FF64DB108C3B" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
|
||
31-32 posterior leg (31
|
||
<taxonomicName id="4DB13E37BA469290B081E5A1D1A068EA" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
|
||
32
|
||
<taxonomicName id="F33AAB3570C8E779BFF818847C550982" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta nigra" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="nigra">Elocomosta nigra</taxonomicName>
|
||
) 33 abdomen of
|
||
<taxonomicName id="9203C956A53A0B77C1701D9CB391013F" class="Insecta" family="Hydrophilidae" genus="Elocomosta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elocomosta lilizheni" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="lilizheni">Elocomosta lilizheni</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |