234 lines
35 KiB
XML
234 lines
35 KiB
XML
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<mods:title id="6E873EEB27FC759A789E756ECDB95119">On the taxonomy of some South American and Australo-Pacific Anthicidae Latreille, 1819 (Coleoptera) with new subfamily and tribe-rank synonymies</mods:title>
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<mods:namePart id="06700EF3D08546E86F7043E2082F59E9">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="B9B255DACEA692EA20306F1529C09D11">Department of Life Sciences, Natural History Museum, SW 7 5 BD London, United Kingdom & Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401 Daugavpils, Latvia & Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004 Rīga, Latvia</mods:affiliation>
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<mods:title id="CE145AE62B35458E18FB626D4CDB6884">Zootaxa</mods:title>
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<mods:date id="D73558B7B7B5ED6A0CB1ED81CEAB140F">2024</mods:date>
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<paragraph id="8BDE36D23612341A0D84D48BBDC6BAD9" blockId="11.[151,508,896,922]" box="[151,508,896,922]" pageId="11" pageNumber="412">
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<taxonomicName id="4C614D513612341A0D84D48BBDC6BAD9" authority="Champion, 1916" authorityName="Champion" authorityYear="1916" box="[151,508,896,922]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAC03612341A0D84D48BBDC6BAD9" bold="true" box="[151,508,896,922]" pageId="11" pageNumber="412">
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<emphasis id="B915EAC03612341A0D84D48BBD1CBAD9" bold="true" box="[151,294,896,922]" italics="true" pageId="11" pageNumber="412">Ictistygnina</emphasis>
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<bibRefCitation id="EFF04B233612341A0C3DD48BBDC6BAD9" author="Champion, G. C." box="[302,508,896,922]" pageId="11" pageNumber="412" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion, 1916</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B65593612341A0D84D4CFB847BA9F" box="[151,1149,964,989]" pageId="11" pageNumber="412" type="reference_group">
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<paragraph id="8BDE36D23612341A0D84D4CFB847BA9F" blockId="11.[151,1149,964,989]" box="[151,1149,964,989]" pageId="11" pageNumber="412">
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<bibRefCitation id="EFF04B233612341A0D84D4CFBDBABA9E" author="Champion, G. C." box="[151,384,964,989]" pageId="11" pageNumber="412" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion (1916: 190)</bibRefCitation>
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,
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<typeStatus id="54DA88703612341A0C98D4CEBD8DBA9E" box="[395,439,965,989]" pageId="11" pageNumber="412">type</typeStatus>
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species
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<taxonomicName id="4C614D513612341A0F1FD4CEBFAFBA9F" authority="Champion, 1916" authorityName="Champion" authorityYear="1916" box="[524,917,964,988]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="filicornis">
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<emphasis id="B915EAC03612341A0F1FD4CEBEDCBA9F" box="[524,742,964,988]" italics="true" pageId="11" pageNumber="412">Ictistygnina filicornis</emphasis>
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<bibRefCitation id="EFF04B233612341A0FFFD4CFBFAFBA9F" author="Champion, G. C." box="[748,917,964,988]" pageId="11" pageNumber="412" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion, 1916</bibRefCitation>
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</taxonomicName>
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[original designation].
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B65593612341A0D84D300BF38BFCB" pageId="11" pageNumber="412" type="description">
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<paragraph id="8BDE36D23612341A0D84D300BF0CBDF6" blockId="11.[151,1437,1034,1565]" pageId="11" pageNumber="412">
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The genus was originally placed in the
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<taxonomicName id="4C614D513612341A0F2CD300BF6BBD66" authority="Latreille, 1825" authorityName="Latreille" authorityYear="1825" box="[575,849,1034,1061]" class="Insecta" family="Lagriidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="family">Lagriidae Latreille, 1825</taxonomicName>
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(
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<bibRefCitation id="EFF04B233612341A0E4DD300B82FBD66" author="Champion, G. C." box="[862,1045,1035,1061]" pageId="11" pageNumber="412" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion 1916</bibRefCitation>
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), now a subfamily of
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<taxonomicName id="4C614D513612341A09EFD300BD01BD0A" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="family">
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Tenebrionidae Latreille,
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<quantity id="4C999B373612341A0C10D324BD01BD0A" box="[259,315,1071,1097]" metricMagnitude="1" metricUnit="m" metricValue="4.57708" pageId="11" pageNumber="412" unit="in" value="1802.0">1802</quantity>
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</taxonomicName>
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in accordance with the present classification (
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<bibRefCitation id="EFF04B233612341A0E28D324B810BD0A" author="Bouchard, P. & Bousquet, Y. & Aalbu, R. L. & Alonso-Zarazaga, M. A. & Merkl, O. & Davies, A. E." box="[827,1066,1071,1097]" pageId="11" pageNumber="412" pagination="1 - 633" refId="ref12179" refString="Bouchard, P., Bousquet, Y., Aalbu, R. L., Alonso-Zarazaga, M. A., Merkl, O. & Davies, A. E. (2021) Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera). ZooKeys, 1050, 1 - 633. https: // doi. org / 10.3897 / zookeys. 1050.64217" type="journal article" year="2021">
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Bouchard
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<emphasis id="B915EAC03612341A0EA3D324BFDBBD0A" box="[944,993,1071,1097]" italics="true" pageId="11" pageNumber="412">et al</emphasis>
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. 2021
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</bibRefCitation>
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). Current position of the genus in
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<taxonomicName id="4C614D513612341A0D84D358BD2DBD2E" authorityName="Borchmann" authorityYear="1936" box="[151,279,1107,1133]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="402" phylum="Arthropoda" rank="tribe" tribe="Ictistygnini">Ictistygnini</taxonomicName>
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was first proposed by
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<bibRefCitation id="EFF04B233612341A0F1FD358BF72BD2E" author="Borchmann, F." box="[524,840,1106,1133]" pageId="11" pageNumber="412" refId="ref12133" refString="Borchmann, F. (1936) Coleoptera Heteromera Fam. Lagriidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 204. Louis Desmet-Verteneuil, Brussels, Belgium, pp. 561, pls. 9." type="journal volume" year="1936">Borchmann (1936: 534‒535</bibRefCitation>
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; as ‘Ictistygninae’), which was followed subsequently by several authors. The genus is considered to be monotypic:
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<taxonomicName id="4C614D513612341A0E47D37CB87FBDD2" authorityName="Champion" authorityYear="1916" box="[852,1093,1143,1169]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="filicornis">
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<emphasis id="B915EAC03612341A0E47D37CB87FBDD2" box="[852,1093,1143,1169]" italics="true" pageId="11" pageNumber="412">Ictistygnina filicornis</emphasis>
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</taxonomicName>
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remains known only from the state of
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<collectingRegion id="49A5F8303612341A0DE3D390BDA8BDF6" box="[240,402,1179,1205]" country="Brazil" name="Rio de Janeiro" pageId="11" pageNumber="412">Rio de Janeiro</collectingRegion>
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in southern
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<collectingCountry id="F37676423612341A0F0ED390BE5BBDF6" box="[541,609,1179,1205]" name="Brazil" pageId="11" pageNumber="412">Brazil</collectingCountry>
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(
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<bibRefCitation id="EFF04B233612341A0F63D390BF13BDF6" author="Champion, G. C." box="[624,809,1179,1205]" pageId="11" pageNumber="412" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion 1916</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BDE36D23612341A0DD4D3B4BE14BC96" blockId="11.[151,1437,1034,1565]" pageId="11" pageNumber="412">
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<bibRefCitation id="EFF04B233612341A0DD4D3B4BDC0BD9A" author="Champion, G. C." box="[199,506,1215,1241]" pageId="11" pageNumber="412" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion (1916: 190‒191)</bibRefCitation>
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defined
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<taxonomicName id="4C614D513612341A0F4BD3B4BEE6BD9B" authorityName="Champion" authorityYear="1916" box="[600,732,1215,1240]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAC03612341A0F4BD3B4BEE6BD9B" box="[600,732,1215,1240]" italics="true" pageId="11" pageNumber="412">Ictistygnina</emphasis>
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</taxonomicName>
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based on the following set of morphological features: ‘narrowly closed anterior coxal cavities’, ‘extremely elongate antennae, large eyes’, ‘difference in the size of the eyes in the two sexes’ and ‘very small second joint to the unusually elongate, filiform antennae’. The new genus was considered to exhibit similarities with the Australian
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<taxonomicName id="4C614D513612341A0F7FD220BED6BC06" authorityName="Borchmann" authorityYear="1936" box="[620,748,1323,1349]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="402" phylum="Arthropoda" rank="tribe" tribe="Ictistygnini">Ictistygnini</taxonomicName>
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(in that time sense) genus
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<taxonomicName id="4C614D513612341A090FD220B91CBC06" authority="Pascoe, 1866" authorityName="Pascoe" authorityYear="1866" box="[1052,1318,1323,1349]" class="Insecta" family="Anthicidae" genus="Ictistygna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAC03612341A090FD220B8B0BC07" box="[1052,1162,1323,1348]" italics="true" pageId="11" pageNumber="412">Ictistygna</emphasis>
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Pascoe, 1866
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</taxonomicName>
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as well as the Calopodinae A.
