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<document id="511C72A0CD04CC4FED5EFEDEB0C2FFBB" ID-DOI="10.11646/zootaxa.4000.1.5" ID-GBIF-Dataset="795cfeae-aecb-48ad-98b4-41f924bbab14" ID-ISSN="1175-5326" ID-Zenodo-Dep="235591" ID-ZooBank="10B33B13-6E22-486A-9B85-C08934401B30" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461185097193" checkinUser="plazi" docAuthor="Shimizu, So &amp; Watanabe, Kyohei" docDate="2015" docId="6568CF31FFE9FF8D16AEF9DCFC38C412" docLanguage="en" docName="zt04000p122.pdf" docOrigin="Zootaxa 4000 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptophion septentrionis Shimizu &amp; Watanabe, sp. nov." docType="treatment" docUuid="4990FD75-A7B1-40F7-8487-94561A826D4D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="118" masterDocId="9951B749FFEFFF8A1639FFC2FF6AC362" masterDocTitle="Discovery of the genus Leptophion Cameron, 1901, from Japan and the Palaearctic region, with description of two new species (Hymenoptera: Ichneumonidae: Ophioninae)" masterLastPageNumber="122" masterPageNumber="111" pageNumber="117" updateTime="1698599408073" updateUser="plazi">
<mods:mods id="9595122FA7F5D3FE611E22992E91BC82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC0428CF11C35A8A1FD77EC40C0B1960">
<mods:title id="29DF579052B93D4C827DABB3C513FF1D">Discovery of the genus Leptophion Cameron, 1901, from Japan and the Palaearctic region, with description of two new species (Hymenoptera: Ichneumonidae: Ophioninae)</mods:title>
</mods:titleInfo>
<mods:name id="1375FCF7C29DD404B8C924060A4B8306" type="personal">
<mods:role id="74EF7A0E4533DA8D7CB6901A756D71F4">
<mods:roleTerm id="7C69571AB308E0546E021470C60AB41B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32ADDA22F2FF81996FA5F54DF8AFDAA9">Shimizu, So</mods:namePart>
</mods:name>
<mods:name id="332403AADE567AA5D8962816F3EB7887" type="personal">
<mods:role id="EEF3503DC61FB960686B3927D7AEFFD9">
<mods:roleTerm id="AA630FB29669EE03B78D432534F5BDFD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F129D442094BFD64A1E931A94D0382F">Watanabe, Kyohei</mods:namePart>
</mods:name>
<mods:typeOfResource id="D07945E40B7EB72583CDFACBBF219B12">text</mods:typeOfResource>
<mods:relatedItem id="6BD2346E459A21E77A28C11522AB27BD" type="host">
<mods:titleInfo id="FF694D8F9A0CB4C00A349652994C8917">
<mods:title id="F0DD7F92376B6E61A559F106847DC498">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5CEAAC718BB44F5C00E3BD55E77309BA">
<mods:date id="76A7FEC3D69CCCF832B25807B573FBB9">2015</mods:date>
<mods:detail id="8F6C2C77384881ADCCC7641C04453166" type="volume">
<mods:number id="6BC1E46BD00E823738E0282C33C58F24">4000</mods:number>
</mods:detail>
<mods:detail id="ED01665AB3E2EA73EC47C71E650CCF69" type="issue">
<mods:number id="A2A40EAF266FA9840B70D5F2CCAA064A">1</mods:number>
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<mods:start id="18F67F506455AE25B4ACDE588616C9BD">111</mods:start>
<mods:end id="C743FE91AE36445D8BE9338954579B6F">122</mods:end>
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<mods:classification id="4D36B2DBC3E84C45F1397E9BC35DC31D">journal article</mods:classification>
<mods:identifier id="542650631C9BC1343595DE3BEA244C99" type="DOI">10.11646/zootaxa.4000.1.5</mods:identifier>
<mods:identifier id="A7E97849D92222FF82630E8C01971A4B" type="GBIF-Dataset">795cfeae-aecb-48ad-98b4-41f924bbab14</mods:identifier>
<mods:identifier id="B018EA8BC11D0241AFC878B572942B14" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BD332A038BA12072EC1AEE9CC8E4E8E4" type="Zenodo-Dep">235591</mods:identifier>
<mods:identifier id="A9EAE03BA88CE97E4D2F1CBF99F4AEB3" type="ZooBank">10B33B13-6E22-486A-9B85-C08934401B30</mods:identifier>
</mods:mods>
<treatment id="6568CF31FFE9FF8D16AEF9DCFC38C412" ID-DOI="http://doi.org/10.5281/zenodo.5669756" ID-GBIF-Taxon="119628366" ID-Zenodo-Dep="5669756" LSID="urn:lsid:zoobank.org:act:4990FD75-A7B1-40F7-8487-94561A826D4D" httpUri="http://treatment.plazi.org/id/6568CF31FFE9FF8D16AEF9DCFC38C412" lastPageId="7" lastPageNumber="118" pageId="6" pageNumber="117">
<subSubSection id="A5DB2DACFFE9FF8C16AEF9DCFE93C51C" pageId="6" pageNumber="117" type="nomenclature">
<paragraph id="ED7E7E27FFE9FF8C16AEF9DCFC56C55B" blockId="6.[151,828,1566,1662]" box="[151,828,1566,1593]" pageId="6" pageNumber="117">
<heading id="B636C94BFFE9FF8C16AEF9DCFC56C55B" bold="true" box="[151,828,1566,1593]" fontSize="11" level="1" pageId="6" pageNumber="117" reason="1">
<emphasis id="DFB5A235FFE9FF8C16AEF9DCFC56C55B" bold="true" box="[151,828,1566,1593]" pageId="6" pageNumber="117">
<taxonomicName id="2AC105A4FFE9FF8C16AEF9DCFDBDC55B" authority="Shimizu &amp; Watanabe" authorityName="Shimizu &amp; Watanabe" box="[151,727,1566,1593]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="septentrionis" status="sp. nov.">
<emphasis id="DFB5A235FFE9FF8C16AEF9DCFEAEC55A" bold="true" box="[151,452,1566,1592]" italics="true" pageId="6" pageNumber="117">Leptophion septentrionis</emphasis>
Shimizu &amp; Watanabe
</taxonomicName>
<taxonomicNameLabel id="C4861F4EFFE9FF8C14E7F9DDFC56C55B" box="[734,828,1567,1593]" pageId="6" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="ED7E7E27FFE9FF8C16AEF980FD32C538" blockId="6.[151,828,1566,1662]" box="[151,600,1601,1626]" pageId="6" pageNumber="117">[Japanese name: Oo-usumon-amebachi]</paragraph>
<paragraph id="ED7E7E27FFE9FF8C16AEF9A7FE93C51C" blockId="6.[151,828,1566,1662]" box="[151,505,1637,1662]" pageId="6" pageNumber="117">
(
<figureCitation id="75FA62A2FFE9FF8C16A6F9A7FF8DC51C" box="[159,231,1637,1662]" captionStart="FIGURES 1, 2" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[151,1429,517,1883]" captionTargetId="figure@4.[151,1436,517,1898]" captionTargetPageId="4" captionText="FIGURES 1, 2. Habitus of Japanese species of Leptophion. 1. L. parvus sp. nov. (holotype). 2. L. septentrionis sp. nov. (holotype)." httpUri="https://zenodo.org/record/235592/files/figure.png" pageId="6" pageNumber="117">Figs 2</figureCitation>
,
<figureCitation id="75FA62A2FFE9FF8C16CAF9A7FEF7C51C" box="[243,413,1637,1662]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="6" pageNumber="117">6–8, 10, 12, 13</figureCitation>
,
<figureCitation id="75FA62A2FFE9FF8C1790F9A7FE9AC51C" box="[425,496,1637,1662]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="6" pageNumber="117">16, 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A5DB2DACFFE9FF8C16AEF96FFC42C46C" pageId="6" pageNumber="117" type="materials_examined">
<paragraph id="ED7E7E27FFE9FF8C16AEF96FFC42C46C" blockId="6.[151,1437,1709,2022]" pageId="6" pageNumber="117">
<emphasis id="DFB5A235FFE9FF8C16AEF96FFEF5C5A4" bold="true" box="[151,415,1709,1734]" pageId="6" pageNumber="117">
<typeStatus id="327AC085FFE9FF8C16AEF96FFFB8C5A4" box="[151,210,1709,1734]" pageId="6" pageNumber="117">Type</typeStatus>
series.
<typeStatus id="327AC085FFE9FF8C1713F96FFEF3C5A4" box="[298,409,1709,1734]" pageId="6" pageNumber="117" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female,
<collectingCountry id="95D63EB7FFE9FF8C143FF96CFD20C5A7" box="[518,586,1710,1733]" name="Japan" pageId="6" pageNumber="117">Japan</collectingCountry>
: Anbo-rindo,
<quantity id="2A39D3C2FFE9FF8C14DEF96FFC44C5A7" box="[743,814,1709,1734]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="117" unit="m" value="350.0">350 m</quantity>
alt., Yakushima Is., Kagoshima Pref.,
<date id="997F58E7FFE9FF8C12D8F96CFA1AC5A4" box="[1249,1392,1709,1734]" pageId="6" pageNumber="117" value="1982-07-22">22. vii. 1982</date>
(K. Konishi &amp; S. Yoshimatsu, light trap).
<emphasis id="DFB5A235FFE9FF8C1473F913FDDCC588" bold="true" box="[586,694,1745,1770]" pageId="6" pageNumber="117">
<typeStatus id="327AC085FFE9FF8C1473F913FDDCC588" box="[586,694,1745,1770]" pageId="6" pageNumber="117" type="paratype">Paratype</typeStatus>
</emphasis>
.
<specimenCount id="FBC7B5AEFFE9FF8C14FFF910FC46C588" box="[710,812,1745,1770]" pageId="6" pageNumber="117" type="female">1 female</specimenCount>
,
<collectingCountry id="95D63EB7FFE9FF8C1500F910FC17C58B" box="[825,893,1746,1769]" name="Japan" pageId="6" pageNumber="117">Japan</collectingCountry>
: Mt.Yuwandake, Amamioshima Is., Kagoshima Pref.,
<date id="997F58E7FFE9FF8C16E2F934FE3FC46C" box="[219,341,1782,1806]" pageId="6" pageNumber="117" value="1983-05-11">11. v. 1983</date>
(S. Yoshimatsu &amp; S. Nomura, light trap).
</paragraph>
</subSubSection>
<subSubSection id="A5DB2DACFFE9FF8D16FEF8DBFEEDC7A7" lastPageId="7" lastPageNumber="118" pageId="6" pageNumber="117" type="description">
<paragraph id="ED7E7E27FFE9FF8C16FEF8DBFBE2C450" blockId="6.[151,1437,1709,2022]" box="[199,1160,1817,1842]" pageId="6" pageNumber="117">
<emphasis id="DFB5A235FFE9FF8C16FEF8DBFE33C450" bold="true" box="[199,345,1817,1842]" pageId="6" pageNumber="117">Description.</emphasis>
Female (n=2). Body length 25.0–27.0 mm (
<typeStatus id="327AC085FFE9FF8C157EF8DBFCC5C450" box="[839,943,1817,1842]" pageId="6" pageNumber="117" type="holotype">holotype</typeStatus>
: 27.0 mm) (
<figureCitation id="75FA62A2FFE9FF8C120FF8DBFB11C450" box="[1078,1147,1817,1842]" captionStart="FIGURES 1, 2" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[151,1429,517,1883]" captionTargetId="figure@4.[151,1436,517,1898]" captionTargetPageId="4" captionText="FIGURES 1, 2. Habitus of Japanese species of Leptophion. 1. L. parvus sp. nov. (holotype). 2. L. septentrionis sp. nov. (holotype)." httpUri="https://zenodo.org/record/235592/files/figure.png" pageId="6" pageNumber="117">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="ED7E7E27FFE9FF8D16FEF8FFFD53C3B7" blockId="6.[151,1437,1709,2022]" lastBlockId="7.[151,1437,151,1904]" lastPageId="7" lastPageNumber="118" pageId="6" pageNumber="117">
Head. Face 0.7–0.8 (
<typeStatus id="327AC085FFE9FF8C17FBF8FFFD40C434" box="[450,554,1853,1878]" pageId="6" pageNumber="117" type="holotype">holotype</typeStatus>
: 0.8) × as long as wide, entirely covered with punctures and hairs, with longitudinal ridge on upper central area (
<figureCitation id="75FA62A2FFE9FF8C1454F8A3FDBFC418" box="[621,725,1889,1914]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="6" pageNumber="117">Figs 6–8</figureCitation>
). Clypeus 0.5 × as long as wide, evenly strongly convex with punctures, in lateral profile of lower margin almost straight (
<figureCitation id="75FA62A2FFE9FF8C1504F847FCC8C4FC" box="[829,930,1925,1950]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="6" pageNumber="117">Figs 6, 7</figureCitation>
). Malar space 0.3 × as long as basal width of mandible. Lateral ocelli adjacent with eye (
<figureCitation id="75FA62A2FFE9FF8C14A9F86BFD91C4A0" box="[656,763,1961,1986]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="6" pageNumber="117">Figs 7, 8</figureCitation>
). Occiput polished with fine white hairs. Temple strongly polished with sparse and short hairs in upper part, and with sparse and long hairs in lower part. Antenna with 67–71 (
<typeStatus id="327AC085FFE8FF8D16A6FF55FE6DC3D2" box="[159,263,151,176]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 71) flagellomeres. First flagellomere 2.1 × as long as second flagellomere, 20th flagellomere 1.9–2.0 (
<typeStatus id="327AC085FFE8FF8D16A6FF7EFE6DC3B7" box="[159,263,188,213]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 1.9) × as long as its width.
