216 lines
29 KiB
XML
216 lines
29 KiB
XML
<document id="776F24E0A8E0287EF7D78BBDB0A426D0" ID-DOI="10.1515/aemnp-2017-0115" ID-ISSN="0374-1036" ID-Zenodo-Dep="5324616" ID-ZooBank="9578B20F-612B-4D7E-A6B3-6FFDBF7FE276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630149621436" checkinUser="marcus" docAuthor="Purchart, Luboš & Kamiński, Marcin Jan" docDate="2017" docId="685887ABFFBA8458FE3FEC975640287D" docLanguage="en" docName="ActaEntMusNatPra.2017.57.suppl.139-163.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57" docSource="http://dx.doi.org/10.1515/aemnp-2017-0115" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Clitobius omanicus Purchart & Kamiński 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="150" masterDocId="9461FFD3FFB08453FFD8EC4C55312B4E" masterDocTitle="A taxonomic review of the genus Clitobius with description of a new species from Oman (Coleoptera: Tenebrionidae)" masterLastPageNumber="163" masterPageNumber="139" pageNumber="149" updateTime="1699164527381" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1E152DD368F7D47B0F31A7EB2CF38B56">A taxonomic review of the genus Clitobius with description of a new species from Oman (Coleoptera: Tenebrionidae)</mods:title>
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<mods:name id="A0455749E0B46F8557DEA49BEBB54B0E" type="personal">
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<mods:namePart id="0E11E1A5C3AFC795C89EF70F0F6FA9D2">Purchart, Luboš</mods:namePart>
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<mods:affiliation id="EE2F891B383FBA97AD8BF1D96801A82C">) &) Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic; e-mails: lubos. purchart @ post. cz, lubos. purchart @ mendelu. cz</mods:affiliation>
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<mods:name id="7D271ACBCC74147599F67F435492114C" type="personal">
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<mods:roleTerm id="B0EB4576F613904464148327F5CA9A0B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A5F04689B17E0BC19EEB765848228955">Kamiński, Marcin Jan</mods:namePart>
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<mods:affiliation id="7A09A3594D6971C6FF5A89D3968FCBDB">) &) Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warsaw, Poland; e-mail: mkaminski @ miiz. waw. pl</mods:affiliation>
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<mods:typeOfResource id="BC9A52FE67B17C0D0B130845326BD6EF">text</mods:typeOfResource>
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<mods:title id="2792FDCE17EDC9260F31252FE1C664BA">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:date id="72A2482458F649D6FC1FDEEF289554C8">2017</mods:date>
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<mods:title id="C1DAE805C6AA7E6EBD0CB9F83E0FBD57">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="69BD4FE2B5BC06ADA24DC358E84B19EF">2017-12-31</mods:number>
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<mods:number id="DEF77A2D4278D1E5687CD52D7070C36C">57</mods:number>
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<mods:start id="34A7F2C8EA5083050169D9C59F40945A">139</mods:start>
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<mods:url id="6A1672431C0419A47F2D3E08970351D5">http://dx.doi.org/10.1515/aemnp-2017-0115</mods:url>
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<mods:classification id="13CCB380755CCD5A5F68A2DBD4C0554C">journal article</mods:classification>
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<mods:identifier id="659A976F58BD006D059B52AC7AB55AFF" type="DOI">10.1515/aemnp-2017-0115</mods:identifier>
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<treatment id="685887ABFFBA8458FE3FEC975640287D" LSID="urn:lsid:plazi:treatment:685887ABFFBA8458FE3FEC975640287D" httpUri="http://treatment.plazi.org/id/685887ABFFBA8458FE3FEC975640287D" lastPageId="11" lastPageNumber="150" pageId="10" pageNumber="149">
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<subSubSection id="A8EB6536FFBA8459FE3FEC9756262BBD" box="[487,791,219,243]" pageId="10" pageNumber="149" type="nomenclature">
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<paragraph id="E04E36BDFFBA8459FE3FEC9756262BBD" blockId="10.[487,791,219,243]" box="[487,791,219,243]" pageId="10" pageNumber="149">
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<heading id="BB0681D1FFBA8459FE3FEC9756262BBD" bold="true" box="[487,791,219,243]" centered="true" fontSize="8" level="2" pageId="10" pageNumber="149" reason="3">
