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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1047.64034" ID-GBIF-Dataset="fe762bd5-f8cc-4886-b09f-9ab5bc7bbcec" ID-PMC="PMC8257564" ID-Pensoft-Pub="1313-2970-1047-61" ID-Pensoft-UUID="23C3761CE1BF5BFAA5301C213FB1D457" ID-PubMed="34248366" ID-ZooBank="835A7E2B82E845B09E3FB8548031A048" ModsDocID="1313-2970-1047-61" checkinTime="1624985784966" checkinUser="pensoft" docAuthor="Falniowski, Andrzej, Lewarne, Brian, Rysiewska, Aleksandra, Osikowski, Artur &amp; Hofman, Sebastian" docDate="2021" docId="E38D4FA4D7D45A4F9432FD2EA6679075" docLanguage="en" docName="ZooKeys 1047: 61-89" docOrigin="ZooKeys 1047" docPubDate="2021-06-28" docSource="http://dx.doi.org/10.3897/zookeys.1047.64034" docTitle="Iglicopsis butoti Falniowski &amp; Hofman 2021, sp. nov." docType="treatment" docUuid="C1A9D0B0-4B10-4977-B69B-7C4C42BB19D3" docUuidSource="ZooBank" docVersion="4" id="23C3761CE1BF5BFAA5301C213FB1D457" lastPageNumber="61" masterDocId="23C3761CE1BF5BFAA5301C213FB1D457" masterDocTitle="Crenobiont, stygophile and stygobiont molluscs in the hydrographic area of the Trebisnjica River Basin" masterLastPageNumber="89" masterPageNumber="61" pageNumber="61" updateTime="1668150568600" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Crenobiont, stygophile and stygobiont molluscs in the hydrographic area of the Trebisnjica River Basin</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Falniowski, Andrzej</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3899-6857</mods:nameIdentifier>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387 Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lewarne, Brian</mods:namePart>
<mods:affiliation>The Devon Karst Research Society, Library &amp; Office, 46, Morley Court, Western Approach, Plymouth, Devon, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rysiewska, Aleksandra</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387 Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Osikowski, Artur</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
<mods:affiliation>Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059 Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hofman, Sebastian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6044-3055</mods:nameIdentifier>
<mods:affiliation>Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387 Krakow, Poland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1047</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>89</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1047.64034</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1047.64034</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1047-61</mods:identifier>
<mods:identifier type="ZooBank">835A7E2B82E845B09E3FB8548031A048</mods:identifier>
<mods:identifier type="Pensoft-UUID">23C3761CE1BF5BFAA5301C213FB1D457</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="183203259" LSID="urn:lsid:zoobank.org:act:C1A9D0B0-4B10-4977-B69B-7C4C42BB19D3" httpUri="http://treatment.plazi.org/id/E38D4FA4D7D45A4F9432FD2EA6679075" lastPageNumber="61" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="http://zoobank.org/C1A9D0B0-4B10-4977-B69B-7C4C42BB19D3" authority="Falniowski &amp; Hofman" authorityName="Falniowski &amp; Hofman" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Iglicopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iglicopsis butoti" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="species" species="butoti" status="sp. nov.">Iglicopsis butoti Falniowski &amp; Hofman</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="61">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Shells: A-L Travunijana vruljakensis M-P Iglicopsis butoti M holotype N 2 F 61 O 2 F 68 P 2 F 69 (extraction numbers, see Table 3). Scale bars: 1 mm." figureDoi="10.3897/zookeys.1047.64034.figure9" httpUri="https://binary.pensoft.net/fig/560481" pageId="0" pageNumber="61">Fig. 9M-P</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="genbank no">
<paragraph pageId="0" pageNumber="61">GenBank no.</paragraph>
<paragraph pageId="0" pageNumber="61">COI: MW879273-MW879275; H3: MW865749-MW865751</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="type material">
<paragraph pageId="0" pageNumber="61">Type materials.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Holotype.</emphasis>
Ethanol-fixed specimen (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Shells: A-L Travunijana vruljakensis M-P Iglicopsis butoti M holotype N 2 F 61 O 2 F 68 P 2 F 69 (extraction numbers, see Table 3). Scale bars: 1 mm." figureDoi="10.3897/zookeys.1047.64034.figure9" httpUri="https://binary.pensoft.net/fig/560481" pageId="0" pageNumber="61">9M</figureCitation>
), Vrelo
<normalizedToken originalValue="„Polički">&quot;Policki</normalizedToken>
Studenac&quot; (Crkvina); a cave spring in the left bank of and adjacent to the
<normalizedToken originalValue="Trebišnjica">Trebisnjica</normalizedToken>
River (
<geoCoordinate degrees="42.71288" direction="north" orientation="latitude" precision="1" value="42.71288">N 42.71288</geoCoordinate>
,
<geoCoordinate degrees="18.36514" direction="east" orientation="longitude" precision="1" value="18.36514">E 18.36514</geoCoordinate>
) (our locality 13, Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of the studied taxa. Localities' numbers correspond to Table 1." figureDoi="10.3897/zookeys.1047.64034.figure4" httpUri="https://binary.pensoft.net/fig/560476" pageId="0" pageNumber="61">4</figureCitation>
) close to Trebinje (Bosnia and Herzegovina), interstitially, 50 cm below the gravel floor of the spring; in the collection of the Department of Malacology of Jagiellonian University, voucher number ZMUJ-M.2651.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Paratypes.</emphasis>
Three paratypes destroyed to extract DNA, one specimen ethanol-fixed, in the collection of the Department of Malacology of Jagiellonian University, ZMUJ-M.2652.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="61">
Shell minute, ovate-conic, distinguishable from
<taxonomicName authorityName="Pesic &amp; Gloer" authorityYear="2013" class="Gastropoda" family="Hydrobiidae" genus="Montenegrospeum" kingdom="Animalia" lsidName="Montenegrospeum" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Montenegrospeum</emphasis>
</taxonomicName>
by a more oval habitus, broader spire and broader flat apex, sometimes showing scalarity at the body whorl; the penis with the left-side outgrowth located more proximally and bi-lobed, and additional flat outgrowth on the right side.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Shell</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Shells: A-L Travunijana vruljakensis M-P Iglicopsis butoti M holotype N 2 F 61 O 2 F 68 P 2 F 69 (extraction numbers, see Table 3). Scale bars: 1 mm." figureDoi="10.3897/zookeys.1047.64034.figure9" httpUri="https://binary.pensoft.net/fig/560481" pageId="0" pageNumber="61">9M-P</figureCitation>
) up to 1.49 mm high and 0.55 mm broad, ovate-conic, whitish, translucent, thin-walled, and consisting of about five whorls, growing regularly and separated by moderately deep suture. Spire high and broad, apex broad and flat, body whorl less than 0.5 of the shell height, Aperture small, prosocline, oval in shape, peristome complete and thin, somewhat swollen, in contact with the wall of the body whorl, in some specimens showing scalarity close to the aperture, umbilicus slit-like. Shell surface smooth, with growth lines hardly visible.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Measurements</emphasis>
of holotype and sequenced and illustrated shells: Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Shell measurements (in mm) of holotype and sequenced and illustrated specimens of Iglicopsis butoti sp. nov. For explanation of the symbols a - β, see Fig. 13 B." httpUri="http://table.plazi.org/id/E30E1986295F1C38B7A415F054EAE2F5" pageId="0" pageNumber="61" tableUuid="E30E1986295F1C38B7A415F054EAE2F5">3</tableCitation>
. Shell variability slight; scalarity and much bigger dimensions of one specimen (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Shells: A-L Travunijana vruljakensis M-P Iglicopsis butoti M holotype N 2 F 61 O 2 F 68 P 2 F 69 (extraction numbers, see Table 3). Scale bars: 1 mm." figureDoi="10.3897/zookeys.1047.64034.figure9" httpUri="https://binary.pensoft.net/fig/560481" pageId="0" pageNumber="61">9P</figureCitation>
) most probably caused by the larval
<taxonomicName authorityName="Pesic &amp; Gloer" authorityYear="2013" class="Trematoda" kingdom="Animalia" lsidName="" pageId="0" pageNumber="61" phylum="Mollusca" rank="class">Trematoda</taxonomicName>
(parasite gigantism).
</paragraph>
<caption ID-Table-UUID="E30E1986295F1C38B7A415F054EAE2F5" httpUri="http://table.plazi.org/id/E30E1986295F1C38B7A415F054EAE2F5" pageId="0" pageNumber="61" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Table 3.</emphasis>
Shell measurements (in mm) of holotype and sequenced and illustrated specimens of
<taxonomicName authorityName="Falniowski &amp; Hofman" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Iglicopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iglicopsis butoti" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="species" species="butoti">
<emphasis italics="true" pageId="0" pageNumber="61">Iglicopsis butoti</emphasis>
</taxonomicName>
sp. nov. For explanation of the symbols
<emphasis italics="true" pageId="0" pageNumber="61">a</emphasis>
-β, see Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A Penis of Iglicopsis butoti, scale bar: 0.1 mm B shell measurements: a - shell height, b - body whorl breadth, c - aperture height, d - spire height, e - aperture breadth, α - apex angle, β - angle between body whorl suture and horizontal surface." figureDoi="10.3897/zookeys.1047.64034.figure13" httpUri="https://binary.pensoft.net/fig/560485" pageId="0" pageNumber="61">13B</figureCitation>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<table pageId="0" pageNumber="61">
<tr pageId="0" pageNumber="61">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="61" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Holotype</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">2F61</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">2F68</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">2F69</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="61">a</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.49</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.87</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="61">b</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.55</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.54</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.54</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.70</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="61">c</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.43</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.39</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.43</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.44</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="61">d</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.80</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.62</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.67</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.93</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="61">e</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.37</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.34</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.35</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">0.44</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<normalizedToken originalValue="α">α</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">90</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">89</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">90</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">90</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<normalizedToken originalValue="β">β</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">18</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">18</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Soft parts morphology and anatomy</emphasis>
. Body white, pigmentless, with no eyes. Ctenidium with nine short lamellae, osphradium elongated. Tectum forming a characteristic broad loop (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Shells: A-L Travunijana vruljakensis M-P Iglicopsis butoti M holotype N 2 F 61 O 2 F 68 P 2 F 69 (extraction numbers, see Table 3). Scale bars: 1 mm." figureDoi="10.3897/zookeys.1047.64034.figure9" httpUri="https://binary.pensoft.net/fig/560481" pageId="0" pageNumber="61">9N</figureCitation>
). Female reproductive organs with unpigmented renal oviduct, bursa copulatrix and two small receptacula seminis; details unknown.
</paragraph>
<paragraph pageId="0" pageNumber="61">
The radula (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Radula of Iglicopsis butoti, scale bars: 10 µm." figureDoi="10.3897/zookeys.1047.64034.figure12" httpUri="https://binary.pensoft.net/fig/560484" pageId="0" pageNumber="61">12</figureCitation>
) with the central tooth cusp formula:
</paragraph>
<paragraph pageId="0" pageNumber="61">(4)3-1-3(4)1-1 or (5)4-1-4(5)1-1</paragraph>
<caption doi="10.3897/zookeys.1047.64034.figure12" httpUri="https://binary.pensoft.net/fig/560484" pageId="0" pageNumber="61" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 12.</emphasis>
Radula of
<taxonomicName authorityName="Falniowski &amp; Hofman" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Iglicopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iglicopsis butoti" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="species" species="butoti">
<emphasis italics="true" pageId="0" pageNumber="61">Iglicopsis butoti</emphasis>
</taxonomicName>
, scale bars: 10
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">Rather long and slender cusps grow regularly to central one. Lateral cusp with 5 - 1 - 5(6) long and massive cusps. Inner marginal tooth with ca 23 slender cusps of nearly invariable length along the tooth edge, outer marginal tooth with 26 broadly triangular cusps.</paragraph>
<paragraph pageId="0" pageNumber="61">
Penis (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A Penis of Iglicopsis butoti, scale bar: 0.1 mm B shell measurements: a - shell height, b - body whorl breadth, c - aperture height, d - spire height, e - aperture breadth, α - apex angle, β - angle between body whorl suture and horizontal surface." figureDoi="10.3897/zookeys.1047.64034.figure13" httpUri="https://binary.pensoft.net/fig/560485" pageId="0" pageNumber="61">13A</figureCitation>
) long, tapering, below the half of its length, proximally, bi-lobed outgrowth on the left side and flat outgrowth at the right side, at the distal part and the vas deferens well visible inside, running in zigzags.
</paragraph>
<caption doi="10.3897/zookeys.1047.64034.figure13" httpUri="https://binary.pensoft.net/fig/560485" pageId="0" pageNumber="61" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 13.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
Penis of
<taxonomicName authorityName="Falniowski &amp; Hofman" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Iglicopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iglicopsis butoti" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="species" species="butoti">
<emphasis italics="true" pageId="0" pageNumber="61">Iglicopsis butoti</emphasis>
</taxonomicName>
, scale bar: 0.1 mm
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
shell measurements:
<emphasis italics="true" pageId="0" pageNumber="61">a</emphasis>
- shell height,
<emphasis italics="true" pageId="0" pageNumber="61">b</emphasis>
- body whorl breadth,
<emphasis italics="true" pageId="0" pageNumber="61">c</emphasis>
- aperture height,
<emphasis italics="true" pageId="0" pageNumber="61">d</emphasis>
- spire height,
<emphasis italics="true" pageId="0" pageNumber="61">e</emphasis>
- aperture breadth,
<normalizedToken originalValue="α">α</normalizedToken>
- apex angle,
<normalizedToken originalValue="β">β</normalizedToken>
- angle between body whorl suture and horizontal surface.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="etymology">
<paragraph pageId="0" pageNumber="61">Derivatio nominis.</paragraph>
<paragraph pageId="0" pageNumber="61">
The genus name refers to the similarity of the shell to the moitessieriid genus
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Iglica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iglica" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Iglica</emphasis>
</taxonomicName>
Wagner, 1927. The specific epithet
<taxonomicName authorityName="Falniowski &amp; Hofman" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Iglicopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iglicopsis butoti" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="species" species="butoti">
<emphasis italics="true" pageId="0" pageNumber="61">Iglicopsis butoti</emphasis>
</taxonomicName>
refers to the memory of Dr Louis J. M. Butot, a Dutch malacologist devoted mostly to the Greek malacofauna, good friend and the mentor of AF.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="distribution">
<paragraph pageId="0" pageNumber="61">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="61">Known from the type locality only.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="molecular relationships">
<paragraph pageId="0" pageNumber="61">Molecular relationships.</paragraph>
<paragraph pageId="0" pageNumber="61">
despite its shell morphology,
<taxonomicName authorityName="Falniowski &amp; Hofman" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Iglicopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iglicopsis" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Iglicopsis</emphasis>
</taxonomicName>
clearly belongs to the
<taxonomicName class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
Stimpson, 1865,
<taxonomicName authorityName="Radoman" authorityYear="1973" lsidName="" pageId="0" pageNumber="61" rank="subFamily" subFamily="Sadlerianinae">Sadlerianinae</taxonomicName>
Szarowska, 2006, and not to the
<taxonomicName authorityName="Bourguignat" authorityYear="1863" class="Gastropoda" family="Moitessieriidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="family">Moitessieriidae</taxonomicName>
Bourguignat, 1863 (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phylogenetic relationships of Travunijana and Iglicopsis based on COI, H 3 and concatenated loci; bootstrap supports given if over 60 %, their values together with Bayesian probabilities." figureDoi="10.3897/zookeys.1047.64034.figure8" httpUri="https://binary.pensoft.net/fig/560480" pageId="0" pageNumber="61">8</figureCitation>
). Its sister species is
<taxonomicName baseAuthorityName="Pesic &amp; Gloeer" baseAuthorityYear="2012" class="Gastropoda" family="Hydrobiidae" genus="Montenegrospeum" kingdom="Animalia" lsidName="Montenegrospeum bogici" order="Littorinimorpha" pageId="0" pageNumber="61" phylum="Mollusca" rank="species" species="bogici">
<emphasis italics="true" pageId="0" pageNumber="61">Montenegrospeum bogici</emphasis>
</taxonomicName>
in the H3 tree (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phylogenetic relationships of Travunijana and Iglicopsis based on COI, H 3 and concatenated loci; bootstrap supports given if over 60 %, their values together with Bayesian probabilities." figureDoi="10.3897/zookeys.1047.64034.figure8" httpUri="https://binary.pensoft.net/fig/560480" pageId="0" pageNumber="61">8</figureCitation>
, bootstrap 95%), and on the tree based on both concatenated loci (but with bootstrap 63% only); in the COI tree the bootstrap does not support its phylogenetic position.
</paragraph>
</subSubSection>
</treatment>
</document>
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