treatments-xml/data/76/3A/87/763A87EEFFCAFF95FEBDFD04FBD179DD.xml

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<mods:title id="1526CEEB9CBF46EA786861A34E0E9F6A">Revision of the New World Heteromeringia (Diptera: Clusiidae: Clusiodinae)</mods:title>
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<treatment id="763A87EEFFCAFF95FEBDFD04FBD179DD" ID-DOI="http://doi.org/10.5281/zenodo.4793466" ID-GBIF-Taxon="182395263" ID-Zenodo-Dep="4793466" LSID="urn:lsid:plazi:treatment:763A87EEFFCAFF95FEBDFD04FBD179DD" httpUri="http://treatment.plazi.org/id/763A87EEFFCAFF95FEBDFD04FBD179DD" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
<subSubSection id="B6896573FFCAFF94FEBDFD04FDBD7FCA" box="[265,598,729,753]" pageId="2" pageNumber="39" type="nomenclature">
<paragraph id="FE2C36F8FFCAFF94FEBDFD04FDBD7FCA" blockId="2.[265,598,729,753]" box="[265,598,729,753]" pageId="2" pageNumber="39">
<heading id="A5648194FFCAFF94FEBDFD04FDBD7FCA" box="[265,598,729,753]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="39" reason="2">
<emphasis id="CCE7EAEAFFCAFF94FEBDFD04FDBD7FCA" bold="true" box="[265,598,729,753]" pageId="2" pageNumber="39">
<taxonomicName id="39934D7BFFCAFF94FEBDFD04FDFB7FC9" ID-CoL="4WJ9" ID-ENA="574050" authority="CZERNY" authorityName="CZERNY" authorityYear="1903" box="[265,528,729,753]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="CCE7EAEAFFCAFF94FEBDFD04FE517FD4" bold="true" box="[265,442,729,753]" italics="true" pageId="2" pageNumber="39">Heteromeringia</emphasis>
CZERNY
</taxonomicName>
,, 1903
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B6896573FFCAFF94FF09FCD5FDE67E72" pageId="2" pageNumber="39" type="reference_group">
<paragraph id="FE2C36F8FFCAFF94FF09FCD5FDE67E72" blockId="2.[189,1179,776,855]" pageId="2" pageNumber="39">
<emphasis id="CCE7EAEAFFCAFF94FF09FCD5FEAC7E38" box="[189,327,776,797]" italics="true" pageId="2" pageNumber="39">HeteromeringiaHeteromeringia</emphasis>
CZERNY, 1903: 72. 
<bibRefCitation id="9A024B09FFCAFF94FE48FCD5FD5C7E38" author="KERTESZ, K." box="[508,695,776,797]" pageId="2" pageNumber="39" pagination="566 - 573" refId="ref19322" refString="KERTESZ, K. 1903: Beitrage zur Kenntniss der Heteroneuriden. - Annales Musei Nationalis Hungarici 1: 566 - 573." type="journal article" year="1903">KERTESZ, 1903: 567</bibRefCitation>
. 
<bibRefCitation id="9A024B09FFCAFF94FD0BFCD5FC9F7E3E" author="JOHNSON, C. W." box="[703,884,776,797]" pageId="2" pageNumber="39" pagination="97 - 101" refId="ref19288" refString="JOHNSON, C. W. 1913: A study of the Clusiodidae, (Heteroneuridae) of the Eastern United States. - Psyche 20 (3): 97 - 101." type="journal article" year="1913">JOHNSON, 1913: 98</bibRefCitation>
. 