</paragraph>
<paragraph id="ED7E7E27FFE8FF8D16FEFF22FA13C2CF" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="118">
Mesosoma. Pronotum with widely wrinkled on posterior part and lower half part (
<figureCitation id="75FA62A2FFE8FF8D1258FF1DFBDEC39A" box="[1121,1204,223,248]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="7" pageNumber="118">Fig. 13</figureCitation>
). Anterior margin of mesoscutum in lateral profile with gently concave (
<figureCitation id="75FA62A2FFE8FF8D14EBFEC6FC4EC27E" box="[722,804,260,285]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="7" pageNumber="118">Fig. 13</figureCitation>
). Mesoscutum smooth, without notaulus, almost entirely covered with hairs. Scutellum with a lateral longitudinal carina developed on basal 0.8. Epicnemium with densely and obliquely wrinkled. Epicnemial carina present, its dorsal end not reaching to anterior margin of mesopleuron. Mesopleuron partly covered with longitudinal wrinkles and entirely covered with long hairs (
<figureCitation id="75FA62A2FFE8FF8D129FFEADFB92C2EA" box="[1190,1272,367,392]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="7" pageNumber="118">Fig. 13</figureCitation>
). Metapleuron polished with hairs. Submetapleural carina present. Propodeum in lateral profile rounded, with confused carina.
</paragraph>
<paragraph id="ED7E7E27FFE8FF8D16FEFE75FE0CC1CA" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="118">
Wings. Fore wing, 17.0–18.0 mm (
<typeStatus id="327AC085FFE8FF8D146AFE75FDD1C2B2" box="[595,699,439,464]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 18.0 mm), with AI, 1.0;
<collectingCountry id="95D63EB7FFE8FF8D15E5FE7AFC90C2AD" box="[988,1018,440,463]" name="Belgium" pageId="7" pageNumber="118">BI</collectingCountry>
, 3.1–3.4 (
<typeStatus id="327AC085FFE8FF8D1252FE75FBB9C2B2" box="[1131,1235,439,464]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 3.4); CI, 0.3;
<collectingCountry id="95D63EB7FFE8FF8D1341FE7AFAF2C2AD" box="[1400,1432,440,463]" name="Iceland" pageId="7" pageNumber="118">DI</collectingCountry>
, 0.5; MI, 2.6–2.8 (
<typeStatus id="327AC085FFE8FF8D1762FE1EFEA9C297" box="[347,451,476,501]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 2.6); ICI, 0.9; SDI, 0.9. Vein 1
<emphasis id="DFB5A235FFE8FF8D1518FE1FFC33C296" box="[801,857,477,500]" italics="true" pageId="7" pageNumber="118">m-cu</emphasis>
strongly sinuate. Vein
<emphasis id="DFB5A235FFE8FF8D1260FE1FFB1FC296" box="[1113,1141,477,500]" italics="true" pageId="7" pageNumber="118">Rs</emphasis>
+2
<emphasis id="DFB5A235FFE8FF8D12AAFE1FFBF4C296" box="[1171,1182,477,500]" italics="true" pageId="7" pageNumber="118">r</emphasis>
apparently geniculate. Discosubmarginal cell with a glabrous area below pterostigma (
<figureCitation id="75FA62A2FFE8FF8D154DFE3DFCA3C17A" box="[884,969,511,536]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="7" pageNumber="118">Fig. 16</figureCitation>
).
<collectingCountry id="95D63EB7FFE8FF8D15D9FE3DFB72C17A" box="[992,1048,511,536]" name="India" pageId="7" pageNumber="118">Hind</collectingCountry>
wing with NI, 1.1–1.3 (
<typeStatus id="327AC085FFE8FF8D1308FE3DFAF3C17A" box="[1329,1433,511,536]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 1.3). Vein
<emphasis id="DFB5A235FFE8FF8D1732FDE7FE4DC15E" box="[267,295,549,572]" italics="true" pageId="7" pageNumber="118">Rs</emphasis>
strongly curved. Marginal cell of hind wing entirely covered with hairs, its proximal part with an area with isolated dense hairs enclosed by glabrous area (
<figureCitation id="75FA62A2FFE8FF8D14C1FD85FC27C102" box="[760,845,583,608]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="7" pageNumber="118">Fig. 17</figureCitation>
). The angle of an intersection of two veins of hind wing,
<emphasis id="DFB5A235FFE8FF8D16E4FDAFFF84C1E6" box="[221,238,621,644]" italics="true" pageId="7" pageNumber="118">R</emphasis>
1 and
<emphasis id="DFB5A235FFE8FF8D170BFDAFFE24C1E6" box="[306,334,621,644]" italics="true" pageId="7" pageNumber="118">Rs</emphasis>
, about 95°. Vein
<emphasis id="DFB5A235FFE8FF8D1428FDAFFD48C1E6" box="[529,546,621,644]" italics="true" pageId="7" pageNumber="118">R</emphasis>
1 with 5–6 ununiformed hamuli (
<typeStatus id="327AC085FFE8FF8D15ACFDAEFC97C1E7" box="[917,1021,620,645]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 6) and penultimate one significantly elongate (
<figureCitation id="75FA62A2FFE8FF8D173CFD4DFE32C1CA" box="[261,344,655,680]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="7" pageNumber="118">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="ED7E7E27FFE8FF8D16FEFD77FC9AC192" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="118">
Legs. Distal pecten of hind tarsal claws longer than true apex of claw and other pectens of hind tarsal claw (
<figureCitation id="75FA62A2FFE8FF8D16A6FD15FF99C192" box="[159,243,727,752]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="7" pageNumber="118">Fig. 12</figureCitation>
). Apex of middle and hind tibial spurs strongly curved as hook-like.
</paragraph>
<paragraph id="ED7E7E27FFE8FF8D16FEFD3FFEFFC0C7" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="118">
Metasoma. PI, 2.5–2.6 (
<typeStatus id="327AC085FFE8FF8D17EAFD3EFD51C077" box="[467,571,764,789]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 2.5); DMI, 1.1–1.3 (
<typeStatus id="327AC085FFE8FF8D1511FD3EFCFAC077" box="[808,912,764,789]" pageId="7" pageNumber="118" type="holotype">holotype</typeStatus>
: 1.1). Anterior part of first tergite polished and smooth. Posterior part of first tergite and entire parts of second to eighth tergites polished, entirely covered with dense hairs. Second sternite to anterior part of third sternite, polished, with sparsely long hairs. Posterior part of third sternite to fifth sternite, polished, with fine microtrichia, long hairs along the ventral margin. Ovipositor sheath with long hairs.
</paragraph>
<paragraph id="ED7E7E27FFE8FF8D16FEFC6DFDCDC7C2" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="118">
Coloration (
<figureCitation id="75FA62A2FFE8FF8D1777FC6DFEF9C0AA" box="[334,403,943,968]" captionStart="FIGURES 1, 2" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[151,1429,517,1883]" captionTargetId="figure@4.[151,1436,517,1898]" captionTargetPageId="4" captionText="FIGURES 1, 2. Habitus of Japanese species of Leptophion. 1. L. parvus sp. nov. (holotype). 2. L. septentrionis sp. nov. (holotype)." httpUri="https://zenodo.org/record/235592/files/figure.png" pageId="7" pageNumber="118">Fig. 2</figureCitation>
). Yellowish brown except for: vittae of mesoscutum, posterior part of mesopleuron, anterior part of metapleuron, and third to eighth metasomal tergites. Posterior area of mesopleuron, upper half area of pleuron, and posterior end of propodeum with dark brown marks. Mesoscutum with three longitudinal vittae, both sides of vitta dark brown, middle vitta pale brown. Metasoma gradually darken toward its posterior end, seventh and eighth metasomal tergites entirely dark or grayish brown. Flagellum yellowish brown, basoventral part of flagellar segments on around center of the antenna, with blackish semicylindrical marks. Proximal part of marginal cell on fore wing with a clouded area (
<figureCitation id="75FA62A2FFE8FF8D147FFB45FDF3C7C2" box="[582,665,1159,1184]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="7" pageNumber="118">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="ED7E7E27FFE8FF8D16FEFB6EFEEDC7A7" blockId="7.[151,1437,151,1904]" box="[199,391,1196,1221]" pageId="7" pageNumber="118">
<emphasis id="DFB5A235FFE8FF8D16FEFB6EFE61C7A7" bold="true" box="[199,267,1196,1221]" pageId="7" pageNumber="118">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A5DB2DACFFE8FF8D16FEFB0DFB68C66E" pageId="7" pageNumber="118" type="distribution">
<paragraph id="ED7E7E27FFE8FF8D16FEFB0DFB75C78A" blockId="7.[151,1437,151,1904]" box="[199,1055,1231,1256]" pageId="7" pageNumber="118">
<emphasis id="DFB5A235FFE8FF8D16FEFB0DFE33C78A" bold="true" box="[199,345,1231,1256]" pageId="7" pageNumber="118">Distribution</emphasis>
.
<collectingCountry id="95D63EB7FFE8FF8D175EFB13FECDC78A" box="[359,423,1233,1256]" name="Japan" pageId="7" pageNumber="118">Japan</collectingCountry>
(Amamioshima Island and Yakushima Island) (
<figureCitation id="75FA62A2FFE8FF8D1586FB0DFB78C78A" box="[959,1042,1231,1256]" captionStart="FIGURE 18" captionStartId="8.[151,250,1730,1752]" captionText="FIGURE 18. Distribution of Leptophion in Japan (Ryukyus)." pageId="7" pageNumber="118">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="ED7E7E27FFE8FF8D16FEFB36FB68C66E" blockId="7.[151,1437,151,1904]" box="[199,1026,1268,1293]" pageId="7" pageNumber="118">
<emphasis id="DFB5A235FFE8FF8D16FEFB36FE29C66F" bold="true" box="[199,323,1268,1293]" pageId="7" pageNumber="118">Bionomics</emphasis>
. Host is unknown. All specimens were collected in light traps.
</paragraph>
</subSubSection>
<subSubSection id="A5DB2DACFFE8FF8D16FEFAD5FDCBC637" pageId="7" pageNumber="118" type="etymology">
<paragraph id="ED7E7E27FFE8FF8D16FEFAD5FDCBC637" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="118">
<emphasis id="DFB5A235FFE8FF8D16FEFAD5FE2CC652" bold="true" box="[199,326,1303,1328]" pageId="7" pageNumber="118">Etymology</emphasis>
. This species has the northernmost distribution within
<taxonomicName id="2AC105A4FFE8FF8D15F3FADAFB23C652" box="[970,1097,1304,1328]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="DFB5A235FFE8FF8D15F3FADAFB23C652" box="[970,1097,1304,1328]" italics="true" pageId="7" pageNumber="118">Leptophion</emphasis>
</taxonomicName>
, so that the specific name is from Latin “septentrio”, which means “north”.
</paragraph>
</subSubSection>
<subSubSection id="A5DB2DACFFE8FF8D16FEFA9DFC38C412" pageId="7" pageNumber="118" type="discussion">
<paragraph id="ED7E7E27FFE8FF8D16FEFA9DFC38C412" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="118">
<emphasis id="DFB5A235FFE8FF8D16FEFA9DFE51C61A" bold="true" box="[199,315,1375,1400]" pageId="7" pageNumber="118">Remarks.</emphasis>
In this genus, nine species have the clouded area on the proximal part of marginal cell of fore wing. Among them,
<taxonomicName id="2AC105A4FFE8FF8D170EFA47FE8CC6FE" box="[311,486,1413,1436]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="septentrionis" status="sp. nov.">
<emphasis id="DFB5A235FFE8FF8D170EFA47FE8CC6FE" box="[311,486,1413,1436]" italics="true" pageId="7" pageNumber="118">L. septentrionis</emphasis>
</taxonomicName>
<emphasis id="DFB5A235FFE8FF8D17D5FA46FD2CC6FF" bold="true" box="[492,582,1412,1437]" pageId="7" pageNumber="118">
<taxonomicNameLabel id="C4861F4EFFE8FF8D17D5FA46FD2CC6FF" box="[492,582,1412,1437]" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="2AC105A4FFE8FF8D152CFA47FCADC6FE" box="[789,967,1412,1436]" class="Insecta" family="Scarabaeidae" genus="Spilophion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="DFB5A235FFE8FF8D152CFA47FCADC6FE" box="[789,967,1412,1436]" italics="true" pageId="7" pageNumber="118">L. maculipennis</emphasis>
</taxonomicName>
(recorded from Australasian and Oriental regions) in the following combination of character states: (1) in coloration, entirely yellowish brown (exceptions: vittae of mesoscutum, posterior part of mesopleuron, anterior part of metapleuron, a part of metasoma and posterior end of propodeum with dark brown marks) and proximal part of marginal cell on fore wing with clouded area (
<figureCitation id="75FA62A2FFE8FF8D1356FA2DFFCDC54E" captionStart="FIGURES 1, 2" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[151,1429,517,1883]" captionTargetId="figure@4.[151,1436,517,1898]" captionTargetPageId="4" captionText="FIGURES 1, 2. Habitus of Japanese species of Leptophion. 1. L. parvus sp. nov. (holotype). 2. L. septentrionis sp. nov. (holotype)." httpUri="https://zenodo.org/record/235592/files/figure.png" pageId="7" pageNumber="118">Fig. 2</figureCitation>
); (2) vein 1
<emphasis id="DFB5A235FFE8FF8D1713F9D7FE08C54E" box="[298,354,1557,1580]" italics="true" pageId="7" pageNumber="118">m-cu</emphasis>
strongly sinuate (
<figureCitation id="75FA62A2FFE8FF8D1412F9D6FD15C54F" box="[555,639,1556,1581]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="7" pageNumber="118">Fig. 16</figureCitation>
); (3) the angle of an intersection of two veins of hind wing,
<emphasis id="DFB5A235FFE8FF8D131AF9D7FA5EC54E" box="[1315,1332,1557,1580]" italics="true" pageId="7" pageNumber="118">R</emphasis>
1 and
<emphasis id="DFB5A235FFE8FF8D1340F9D7FAFFC54E" box="[1401,1429,1557,1580]" italics="true" pageId="7" pageNumber="118">Rs</emphasis>
, around 90° (
<figureCitation id="75FA62A2FFE8FF8D171FF9F5FE11C532" box="[294,379,1591,1616]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="7" pageNumber="118">Fig. 17</figureCitation>
); and (4) proximal part on marginal cell of hind wing with an area with isolated dense hairs enclosed by glabrous area (
<figureCitation id="75FA62A2FFE8FF8D17EDF99EFD43C516" box="[468,553,1628,1653]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="7" pageNumber="118">Fig. 17</figureCitation>
), but
<taxonomicName id="2AC105A4FFE8FF8D1456F99FFC4BC516" box="[623,801,1629,1652]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="septentrionis" status="sp. nov.">
<emphasis id="DFB5A235FFE8FF8D1456F99FFC4BC516" box="[623,801,1629,1652]" italics="true" pageId="7" pageNumber="118">L. septentrionis</emphasis>
</taxonomicName>
<emphasis id="DFB5A235FFE8FF8D1513F99EFCEDC517" bold="true" box="[810,903,1628,1653]" pageId="7" pageNumber="118">
<taxonomicNameLabel id="C4861F4EFFE8FF8D1513F99EFCEDC517" box="[810,903,1628,1653]" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="2AC105A4FFE8FF8D121AF99FFBB3C516" box="[1059,1241,1628,1652]" class="Insecta" family="Scarabaeidae" genus="Spilophion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="DFB5A235FFE8FF8D121AF99FFBB3C516" box="[1059,1241,1628,1652]" italics="true" pageId="7" pageNumber="118">L. maculipennis</emphasis>
</taxonomicName>
in the following character states: (1) mesopleuron with longitudinal wrinkles (mesopleuron strongly polished and without wrinkles in
<taxonomicName id="2AC105A4FFE8FF8D168EF967FE07C5DE" box="[183,365,1700,1724]" class="Insecta" family="Scarabaeidae" genus="Spilophion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="DFB5A235FFE8FF8D168EF967FE07C5DE" box="[183,365,1700,1724]" italics="true" pageId="7" pageNumber="118">L. maculipennis</emphasis>
</taxonomicName>
) (
<figureCitation id="75FA62A2FFE8FF8D17B0F966FEB4C5DE" box="[393,478,1700,1725]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="7" pageNumber="118">Fig. 13</figureCitation>
) and (2) AI=1.0 (AI=
<quantity id="2A39D3C2FFE8FF8D14D8F966FC3EC5DF" box="[737,852,1700,1725]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.318" metricValueMin="3.302" pageId="7" pageNumber="118" unit="in" value="1.5" valueMax="1.7" valueMin="1.3">1.3–1.7 in</quantity>
<taxonomicName id="2AC105A4FFE8FF8D1567F967FB7EC5DE" box="[862,1044,1700,1724]" class="Insecta" family="Scarabaeidae" genus="Spilophion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="DFB5A235FFE8FF8D1567F967FB7EC5DE" box="[862,1044,1700,1724]" italics="true" pageId="7" pageNumber="118">L. maculipennis</emphasis>
</taxonomicName>
).
<taxonomicName id="2AC105A4FFE8FF8D1216F967FB8BC5DE" box="[1071,1249,1701,1724]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="septentrionis" status="sp. nov.">
<emphasis id="DFB5A235FFE8FF8D1216F967FB8BC5DE" box="[1071,1249,1701,1724]" italics="true" pageId="7" pageNumber="118">L. septentrionis</emphasis>
</taxonomicName>
<emphasis id="DFB5A235FFE8FF8D12D2F966FA2DC5DF" bold="true" box="[1259,1351,1700,1725]" pageId="7" pageNumber="118">
<taxonomicNameLabel id="C4861F4EFFE8FF8D12D2F966FA2DC5DF" box="[1259,1351,1700,1725]" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also similar to
<taxonomicName id="2AC105A4FFE8FF8D1730F90BFE18C582" box="[265,370,1737,1760]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="DFB5A235FFE8FF8D1730F90BFE18C582" box="[265,370,1737,1760]" italics="true" pageId="7" pageNumber="118">L. parvus</emphasis>
</taxonomicName>
<emphasis id="DFB5A235FFE8FF8D1743F905FEB9C582" bold="true" box="[378,467,1735,1760]" pageId="7" pageNumber="118">
<taxonomicNameLabel id="C4861F4EFFE8FF8D1743F905FEB9C582" box="[378,467,1735,1760]" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but it can be distinguished by the following combination of character states: (1) body large and the length of fore wing, 17.0–18.0 mm (body small and the length of fore wing,
<quantity id="2A39D3C2FFE8FF8D12BAF92EFB8DC466" box="[1155,1255,1772,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="118" unit="mm" value="13.5">13.5 mm</quantity>
in
<taxonomicName id="2AC105A4FFE8FF8D1333F92FFA19C466" box="[1290,1395,1773,1796]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="DFB5A235FFE8FF8D1333F92FFA19C466" box="[1290,1395,1773,1796]" italics="true" pageId="7" pageNumber="118">L. parvus</emphasis>
</taxonomicName>
<emphasis id="DFB5A235FFE8FF8D1343F92EFFA2C44A" bold="true" pageId="7" pageNumber="118">
<taxonomicNameLabel id="C4861F4EFFE8FF8D1343F92EFFA2C44A" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); (2) mesopleuron with longitudinal wrinkles (mesopleuron strongly polished and without wrinkles in
<taxonomicName id="2AC105A4FFE8FF8D13BCF8D3FF89C42E" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="DFB5A235FFE8FF8D13BCF8D3FF89C42E" italics="true" pageId="7" pageNumber="118">L. parvus</emphasis>
</taxonomicName>
<emphasis id="DFB5A235FFE8FF8D16D2F8F6FE2CC42F" bold="true" box="[235,326,1844,1869]" pageId="7" pageNumber="118">
<taxonomicNameLabel id="C4861F4EFFE8FF8D16D2F8F6FE2CC42F" box="[235,326,1844,1869]" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (
<figureCitation id="75FA62A2FFE8FF8D1766F8F6FED8C42E" box="[351,434,1844,1869]" captionStart="FIGURES 3 – 13" captionStartId="5.[151,264,1716,1739]" captionTargetBox="[157,1399,198,1667]" captionTargetId="figure@5.[151,1435,193,1695]" captionTargetPageId="5" captionText="FIGURES 3 – 13. Details of Japanese species of Leptophion. 3 – 5, 9, 11. L. parvus sp. nov. (holotype): 3. head, frontal view; 4. head, lateral view; 5. head, dorsal view; 9. hamuli of right hind wing, dorsal view; 11. pecten of right hind tarsal claw, lateral view. 6 – 8, 10, 12, 13. L. septentrionis sp. nov. (6 – 8, 10, 13. paratype; 12. holotype): 6. head, frontal view; 7. head, lateral view; 8. head, dorsal view; 10. hamuli of right hind wing, dorsolateral view; 12. pecten of right hind tarsal claw, lateral view; 13. mesosoma (surface sculptures on mesonotum, scutellum, and postscutellum omitted), lateral view." httpUri="https://zenodo.org/record/235593/files/figure.png" pageId="7" pageNumber="118">Fig. 13</figureCitation>
); and (3) pterostigma of fore wing with a clouded membranous area (pterostigma with a transpicuous membranous area in
<taxonomicName id="2AC105A4FFE8FF8D142CF89BFD15C412" box="[533,639,1881,1904]" class="Insecta" family="Ichneumonidae" genus="Leptophion" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="DFB5A235FFE8FF8D142CF89BFD15C412" box="[533,639,1881,1904]" italics="true" pageId="7" pageNumber="118">L. parvus</emphasis>
</taxonomicName>
<emphasis id="DFB5A235FFE8FF8D14BFF895FDB5C412" bold="true" box="[646,735,1879,1904]" pageId="7" pageNumber="118">
<taxonomicNameLabel id="C4861F4EFFE8FF8D14BFF895FDB5C412" box="[646,735,1879,1904]" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (
<figureCitation id="75FA62A2FFE8FF8D14C1F895FC2FC412" box="[760,837,1879,1904]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1011,1034]" captionTargetBox="[151,1436,195,990]" captionTargetId="figure@6.[151,1437,193,990]" captionTargetPageId="6" captionText="FIGURES 14 – 17. Wing cells of Japanese species of Leptophion. 14 – 15. L. parvus sp. nov. (holotype): 14. discosubmarginal cell of fore wing; 15. marginal cell of hind wing. 16 – 17. L. septentrionis sp. nov. (holotype): 16. discosubmarginal cell of fore wing; 17. marginal cell of hind wing." httpUri="https://zenodo.org/record/235594/files/figure.png" pageId="7" pageNumber="118">Fig.14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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