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<document id="46BBFF527BCD6BD75E9A8B7A3BAF3488" ID-CLB-Dataset="46846" ID-DOI="10.11646/zootaxa.4072.4.1" ID-GBIF-Dataset="36996e71-2028-43cb-a42e-aebb74b47cf2" ID-ISSN="1175-5326" ID-Zenodo-Dep="266752" ID-ZooBank="232286BA-A338-468E-842D-8A81F8269551" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460358657892" checkinUser="plazi" docAuthor="Myers, Alan A. &amp; Nithyanandan, Manickam" docDate="2016" docId="03DB87CCFFAAFFD5B0C0FF117DE1679E" docLanguage="en" docName="zootaxa.4072.4.1.pdf" docOrigin="Zootaxa 4072 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceradocus (Denticeradocus) alama Myers &amp; Nithyanandan, 2016, sp. nov." docType="treatment" docVersion="14" lastPageNumber="424" masterDocId="FFE2FFB4FFBEFFC2B057FF867C20635B" masterDocTitle="The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea)" masterLastPageNumber="429" masterPageNumber="401" pageNumber="421" updateTime="1698294210271" updateUser="ExternalLinkService">
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<mods:title id="32F95CF76A146B9DFF41E7CE025A7D2B">The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea)</mods:title>
</mods:titleInfo>
<mods:name id="C9A5B88342BA4701F0CB5AA0FD807402" type="personal">
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<mods:namePart id="74EBD9291BC8A44B17C018D90BD01A05">Myers, Alan A.</mods:namePart>
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<mods:roleTerm id="49C8C4472087B63F0A96590A632435F7">Author</mods:roleTerm>
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<mods:namePart id="8F1AA4C8658ED1D4138F14C27298872B">Nithyanandan, Manickam</mods:namePart>
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<mods:typeOfResource id="225472CB9A68B694729CEFE396DC3ABF">text</mods:typeOfResource>
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<mods:title id="80BA35D6A8EC57CC3F7BD58AD63FEF5D">Zootaxa</mods:title>
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<mods:date id="A371E156625F335DA0F50C3D3AFAEEAD">2016</mods:date>
<mods:detail id="FC893780C22AB47173CE5C6B144E09F2" type="volume">
<mods:number id="BC77D812E40E1C0C214EFC29AB1C2BD5">4072</mods:number>
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<mods:identifier id="D577CAA07C0568EDC7ABE3550014F67A" type="DOI">10.11646/zootaxa.4072.4.1</mods:identifier>
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<treatment id="03DB87CCFFAAFFD5B0C0FF117DE1679E" ID-DOI="http://doi.org/10.5281/zenodo.4654089" ID-GBIF-Taxon="119409280" ID-Zenodo-Dep="4654089" LSID="urn:lsid:plazi:treatment:03DB87CCFFAAFFD5B0C0FF117DE1679E" httpUri="http://treatment.plazi.org/id/03DB87CCFFAAFFD5B0C0FF117DE1679E" lastPageId="23" lastPageNumber="424" pageId="20" pageNumber="421">
<subSubSection id="C3686551FFAAFFD6B0C0FF117D066388" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="8BCD36DAFFAAFFD6B0C0FF117E8663E9" blockId="20.[151,678,151,211]" box="[151,678,151,178]" pageId="20" pageNumber="421">
<heading id="D08581B6FFAAFFD6B0C0FF117E8663E9" bold="true" box="[151,678,151,178]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<emphasis id="B906EAC8FFAAFFD6B0C0FF117E8663E9" bold="true" box="[151,678,151,178]" pageId="20" pageNumber="421">
<taxonomicName id="4C724D59FFAAFFD6B0C0FF117E6263EA" box="[151,578,151,177]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="alama" status="sp. nov." subGenus="Denticeradocus">
<emphasis id="B906EAC8FFAAFFD6B0C0FF117E6263EA" bold="true" box="[151,578,151,177]" italics="true" pageId="20" pageNumber="421">Ceradocus (Denticeradocus) alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFAAFFD6B21EFF1E7E8663E9" box="[585,678,152,178]" pageId="20" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCD36DAFFAAFFD6B0C0FF3C7D066388" blockId="20.[151,678,151,211]" box="[151,294,186,211]" pageId="20" pageNumber="421">
(
<figureCitation id="13492A5FFFAAFFD6B0C8FF3C7D3E6388" box="[159,286,186,211]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStartId-0="20.[151,250,2007,2029]" captionStartId-1="21.[151,250,1799,1821]" captionStartId-2="22.[151,250,1945,1967]" captionTargetBox-0="[156,1421,397,1964]" captionTargetBox-1="[177,1405,191,1767]" captionTargetBox-2="[185,1403,275,1908]" captionTargetId-0="figure@20.[151,1436,387,1985]" captionTargetId-1="figure@21.[169,1417,179,1788]" captionTargetId-2="figure@22.[167,1419,265,1924]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURE 16. Ceradocus (Denticeradocus) alama sp. nov., male, 7.0 mm, female, 6.5 mm, Sea City, Kuwait." captionText-1="FIGURE 17. Ceradocus (Denticeradocus) alama sp. nov., male, 7.0 mm, Sea City, Kuwait." captionText-2="FIGURE 18. Ceradocus (Denticeradocus) alama sp. nov., male, 7.0 mm, Sea City, Kuwait." httpUri-0="https://zenodo.org/record/266768/files/figure.png" httpUri-1="https://zenodo.org/record/266769/files/figure.png" httpUri-2="https://zenodo.org/record/266770/files/figure.png" pageId="20" pageNumber="421">Figs 1618</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3686551FFAAFFD7B0C0FE847D1864B6" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421" type="materials_examined">
<paragraph id="8BCD36DAFFAAFFD6B0C0FE847F696238" blockId="20.[151,1436,258,355]" pageId="20" pageNumber="421">
<emphasis id="B906EAC8FFAAFFD6B0C0FE847D666240" bold="true" box="[151,326,258,283]" pageId="20" pageNumber="421">
<typeStatus id="54C98878FFAAFFD6B0C0FE847CF26240" box="[151,210,258,283]" pageId="20" pageNumber="421">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C98878FFAAFFD6B11AFE847D956240" box="[333,437,258,283]" pageId="20" pageNumber="421" type="holotype">Holotype</typeStatus>
male, 7.0 mm (
<collectionCode id="ED63AE1FFFAAFFD6B23FFE857EF46240" LSID="urn:lsid:biocol.org:col:34665" box="[616,724,259,283]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="20" pageNumber="421">NHMUK</collectionCode>
2015. 3198) North Gate, collected from seaweed and hydroids growing on concrete sides of north tidal gate phase A3 by diving. D.K. Raja,
<date id="FFCC101AFFAAFFD6B456FEA178826264" box="[1025,1186,294,319]" pageId="20" pageNumber="421" value="2014-04-19">19 April 2014</date>
.
<typeStatus id="54C98878FFAAFFD6B4FAFEA1793D6264" box="[1197,1309,295,319]" pageId="20" pageNumber="421" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D74FD53FFAAFFD6B57EFEA079A26264" box="[1321,1410,294,319]" pageId="20" pageNumber="421" type="male">6 males</specimenCount>
,
<specimenCount id="9D74FD53FFAAFFD6B5D9FEA17CCE6238" pageId="20" pageNumber="421" type="female">9 females</specimenCount>
(
<collectionCode id="ED63AE1FFFAAFFD6B0ABFECD7D486238" LSID="urn:lsid:biocol.org:col:34665" box="[252,360,331,355]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="20" pageNumber="421">NHMUK</collectionCode>
2015. 31993208), same data as
<typeStatus id="54C98878FFAAFFD6B288FECC7F656238" box="[735,837,330,355]" pageId="20" pageNumber="421" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF0D6652FFAAFFD6B0C0F85178CB64B6" box="[151,1259,2007,2029]" httpUri="https://zenodo.org/record/266768/files/figure.png" pageId="20" pageNumber="421" targetBox="[156,1421,397,1964]" targetPageId="20">
<paragraph id="8BCD36DAFFAAFFD6B0C0F85178CB64B6" blockId="20.[151,1259,2007,2029]" box="[151,1259,2007,2029]" pageId="20" pageNumber="421">
<emphasis id="B906EAC8FFAAFFD6B0C0F8517D3F64B7" bold="true" box="[151,287,2007,2029]" pageId="20" pageNumber="421">FIGURE 16.</emphasis>
<taxonomicName id="4C724D59FFAAFFD6B172F8517EB064B6" box="[293,656,2007,2029]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="alama" status="sp. nov." subGenus="Denticeradocus">
<emphasis id="B906EAC8FFAAFFD6B172F8517DB364B6" box="[293,403,2007,2029]" italics="true" pageId="20" pageNumber="421">Ceradocus</emphasis>
(
<emphasis id="B906EAC8FFAAFFD6B1F5F8517E6264B6" box="[418,578,2007,2029]" italics="true" pageId="20" pageNumber="421">Denticeradocus</emphasis>
)
<emphasis id="B906EAC8FFAAFFD6B206F8517EB064B6" box="[593,656,2007,2029]" italics="true" pageId="20" pageNumber="421">alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFAAFFD6B2C1F8517EC764B7" box="[662,743,2007,2028]" pageId="20" pageNumber="421" rank="species">
<emphasis id="B906EAC8FFAAFFD6B2C1F8517EC764B7" bold="true" box="[662,743,2007,2028]" pageId="20" pageNumber="421">sp. nov.</emphasis>
</taxonomicNameLabel>
, male, 7.0 mm, female, 6.5 mm, Sea City, Kuwait.
</paragraph>
</caption>
<caption id="DF0D6652FFABFFD7B0C0F881781F6446" box="[151,1087,1799,1821]" httpUri="https://zenodo.org/record/266769/files/figure.png" pageId="21" pageNumber="422" targetBox="[177,1405,191,1767]" targetPageId="21">
<paragraph id="8BCD36DAFFABFFD7B0C0F881781F6446" blockId="21.[151,1087,1799,1821]" box="[151,1087,1799,1821]" pageId="21" pageNumber="422">
<emphasis id="B906EAC8FFABFFD7B0C0F8817D3F6447" bold="true" box="[151,287,1799,1821]" pageId="21" pageNumber="422">FIGURE 17.</emphasis>
<taxonomicName id="4C724D59FFABFFD7B172F8817EB06446" box="[293,656,1799,1821]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="alama" status="sp. nov." subGenus="Denticeradocus">
<emphasis id="B906EAC8FFABFFD7B172F8817DB36446" box="[293,403,1799,1821]" italics="true" pageId="21" pageNumber="422">Ceradocus</emphasis>
(
<emphasis id="B906EAC8FFABFFD7B1F5F8817E626446" box="[418,578,1799,1821]" italics="true" pageId="21" pageNumber="422">Denticeradocus</emphasis>
)
<emphasis id="B906EAC8FFABFFD7B206F8817EB06446" box="[593,656,1799,1821]" italics="true" pageId="21" pageNumber="422">alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFABFFD7B2C1F88E7EC76446" box="[662,743,1800,1821]" pageId="21" pageNumber="422" rank="species">
<emphasis id="B906EAC8FFABFFD7B2C1F88E7EC76446" bold="true" box="[662,743,1800,1821]" pageId="21" pageNumber="422">sp. nov.</emphasis>
</taxonomicNameLabel>
, male, 7.0 mm, Sea City, Kuwait.
</paragraph>
</caption>
<paragraph id="8BCD36DAFFABFFD7B090F8C27D1864B6" blockId="21.[151,1437,1860,2030]" pageId="21" pageNumber="422">
<emphasis id="B906EAC8FFABFFD7B090F8C27DA46406" bold="true" box="[199,388,1860,1885]" pageId="21" pageNumber="422">Other material.</emphasis>
<specimenCount id="9D74FD53FFABFFD7B1DBF8C37DC76406" box="[396,487,1860,1885]" pageId="21" pageNumber="422" type="male">9 males</specimenCount>
,
<specimenCount id="9D74FD53FFABFFD7B1A2F8C07E436406" box="[501,611,1860,1885]" pageId="21" pageNumber="422" type="female">7 females</specimenCount>
(
<collectionCode id="ED63AE1FFFABFFD7B224F8C37EFF6406" LSID="urn:lsid:biocol.org:col:34665" box="[627,735,1861,1885]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="21" pageNumber="422">NHMUK</collectionCode>
2015. 32093218), llBS, grab on bare sand benthos in phase A1, M. Nithyanandan,
<date id="FFCC101AFFABFFD7B1CBF8EE7E1764DA" box="[412,567,1896,1921]" pageId="21" pageNumber="422" value="2014-05-25">25 May 2014</date>
;
<specimenCount id="9D74FD53FFABFFD7B213F8EF7EBE64DA" box="[580,670,1896,1921]" pageId="21" pageNumber="422" type="male">7 males</specimenCount>
,
<specimenCount id="9D74FD53FFABFFD7B2FDF8EF7F3864DA" box="[682,792,1896,1921]" pageId="21" pageNumber="422" type="female">9 females</specimenCount>
(
<collectionCode id="ED63AE1FFFABFFD7B370F8EF7FB364DA" LSID="urn:lsid:biocol.org:col:34665" box="[807,915,1897,1921]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="21" pageNumber="422">NHMUK</collectionCode>
2015. 32193228), subtidal seagrass in phase A1 Sea City, Ocklemann sledge and small grab, M. Nithyanandan,
<date id="FFCC101AFFABFFD7B3FFF80B784D64FE" box="[936,1133,1933,1958]" pageId="21" pageNumber="422" value="2014-10-14">14 October 2014</date>
;
<specimenCount id="9D74FD53FFABFFD7B42BF80B78F564FD" box="[1148,1237,1933,1958]" pageId="21" pageNumber="422" type="male">2 males</specimenCount>
(
<collectionCode id="ED63AE1FFFABFFD7B4B0F80B797364FE" LSID="urn:lsid:biocol.org:col:34665" box="[1255,1363,1933,1957]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="21" pageNumber="422">NHMUK</collectionCode>
2015. 32353236), from sandy benthos in phase A3 Sea City waterways, Ocklemann sledge, M. Nithyanandan,
<date id="FFCC101AFFABFFD7B5D7F8377D1464B6" pageId="21" pageNumber="422" value="2013-10-20">20 October 2013</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3686551FFA8FFD5B090FF117E9D604E" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423" type="etymology">
<paragraph id="8BCD36DAFFA8FFD4B090FF117EF1638F" blockId="22.[199,906,151,213]" pageId="22" pageNumber="423">
<emphasis id="B906EAC8FFA8FFD4B090FF117D6E63EB" bold="true" box="[199,334,151,176]" pageId="22" pageNumber="423">Etymology.</emphasis>
Named after a creek (Al-Ama) in the
<typeStatus id="54C98878FFA8FFD4B2AEFF1E7F0963EB" box="[761,809,152,176]" pageId="22" pageNumber="423">type</typeStatus>
locality.
<emphasis id="B906EAC8FFA8FFD4B090FF3A7D79638E" bold="true" box="[199,345,188,213]" pageId="22" pageNumber="423">Description.</emphasis>
Based on male
<typeStatus id="54C98878FFA8FFD4B25CFF3A7E4D638E" box="[523,621,188,213]" pageId="22" pageNumber="423" type="holotype">holotype</typeStatus>
7.0 mm.
</paragraph>
<caption id="DF0D6652FFA8FFD4B0C0F81F781F64F4" box="[151,1087,1945,1967]" httpUri="https://zenodo.org/record/266770/files/figure.png" pageId="22" pageNumber="423" targetBox="[185,1403,275,1908]" targetPageId="22">
<paragraph id="8BCD36DAFFA8FFD4B0C0F81F781F64F4" blockId="22.[151,1087,1945,1967]" box="[151,1087,1945,1967]" pageId="22" pageNumber="423">
<emphasis id="B906EAC8FFA8FFD4B0C0F81F7D3F64F5" bold="true" box="[151,287,1945,1967]" pageId="22" pageNumber="423">FIGURE 18.</emphasis>
<taxonomicName id="4C724D59FFA8FFD4B172F81F7EB064F4" box="[293,656,1945,1967]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="alama" status="sp. nov." subGenus="Denticeradocus">
<emphasis id="B906EAC8FFA8FFD4B172F81F7DB364F4" box="[293,403,1945,1967]" italics="true" pageId="22" pageNumber="423">Ceradocus</emphasis>
(
<emphasis id="B906EAC8FFA8FFD4B1F5F81F7E6264F4" box="[418,578,1945,1967]" italics="true" pageId="22" pageNumber="423">Denticeradocus</emphasis>
)
<emphasis id="B906EAC8FFA8FFD4B206F81F7EB064F4" box="[593,656,1945,1967]" italics="true" pageId="22" pageNumber="423">alama</emphasis>
</taxonomicName>
<emphasis id="B906EAC8FFA8FFD4B2C1F81C7EC764F4" bold="true" box="[662,743,1946,1967]" pageId="22" pageNumber="423">
<taxonomicNameLabel id="A23557B3FFA8FFD4B2C1F81C7EC764F4" box="[662,743,1946,1967]" pageId="22" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, 7.0 mm, Sea City, Kuwait.
</paragraph>
</caption>
<paragraph id="8BCD36DAFFA9FFD5B090FF1179B46264" blockId="23.[151,1436,151,1221]" pageId="23" pageNumber="424">
<emphasis id="B906EAC8FFA9FFD5B090FF117D2E63EB" bold="true" box="[199,270,151,176]" pageId="23" pageNumber="424">Head.</emphasis>
<emphasis id="B906EAC8FFA9FFD5B142FF1F7D5763EB" box="[277,375,151,176]" italics="true" pageId="23" pageNumber="424">Eye lobe</emphasis>
rounded with sub-ocular notch.
<emphasis id="B906EAC8FFA9FFD5B2B2FF1F7F2F63EB" box="[741,783,153,176]" italics="true" pageId="23" pageNumber="424">Eye</emphasis>
medium size, subround.
<emphasis id="B906EAC8FFA9FFD5B47CFF1E78BE63EB" box="[1067,1182,152,176]" italics="true" pageId="23" pageNumber="424">Antenna 1</emphasis>
about two thirds body length; peduncular article 2 longer than 1, article 3 very short; accessory flagellum with 5 articles; primary flagellum longer than peduncle with 26 articles.
<emphasis id="B906EAC8FFA9FFD5B2E2FF667F0863A3" box="[693,808,224,248]" italics="true" pageId="23" pageNumber="424">Antenna 2</emphasis>
a little shorter than antenna 1; peduncular articles 4 and 5 subequal; flagellum with 24 articles.
<emphasis id="B906EAC8FFA9FFD5B20EFE827EE86247" box="[601,712,260,285]" italics="true" pageId="23" pageNumber="424">Maxilla 1</emphasis>
inner plate posterior margin fringed with setae.
<emphasis id="B906EAC8FFA9FFD5B4A0FE8279426246" box="[1271,1378,260,285]" italics="true" pageId="23" pageNumber="424">Mandible</emphasis>
palp article 1 with acute spine at posterodistal corner; article 3 much shorter than 2 with long distal and marginal setae.
</paragraph>
<paragraph id="8BCD36DAFFA9FFD5B090FECA79616143" blockId="23.[151,1436,151,1221]" pageId="23" pageNumber="424">
<emphasis id="B906EAC8FFA9FFD5B090FECA7D02623E" bold="true" box="[199,290,332,357]" pageId="23" pageNumber="424">Pereon.</emphasis>
<emphasis id="B906EAC8FFA9FFD5B17AFECA7DE5623F" box="[301,453,332,357]" italics="true" pageId="23" pageNumber="424">Gnathopod 1</emphasis>
coxa with very acute anterodistal corner; carpus and propodus subequal in length, strongly setose.
<emphasis id="B906EAC8FFA9FFD5B11CFEE97E3A62D3" box="[331,538,367,392]" italics="true" pageId="23" pageNumber="424">Right gnathopod 2</emphasis>
greatly enlarged; coxa subquadrate; basis stout with spine at anterodistal corner; carpus short, cup-shaped; propodus massive, subovoid, palm with short overhanging shelf, followed by a shallow U-shaped excavation, followed by an oblique undefined margin; dactylus stout, curved, fitting palm.
<emphasis id="B906EAC8FFA9FFD5B527FE3E7D0662AF" italics="true" pageId="23" pageNumber="424">Left gnathopod 2</emphasis>
much smaller than right gnathopod 2, similar in form and size to that of female.
<emphasis id="B906EAC8FFA9FFD5B480FE5A797A62AE" box="[1239,1370,476,501]" italics="true" pageId="23" pageNumber="424">Pereopod 6</emphasis>
basis posterodistal margin with multiple spines.
<emphasis id="B906EAC8FFA9FFD5B223FE797ED5614C" box="[628,757,511,536]" italics="true" pageId="23" pageNumber="424">Pereopod 7</emphasis>
basis posterodistal margin with a single acute spine.
</paragraph>
<paragraph id="8BCD36DAFFA9FFD5B090FDA2787261AB" blockId="23.[151,1436,151,1221]" pageId="23" pageNumber="424">
<emphasis id="B906EAC8FFA9FFD5B090FDA27D316166" bold="true" box="[199,273,548,573]" pageId="23" pageNumber="424">Pleon.</emphasis>
<emphasis id="B906EAC8FFA9FFD5B14DFDA27DC06167" box="[282,480,548,573]" italics="true" pageId="23" pageNumber="424">Pleonal segments</emphasis>
13 strongly dorsally spinous with setae inserted between the spines.
<emphasis id="B906EAC8FFA9FFD5B55FFDA379BC6167" box="[1288,1436,548,572]" italics="true" pageId="23" pageNumber="424">Epimera 13</emphasis>
posterior distal margins strongly spinous, with setae inserted between the spines, posterior margins with robust setae.
<emphasis id="B906EAC8FFA9FFD5B089FDEB7D9261DF" box="[222,434,621,644]" italics="true" pageId="23" pageNumber="424">Urosome segments</emphasis>
12 dorsally serrated with setae inserted between the spines.
<emphasis id="B906EAC8FFA9FFD5B425FDEA78FE61DF" box="[1138,1246,620,645]" italics="true" pageId="23" pageNumber="424">Uropod 1</emphasis>
peduncle a little longer than rami, with basofacial robust seta and with ventrodistal spine; inner ramus marginally longer than outer, slender with marginal and distal robust setae.
<emphasis id="B906EAC8FFA9FFD5B2CDFD327F256197" box="[666,773,692,717]" italics="true" pageId="23" pageNumber="424">Uropod 2</emphasis>
peduncle a little shorter than rami; inner ramus marginally longer than outer.
<emphasis id="B906EAC8FFA9FFD5B134FD517DEE61AB" box="[355,462,727,752]" italics="true" pageId="23" pageNumber="424">Uropod 3</emphasis>
peduncle short; rami enlarged, broad, subequal in length.
</paragraph>
<paragraph id="8BCD36DAFFA9FFD5B090FD7A7E9D604E" blockId="23.[151,1436,151,1221]" box="[199,701,764,789]" pageId="23" pageNumber="424">
<taxonomicName id="4C724D59FFA9FFD5B090FD7A7D2D604E" box="[199,269,764,789]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="23" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAC8FFA9FFD5B090FD7A7D2D604E" box="[199,269,764,789]" italics="true" pageId="23" pageNumber="424">Telson</emphasis>
</taxonomicName>
deeply cleft, lobes strongly divergent.
</paragraph>
</subSubSection>
<subSubSection id="C3686551FFA9FFD5B090FC99783360DB" pageId="23" pageNumber="424" type="description">
<paragraph id="8BCD36DAFFA9FFD5B090FC99783360DB" blockId="23.[151,1436,151,1221]" pageId="23" pageNumber="424">
<emphasis id="B906EAC8FFA9FFD5B090FC997D3E6063" bold="true" box="[199,286,799,824]" pageId="23" pageNumber="424">Female</emphasis>
(sexually dimorphic characters)
<emphasis id="B906EAC8FFA9FFD5B2F2FC997F1E6063" box="[677,830,799,824]" italics="true" pageId="23" pageNumber="424">Gnathopod 2</emphasis>
coxa elongate, rounded, unproduced; basis weakly expanded anteromedially; carpus well developed, about two thirds length of propodus; propodus elongate subovoid, palm oblique, defined by a robust seta; dactylus slender, fitting palm.
</paragraph>
</subSubSection>
<subSubSection id="C3686551FFA9FFD5B090FC0A7F2867FB" pageId="23" pageNumber="424" type="discussion">
<paragraph id="8BCD36DAFFA9FFD5B090FC0A7F2867FB" blockId="23.[151,1436,151,1221]" pageId="23" pageNumber="424">
<emphasis id="B906EAC8FFA9FFD5B090FC0A7D1B60FE" bold="true" box="[199,315,908,933]" pageId="23" pageNumber="424">Remarks.</emphasis>
<taxonomicName id="4C724D59FFA9FFD5B115FC0A7EF360FE" box="[322,723,908,933]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="alama" status="sp. nov." subGenus="Denticeradocus">
<emphasis id="B906EAC8FFA9FFD5B115FC0A7D9B60FE" box="[322,443,908,933]" italics="true" pageId="23" pageNumber="424">Ceradocus</emphasis>
(
<emphasis id="B906EAC8FFA9FFD5B19BFC0A7E5D60FE" box="[460,637,908,933]" italics="true" pageId="23" pageNumber="424">Denticeradocus</emphasis>
)
<emphasis id="B906EAC8FFA9FFD5B2D9FC0A7EF360FE" box="[654,723,908,933]" italics="true" pageId="23" pageNumber="424">alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFA9FFD5B28BFC0A7F1560FF" box="[732,821,908,933]" pageId="23" pageNumber="424" rank="species">
<emphasis id="B906EAC8FFA9FFD5B28BFC0A7F0E60FE" bold="true" box="[732,814,908,933]" pageId="23" pageNumber="424">sp. nov</emphasis>
.
</taxonomicNameLabel>
resembles
<taxonomicName id="4C724D59FFA9FFD5B3E3FC0B78DD60FE" authority="Bate, 1862" authorityName="Bate" authorityYear="1862" box="[948,1277,908,933]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="serratus" subGenus="Denticeradocus">
<emphasis id="B906EAC8FFA9FFD5B3E3FC0B7FEE60FF" box="[948,974,909,932]" italics="true" pageId="23" pageNumber="424">C.</emphasis>
(
<emphasis id="B906EAC8FFA9FFD5B388FC0B7FD360FF" box="[991,1011,909,932]" italics="true" pageId="23" pageNumber="424">D</emphasis>
.)
<emphasis id="B906EAC8FFA9FFD5B45CFC0B784760FF" box="[1035,1127,909,932]" italics="true" pageId="23" pageNumber="424">serratus</emphasis>
(
<bibRefCitation id="EFE34B2BFFA9FFD5B421FC0A78D560FE" author="Bate" box="[1142,1269,908,933]" pageId="23" pageNumber="424" refString="Bate, C. S. (1862) Catalogue of the Specimens of Amphipodous Crustacea in the Collection of the British Museum. Trustees, British Museum, London, 399 pp." type="book" year="1862">Bate, 1862</bibRefCitation>
)
</taxonomicName>
, as figured by
<bibRefCitation id="EFE34B2BFFA9FFD5B0C0FC367D476093" author="Barnard" box="[151,359,943,968]" pageId="23" pageNumber="424" refString="Barnard, J. L. (1972) Gammaridean Amphipoda of Australia, Part I. Smithsonian Contributions to Zoology, 103, 1 - 333. http: // dx. doi. org / 10.5479 / si. 00810282.103" type="journal article" year="1972">J.L. Barnard 1972</bibRefCitation>
(not
<taxonomicName id="4C724D59FFA9FFD5B1FFFC377F326093" authority="Bate, 1862" authorityName="Bate" authorityYear="1862" box="[424,786,943,968]" class="Malacostraca" family="Melitidae" genus="Megamoera" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="serrata">
<emphasis id="B906EAC8FFA9FFD5B1FFFC377EAA6093" box="[424,650,945,968]" italics="true" pageId="23" pageNumber="424">Megamoera serrata</emphasis>
<bibRefCitation id="EFE34B2BFFA9FFD5B2C3FC367F326093" author="Bate" box="[660,786,943,968]" pageId="23" pageNumber="424" refString="Bate, C. S. (1862) Catalogue of the Specimens of Amphipodous Crustacea in the Collection of the British Museum. Trustees, British Museum, London, 399 pp." type="book" year="1862">Bate, 1862</bibRefCitation>
</taxonomicName>
) from
<collectingCountry id="F365764AFFA9FFD5B334FC297FEA6093" box="[867,970,943,968]" name="Australia" pageId="23" pageNumber="424">Australia</collectingCountry>
in the male large gnathopod 2 having a short palmar shelf and in not having a posterodistal defining spine. That species, however, does not have an excavation in the palm of the large male gnathopod 2. It is similar to
<taxonomicName id="4C724D59FFA9FFD5B3CBFC7E786D674B" box="[924,1101,1016,1040]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="serratus" subGenus="Denticeradocus">
<emphasis id="B906EAC8FFA9FFD5B3CBFC7E7F8F6754" box="[924,943,1016,1039]" italics="true" pageId="23" pageNumber="424">C</emphasis>
. (
<emphasis id="B906EAC8FFA9FFD5B392FC7F7FF9674B" box="[965,985,1017,1040]" italics="true" pageId="23" pageNumber="424">D</emphasis>
.)
<emphasis id="B906EAC8FFA9FFD5B3A6FC7F786D674B" box="[1009,1101,1017,1040]" italics="true" pageId="23" pageNumber="424">serratus</emphasis>
</taxonomicName>
as figured by
<bibRefCitation id="EFE34B2BFFA9FFD5B4BAFC7179BC674B" author="Ledoyer" box="[1261,1436,1015,1040]" pageId="23" pageNumber="424" refString="Ledoyer, M. (1983) Crustaces amphipodes gammariens. Familles des Acanthonotozomatidae a Gammaridae. Faune de Madagascar, 59 (1), 1 - 598." type="journal article" year="1983">Ledoyer (1983)</bibRefCitation>
(also not
<taxonomicName id="4C724D59FFA9FFD5B155FB9B7E49676E" authority="Bate, 1862" authorityName="Bate" authorityYear="1862" box="[258,617,1052,1077]" class="Malacostraca" family="Melitidae" genus="Megamoera" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="serrata">
<emphasis id="B906EAC8FFA9FFD5B155FB9B7DC3676F" box="[258,483,1053,1076]" italics="true" pageId="23" pageNumber="424">Megamoera serrata</emphasis>
<bibRefCitation id="EFE34B2BFFA9FFD5B1BCFB9B7E49676E" author="Bate" box="[491,617,1052,1077]" pageId="23" pageNumber="424" refString="Bate, C. S. (1862) Catalogue of the Specimens of Amphipodous Crustacea in the Collection of the British Museum. Trustees, British Museum, London, 399 pp." type="book" year="1862">Bate, 1862</bibRefCitation>
</taxonomicName>
), but that species has a palmar defining corner lacking in
<emphasis id="B906EAC8FFA9FFD5B4AAFB9B7930676F" box="[1277,1296,1053,1076]" italics="true" pageId="23" pageNumber="424">C</emphasis>
. (
<emphasis id="B906EAC8FFA9FFD5B57EFB9B791D676F" box="[1321,1341,1053,1076]" italics="true" pageId="23" pageNumber="424">D</emphasis>
).
<taxonomicName id="4C724D59FFA9FFD5B501FB9A79BC676E" box="[1366,1436,1052,1077]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="alama" status="sp. nov.">
<emphasis id="B906EAC8FFA9FFD5B501FB9A79BC676E" box="[1366,1436,1052,1077]" italics="true" pageId="23" pageNumber="424">alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFA9FFD5B0C0FBB97CD46703" box="[151,244,1087,1112]" pageId="23" pageNumber="424" rank="species">
<emphasis id="B906EAC8FFA9FFD5B0C0FBB97CD46703" bold="true" box="[151,244,1087,1112]" pageId="23" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
Like
<emphasis id="B906EAC8FFA9FFD5B16BFBC67D6F670C" box="[316,335,1088,1111]" italics="true" pageId="23" pageNumber="424">C</emphasis>
. (
<emphasis id="B906EAC8FFA9FFD5B13DFBC77D5E6703" box="[362,382,1089,1112]" italics="true" pageId="23" pageNumber="424">D</emphasis>
).
<taxonomicName id="4C724D59FFA9FFD5B1CEFBB97DFF6703" box="[409,479,1087,1112]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="alama" status="sp. nov.">
<emphasis id="B906EAC8FFA9FFD5B1CEFBB97DFF6703" box="[409,479,1087,1112]" italics="true" pageId="23" pageNumber="424">alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFA9FFD5B1BEFBB97E65670C" box="[489,581,1087,1112]" pageId="23" pageNumber="424" rank="species">
<emphasis id="B906EAC8FFA9FFD5B1BEFBB97E1F6703" bold="true" box="[489,575,1087,1112]" pageId="23" pageNumber="424">sp. nov</emphasis>
.
</taxonomicNameLabel>
, the
<emphasis id="B906EAC8FFA9FFD5B2D3FBC67EB7670C" box="[644,663,1088,1111]" italics="true" pageId="23" pageNumber="424">C</emphasis>
. (
<emphasis id="B906EAC8FFA9FFD5B2E5FBC77EE66703" box="[690,710,1089,1112]" italics="true" pageId="23" pageNumber="424">D</emphasis>
).
<taxonomicName id="4C724D59FFA9FFD5B2B6FBC77F1D6703" box="[737,829,1089,1112]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="B906EAC8FFA9FFD5B2B6FBC77F1D6703" box="[737,829,1089,1112]" italics="true" pageId="23" pageNumber="424">serratus</emphasis>
</taxonomicName>
of
<bibRefCitation id="EFE34B2BFFA9FFD5B33EFBB9782F6703" author="Ledoyer" box="[873,1039,1087,1112]" pageId="23" pageNumber="424" refString="Ledoyer, M. (1983) Crustaces amphipodes gammariens. Familles des Acanthonotozomatidae a Gammaridae. Faune de Madagascar, 59 (1), 1 - 598." type="journal article" year="1983">Ledoyer, 1983</bibRefCitation>
has a palmar excavation, but that excavation is much narrower than in
<emphasis id="B906EAC8FFA9FFD5B26EFBE37E6C6727" box="[569,588,1125,1148]" italics="true" pageId="23" pageNumber="424">C</emphasis>
. (
<emphasis id="B906EAC8FFA9FFD5B233FBE37E586727" box="[612,632,1125,1148]" italics="true" pageId="23" pageNumber="424">D</emphasis>
).
<taxonomicName id="4C724D59FFA9FFD5B2C7FBE27EF66726" box="[656,726,1124,1149]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="alama" status="sp. nov.">
<emphasis id="B906EAC8FFA9FFD5B2C7FBE27EF66726" box="[656,726,1124,1149]" italics="true" pageId="23" pageNumber="424">alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFA9FFD5B28AFBE27F176726" box="[733,823,1124,1149]" pageId="23" pageNumber="424" rank="species">
<emphasis id="B906EAC8FFA9FFD5B28AFBE27F176726" bold="true" box="[733,823,1124,1149]" pageId="23" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="4C724D59FFA9FFD5B369FBE278716727" box="[830,1105,1124,1149]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="serratus" subGenus="Denticeradocus">
<emphasis id="B906EAC8FFA9FFD5B369FBE27F976726" box="[830,951,1124,1149]" italics="true" pageId="23" pageNumber="424">Ceradocus</emphasis>
(
<emphasis id="B906EAC8FFA9FFD5B39EFBE37FFD6727" box="[969,989,1125,1148]" italics="true" pageId="23" pageNumber="424">D</emphasis>
.)
<emphasis id="B906EAC8FFA9FFD5B3A2FBE378716727" box="[1013,1105,1125,1148]" italics="true" pageId="23" pageNumber="424">serratus</emphasis>
</taxonomicName>
of
<bibRefCitation id="EFE34B2BFFA9FFD5B420FBE279066727" author="Ledoyer" box="[1143,1318,1124,1149]" pageId="23" pageNumber="424" refString="Ledoyer, M. (1983) Crustaces amphipodes gammariens. Familles des Acanthonotozomatidae a Gammaridae. Faune de Madagascar, 59 (1), 1 - 598." type="journal article" year="1983">Ledoyer (1983)</bibRefCitation>
also lacks the palmar shelf characteristic of
<emphasis id="B906EAC8FFA9FFD5B25AFB0E7E0067C4" box="[525,544,1160,1183]" italics="true" pageId="23" pageNumber="424">C</emphasis>
. (
<emphasis id="B906EAC8FFA9FFD5B260FB0F7E6B67FB" box="[567,587,1161,1184]" italics="true" pageId="23" pageNumber="424">D</emphasis>
).
<taxonomicName id="4C724D59FFA9FFD5B235FB017E8867FB" box="[610,680,1159,1184]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="alama" status="sp. nov.">
<emphasis id="B906EAC8FFA9FFD5B235FB017E8867FB" box="[610,680,1159,1184]" italics="true" pageId="23" pageNumber="424">alama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557B3FFA9FFD5B2F8FB017F2867FB" box="[687,776,1159,1184]" pageId="23" pageNumber="424" rank="species">
<emphasis id="B906EAC8FFA9FFD5B2F8FB017F2867FB" bold="true" box="[687,776,1159,1184]" pageId="23" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3686551FFA9FFD5B090FB2A7DE1679E" box="[199,449,1196,1221]" pageId="23" pageNumber="424" type="distribution">
<paragraph id="8BCD36DAFFA9FFD5B090FB2A7DE1679E" blockId="23.[151,1436,151,1221]" box="[199,449,1196,1221]" pageId="23" pageNumber="424">
<emphasis id="B906EAC8FFA9FFD5B090FB2A7D40679E" bold="true" box="[199,352,1196,1221]" pageId="23" pageNumber="424">Distribution.</emphasis>
<collectingCountry id="F365764AFFA9FFD5B130FB2A7D9D679E" box="[359,445,1196,1221]" name="Kuwait" pageId="23" pageNumber="424">Kuwait</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>