<bibRefCitation id="9A024B09FFCAFF94FCCFFCD5FBC07E3E" author="MALLOCH, J. R." box="[891,1067,776,795]" pageId="2" pageNumber="39" pagination="2 - 8" refId="ref19482" refString="MALLOCH, J. R. 1918: A revision of the dipterous family Clusiodidae (Heteroneuridae). - Proceedings of the Entomological Society of Washington 20 (1): 2 - 8." type="journal article" year="1918">MALLOCH, 1918: 7</bibRefCitation>
. MELANDER &amp; 
<bibRefCitation id="9A024B09FFCAFF94FF63FCF9FE9B7E12" author="ARGO, N. G." box="[215,368,804,823]" pageId="2" pageNumber="39" pagination="1 - 54" refId="ref19616" refString="MELANDER ,, A. L. &amp; ARGO, N. G. 1924: Revision of the two-winged flies of the family Clusiidae. - Proceedings of the United States Natural History Museum 64: 1 - 54." type="journal article" year="1924">ARGO, 1924: 28</bibRefCitation>
. 
<bibRefCitation id="9A024B09FFCAFF94FECEFCF9FDFC7E12" author="MCAC LPINE, D. K." box="[378,535,804,823]" pageId="2" pageNumber="39" pagination="63 - 94" refId="ref19547" refString="MCAC LPINE, D. K. 1960: A review of the Australian species of Clusiidae (Diptera: Acalyptrata). - Records of the Australian Museum 25: 63 - 94." type="journal article" year="1960">MCAC LPINE 1960</bibRefCitation>
, 83-85. 
<bibRefCitation id="9A024B09FFCAFF94FDD2FCF9FCBB7E1F" author="SASAKAWA, M." box="[614,848,804,826]" pageId="2" pageNumber="39" pagination="61 - 100" refId="ref19696" refString="SASAKAWA, M. 1966: Studies on the Oriental and Pacific Clusiidae (Diptera) pt. 1. Genus Heteromeringia CZERNY, with one new related genus. - Pacific Insects 8 (1): 61 - 100." type="journal article" year="1966">SASAKAWA, 1966: 61-100</bibRefCitation>
. 
<typeStatus id="2128885AFFCAFF94FCEDFCF9FC6D7E1C" box="[857,902,804,825]" pageId="2" pageNumber="39">Type</typeStatus>
species: 
<taxonomicName id="39934D7BFFCAFF94FC6CFCF8FE9D7E72" ID-CoL="6M4VT" authority="(LOEW, 1864)" authorityName="" baseAuthorityName="LOEW" baseAuthorityYear="1864" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="nigrimana">
<emphasis id="CCE7EAEAFFCAFF94FC6CFCF8FF047E77" italics="true" pageId="2" pageNumber="39">Heteromeringia nigrimana</emphasis>
(LOEW, 1864)
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="B6896573FFCAFF94FF0AFCB1FB9C7961" pageId="2" pageNumber="39" type="diagnosis">
<paragraph id="FE2C36F8FFCAFF94FF0AFCB1FECE7EA1" blockId="2.[190,293,876,900]" box="[190,293,876,900]" pageId="2" pageNumber="39">
<emphasis id="CCE7EAEAFFCAFF94FF0AFCB1FECE7EA1" bold="true" box="[190,293,876,900]" pageId="2" pageNumber="39">Diagnosis</emphasis>
</paragraph>
<paragraph id="FE2C36F8FFCAFF94FF0AFC4EFB9C7961" blockId="2.[190,1180,915,1092]" pageId="2" pageNumber="39">
Outer angular extension on pedicel blunt and obtuse. Scutellum flattened and wrinkled dorsally. Subcostal break indistinct. Three or four fronto-orbital bristles, reclinate except for inclinate anterior bristle. Interfrontal bristle absent. One small lateral scutellar bristle (
<figureCitation id="66A82A7DFFCAFF94FC66FC0DFBF97ECD" box="[978,1042,976,1000]" captionStart="Fig" captionStartId="31.[107,138,636,658]" captionTargetBox="[311,909,180,616]" captionTargetId="figure-126@31.[307,912,174,622]" captionTargetPageId="31" captionText="Fig. 8: Heteromeringia czernyi KERTESZ, left lateral with detail of anepisternum and arrow indicating diagnostic upturned bristle. Fig. 9: H. fumipennisfumipennis MELANDER &amp; ARGO, anepisternum with arrow indicating diagnostic lateral disc." figureDoi="http://doi.org/10.5281/zenodo.4751972" httpUri="https://zenodo.org/record/4751972/files/figure.png" pageId="2" pageNumber="39">Fig. 8</figureCitation>
). Acrostichal bristle absent. Tibiae without dorsal preapical bristles. Distiphallus elongate, thin, with one pair of heavily-sclerotized lateral “ribs” and usually with accessory distal sclerites (
<figureCitation id="66A82A7DFFCAFF94FC7DFBD3FBFF7903" box="[969,1044,1038,1062]" captionStart="Figs 10-12" captionStartId="31.[107,144,1550,1572]" captionTargetBox="[316,908,811,1525]" captionTargetId="figure-123@31.[307,912,805,1538]" captionTargetPageId="31" captionText="Figs 10-12: Heteromeringia fucatafucata HENDEL, genitalia. 10 external, left lateral; 11 external, posterior; 12 internal, left lateral. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751979" httpUri="https://zenodo.org/record/4751979/files/figure.png" pageId="2" pageNumber="39">Fig. 12</figureCitation>
). Ejaculatory apodeme elongate with apex fan-shaped (shown from different angles in Figs 24, 36 and 53).
</paragraph>
</subSubSection>
<subSubSection id="B6896573FFCAFF95FF0AFBBCFBD179DD" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39" type="description">
<paragraph id="FE2C36F8FFCAFF94FF0AFBBCFE60795C" blockId="2.[190,395,1121,1145]" box="[190,395,1121,1145]" pageId="2" pageNumber="39">
<emphasis id="CCE7EAEAFFCAFF94FF0AFBBCFE60795C" bold="true" box="[190,395,1121,1145]" pageId="2" pageNumber="39">Generic description</emphasis>
</paragraph>
<paragraph id="FE2C36F8FFCAFF94FF0AFB55FBB47853" blockId="2.[189,1180,1160,1398]" pageId="2" pageNumber="39">
<emphasis id="CCE7EAEAFFCAFF94FF0AFB55FEFE79BF" bold="true" box="[190,277,1160,1178]" pageId="2" pageNumber="39">General.</emphasis>
Body length 2.4-5.0 mm. Colour predominantly brown to black, sometimes yellow with variable dark pattern. Face flat and microsetulose. Arista sparsely plumose to pubescent. Outer angular extension on pedicel blunt and obtuse; inner extension absent. Antenna arising at midpoint of head. Frons almost always with medial pilose patch on anterior half or more (often reduced in females). Postcranium concave. Scutellum flattened dorsally and longitudinally wrinkled. Subcostal break indistinct. Wing clear to variably patterned. M 
<subScript id="621734BDFFCAFF94FC0EFAEFFC3B7819" attach="left" box="[954,976,1330,1340]" fontSize="6" pageId="2" pageNumber="39">1+2</subScript>
ratio (length of the ultimate section of vein M 
<subScript id="621734BDFFCAFF94FE7BFA8CFE38787E" attach="left" box="[463,467,1361,1371]" fontSize="6" pageId="2" pageNumber="39">1</subScript>
divided by length of penultimate section) 4.0-12.0 (usually 4.5-8.0). Fore tarsi usually with slight lateral compression (pronounced in some Old World species).
</paragraph>
<paragraph id="FE2C36F8FFCAFF94FF0AFA49FEDF7889" blockId="2.[190,308,1428,1452]" box="[190,308,1428,1452]" pageId="2" pageNumber="39">
<emphasis id="CCE7EAEAFFCAFF94FF0AFA49FEDF7889" bold="true" box="[190,308,1428,1452]" pageId="2" pageNumber="39">Chaetotaxy</emphasis>
</paragraph>
<paragraph id="FE2C36F8FFCAFF95FF0AFA67FD657C5F" blockId="2.[190,1180,1466,1643]" lastBlockId="3.[107,1096,175,378]" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
Bristles black to brown. Three or four fronto-orbital bristles, reclinate except for inclinate anterior pair; anterior two bristles closely spaced near anterior margin of frons and hind pair positioned near posterior margin; fourth bristle sometimes present between anterior two pairs but less than 2/3 length of other bristles. Postvertical and ocellar bristles divergent, minute to large. Interfrontal bristle absent. Vibrissa well-developed. Usually five to seven short genal bristles (anterior bristle sometimes vibrissa-like). Two subgenal bristles. Pedicel with one outstanding dorsal bristle. Occiput with row of short bristles behind eye margin. Two postsutural dorsocentral bristles, with anterior bristle usually 3/5-4/5 times length of posterior bristle; sometimes small additional bristle directly in front of anterior dorsocentral. Scutellum with one small lateral bristle and one long incurved apical bristle (
<figureCitation id="66A82A7DFFCBFF95FED6FED6FE487C06" box="[354,419,267,291]" captionStart="Fig" captionStartId="31.[107,138,636,658]" captionTargetBox="[311,909,180,616]" captionTargetId="figure-126@31.[307,912,174,622]" captionTargetPageId="31" captionText="Fig. 8: Heteromeringia czernyi KERTESZ, left lateral with detail of anepisternum and arrow indicating diagnostic upturned bristle. Fig. 9: H. fumipennisfumipennis MELANDER &amp; ARGO, anepisternum with arrow indicating diagnostic lateral disc." figureDoi="http://doi.org/10.5281/zenodo.4751972" httpUri="https://zenodo.org/record/4751972/files/figure.png" pageId="3" pageNumber="40">Fig. 8</figureCitation>
). Acrostichal bristle absent. One postpronotal, two notopleural, two postsutural intra-alar and sometimes one small to minute intra post-alar bristle. 1-2 (
<figureCitation id="66A82A7DFFCBFF95FBB7FEF7FBD47C67" box="[1027,1087,298,322]" captionStart="Fig" captionStartId="31.[107,138,636,658]" captionTargetBox="[311,909,180,616]" captionTargetId="figure-126@31.[307,912,174,622]" captionTargetPageId="31" captionText="Fig. 8: Heteromeringia czernyi KERTESZ, left lateral with detail of anepisternum and arrow indicating diagnostic upturned bristle. Fig. 9: H. fumipennisfumipennis MELANDER &amp; ARGO, anepisternum with arrow indicating diagnostic lateral disc." figureDoi="http://doi.org/10.5281/zenodo.4751972" httpUri="https://zenodo.org/record/4751972/files/figure.png" pageId="3" pageNumber="40">Fig. 8</figureCitation>
) strong anepisternal and 1 katepisternal bristle. Tibiae without dorsal preapical bristles. Male fore and mid femora with ventral row of ctenidial bristles.
</paragraph>
<paragraph id="FE2C36F8FFCBFF95FFDEFE40FE437C8A" blockId="3.[106,424,413,431]" box="[106,424,413,431]" pageId="3" pageNumber="40">
<emphasis id="CCE7EAEAFFCBFF95FFDEFE40FE437C8A" bold="true" box="[106,424,413,431]" pageId="3" pageNumber="40">Male abdomen and terminalia</emphasis>
</paragraph>
<paragraph id="FE2C36F8FFCBFF95FFDEFE1EFBCA7EC1" blockId="3.[106,1097,450,996]" pageId="3" pageNumber="40">
Tergite 1 narrow and fused to tergite 2; tergites 3-6 and sternites 2-5 well-developed, separate; sternite 1 reduced to thin strip; sternites 2-5 well-developed. Sternites 6- 8 forming complete asymmetrical ring (= “annulus”); sternite 8 setose and dorsal with right lateral extension; sternite 7 on left side between sternites 6 and 8, heavily sclerotized anteriorly; sternite 6 fused to sternite 7 laterally, heavily sclerotized anteriorly, and weakly attached to sternite 8 ventrally; sternites 6 and 7 variably differentiated ventrally into weakly sclerotized, folded and/or widened regions, forming enlarged membranous “pocket” to enclose distiphallus when it is at rest. Spiracles in membrane ventral to tergites 1-5, (often) dorsolaterally between sternites 6 and 7, and ventrally between sternites 6 and 8. Epandrium dome-shaped, evenly bristled (
<figureCitation id="66A82A7DFFCBFF95FC9CFD65FC2A7FF5" box="[808,961,696,720]" captionStart="Figs 10-12" captionStartId="31.[107,144,1550,1572]" captionTargetBox="[316,908,811,1525]" captionTargetId="figure-123@31.[307,912,805,1538]" captionTargetPageId="31" captionText="Figs 10-12: Heteromeringia fucatafucata HENDEL, genitalia. 10 external, left lateral; 11 external, posterior; 12 internal, left lateral. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751979" httpUri="https://zenodo.org/record/4751979/files/figure.png" pageId="3" pageNumber="40">Figs 10 and 11</figureCitation>
), and usually much narrower than abdomen. Surstylus variable in shape (
<figureCitation id="66A82A7DFFCBFF95FD6FFD0AFCC07FCA" box="[731,811,727,751]" captionStart="Figs 10-12" captionStartId="31.[107,144,1550,1572]" captionTargetBox="[316,908,811,1525]" captionTargetId="figure-123@31.[307,912,805,1538]" captionTargetPageId="31" captionText="Figs 10-12: Heteromeringia fucatafucata HENDEL, genitalia. 10 external, left lateral; 11 external, posterior; 12 internal, left lateral. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751979" httpUri="https://zenodo.org/record/4751979/files/figure.png" pageId="3" pageNumber="40">Figs 10</figureCitation>
, 
<figureCitation id="66A82A7DFFCBFF95FC8CFD0AFCBE7FCA" box="[824,853,727,751]" captionStart="Figs 16-18" captionStartId="32.[324,361,1515,1537]" captionTargetBox="[338,1035,1046,1493]" captionTargetId="figure-185@32.[325,1043,1039,1504]" captionTargetPageId="32" captionText="Figs 16-18: Heteromeringia nanellananella sp. n., genitalia. 16 internal, left lateral; 17 external, left lateral; 18 external, posterior. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751990" httpUri="https://zenodo.org/record/4751990/files/figure.png" pageId="3" pageNumber="40">17</figureCitation>
, 
<figureCitation id="66A82A7DFFCBFF95FCD6FD0AFC967FCA" box="[866,893,727,751]" captionStart="Figs 31-33" captionStartId="34.[190,227,904,926]" captionTargetBox="[382,978,182,873]" captionTargetId="figure-101@34.[380,985,174,892]" captionTargetPageId="34" captionText="Figs 31-33: Heteromeringia czernyi KERTESZ, external terminalia. 31 left lateral; 32 posterior; 33 right surstylus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.4752008" httpUri="https://zenodo.org/record/4752008/files/figure.png" pageId="3" pageNumber="40">33</figureCitation>
and 
<figureCitation id="66A82A7DFFCBFF95FC00FD0BFC3A7FCB" box="[948,977,726,750]" captionStart="Figs 51-53" captionStartId="37.[107,144,1600,1622]" captionTargetBox="[310,897,940,1580]" captionTargetId="figure-104@37.[299,904,928,1587]" captionTargetPageId="37" captionText="Figs 51-53: Heteromeringia apholis sp. n., genitalia. 51 external, left lateral; 52 external, posterior; 53 internal, left lateral." figureDoi="http://doi.org/10.5281/zenodo.4752032" httpUri="https://zenodo.org/record/4752032/files/figure.png" pageId="3" pageNumber="40">51</figureCitation>
), but often less than 2/3 length of epandrium and rounded (very thin if longer); small rounded bristles apical on inner face. Cerci variable (
<figureCitation id="66A82A7DFFCBFF95FE71FCC9FDFF7E09" box="[453,532,788,812]" captionStart="Figs 10-12" captionStartId="31.[107,144,1550,1572]" captionTargetBox="[316,908,811,1525]" captionTargetId="figure-123@31.[307,912,805,1538]" captionTargetPageId="31" captionText="Figs 10-12: Heteromeringia fucatafucata HENDEL, genitalia. 10 external, left lateral; 11 external, posterior; 12 internal, left lateral. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751979" httpUri="https://zenodo.org/record/4751979/files/figure.png" pageId="3" pageNumber="40">Figs 11</figureCitation>
, 
<figureCitation id="66A82A7DFFCBFF95FD96FCC9FDD47E09" box="[546,575,788,812]" captionStart="Figs 13-15" captionStartId="32.[190,227,869,891]" captionTargetBox="[192,1172,186,854]" captionTargetPageId="32" captionText="Figs 13-15: Heteromeringia fumipennis MELANDER Figs 19-21: Heteromeringia nigripes MELANDERELANDER &amp; &amp; ARGO, genitalia. 13 external, left lateral; 14 exter- ARGO, genitalia. 19 external, left lateral; 20 external, posterior; 15 internal, left lateral. Ejaculatory nal, posterior; 21 internal, left lateral. Ejaculatory apodeme omitted. apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751986" httpUri="https://zenodo.org/record/4751986/files/figure.png" pageId="3" pageNumber="40">14</figureCitation>
, 
<figureCitation id="66A82A7DFFCBFF95FDF8FCC9FD827E09" box="[588,617,788,812]" captionStart="Figs 16-18" captionStartId="32.[324,361,1515,1537]" captionTargetBox="[338,1035,1046,1493]" captionTargetId="figure-185@32.[325,1043,1039,1504]" captionTargetPageId="32" captionText="Figs 16-18: Heteromeringia nanellananella sp. n., genitalia. 16 internal, left lateral; 17 external, left lateral; 18 external, posterior. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751990" httpUri="https://zenodo.org/record/4751990/files/figure.png" pageId="3" pageNumber="40">18</figureCitation>
, 
<figureCitation id="66A82A7DFFCBFF95FDC2FCC9FD7A7E09" box="[630,657,788,812]" captionStart="Figs 31-33" captionStartId="34.[190,227,904,926]" captionTargetBox="[382,978,182,873]" captionTargetId="figure-101@34.[380,985,174,892]" captionTargetPageId="34" captionText="Figs 31-33: Heteromeringia czernyi KERTESZ, external terminalia. 31 left lateral; 32 posterior; 33 right surstylus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.4752008" httpUri="https://zenodo.org/record/4752008/files/figure.png" pageId="3" pageNumber="40">32</figureCitation>
and 
<figureCitation id="66A82A7DFFCBFF95FD7CFCC9FD0E7E09" box="[712,741,788,812]" captionStart="Figs 45-47" captionStartId="36.[190,227,1378,1400]" captionTargetBox="[386,983,908,1346]" captionTargetId="figure-103@36.[382,987,897,1364]" captionTargetPageId="36" captionText="Figs 45-47: Heteromeringia quadriseta sp. n., genitalia. 45 internal, left lateral; 46 external, left lateral; 47 external, posterior. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4752026" httpUri="https://zenodo.org/record/4752026/files/figure.png" pageId="3" pageNumber="40">47</figureCitation>
), but usually less than half length of epandrium and often united. Hypandrium fused to pregonite (shape globose to triangular in profile) (
<figureCitation id="66A82A7DFFCBFF95FF77FC8CFEE57E4C" box="[195,270,849,873]" captionStart="Figs 10-12" captionStartId="31.[107,144,1550,1572]" captionTargetBox="[316,908,811,1525]" captionTargetId="figure-123@31.[307,912,805,1538]" captionTargetPageId="31" captionText="Figs 10-12: Heteromeringia fucatafucata HENDEL, genitalia. 10 external, left lateral; 11 external, posterior; 12 internal, left lateral. Ejaculatory apodeme omitted." figureDoi="http://doi.org/10.5281/zenodo.4751979" httpUri="https://zenodo.org/record/4751979/files/figure.png" pageId="3" pageNumber="40">Fig. 12</figureCitation>
), sparsely setose (bristles usually short and stout) and sometimes with distal setulae; hypandrial arm short and stout. Postgonite, epiphallus and lateral lobe of distiphallus absent. Phallapodeme 0.5-1.0 times length of hypandrium + pregonite. Basiphallus fused to distiphallus anteriorly. Distiphallus very elongate, dark, thin, with one pair of heavily-sclerotized lateral “ribs”, and often with distal accessory sclerites. Ejaculatory apodeme elongate with apex fan-shaped.
</paragraph>
<paragraph id="FE2C36F8FFCBFF95FFDFFBDFFE567931" blockId="3.[107,445,1026,1044]" box="[107,445,1026,1044]" pageId="3" pageNumber="40">
<emphasis id="CCE7EAEAFFCBFF95FFDFFBDFFE567931" bold="true" box="[107,445,1026,1044]" pageId="3" pageNumber="40">Female abdomen and terminalia</emphasis>
</paragraph>
<paragraph id="FE2C36F8FFCBFF95FFDEFBFAFBD179DD" blockId="3.[106,1096,1063,1272]" pageId="3" pageNumber="40">
Tergites 1 and 2 fused; tergites 3-6 and sternites 2-6 complete; sternite 1 reduced to thin anterior strip. Segments 7 and 8 narrowed into long, thin tube; terminal segments separated by long membranous area. Spiracles in membrane below tergites on segments 1 to 7. Cerci as long as sternite 10, length four times width. Spermatheca (one pair) pigmented (
<figureCitation id="66A82A7DFFCBFF95FD54FB5EFCA479B9" box="[736,847,1155,1180]" captionStart="Figs 28-30" captionStartId="33.[107,144,1539,1561]" captionTargetBox="[122,1080,1018,1517]" captionTargetId="figure-220@33.[107,1090,1011,1525]" captionTargetPageId="33" captionText="Figs 28-30: Female internal genitalia (spermathecae and ventral receptacle). 28 Heteromeringia fumipennis MELANDER &amp; ARGO; 29 H. nitida JOHNSON; 30 H. nigripesnigripes MELANDERELANDER &amp; ARGO." figureDoi="http://doi.org/10.5281/zenodo.4752002" httpUri="https://zenodo.org/record/4752002/files/figure.png" pageId="3" pageNumber="40">Figs 28-30</figureCitation>
and 
<figureCitation id="66A82A7DFFCBFF95FC31FB5EFC2D79BE" box="[901,966,1155,1179]" captionStart="Figs 48-50" captionStartId="37.[107,144,778,800]" captionTargetBox="[130,1083,195,747]" captionTargetId="figure-107@37.[111,1094,176,765]" captionTargetPageId="37" captionText="Figs 48-50: Female internal genitalia (spermathecae and ventral receptacle). 48 Heteromeringia czernyi KERTESZ; 49 H. flavifronsflavifrons HENNIG; 50 H. lateralis sp. n." figureDoi="http://doi.org/10.5281/zenodo.4752028" httpUri="https://zenodo.org/record/4752028/files/figure.png" pageId="3" pageNumber="40">48-50</figureCitation>
); shape variable, but usually small, transversely wrinkled, cylindrical and telescoped. Genital chamber membranous and elongate. Spermathecal duct weakly sclerotized. Ventral receptacle sac-like (widest subapically), recurved, and with long, sometimes loosely coiled, subterminal flagellum (
<figureCitation id="66A82A7DFFCBFF95FC56FB02FBC479DD" box="[994,1071,1247,1272]" captionStart="Figs 28-30" captionStartId="33.[107,144,1539,1561]" captionTargetBox="[122,1080,1018,1517]" captionTargetId="figure-220@33.[107,1090,1011,1525]" captionTargetPageId="33" captionText="Figs 28-30: Female internal genitalia (spermathecae and ventral receptacle). 28 Heteromeringia fumipennis MELANDER &amp; ARGO; 29 H. nitida JOHNSON; 30 H. nigripesnigripes MELANDERELANDER &amp; ARGO." figureDoi="http://doi.org/10.5281/zenodo.4752002" httpUri="https://zenodo.org/record/4752002/files/figure.png" pageId="3" pageNumber="40">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>