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<collectingRegion id="49A5F8303612341A0C6AD244BD86BC2A" box="[377,444,1359,1385]" country="New Zealand" name="West Coast" pageId="11" pageNumber="412">Costa</collectingRegion>
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, 1852 genus
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<taxonomicName id="4C614D513612341A0F41D244BF5DBC2A" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[594,871,1358,1385]" class="Insecta" family="Oedemeridae" genus="Calopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAC03612341A0F41D244BE95BC2A" box="[594,687,1359,1385]" italics="true" pageId="11" pageNumber="412">Calopus</emphasis>
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Fabricius, 1775
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</taxonomicName>
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(
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<taxonomicName id="4C614D513612341A0E6AD244B887BC2A" authority="Latreille, 1810" authorityName="Latreille" authorityYear="1810" box="[889,1213,1359,1385]" class="Insecta" family="Oedemeridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="family">Oedemeridae Latreille, 1810</taxonomicName>
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) (
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<bibRefCitation id="EFF04B233612341A09C6D244B9AABC2A" author="Champion, G. C." box="[1237,1424,1359,1385]" pageId="11" pageNumber="412" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion 1916</bibRefCitation>
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||
).
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<bibRefCitation id="EFF04B233612341A0D84D278BD91BCCE" author="Borchmann, F." box="[151,427,1394,1421]" pageId="11" pageNumber="412" refId="ref12133" refString="Borchmann, F. (1936) Coleoptera Heteromera Fam. Lagriidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 204. Louis Desmet-Verteneuil, Brussels, Belgium, pp. 561, pls. 9." type="journal volume" year="1936">Borchmann (1936: 534)</bibRefCitation>
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erected a new subfamily ‘Ictistygninae’ in what was
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<taxonomicName id="4C614D513612341A0909D278B8BFBCCE" authorityName="Latreille" authorityYear="1825" box="[1050,1157,1395,1421]" class="Insecta" family="Lagriidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="family">Lagriidae</taxonomicName>
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at that time, and which was downgraded to the rank of a tribe,
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<taxonomicName id="4C614D513612341A0F5CD29CBEEBBCF2" authorityName="Borchmann" authorityYear="1936" box="[591,721,1431,1457]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="402" phylum="Arthropoda" rank="tribe" tribe="Ictistygnini">Ictistygnini</taxonomicName>
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, in
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<taxonomicName id="4C614D513612341A0FE9D29CBFA8BCF2" authorityName="LeConte" authorityYear="1862" box="[762,914,1431,1457]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="subFamily" subFamily="Eurygeniinae">Eurygeniinae</taxonomicName>
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,
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<taxonomicName id="4C614D513612341A0E8FD29CB82DBCF2" authorityName="Latreille" authorityYear="1819" box="[924,1047,1431,1457]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
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by
|
||
<bibRefCitation id="EFF04B233612341A0953D29CB90FBCF2" author="Abdullah, M." box="[1088,1333,1431,1457]" pageId="11" pageNumber="412" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah (1969: 340)</bibRefCitation>
|
||
, where it remains up to now (
|
||
<bibRefCitation id="EFF04B233612341A0C66D2B0BE18BC96" author="Chandler, D. S." box="[373,546,1467,1493]" pageId="11" pageNumber="412" refId="ref12485" refString="Chandler, D. S. (2010) 11.26. Anthicidae Latreille, 1819. In: Leschen, R. A. B., Beutel, R. G. & Lawrence, J. F. (Eds.), Coleoptera, Beetles. Vol. 2. Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). Arthropoda Insecta. Handbook of Zoology. De Gruyter, Berlin and New York, pp. 729 - 741. https: // doi. org / 10.1515 / 9783110911213.729" type="book" year="2010">Chandler 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDE36D23612341A0DD4D2D4B85FBF5E" blockId="11.[151,1437,1034,1565]" pageId="11" pageNumber="412">
|
||
In this paper I do not aim to review all features of eurygeniine genera or provide a more comprehensive comparative diagnosis for
|
||
<taxonomicName id="4C614D513612341A0CD2D108BE7FBF5F" authorityName="Champion" authorityYear="1916" box="[449,581,1539,1564]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAC03612341A0CD2D108BE7FBF5F" box="[449,581,1539,1564]" italics="true" pageId="11" pageNumber="412">Ictistygnina</emphasis>
|
||
</taxonomicName>
|
||
but provide only the redescription of this group.
|
||
</paragraph>
|
||
<paragraph id="8BDE36D23612341A0D84D165BF38BFCB" blockId="11.[151,770,1646,1673]" box="[151,770,1646,1673]" pageId="11" pageNumber="412">
|
||
<heading id="D09681BE3612341A0D84D165BF38BFCB" bold="true" box="[151,770,1646,1673]" fontSize="11" level="1" pageId="11" pageNumber="412" reason="1">
|
||
<emphasis id="B915EAC03612341A0D84D165BF38BFCB" bold="true" box="[151,770,1646,1673]" pageId="11" pageNumber="412">
|
||
Supplementary description of
|
||
<taxonomicName id="4C614D513612341A0F01D164BE9BBFCA" authorityName="Champion" authorityYear="1916" box="[530,673,1647,1673]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAC03612341A0F01D164BE9BBFCA" bold="true" box="[530,673,1647,1673]" italics="true" pageId="11" pageNumber="412">Ictistygnina</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="135A2A573612341A0FA2D165BEC3BFCB" box="[689,761,1646,1672]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[192,1396,182,1920]" captionTargetId="figure-40@12.[192,1396,181,1920]" captionTargetPageId="12" captionText="FIGURE 5. Ictistygnina filicornis Champion, 1916. A—Syntype ♂, habitus, dorsal view; B—ditto, forebody, antero-dorsal view: C—Syntype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13628179" httpUri="https://zenodo.org/record/13628179/files/figure.png" pageId="11" pageNumber="412">Fig. 5</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B65593612341A0D84D1BCBCD3BEEA" pageId="11" pageNumber="412" type="materials_examined">
|
||
<paragraph id="8BDE36D23612341A0D84D1BCBCD3BEEA" blockId="11.[151,1437,1718,1961]" pageId="11" pageNumber="412">
|
||
<emphasis id="B915EAC03612341A0D84D1BCBD80BF93" bold="true" box="[151,442,1718,1745]" pageId="11" pageNumber="412">Type material examined.</emphasis>
|
||
<typeStatus id="54DA88703612341A0CADD1BCBE20BF92" box="[446,538,1719,1745]" pageId="11" pageNumber="412" type="syntype">Syntype</typeStatus>
|
||
<specimenCount id="9D67FD5B3612341A0F0DD1BCBE7BBF92" box="[542,577,1719,1745]" count="1" pageId="11" pageNumber="412" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED70AE173612341A0F55D1BCBEA0BF92" box="[582,666,1719,1745]" pageId="11" pageNumber="412">BMNH</collectionCode>
|
||
(
|
||
<figureCitation id="135A2A573612341A0FB5D1BCBF1ABF92" box="[678,800,1718,1745]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[192,1396,182,1920]" captionTargetId="figure-40@12.[192,1396,181,1920]" captionTargetPageId="12" captionText="FIGURE 5. Ictistygnina filicornis Champion, 1916. A—Syntype ♂, habitus, dorsal view; B—ditto, forebody, antero-dorsal view: C—Syntype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13628179" httpUri="https://zenodo.org/record/13628179/files/figure.png" pageId="11" pageNumber="412">Fig. 5A‒B</figureCitation>
|
||
): Fry RioJan [handwritten] // Fry Coll. 1905.100. [printed] // Sp. figured. [printed] //
|
||
<specimenCount id="9D67FD5B3612341A0CD0D1D0BDE2BFB6" box="[451,472,1755,1781]" count="1" pageId="11" pageNumber="412" type="male">♂</specimenCount>
|
||
[printed] //
|
||
<taxonomicName id="4C614D513612341A0F76D1D0BFA3BFB6" authority=", Ch." authorityName="Ch." box="[613,921,1755,1781]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="filicornis">Ictistygnina filicornis, Ch.</taxonomicName>
|
||
[handwritten] // dorsal BM 17 [handwritten] //
|
||
<typeStatus id="54DA88703612341A0DBCD1F4BD17BE5A" box="[175,301,1791,1817]" pageId="11" pageNumber="412" type="syntype">SYNTYPE</typeStatus>
|
||
[printed, label circular, blue frame];
|
||
<typeStatus id="54DA88703612341A0FC7D1F4BF11BE5A" box="[724,811,1791,1817]" pageId="11" pageNumber="412" type="syntype">syntype</typeStatus>
|
||
<specimenCount id="9D67FD5B3612341A0E27D1F4BF6DBE5A" box="[820,855,1791,1817]" count="1" pageId="11" pageNumber="412" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED70AE173612341A0E73D1F4BF8EBE5A" box="[864,948,1791,1817]" pageId="11" pageNumber="412">BMNH</collectionCode>
|
||
(
|
||
<figureCitation id="135A2A573612341A0ED7D1F4B878BE5A" box="[964,1090,1790,1817]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[192,1396,182,1920]" captionTargetId="figure-40@12.[192,1396,181,1920]" captionTargetPageId="12" captionText="FIGURE 5. Ictistygnina filicornis Champion, 1916. A—Syntype ♂, habitus, dorsal view; B—ditto, forebody, antero-dorsal view: C—Syntype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13628179" httpUri="https://zenodo.org/record/13628179/files/figure.png" pageId="11" pageNumber="412">Fig. 5C‒D</figureCitation>
|
||
): Fry
|
||
<superScript id="7C149B9A3612341A0995D1F4B8E6BE4E" attach="left" box="[1158,1244,1791,1817]" fontSize="6" pageId="11" pageNumber="412">RioJano</superScript>
|
||
. [handwritten] // Fry Coll. 1905.100. [printed] //
|
||
<specimenCount id="9D67FD5B3612341A0CE9D028BE35BE7E" box="[506,527,1827,1853]" count="1" pageId="11" pageNumber="412" type="female">♀</specimenCount>
|
||
[printed] //
|
||
<taxonomicName id="4C614D513612341A0F85D028BF87BE7E" authority=", Ch." authorityName="Ch." box="[662,957,1827,1853]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="filicornis">Ictistygnina filicornis, Ch.</taxonomicName>
|
||
[handwritten] //
|
||
<typeStatus id="54DA88703612341A0969D028B8C2BE7E" box="[1146,1272,1827,1853]" pageId="11" pageNumber="412" type="syntype">SYNTYPE</typeStatus>
|
||
[printed, label circular, blue frame];
|
||
<typeStatus id="54DA88703612341A0C83D04CBDDDBE22" box="[400,487,1863,1889]" pageId="11" pageNumber="412" type="syntype">syntype</typeStatus>
|
||
<specimenCount id="9D67FD5B3612341A0CE3D04CBE29BE22" box="[496,531,1863,1889]" count="1" pageId="11" pageNumber="412" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED70AE173612341A0F0FD04CBE4EBE22" box="[540,628,1863,1889]" pageId="11" pageNumber="412">BMNH</collectionCode>
|
||
: Fry RioJan [handwritten] // 2992 [handwritten] // Fry Coll. 1905.100. [printed] //
|
||
<specimenCount id="9D67FD5B3612341A0C33D060BD0FBEC6" box="[288,309,1899,1925]" count="1" pageId="11" pageNumber="412" type="male">♂</specimenCount>
|
||
[printed] //
|
||
<taxonomicName id="4C614D513612341A0CDAD060BF3EBEC6" authority=", Ch." authorityName="Ch." box="[457,772,1899,1925]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="filicornis">Ictistygnina filicornis, Ch.</taxonomicName>
|
||
[handwritten] //
|
||
<typeStatus id="54DA88703612341A0EDDD060B876BEC6" box="[974,1100,1899,1925]" pageId="11" pageNumber="412" type="syntype">SYNTYPE</typeStatus>
|
||
[printed, label circular, blue frame].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1E665A3615341D0D84D09CB81CBE97" ID-DOI="http://doi.org/10.5281/zenodo.13628179" ID-Zenodo-Dep="13628179" httpUri="https://zenodo.org/record/13628179/files/figure.png" pageId="12" pageNumber="413" startId="12.[151,250,1943,1967]" targetBox="[192,1396,182,1920]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BDE36D23615341D0D84D09CB81CBE97" blockId="12.[151,1437,1943,2004]" pageId="12" pageNumber="413">
|
||
<emphasis id="B915EAC03615341D0D84D09CBD2FBEF3" bold="true" box="[151,277,1943,1968]" pageId="12" pageNumber="413">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C614D513615341D0C0DD093BE88BEF3" authority="Champion, 1916" authorityName="Champion" authorityYear="1916" box="[286,690,1944,1968]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="filicornis">
|
||
<emphasis id="B915EAC03615341D0C0DD093BDC1BEF3" box="[286,507,1944,1968]" italics="true" pageId="12" pageNumber="413">Ictistygnina filicornis</emphasis>
|
||
<bibRefCitation id="EFF04B233615341D0F10D093BE88BEF3" author="Champion, G. C." box="[515,690,1944,1968]" pageId="12" pageNumber="413" pagination="181 - 242" refId="ref12339" refString="Champion, G. C. (1916) On new and little-known Lagriidae and Pedilidae. Transactions of the Royal Entomological Society of London, 64 (2), 181 - 242, pls. lxii - lvix. https: // doi. org / 10.1111 / j. 1365 - 2311.1916. tb 03127. x" type="journal article" year="1916">Champion, 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
. A—Syntype ♂, habitus, dorsal view; B—ditto, forebody, antero-dorsal view: C—Syntype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale].
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37B65593614341C0DD4D79CBFBBBF4A" pageId="13" pageNumber="414" type="description">
|
||
<paragraph id="8BDE36D23614341C0DD4D79CBDCCBC16" blockId="13.[151,1437,151,1545]" pageId="13" pageNumber="414">
|
||
Frontoclypeal suture not observed. Mandible apex unidentate, rounded (broadly scooped). Right mandible with an indistinct indentation at inner edge at some distance from apex. Terminal maxillary palpomere subcultriform. Compound eye nearly holoptic, occupying nearly whole head side posterior to antennal insertion, shallowly notched at anterior margin at antennal insertion; posterior eye margin subtruncate to shallowly emarginate.Antennal insertion touching anterior margin of a compound eye in male, slightly distant from it in female. Antenna extraordinarily long and slender, in male extending beyond elytral apex when directed posteriad, in female extending to slightly beyond metacoxal cavity. Antennomeres 3‒11 with moderately long, dense, suberect setae. Antennomere two strongly shortened, slightly transverse to about as long as wide. Terminal antennomere not modified, long and slender. Cranial ‘neck’ about one third head width across compound eyes. Pronotum about as wide as head across compound eyes, about as long (excluding anterior flange) as wide (about 1.2× as long as wide including anterior flange), lateral margins broadly rounded. Anterior pronotal flange distinct and broad dorsally, nearly same wide ventrally as dorsally, dorsally somewhat elevated (compared to a pronotal disc) and projected anteriad to conceal most of cranial ‘neck’. Suture of anterior flange incision-like deep and narrow laterally, shallow and rather broad dorsally. Basal pronotal margin dorsally with a deep and rather wide sulcus which widens on lateral margins. Elytra flattened, strongly elongate. Epipleuron narrow, incomplete. At least distal half of each tibia spinose. Tibial terminal spurs paired, elongate, straight to barely curved, margins smooth, apically acutely angulate. Penultimate tarsomere short, distinctly bilobate. Pretarsal claws rather long, each with an indistinct basal indentation. Procoxae elongate, contiguous. Procoxal cavity narrowly open externally behind (propleura narrowly separated or come in contact or even slightly overlap depending on the condition of the specimen but are never fused together in the sense used in the contemporary morphological terminology of Tenebrionoidea), closed internally. Postcoxal plate not present. Mesanepisterna meet in front of a triangular mesoventrite, shortly produced anteriorly at place of meeting, building a flat, acutely angulate anterior projection, with a distinct short longitudinal suture at place of meeting. Mesanepisternum expanded laterally, separated from mesoventrite by a distinct, nearly straight suture; a distinct, flat, rugose ventral sulcus separates anterior portion of mesanepisternum from its main portion; less prominent postmarginal and distinct accessory carina present, separated by ventral sulcus (both carinae, separated, by much narrower ventral sulcus, appear also in
|
||
<taxonomicName id="4C614D513614341C0F46D310BEF9BD77" authorityName="Pascoe" authorityYear="1866" box="[597,707,1051,1076]" class="Insecta" family="Anthicidae" genus="Ictistygna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAC03614341C0F46D310BEF9BD77" box="[597,707,1051,1076]" italics="true" pageId="13" pageNumber="414">Ictistygna</emphasis>
|
||
</taxonomicName>
|
||
); procoxal rest not present; area between anterior and main part of mesanepisternum glossy and glabrous, slightly globose in ventral aspect; posterior margin of mesanepisternum with a row of long, dense setae which continues over along posterior and lateral (external) margins of mesepimeron; most of mesanepisternum glabrous, with sporadic setae. Intercoxal process of mesoventrite narrow, long, elevated in ventral aspect, meets an acute anterior process of metaventrite at about posterior third of mesocoxal cavity; mesocoxal cavities separated. Mesepimeron distinct, broadly subtriangular; long setae not visible in dorsal view. Metaventrite longer than mesosternum, discrimen not observed. Metanepisternum long, narrow. Metacoxal cavities narrowly separated by an acute anterior process of first visible abdominal segment. Male morphological sternite VII emarginate at posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BDE36D23614341C0DD4D255BFBBBF4A" blockId="13.[151,1437,151,1545]" pageId="13" pageNumber="414">
|
||
<emphasis id="B915EAC03614341C0DD4D255BD8DBC3B" bold="true" box="[199,439,1374,1400]" pageId="13" pageNumber="414">Sexual dimorphism.</emphasis>
|
||
<taxonomicName id="4C614D513614341C0CADD254BE78BC3B" authorityName="Champion" authorityYear="1916" box="[446,578,1375,1400]" class="Insecta" family="Anthicidae" genus="Ictistygnina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAC03614341C0CADD254BE78BC3B" box="[446,578,1375,1400]" italics="true" pageId="13" pageNumber="414">Ictistygnina</emphasis>
|
||
</taxonomicName>
|
||
exhibits a strong sexual dimorphism (
|
||
<figureCitation id="135A2A573614341C0EFCD254B809BC3A" box="[1007,1075,1374,1401]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[192,1396,182,1920]" captionTargetId="figure-40@12.[192,1396,181,1920]" captionTargetPageId="12" captionText="FIGURE 5. Ictistygnina filicornis Champion, 1916. A—Syntype ♂, habitus, dorsal view; B—ditto, forebody, antero-dorsal view: C—Syntype ♀, habitus, dorsal view; D—ditto, forebody, dorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13628179" httpUri="https://zenodo.org/record/13628179/files/figure.png" pageId="13" pageNumber="414">Fig. 5</figureCitation>
|
||
). Female frons is comparatively wider, compound eyes are smaller (and, therefore, cranial ‘neck’ is wider than that of male, about half head width, postocular area longer than in male. Female antenna significantly shorter than that of male, not extending to elytral apex. Female terminal maxillary palpomere is less elongate, more strongly securiform (but still subcultriform in the sense of the commonly used anthicid morphological terminology).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |