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<document id="0A13A3DCF03139777125881AF0CD1374" ID-DOI="10.11646/zootaxa.5418.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10726419" ID-ZooBank="928EFF22-3EE9-4C29-92E0-4490C3FF0490" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709201492302" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2024" docId="AB0F87F379321918FF6EF886FB11FE48" docLanguage="en" docName="zootaxa.5418.3.5.pdf" docOrigin="Zootaxa 5418 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cirrothaumatia tornocarpa" docType="treatment" docVersion="2" lastPageNumber="274" masterDocId="5736FF8B7937191EFFF9FFD0FFD6FF99" masterDocTitle="Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)" masterLastPageNumber="278" masterPageNumber="268" pageNumber="273" updateTime="1709201945160" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B8293E2E26DC1F51033F145C25AFA550" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9F9DA897035B61178D00A6BFA8D302BD">Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)</mods:title>
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<mods:namePart id="26B86C7F9C56AF8F812284D00D891F67">Heppner, John B.</mods:namePart>
<mods:affiliation id="C93782177B4A77731D9F9631D8DD95B6">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA.</mods:affiliation>
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<mods:namePart id="51D61A9D1267193D0A62804F5FB45705">Bae, Yang-Seop</mods:namePart>
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<mods:affiliation id="8758F20D9F51E156F3851F79EB280382">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="72F6264DB07FBC6AF4F0507CDF6FB0E4" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<treatment id="AB0F87F379321918FF6EF886FB11FE48" ID-DOI="http://doi.org/10.5281/zenodo.10726443" ID-Zenodo-Dep="10726443" LSID="urn:lsid:plazi:treatment:AB0F87F379321918FF6EF886FB11FE48" httpUri="http://treatment.plazi.org/id/AB0F87F379321918FF6EF886FB11FE48" lastPageId="6" lastPageNumber="274" pageId="5" pageNumber="273">
<subSubSection id="6BBC656E7932191BFF6EF886FD79F8E9" box="[151,687,1878,1905]" pageId="5" pageNumber="273" type="nomenclature">
<paragraph id="231936E57932191BFF6EF886FD79F8E9" blockId="5.[151,687,1878,1941]" box="[151,687,1878,1905]" pageId="5" pageNumber="273">
<heading id="785181897932191BFF6EF886FD79F8E9" bold="true" box="[151,687,1878,1905]" fontSize="11" level="1" pageId="5" pageNumber="273" reason="1">
<taxonomicName id="E4A64D667932191BFF6EF886FD79F8E9" ID-CoL="96CHX" authority="(Meyrick, 1932)" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[151,687,1878,1905]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="tornocarpa">
<emphasis id="11D2EAF77932191BFF6EF886FD79F8E9" bold="true" box="[151,687,1878,1905]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF6EF886FE34F8E8" bold="true" box="[151,482,1878,1905]" italics="true" pageId="5" pageNumber="273">Cirrothaumatia tornocarpa</emphasis>
(Meyrick, 1932)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF6EF8ABFCDAF844" pageId="5" pageNumber="273" type="description">
<paragraph id="231936E57932191BFF6EF8ABFE9CF80C" blockId="5.[151,687,1878,1941]" box="[151,330,1915,1941]" pageId="5" pageNumber="273">
(
<figureCitation id="BB9D2A607932191BFF66F8ABFF38F80C" box="[159,238,1915,1941]" captionStart="FIGURES 7–9" captionStartId="6.[151,264,1828,1852]" captionTargetBox="[156,1426,490,1796]" captionTargetId="figure-133@6.[151,1436,489,1805]" captionTargetPageId="6" captionText="FIGURES 7–9. Genital features of related species of Cirrothaumatia: 7) C. tornosema (Clarke), female holotype (Guatemala, USNM) and male (a, shown without left valva; valval scale tuft shown intact on left valval base: Costa Rica, VOB); 8) C. tornocarpa (Meyrick), female holotype (Panama, NHMV); 9) C. vesta (Clarke), female paratype and male holotype (a, with aedeagus to right) (Venezuela, USNM) (Fig. 7, 9 and 9a, after Clarke (1968); Fig. 7a, after Razowski (1964); Fig. 8, after Razowski and Becker (1986))." figureDoi="http://doi.org/10.5281/zenodo.10726429" httpUri="https://zenodo.org/record/10726429/files/figure.png" pageId="5" pageNumber="273">Figs. 8</figureCitation>
,
<figureCitation id="BB9D2A607932191BFF02F8ABFE97F80C" box="[251,321,1915,1941]" captionStart-0="FIGURES 10–17" captionStart-1="FIGURE 18" captionStartId-0="7.[151,264,1718,1742]" captionStartId-1="8.[151,250,1873,1897]" captionTargetBox-0="[189,1399,181,1694]" captionTargetBox-1="[218,1370,665,1850]" captionTargetId-0="figure-28@7.[189,1399,181,1694]" captionTargetId-1="figure-209@8.[218,1370,665,1850]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." captionText-1="FIGURE 18. Distribution map of known localities for Cirrothaumatia species (NGS base map): C. huemboana n. sp., yellow square (Peru:Amazonas). C. moyobamba n. sp., black-bordered white square (Peru: San Martín). C. pichita n. sp., white square (Peru: Junín). C. tornocarpa (Meyrick), red square (Panama: Chiriquí). C. tornosema (Clarke), black squares (Mexico:Veracruz; Guatemala: Quetzaltenango; Costa Rica: Cartago, Heredia, and Puntarenas; and Ecuador: Pichincha). C. vesta (Clarke), blue square (Venezuela: Aragua)." figureDoi-0="http://doi.org/10.5281/zenodo.10726433" figureDoi-1="http://doi.org/10.5281/zenodo.10726435" httpUri-0="https://zenodo.org/record/10726433/files/figure.png" httpUri-1="https://zenodo.org/record/10726435/files/figure.png" pageId="5" pageNumber="273">17–18</figureCitation>
)
</paragraph>
<paragraph id="231936E57932191BFF6EF812FCDAF844" blockId="5.[151,780,1986,2013]" box="[151,780,1986,2013]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF6EF812FEEAF845" bold="true" box="[151,316,1986,2012]" pageId="5" pageNumber="273">Type-locality:</emphasis>
Lino, [Boquete Dist.,
<collectingRegion id="E162F8077932191BFDCEF813FD4CF844" box="[567,666,1987,2013]" country="Panama" name="Chiriqui" pageId="5" pageNumber="273">Chiriquí</collectingRegion>
],
<collectingCountry id="5BB176757932191BFD52F813FCD1F844" box="[683,775,1987,2013]" name="Panama" pageId="5" pageNumber="273">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E79311918FF3EFF46FD3CFF60" pageId="6" pageNumber="274" type="diagnosis">
<paragraph id="231936E579311918FF3EFF46FD3CFF60" blockId="6.[151,1437,150,465]" pageId="6" pageNumber="274">
<emphasis id="11D2EAF779311918FF3EFF46FEECFF29" bold="true" box="[199,314,150,176]" pageId="6" pageNumber="274">Diagnosis</emphasis>
. Wing expanse
<quantity id="E45E9B0079311918FE17FF47FDE8FF28" box="[494,574,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="274" unit="mm" value="12.0">12 mm</quantity>
female (n = 1). The female genitalia of this species are distinct, with a small semicircular signum on the bursa wall (
<figureCitation id="BB9D2A6079311918FDB4FF6BFD46FF4C" box="[589,656,187,213]" captionStart="FIGURES 7–9" captionStartId="6.[151,264,1828,1852]" captionTargetBox="[156,1426,490,1796]" captionTargetId="figure-133@6.[151,1436,489,1805]" captionTargetPageId="6" captionText="FIGURES 7–9. Genital features of related species of Cirrothaumatia: 7) C. tornosema (Clarke), female holotype (Guatemala, USNM) and male (a, shown without left valva; valval scale tuft shown intact on left valval base: Costa Rica, VOB); 8) C. tornocarpa (Meyrick), female holotype (Panama, NHMV); 9) C. vesta (Clarke), female paratype and male holotype (a, with aedeagus to right) (Venezuela, USNM) (Fig. 7, 9 and 9a, after Clarke (1968); Fig. 7a, after Razowski (1964); Fig. 8, after Razowski and Becker (1986))." figureDoi="http://doi.org/10.5281/zenodo.10726429" httpUri="https://zenodo.org/record/10726429/files/figure.png" pageId="6" pageNumber="274">Fig. 8</figureCitation>
), and the absence of the lateral carina-like cestum of most of the other species in the genus; it also lacks an accessory bursa.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E79311918FF3EFED3FED1FED8" pageId="6" pageNumber="274" type="materials_examined">
<paragraph id="231936E579311918FF3EFED3FED1FED8" blockId="6.[151,1437,150,465]" pageId="6" pageNumber="274">
<materialsCitation id="93CE3CB879311918FF3EFED3FED1FED8" collectionCode="NHMV" country="Panama" elevation="800" location="Female" municipality="Lino" pageId="6" pageNumber="274" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<emphasis id="11D2EAF779311918FF3EFED3FEE2FE84" bold="true" box="[199,308,259,285]" pageId="6" pageNumber="274">
<typeStatus id="FC1D884779311918FF3EFED3FEE2FE84" box="[199,308,259,285]" pageId="6" pageNumber="274" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="2679603E79311918FEBDFED3FE4FFE84" LSID="urn:lsid:plazi:treatment:AB0F87F379321918FF6EF886FB11FE48:2679603E79311918FEBDFED3FE4FFE84" box="[324,409,259,285]" country="Panama" municipality="Lino" name="Female" pageId="6" pageNumber="274" stateProvince="Chiriqui">Female</location>
,
<collectingMunicipality id="C37DAC9F79311918FE5EFED3FE0AFE84" box="[423,476,259,285]" pageId="6" pageNumber="274">Lino</collectingMunicipality>
(
<quantity id="E45E9B0079311918FE14FED3FDEDFE84" box="[493,571,259,285]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="274" unit="m" value="800.0">
<elevation id="A88BD1D679311918FE14FED3FDEDFE84" box="[493,571,259,285]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="274" unit="m" value="800.0">800 m</elevation>
</quantity>
), [Boquete Dist.,
<collectingRegion id="E162F80779311918FCFEFED3FCBCFE84" box="[775,874,259,285]" country="Panama" name="Chiriqui" pageId="6" pageNumber="274">Chiriquí</collectingRegion>
],
<collectingCountry id="5BB1767579311918FC87FED3FC0CFE84" box="[894,986,259,285]" name="Panama" pageId="6" pageNumber="274">Panama</collectingCountry>
, [1930?] (gen. slide J. Razowski-4230) (
<collectionCode id="45B7AE2079311918FF66FEF7FF2DFED8" box="[159,251,295,321]" pageId="6" pageNumber="274">NHMV</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E79311918FF3EFE9AFE70FEFC" box="[199,422,330,357]" pageId="6" pageNumber="274" type="biology_ecology">
<paragraph id="231936E579311918FF3EFE9AFE70FEFC" blockId="6.[151,1437,150,465]" box="[199,422,330,357]" pageId="6" pageNumber="274">
<emphasis id="11D2EAF779311918FF3EFE9AFEF7FEFD" bold="true" box="[199,289,330,356]" pageId="6" pageNumber="274">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E79311918FF3EFEBEFCBAFE10" box="[199,876,366,393]" pageId="6" pageNumber="274" type="distribution">
<paragraph id="231936E579311918FF3EFEBEFCBAFE10" blockId="6.[151,1437,150,465]" box="[199,876,366,393]" pageId="6" pageNumber="274">
<emphasis id="11D2EAF779311918FF3EFEBEFE8FFE11" bold="true" box="[199,345,366,392]" pageId="6" pageNumber="274">Distribution</emphasis>
. Known only from western
<collectingCountry id="5BB1767579311918FD6CFEBFFD38FE10" box="[661,750,367,393]" name="Panama" pageId="6" pageNumber="274">Panama</collectingCountry>
(
<collectingRegion id="E162F80779311918FD04FEBFFCB7FE10" box="[765,865,367,393]" country="Panama" name="Chiriqui" pageId="6" pageNumber="274">Chiriquí</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E79311918FF3EFE42FB11FE48" pageId="6" pageNumber="274" type="discussion">
<paragraph id="231936E579311918FF3EFE42FB11FE48" blockId="6.[151,1437,150,465]" pageId="6" pageNumber="274">
<emphasis id="11D2EAF779311918FF3EFE42FE93FE35" bold="true" box="[199,325,402,428]" pageId="6" pageNumber="274">Discussion</emphasis>
. While the wing maculation is similar to other species of the genus, the female genitalia do not conform, hence, the male is necessary to verify whether this species belongs in
<taxonomicName id="E4A64D6679311918FBE8FE67FB16FE48" authorityName="Razowski &amp; Becker" authorityYear="1986" box="[1041,1216,439,465]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="11D2EAF779311918FBE8FE67FB16FE48" box="[1041,1216,439,465]" italics="true" pageId="6" pageNumber="274">Cirrothaumatia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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