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<emphasis id="D285EAAFFFBA8459FE3FEC9756262BBD" bold="true" box="[487,791,219,243]" pageId="10" pageNumber="149">
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<taxonomicName id="27F14D3EFFBA8459FE3FEC97578A2BBD" authority="Purchart & Kamiński, 2017" authorityName="Purchart & Kamiński" authorityYear="2017" box="[487,699,219,243]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="149" phylum="Arthropoda" rank="species" species="omanicus" status="sp. nov.">
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<emphasis id="D285EAAFFFBA8459FE3FEC97578A2BBD" bold="true" box="[487,699,219,243]" italics="true" pageId="10" pageNumber="149">Clitobius omanicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C9B657D4FFBA8459FD1AEC9056262BBD" box="[706,791,220,243]" pageId="10" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A8EB6536FFBA8459FDF8ECB657EC2A43" box="[544,733,250,269]" pageId="10" pageNumber="149" type="description">
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<paragraph id="E04E36BDFFBA8459FDF8ECB657EC2A43" blockId="10.[544,733,250,269]" box="[544,733,250,269]" pageId="10" pageNumber="149">
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(
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<figureCitation id="78CA2A38FFBA8459FDFEECB6575A2A43" box="[550,619,250,269]" captionStart="Fig" captionStartId="5.[115,146,891,910]" captionTargetBox="[113,1089,236,871]" captionTargetId="figure-377@5.[112,1089,211,882]" captionTargetPageId="5" captionText="Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5324618" httpUri="https://zenodo.org/record/5324618/files/figure.png" pageId="10" pageNumber="149">Figs 1B</figureCitation>
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,
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<figureCitation id="78CA2A38FFBA8459FDADECB657F32A43" box="[629,706,250,269]" captionStart="Fig" captionStartId="7.[113,144,1299,1318]" captionTargetBox="[130,1074,221,1287]" captionTargetId="figure-13@7.[130,1074,219,1290]" captionTargetPageId="7" captionText="Fig. 2. Detailed morphology of Clitobius obesus (Waterhouse, 1881) (A–C, F) and C. omanicus sp. nov. (D, E, G). Abbreviations: a – trochanter base; b – femur base; c – rounded posterior angle of pronotal disc; d – not depressed sides of pronotal disc; e – convex intervals; f – ventral; g – dorsal; h – lateral view." figureDoi="http://doi.org/10.5281/zenodo.5324620" httpUri="https://zenodo.org/record/5324620/files/figure.png" pageId="10" pageNumber="149">2D, E, G</figureCitation>
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,
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<figureCitation id="78CA2A38FFBA8459FD14ECB657E62A43" box="[716,727,250,269]" captionStart="Fig" captionStartId="14.[149,180,1540,1559]" captionTargetBox="[151,1126,221,1457]" captionTargetId="figure-10@14.[151,1127,220,1457]" captionTargetPageId="14" captionText="Fig. 4. Analysed localities and potential distribution of Clitobius ovatus (Erichson, 1843) (A) and C. oblongiusculus (Fairmaire, 1875) (B). Average AUC and habitat suitability threshold values for each analysis are presented." figureDoi="http://doi.org/10.5281/zenodo.5324624" httpUri="https://zenodo.org/record/5324624/files/figure.png" pageId="10" pageNumber="149">4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A8EB6536FFBA8459FF4FED68546B2AC9" pageId="10" pageNumber="149" type="materials_examined">
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<paragraph id="E04E36BDFFBA8459FF4FED6857722A79" blockId="10.[151,1128,292,391]" box="[151,579,292,311]" pageId="10" pageNumber="149">
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<emphasis id="D285EAAFFFBA8459FF4FED6854212A79" bold="true" box="[151,272,292,311]" pageId="10" pageNumber="149">
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<typeStatus id="3F4A881FFFBA8459FF4FED6855F52A79" box="[151,196,292,311]" pageId="10" pageNumber="149">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="98E6762DFFBA8459FECDED68547A2A79" box="[277,331,292,311]" name="Oman" pageId="10" pageNumber="149">Oman</collectingCountry>
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,
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<collectingRegion id="2235F85FFFBA8459FE8CED6854D52A79" box="[340,484,292,311]" country="Oman" name="Zufar" pageId="10" pageNumber="149">Dhofar Province</collectingRegion>
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, Wadi Ain.
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</paragraph>
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<paragraph id="E04E36BDFFBA8459FF4FED72546B2AC9" blockId="10.[151,1128,292,391]" pageId="10" pageNumber="149">
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<emphasis id="D285EAAFFFBA8459FF4FED72542A2A1F" bold="true" box="[151,283,318,337]" pageId="10" pageNumber="149">Type material.</emphasis>
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<materialsCitation id="50993CE0FFBA8459FEC7ED7257B42A22" collectingDate="2012-03-29" collectionCode="NMPC" country="Oman" elevation="59" latitude="17.016666" location="Mirbat" longLatPrecision="1280" longitude="54.783333" pageId="10" pageNumber="149" specimenCount="1" stateProvince="Zufar" typeStatus="holotype">
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<typeStatus id="3F4A881FFFBA8459FEC7ED7254422A1E" box="[287,371,318,337]" pageId="10" pageNumber="149" type="holotype">HOLOTYPE</typeStatus>
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, J (
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<collectionCode id="86E0AE78FFBA8459FE4FED7254E92A1F" box="[407,472,318,337]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="149">NMPC</collectionCode>
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), labelled:
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||
<collectingCountry id="98E6762DFFBA8459FDEBED7257472A1F" box="[563,630,318,337]" name="Oman" pageId="10" pageNumber="149">OMAN</collectingCountry>
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,
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<collectingRegion id="2235F85FFFBA8459FDA6ED7257DC2A1F" box="[638,749,318,337]" country="Oman" name="Zufar" pageId="10" pageNumber="149">Dhofar Prov.</collectingRegion>
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/ WADI AIN,
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<elevation id="6BDCD18EFFBA8459FCBDED72568D2A1F" box="[869,956,318,337]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="10" pageNumber="149" unit="m" value="59.0">
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<quantity id="27099B58FFBA8459FCBDED7256A12A1F" box="[869,912,318,337]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="10" pageNumber="149" unit="m" value="59.0">59 m</quantity>
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a.s.l.
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||
</elevation>
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/
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||
<quantity id="27099B58FFBA8459FC13ED7251312A1F" box="[971,1024,318,337]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="149" unit="km" value="13.0">13 km</quantity>
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E of
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<location id="E52E6066FFBA8459FBF4ED7251572A1F" LSID="urn:lsid:plazi:treatment:685887ABFFBA8458FE3FEC975640287D:E52E6066FFBA8459FBF4ED7251572A1F" box="[1068,1126,318,337]" country="Oman" latitude="17.016666" longLatPrecision="1280" longitude="54.783333" name="Mirbat" pageId="10" pageNumber="149" stateProvince="Zufar">Mirbat</location>
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/
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<geoCoordinate id="85C5507AFFBA8459FF7CED1555DF2A22" box="[164,238,345,364]" degrees="17" direction="north" minutes="01" orientation="latitude" pageId="10" pageNumber="149" precision="925" value="17.016666">17°01′N</geoCoordinate>
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;
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<geoCoordinate id="85C5507AFFBA8459FF20ED15540D2A22" box="[248,316,345,364]" degrees="54" direction="east" minutes="47" orientation="longitude" pageId="10" pageNumber="149" precision="925" value="54.783333">54°47′E</geoCoordinate>
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[GPS] /
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<date id="944F107DFFBA8459FE53ED1554C62A22" box="[395,503,345,364]" pageId="10" pageNumber="149" value="2012-03-29">
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<collectingDate id="840BE995FFBA8459FE53ED1554C62A22" box="[395,503,345,364]" pageId="10" pageNumber="149" value="2012-03-29">29. iii. 2012</collectingDate>
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</date>
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, A. REITER lgt
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</materialsCitation>
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.
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<typeStatus id="3F4A881FFFBA8459FD57ED1557D62A24" box="[655,743,345,364]" pageId="10" pageNumber="149" type="paratype">PARATYPES</typeStatus>
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(1 J
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<specimenCount id="F6F7FD34FFBA8459FCC3ED15567C2A25" box="[795,845,338,364]" pageId="10" pageNumber="149" type="female">2 ♀♀</specimenCount>
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<collectionCode id="86E0AE78FFBA8459FC8BED1556A32A22" box="[851,914,345,364]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="149">NMPC</collectionCode>
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, 1 J
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<collectionCode id="86E0AE78FFBA8459FC1AED1556CB2A22" box="[962,1018,345,364]" pageId="10" pageNumber="149">LPCB</collectionCode>
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, 1 J
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<collectionCode id="86E0AE78FFBA8459FBF3ED15516C2A22" box="[1067,1117,345,364]" pageId="10" pageNumber="149">MIIZ</collectionCode>
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): same data as
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<typeStatus id="3F4A881FFFBA8459FED0ED3854642AC9" box="[264,341,372,391]" pageId="10" pageNumber="149" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="A8EB6536FFBA8458FF4FEDD3568929BD" lastPageId="11" lastPageNumber="150" pageId="10" pageNumber="149" type="description">
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<paragraph id="E04E36BDFFBA8459FF4FEDD356BC2AF8" blockId="10.[151,1127,415,1526]" box="[151,909,415,438]" pageId="10" pageNumber="149">
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<emphasis id="D285EAAFFFBA8459FF4FEDD354132AF8" bold="true" box="[151,290,415,438]" pageId="10" pageNumber="149">Description.</emphasis>
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Body length =
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<quantity id="27099B58FFBA8459FE10EDD357792AF8" box="[456,584,415,438]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.5" metricValueMin="5.8" pageId="10" pageNumber="149" unit="mm" value="6.15" valueMax="6.5" valueMin="5.8">5.8–6.5 mm</quantity>
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||
(n = 6). Habitus as in
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<figureCitation id="78CA2A38FFBA8459FCEFEDD356B82AF8" box="[823,905,415,438]" captionStart="Fig" captionStartId="5.[115,146,891,910]" captionTargetBox="[113,1089,236,871]" captionTargetId="figure-377@5.[112,1089,211,882]" captionTargetPageId="5" captionText="Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5324618" httpUri="https://zenodo.org/record/5324618/files/figure.png" pageId="10" pageNumber="149">Fig. 1B</figureCitation>
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.
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</paragraph>
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||
<paragraph id="E04E36BDFFBA8459FF6DEDF3571F2818" blockId="10.[151,1127,415,1526]" pageId="10" pageNumber="149">Dorsal side of head dull, with fine, but often merging punctures. Frontoclypeal suture extremely fine. Canthus rounded in basal part. Clypeal emargination relatively shallow (clypeal emargination width / depth ratio ca. 10.0). Labrum wide (width / length ratio ca. 1.8); delicately emarginated in middle. Sides of mentum subparallel. Mentum with median keel; lateral wings directed towards body front; not covering cardines or basistipites. Submentum pentagonal. Apical maxillary palpomere trapezoidal; with relatively large sensory field (occupying the whole proximal edge). Maxillary palpomere II elongated (length to width ratio ca. 3.0). Palpifer with bunch of setae located near base. Anterior tentorial pits circular. Dorsally eye not shielded by longitudinal keel. Eye partly divided by canthus (three rows of ocelli visible between canthus and gena); ventral portion contains slightly more ocelli than dorsal one. Antenna relatively long (length of antenna / maximal length of pronotum ca. 1.3); antennomere III slender (ratio of its length to length of antennomere II ca. 1.5); antennomeres VII–XI forming a loosely joined club.</paragraph>
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<paragraph id="E04E36BDFFBA8459FF6DEF1357D12FD8" blockId="10.[151,1127,415,1526]" pageId="10" pageNumber="149">Pronotum narrower than elytra (width ratio elytra / pronotum ca. 1.2). Length to width ratio of pronotal disc ca. 0.6. Pronotal disc dull; with conspicuous microsculpture (magnification 40×); covered with fine and shallow punctures (1–2 diameters apart); covered with short and extremely fine setae. Lateral sides of pronotal disc rounded; widest in basal third; without apophyseal depressions. Lateral emargination of pronotum narrow and complete; basal emargination complete (clearly visible, relatively broad); anterior slightly interrupted in middle. Middle of prosternum and basal part of hypomeron covered with coarse setae. The latter structure flattened near lateral edges. Intercoxal process of prosternum not depressed apically, with sharp tip slightly projecting outwards (lateral view). Procoxae situated closely (width ratio of procoxa and intercoxal process ca. 1.3).</paragraph>
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<paragraph id="E04E36BDFFBA8459FF6DE8D357A22E18" blockId="10.[151,1127,415,1526]" pageId="10" pageNumber="149">Elytra dull; glabrous. Elytral striae impressed on whole length, visible as nine rows of conspicuous punctures on each elytron (2 diameters apart). Intervals flat, with fine punctures (3–4 diameters apart). Elytral base sinusoidal; not emarginate. Epipleuron straight in most of its length, terminating at level of abdominal ventrite V; situated ventrally, only basal part visible in dorsal aspect near elytral slope. Elytral humerus obtuse. Scutellum relatively large; triangular. Metathoracic wings fully developed.</paragraph>
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<paragraph id="E04E36BDFFBA8459FF6DE91357DB2EB8" blockId="10.[151,1127,415,1526]" pageId="10" pageNumber="149">Mesoventrite without processes. Metaventrite relatively elongated (length ratio cavity of metacoxa / metaventrite between insertions of meso- and metacoxae ca. 0.9); without processes, however in apical part slightly elevated; with median carina reaching base of elevated part. Process of abdominal ventrite I narrow and sharp (ratio of distances between mexo- and metacoxae ca. 0.5). Abdominal ventrite V not bordered.</paragraph>
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<paragraph id="E04E36BDFFBB8458FF48EC90540E2A7D" blockId="11.[113,1089,220,819]" pageId="11" pageNumber="150">
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Protrochanters of opatrinoid
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||
<typeStatus id="3F4A881FFFBB8458FE1AEC9054C12BBD" box="[450,496,220,243]" pageId="11" pageNumber="150">type</typeStatus>
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(with elongated base). Meso- and metatrochanters only slightly covered by respective femora. Metacoxa elongated. Male and female tarsi, tibiae and femora simple.
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</paragraph>
|
||
<paragraph id="E04E36BDFFBB8458FF48ED70568929BD" blockId="11.[113,1089,220,819]" pageId="11" pageNumber="150">
|
||
Aedeagal tegmen as in
|
||
<figureCitation id="78CA2A38FFBB8458FE50ED7054EC2A1D" box="[392,477,316,339]" captionStart="Fig" captionStartId="5.[115,146,891,910]" captionTargetBox="[113,1089,236,871]" captionTargetId="figure-377@5.[112,1089,211,882]" captionTargetPageId="5" captionText="Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5324618" httpUri="https://zenodo.org/record/5324618/files/figure.png" pageId="11" pageNumber="150">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="78CA2A38FFBB8458FE30ED70573C2A1D" box="[488,525,316,339]" captionStart="Fig" captionStartId="7.[113,144,1299,1318]" captionTargetBox="[130,1074,221,1287]" captionTargetId="figure-13@7.[130,1074,219,1290]" captionTargetPageId="7" captionText="Fig. 2. Detailed morphology of Clitobius obesus (Waterhouse, 1881) (A–C, F) and C. omanicus sp. nov. (D, E, G). Abbreviations: a – trochanter base; b – femur base; c – rounded posterior angle of pronotal disc; d – not depressed sides of pronotal disc; e – convex intervals; f – ventral; g – dorsal; h – lateral view." figureDoi="http://doi.org/10.5281/zenodo.5324620" httpUri="https://zenodo.org/record/5324620/files/figure.png" pageId="11" pageNumber="150">2G</figureCitation>
|
||
. Parameres evenly narrowing towards apex; tip sinusoidal (lateral view). Clavae absent. Penis basally covered by fused parameres. Ovipositor with paraprocts equal in length to coxites. Paraproct does not shield valvifer and other lobes (directed basally). Valvifer wide and short; third and fourth lobes of coxities elongated. Gonostylus situated laterally. Vagina and bursa copulatrix without sclerites. Spiculum ventrale short.
|
||
<emphasis id="D285EAAFFFBB8458FFA9ED90545F2ABD" bold="true" box="[113,366,476,499]" pageId="11" pageNumber="150">Differential diagnosis.</emphasis>
|
||
This species is similar to
|
||
<taxonomicName id="27F14D3EFFBB8458FD4BED9056912ABD" baseAuthorityName="Fairmaire" baseAuthorityYear="1875" box="[659,928,476,499]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="150" phylum="Arthropoda" rank="species" species="oblongiusculus">
|
||
<emphasis id="D285EAAFFFBB8458FD4BED9056912ABD" box="[659,928,476,499]" italics="true" pageId="11" pageNumber="150">Clitobius oblongiusculus</emphasis>
|
||
</taxonomicName>
|
||
in sharing the following features: dorsal side of the body glabrous or covered with fine setae (
|
||
<figureCitation id="78CA2A38FFBB8458FC06EDB05100295D" box="[990,1073,508,531]" captionStart="Fig" captionStartId="5.[115,146,891,910]" captionTargetBox="[113,1089,236,871]" captionTargetId="figure-377@5.[112,1089,211,882]" captionTargetPageId="5" captionText="Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5324618" httpUri="https://zenodo.org/record/5324618/files/figure.png" pageId="11" pageNumber="150">Fig. 1B</figureCitation>
|
||
), metathoracic wings fully developed, pronotum widest in the middle or near the base (
|
||
<figureCitation id="78CA2A38FFBB8458FBCDEE5055A0291D" captionStart="Fig" captionStartId="5.[115,146,891,910]" captionTargetBox="[113,1089,236,871]" captionTargetId="figure-377@5.[112,1089,211,882]" captionTargetPageId="5" captionText="Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5324618" httpUri="https://zenodo.org/record/5324618/files/figure.png" pageId="11" pageNumber="150">Fig. 1B</figureCitation>
|
||
), elytral humeri not protruding outwards (
|
||
<figureCitation id="78CA2A38FFBB8458FD93EE7057AA291D" box="[587,667,572,595]" captionStart="Fig" captionStartId="5.[115,146,891,910]" captionTargetBox="[113,1089,236,871]" captionTargetId="figure-377@5.[112,1089,211,882]" captionTargetPageId="5" captionText="Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5324618" httpUri="https://zenodo.org/record/5324618/files/figure.png" pageId="11" pageNumber="150">Fig. 1B</figureCitation>
|
||
), protibiae slender, without a prominent apical tooth (
|
||
<figureCitation id="78CA2A38FFBB8458FF26EE10547E293D" box="[254,335,604,627]" captionStart="Fig" captionStartId="5.[115,146,891,910]" captionTargetBox="[113,1089,236,871]" captionTargetId="figure-377@5.[112,1089,211,882]" captionTargetPageId="5" captionText="Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5324618" httpUri="https://zenodo.org/record/5324618/files/figure.png" pageId="11" pageNumber="150">Fig. 1B</figureCitation>
|
||
), and male protarsi not widened. Both species can be distinguished by different body shape (
|
||
<taxonomicName id="27F14D3EFFBB8458FE86EE30572E29DD" baseAuthorityName="Fairmaire" baseAuthorityYear="1875" box="[350,543,636,659]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="150" phylum="Arthropoda" rank="species" species="oblongiusculus">
|
||
<emphasis id="D285EAAFFFBB8458FE86EE30572E29DD" box="[350,543,636,659]" italics="true" pageId="11" pageNumber="150">C. oblongiusculus</emphasis>
|
||
</taxonomicName>
|
||
– elongate;
|
||
<taxonomicName id="27F14D3EFFBB8458FD7DEE30561A29DD" authority="Purchart & Kamiński, 2017" authorityName="Purchart & Kamiński" authorityYear="2017" box="[677,811,636,659]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="150" phylum="Arthropoda" rank="species" species="omanicus" status="sp. nov.">
|
||
<emphasis id="D285EAAFFFBB8458FD7DEE30561A29DD" box="[677,811,636,659]" italics="true" pageId="11" pageNumber="150">C. omanicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C9B657D4FFBB8458FCEAEE3056B729DD" box="[818,902,636,659]" pageId="11" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
|
||
– oval), shape of antennomere III (
|
||
<taxonomicName id="27F14D3EFFBB8458FEF5EED054DC29FD" baseAuthorityName="Fairmaire" baseAuthorityYear="1875" box="[301,493,668,691]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="150" phylum="Arthropoda" rank="species" species="oblongiusculus">
|
||
<emphasis id="D285EAAFFFBB8458FEF5EED054DC29FD" box="[301,493,668,691]" italics="true" pageId="11" pageNumber="150">C. oblongiusculus</emphasis>
|
||
</taxonomicName>
|
||
– short;
|
||
<taxonomicName id="27F14D3EFFBB8458FD95EED057E229FD" authority="Purchart & Kamiński, 2017" authorityName="Purchart & Kamiński" authorityYear="2017" box="[589,723,668,691]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="150" phylum="Arthropoda" rank="species" species="omanicus" status="sp. nov.">
|
||
<emphasis id="D285EAAFFFBB8458FD95EED057E229FD" box="[589,723,668,691]" italics="true" pageId="11" pageNumber="150">C. omanicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C9B657D4FFBB8458FD02EED0561C29FD" box="[730,813,668,691]" pageId="11" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
|
||
– elongate), and shape of the posterior pronotal angles (
|
||
<taxonomicName id="27F14D3EFFBB8458FE6DEEF05747299D" baseAuthorityName="Fairmaire" baseAuthorityYear="1875" box="[437,630,700,723]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="150" phylum="Arthropoda" rank="species" species="oblongiusculus">
|
||
<emphasis id="D285EAAFFFBB8458FE6DEEF05747299D" box="[437,630,700,723]" italics="true" pageId="11" pageNumber="150">C. oblongiusculus</emphasis>
|
||
</taxonomicName>
|
||
– sharp;
|
||
<taxonomicName id="27F14D3EFFBB8458FD04EEF05653299D" authority="Purchart & Kamiński, 2017" authorityName="Purchart & Kamiński" authorityYear="2017" box="[732,866,700,723]" class="Insecta" family="Tenebrionidae" genus="Clitobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="150" phylum="Arthropoda" rank="species" species="omanicus" status="sp. nov.">
|
||
<emphasis id="D285EAAFFFBB8458FD04EEF05653299D" box="[732,866,700,723]" italics="true" pageId="11" pageNumber="150">C. omanicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C9B657D4FFBB8458FCB2EEF0568F299D" box="[874,958,700,723]" pageId="11" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
|
||
– rounded). For details and additional characters, see the identification key attached below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8EB6536FFBB8458FFA9EEB0510D285D" box="[113,1084,764,787]" pageId="11" pageNumber="150" type="etymology">
|
||
<paragraph id="E04E36BDFFBB8458FFA9EEB0510D285D" blockId="11.[113,1089,220,819]" box="[113,1084,764,787]" pageId="11" pageNumber="150">
|
||
<emphasis id="D285EAAFFFBB8458FFA9EEB055C3285D" bold="true" box="[113,242,764,787]" pageId="11" pageNumber="150">Etymology.</emphasis>
|
||
The name refers to country name
|
||
<collectingCountry id="98E6762DFFBB8458FDB8EEB05795285D" box="[608,676,764,787]" name="Oman" pageId="11" pageNumber="150">Oman</collectingCountry>
|
||
, where the new species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8EB6536FFBB8458FFA9EF505640287D" box="[113,881,796,819]" pageId="11" pageNumber="150" type="distribution">
|
||
<paragraph id="E04E36BDFFBB8458FFA9EF505640287D" blockId="11.[113,1089,220,819]" box="[113,881,796,819]" pageId="11" pageNumber="150">
|
||
<emphasis id="D285EAAFFFBB8458FFA9EF505435287D" bold="true" box="[113,260,796,819]" pageId="11" pageNumber="150">Distribution.</emphasis>
|
||
<collectingCountry id="98E6762DFFBB8458FED2EF50547A287D" box="[266,331,796,819]" name="Oman" pageId="11" pageNumber="150">Oman</collectingCountry>
|
||
– so far known from the
|
||
<typeStatus id="3F4A881FFFBB8458FD81EF5057B6287D" box="[601,647,796,819]" pageId="11" pageNumber="150">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="78CA2A38FFBB8458FCFBEF505655287D" box="[803,868,796,819]" captionStart="Fig" captionStartId="15.[113,144,719,738]" captionTargetBox="[117,1084,221,697]" captionTargetId="figure-13@15.[113,1088,217,701]" captionTargetPageId="15" captionText="Fig. 5. Distribution of chosen species representing Clitobius: taxa restricted to the Horn of Africa and its surroundings (A), and Southern Africa (B)." figureDoi="http://doi.org/10.5281/zenodo.5324626" httpUri="https://zenodo.org/record/5324626/files/figure.png" pageId="11" pageNumber="150">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |