307 lines
45 KiB
XML
307 lines
45 KiB
XML
<document id="ADAA4A6BD131FCCA7F19C578BD5824D5" ID-CLB-Dataset="56366" ID-DOI="10.11646/zootaxa.5306.2.6" ID-GBIF-Dataset="05c564eb-3640-4ffb-903d-06d5219856e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8058811" ID-ZooBank="7D456B11-BBF6-4816-AB81-5D02ACE02DAD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1687249692376" checkinUser="plazi" docAuthor="Szawaryn, Karol & Czerwiński, Tomasz" docDate="2023" docId="CD03879FFFD7FF88158AFD991DDA69D2" docLanguage="en" docName="zootaxa.5306.2.6.pdf" docOrigin="Zootaxa 5306 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5306.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Merma antakotako Szawaryn & Czerwiński 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="271" masterDocId="313AFFE7FFD3FF8D151DFFDE196E6E02" masterDocTitle="Three new species of Merma Weise (Coleoptera: Coccinellidae: Epilachnini) from Madagascar" masterLastPageNumber="276" masterPageNumber="266" pageNumber="270" updateTime="1687345819786" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="B9C00ED7A7E343641C88FE76A5B73C93" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="773497CC6D4FDF1CB27EF1A85BBA4265">
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<mods:title id="B950B40D00870533E91F9D09169BEECD">Three new species of Merma Weise (Coleoptera: Coccinellidae: Epilachnini) from Madagascar</mods:title>
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<mods:name id="FDD341753C6DBD94FF7CD3B363575A42" type="personal">
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<mods:roleTerm id="6E863EAB7E5808309F1CE41D9FE4F52E">Author</mods:roleTerm>
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<mods:namePart id="15B82AC3811F5054DF4AC8083AA691AF">Szawaryn, Karol</mods:namePart>
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<mods:nameIdentifier id="A50417815AFEA19EBBDC220B64517047" type="ORCID">0000-0002-9329-4268</mods:nameIdentifier>
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<mods:affiliation id="9939FB3AD75A4DF3F6142C1DCF19E617">k. szawaryn @ gmail. com; https: // orcid. org / 0000 - 0002 - 9329 - 4268</mods:affiliation>
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<mods:nameIdentifier id="A7FB36761544A7FDC057FCE24DF1BD96" type="email">k.szawaryn@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="70EE3E9AFDB4944382896A5DBE51A844">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="30713119C8DE9B8DC847D5A4603F2CE1">Czerwiński, Tomasz</mods:namePart>
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<mods:nameIdentifier id="F6E07F1E96FB3DB4F78F8218F60D5F51" type="ORCID">0000-0002-0598-7858</mods:nameIdentifier>
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<mods:affiliation id="D6E21E926711DA197D17C4EA250A33BD">tczerwinski @ miiz. waw. pl; https: // orcid. org / 0000 - 0002 - 0598 - 7858</mods:affiliation>
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<mods:nameIdentifier id="3CBC65234F53A1F3D960BEAB696B5FD5" type="email">tczerwinski@miiz.waw.pl</mods:nameIdentifier>
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<mods:title id="B7FBC9F040D0491AC09C1F8F72AC4774">Zootaxa</mods:title>
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<mods:part id="943C30454A437F83D13C0E5AC43C7FBA">
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<mods:date id="9C22C2D04814280EF2B21F751294D9B8">2023</mods:date>
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<mods:number id="3FD7E077FA6F42A3B6D8892143E285F7">2023-06-20</mods:number>
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<mods:number id="70EFC22ABA46154435B41A0C38141A6A">5306</mods:number>
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<mods:url id="F2ECD4FA1B1D0674243C33921350DE62">http://dx.doi.org/10.11646/zootaxa.5306.2.6</mods:url>
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<mods:classification id="54D39E410A2BBF229E363C581F57A433">journal article</mods:classification>
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<mods:identifier id="ED506D88380E2BBB96ABF533630F6C23" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D41391950062EA4B0B0495AFC937467C" type="Zenodo-Dep">8058811</mods:identifier>
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<treatment id="CD03879FFFD7FF88158AFD991DDA69D2" ID-DOI="http://doi.org/10.5281/zenodo.8058891" ID-GBIF-Taxon="209696206" ID-Zenodo-Dep="8058891" LSID="urn:lsid:plazi:treatment:CD03879FFFD7FF88158AFD991DDA69D2" httpUri="http://treatment.plazi.org/id/CD03879FFFD7FF88158AFD991DDA69D2" lastPageId="5" lastPageNumber="271" pageId="4" pageNumber="270">
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<subSubSection id="0DB06502FFD7FF89158AFD99188D6C63" box="[151,483,582,609]" pageId="4" pageNumber="270" type="nomenclature">
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<paragraph id="45153689FFD7FF89158AFD99188D6C63" blockId="4.[151,994,582,681]" box="[151,483,582,609]" pageId="4" pageNumber="270">
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<heading id="1E5D81E5FFD7FF89158AFD99188D6C63" bold="true" box="[151,483,582,609]" fontSize="11" level="1" pageId="4" pageNumber="270" reason="1">
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<emphasis id="77DEEA9BFFD7FF89158AFD99188D6C63" bold="true" box="[151,483,582,609]" pageId="4" pageNumber="270">
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<taxonomicName id="82AA4D0AFFD7FF89158AFD9918106C63" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[151,382,582,609]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="antakotako" status="sp. nov.">
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<emphasis id="77DEEA9BFFD7FF89158AFD9918106C63" bold="true" box="[151,382,582,609]" italics="true" pageId="4" pageNumber="270">Merma antakotako</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6CED57E0FFD7FF891498FD99188D6C63" box="[389,483,583,609]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="0DB06502FFD7FF89158AFDB51A8C6CAB" pageId="4" pageNumber="270" type="description">
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<paragraph id="45153689FFD7FF89158AFDB5182A6C87" blockId="4.[151,994,582,681]" box="[151,324,619,645]" pageId="4" pageNumber="270">
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<figureCitation id="DD912A0CFFD7FF89158AFDB518606C87" box="[151,270,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1512,1536]" captionTargetBox="[175,1428,187,1482]" captionTargetId="figure-17@2.[153,1434,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Merma species. A–C) M. ankarana sp. nov., male, holotype (MNHN); A) dorsal; B) lateral; C) frontal; D–F) M. antakotako sp. nov., male, paratype (MNHN); D) dorsal; E) lateral; F) frontal; G–I) M. fisheri sp. nov., male, holotype (CAS); G) dorsal; H) lateral; I) frontal." figureDoi="http://doi.org/10.5281/zenodo.8058813" httpUri="https://zenodo.org/record/8058813/files/figure.png" pageId="4" pageNumber="270">Figs 1D–F</figureCitation>
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,
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<figureCitation id="DD912A0CFFD7FF891407FDB518476C87" box="[282,297,619,645]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">3</figureCitation>
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,
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<figureCitation id="DD912A0CFFD7FF89142BFDB5182A6C87" box="[310,324,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="4" pageNumber="270">4</figureCitation>
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</paragraph>
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||
<paragraph id="45153689FFD7FF89158AFD501A8C6CAB" blockId="4.[151,994,582,681]" box="[151,994,654,681]" pageId="4" pageNumber="270">
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<uri id="313B3A8BFFD7FF89158AFD501A8C6CAB" box="[151,994,654,681]" pageId="4" pageNumber="270">
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urn:lsid:zoobank.org:act:
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<uuid id="310C0C5CFFD7FF8914BBFD501A8C6CAB" box="[422,994,654,681]" pageId="4" pageNumber="270">75776EBF-DE80-4BC5-ADC2-DBA5F8FDA779</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0DB06502FFD7FF89158AFD081C356A13" pageId="4" pageNumber="270" type="diagnosis">
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<paragraph id="45153689FFD7FF89158AFD081C356A13" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="270">
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<emphasis id="77DEEA9BFFD7FF89158AFD08187C6CF2" bold="true" box="[151,274,726,752]" pageId="4" pageNumber="270">Diagnosis.</emphasis>
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Coloration of
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<taxonomicName id="82AA4D0AFFD7FF8914A0FD091B0F6CF3" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[445,609,727,753]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="antakotako" status="sp. nov.">
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||
<emphasis id="77DEEA9BFFD7FF8914A0FD0918BA6CF2" box="[445,468,727,752]" italics="true" pageId="4" pageNumber="270">M</emphasis>
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.
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<emphasis id="77DEEA9BFFD7FF8914F8FD091B0F6CF3" box="[485,609,727,753]" italics="true" pageId="4" pageNumber="270">antakotako</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6CED57E0FFD7FF891776FD091BAA6CF3" box="[619,708,727,753]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
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is identical to all other Malagasy species of the genus
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<taxonomicName id="82AA4D0AFFD7FF891059FD091CFB6CF2" authorityName="Weise" authorityYear="1898" box="[1348,1429,727,752]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="genus">
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<emphasis id="77DEEA9BFFD7FF891059FD091CFB6CF2" box="[1348,1429,727,752]" italics="true" pageId="4" pageNumber="270">Merma</emphasis>
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</taxonomicName>
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. From
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<taxonomicName id="82AA4D0AFFD7FF8915C7FD25183F6D17" baseAuthorityName="Sicard" baseAuthorityYear="1908" box="[218,337,763,789]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="hovana">
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<emphasis id="77DEEA9BFFD7FF8915C7FD25199F6D16" box="[218,241,763,788]" italics="true" pageId="4" pageNumber="270">M</emphasis>
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.
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<emphasis id="77DEEA9BFFD7FF8915E2FD25183F6D17" box="[255,337,763,789]" italics="true" pageId="4" pageNumber="270">hovana</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="82AA4D0AFFD7FF89149AFD251B286D17" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[391,582,763,789]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="quadriguttata">
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<emphasis id="77DEEA9BFFD7FF89149AFD2518F06D16" box="[391,414,763,788]" italics="true" pageId="4" pageNumber="270">M</emphasis>
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.
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<emphasis id="77DEEA9BFFD7FF8914B1FD251B286D17" box="[428,582,763,789]" italics="true" pageId="4" pageNumber="270">quadriguttata</emphasis>
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</taxonomicName>
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it can be easily separated by its larger body size, tarsal claws with additional subquadrate basal tooth and emarginate apex of ventrite
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<quantity id="82529B6CFFD7FF891628FCC01A0B6D3B" box="[821,869,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="270" unit="in" value="5.0">5 in</quantity>
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male (ventrite
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<quantity id="82529B6CFFD7FF891100FCC01D236D3B" box="[1053,1101,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="270" unit="in" value="5.0">5 in</quantity>
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males of
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<taxonomicName id="82AA4D0AFFD7FF8911D4FCC11C506D3B" baseAuthorityName="Sicard" baseAuthorityYear="1908" box="[1225,1342,799,825]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="hovana">
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<emphasis id="77DEEA9BFFD7FF8911D4FCC11C506D3B" box="[1225,1342,799,825]" italics="true" pageId="4" pageNumber="270">M hovana</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="82AA4D0AFFD7FF891063FCC1185F6D5F" baseAuthorityName="Sicard" baseAuthorityYear="1909" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="quadriguttata">
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<emphasis id="77DEEA9BFFD7FF891063FCC1185F6D5F" italics="true" pageId="4" pageNumber="270">M. quadriguttata</emphasis>
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</taxonomicName>
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have arcuate apex).
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<taxonomicName id="82AA4D0AFFD7FF89170AFC9D1B856D5F" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[535,747,835,861]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="antakotako" status="sp. nov.">
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<emphasis id="77DEEA9BFFD7FF89170AFC9D1B856D5F" box="[535,747,835,861]" italics="true" pageId="4" pageNumber="270">Merma antakotako</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6CED57E0FFD7FF8917EFFC9D1A266D5F" box="[754,840,835,861]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
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has distinctly explanate lateral elytral margins, while in
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<taxonomicName id="82AA4D0AFFD7FF8915A5FCB918226D83" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[184,332,871,897]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
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<emphasis id="77DEEA9BFFD7FF8915A5FCB918226D83" box="[184,332,871,897]" italics="true" pageId="4" pageNumber="270">M. ankarana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6CED57E0FFD7FF89144AFCB918DC6D83" box="[343,434,871,897]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="82AA4D0AFFD7FF8914EDFCB91B0F6D83" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[496,609,871,897]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
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<emphasis id="77DEEA9BFFD7FF8914EDFCB91B0F6D83" box="[496,609,871,897]" italics="true" pageId="4" pageNumber="270">M. fisheri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6CED57E0FFD7FF891771FCB91BA86D83" box="[620,710,871,897]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
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it is narrowly explanate. With
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<taxonomicName id="82AA4D0AFFD7FF89112AFCB91DA56D83" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[1079,1227,871,897]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
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<emphasis id="77DEEA9BFFD7FF89112AFCB91DA56D83" box="[1079,1227,871,897]" italics="true" pageId="4" pageNumber="270">M. ankarana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6CED57E0FFD7FF8911CAFCB91C5F6D83" box="[1239,1329,871,897]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
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it shares emarginate posterior margin of ventrite
|
||
<quantity id="82529B6CFFD7FF89177BFC541BFB6DA7" box="[614,661,906,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="270" unit="in" value="5.0">5 in</quantity>
|
||
male, while in
|
||
<taxonomicName id="82AA4D0AFFD7FF891652FC551AAE6DA7" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[847,960,907,933]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
|
||
<emphasis id="77DEEA9BFFD7FF891652FC551AAE6DA7" box="[847,960,907,933]" italics="true" pageId="4" pageNumber="270">M. fisheri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6CED57E0FFD7FF8916D7FC551D4A6DA7" box="[970,1060,907,933]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
|
||
it is rounded. The most reliable character to separate that species are structures of the male genitalia: penis guide in lateral view with inner margin almost straight/ slightly sinusoidal in middle, with small lateral thorn-like projections at level of 1/3 of its length (
|
||
<figureCitation id="DD912A0CFFD7FF891582FC2919986A13" box="[159,246,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="4" pageNumber="270">Fig. 4B</figureCitation>
|
||
, arrow); tip of penis with broad, subtriangular lateral projection, and a single row of small denticles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DB06502FFD7FF8915DAFBC51CFA6A37" box="[199,1428,1051,1077]" pageId="4" pageNumber="270" type="etymology">
|
||
<paragraph id="45153689FFD7FF8915DAFBC51CFA6A37" blockId="4.[151,1437,726,2013]" box="[199,1428,1051,1077]" pageId="4" pageNumber="270">
|
||
<emphasis id="77DEEA9BFFD7FF8915DAFBC518206A37" bold="true" box="[199,334,1051,1077]" pageId="4" pageNumber="270">Etymology.</emphasis>
|
||
Specific epithet is given after the Antakotako village where the
|
||
<typeStatus id="9A11882BFFD7FF891100FBC51D136A37" box="[1053,1149,1051,1077]" pageId="4" pageNumber="270">paratype</typeStatus>
|
||
specimen was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DB06502FFD7FF8815DAFBE01DA36961" lastPageId="5" lastPageNumber="271" pageId="4" pageNumber="270" type="description">
|
||
<paragraph id="45153689FFD7FF8915DAFBE01DA76AC7" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="270">
|
||
<emphasis id="77DEEA9BFFD7FF8915DAFBE018376A5A" bold="true" box="[199,345,1086,1112]" pageId="4" pageNumber="270">Description.</emphasis>
|
||
Length 5.8–6.0 mm, width 5.0–
|
||
<quantity id="82529B6CFFD7FF8917A1FBE01A7C6A5B" box="[700,786,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="270" unit="mm" value="5.1">5.1 mm</quantity>
|
||
; length/width ratio = 1.16–1.17. Body (
|
||
<figureCitation id="DD912A0CFFD7FF8911D9FBE11C596A5B" box="[1220,1335,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,250,1512,1536]" captionTargetBox="[175,1428,187,1482]" captionTargetId="figure-17@2.[153,1434,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Merma species. A–C) M. ankarana sp. nov., male, holotype (MNHN); A) dorsal; B) lateral; C) frontal; D–F) M. antakotako sp. nov., male, paratype (MNHN); D) dorsal; E) lateral; F) frontal; G–I) M. fisheri sp. nov., male, holotype (CAS); G) dorsal; H) lateral; I) frontal." figureDoi="http://doi.org/10.5281/zenodo.8058813" httpUri="https://zenodo.org/record/8058813/files/figure.png" pageId="4" pageNumber="270">Fig. 1D–F</figureCitation>
|
||
) roundly oval, convex, dorsum pubescent. Head, mouthparts, legs and abdomen reddish-orange. Pronotum dark brown with anterior corners and lateral margins pale orange. Elytra dark brown, with two orange or reddish maculae on each elytron, anterior macula placed below humeral callus, posterior macula in apical part of elytron.
|
||
</paragraph>
|
||
<paragraph id="45153689FFD7FF8915DAFB111CE26853" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="270">
|
||
Head transverse, partially concealed in prothorax. Interocular distance about 0.6 times of head width. Inner eye orbits slightly emarginate antero-medially, closest in the middle. Antenna (
|
||
<figureCitation id="DD912A0CFFD7FF8916C6FB2D1D5F6B0F" box="[987,1073,1267,1293]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3B</figureCitation>
|
||
) composed of 11 antennomeres, about as long as 0.6 head width; scape large, elongate, slightly swollen; pedicel distinctly narrower than scape, elongate; antennomere 3 elongate, 1.3 times longer than pedicel; antennomeres 4–7 elongate, about 1.5 times as long as its width; antennomere 8 as long as wide; antennomeres 9–11 forming a distinct asymmetrical club. Ventral antennal (
|
||
<figureCitation id="DD912A0CFFD7FF891419FA5D18366B9F" box="[260,344,1411,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3C</figureCitation>
|
||
) grooves distinct in anterior part, moderately long and straight, along inner margin of eye. Clypeus short, parallel-sided, its anterior margin emarginate, smooth without groove. Labrum transverse, short, anterior margin slightly emarginate, membranous. Prothorax. Hypomeron smooth, simply / finely punctate. Prosternal process (
|
||
<figureCitation id="DD912A0CFFD7FF891072FA1519D9680B" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3E</figureCitation>
|
||
) smooth, without carinae, bordered laterally. Prosternum in front of coxa about as long as coxal longitudinal diameter; anterior margin, uniformly arcuate, bordered; posterior margin with bordering line starting from the base of prosternal process, laterally reaching notosternal suture. Surface of prosternum covered with sparse small punctae.
|
||
</paragraph>
|
||
<paragraph id="45153689FFD7FF8915DAF9851DF6692B" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="270">
|
||
Pterothorax.Mesoventral process covered with small tubercles along lateral margins, about as wide as mesocoxal diameter; meso-metaventral suture straight. Elytra with punctate of two sizes; lateral margins distinctly explanate, entirely visible from above. Epipleuron complete (
|
||
<figureCitation id="DD912A0CFFD7FF8917CFF97D1A4368BF" box="[722,813,1699,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3A</figureCitation>
|
||
), broad, with distinct depressions to accommodate mid and hind legs in repose, inner margin with broad bordering line present in median and apical part. Mesoventrite covered with sparse and small punctures, with just a few larger in median part. Metaventral postcoxal lines (
|
||
<figureCitation id="DD912A0CFFD7FF89105DF9351CFA6907" box="[1344,1428,1771,1797]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3E</figureCitation>
|
||
) joined on metaventral process forming straight line, recurved and then descending laterally.
|
||
</paragraph>
|
||
<paragraph id="45153689FFD7FF8915DAF8ED18C369DF" blockId="4.[151,1437,726,2013]" pageId="4" pageNumber="270">
|
||
Legs (
|
||
<figureCitation id="DD912A0CFFD7FF891410F8ED1808694F" box="[269,358,1843,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3A</figureCitation>
|
||
) short and stout with apices of mid and hind femora not protruding from outer margin of elytral epipleuron. Trochanters angulately produced, with bordering line on their outer surfaces, and cavities on their inner surfaces for receiving tip of tibiae in repose. Protibiae slightly widened, all tibiae on outer edge with oblique carina near apex (
|
||
<figureCitation id="DD912A0CFFD7FF891409F841180369BB" box="[276,365,1951,1977]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3D</figureCitation>
|
||
), without distinct apical spurs. Tarsal claws (
|
||
<figureCitation id="DD912A0CFFD7FF891675F8411AD169BB" box="[872,959,1951,1977]" captionStart="FIGURE 3" captionStartId="5.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-34@5.[151,1436,181,1375]" captionTargetPageId="5" captionText="FIGURE 3. Merma antakotako sp. nov., male, paratype (MNHN): A) habitus, ventral; B) antenna; C) mouthparts; D) apex of hind tibia and tarsus; E) pro-, meso- and metathotax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="4" pageNumber="270">Fig. 3D</figureCitation>
|
||
) double with additional subquadrate basal tooth, empodium absent.
|
||
</paragraph>
|
||
<caption id="11D56601FFD6FF88158AFAA91A566BB1" ID-DOI="http://doi.org/10.5281/zenodo.8058819" ID-Zenodo-Dep="8058819" httpUri="https://zenodo.org/record/8058819/files/figure.png" pageId="5" pageNumber="271" startId="5.[151,250,1399,1423]" targetBox="[151,1436,181,1375]" targetPageId="5" targetType="figure">
|
||
<paragraph id="45153689FFD6FF88158AFAA91A566BB1" blockId="5.[151,1436,1399,1459]" pageId="5" pageNumber="271">
|
||
<emphasis id="77DEEA9BFFD6FF88158AFAA9187D6B8D" bold="true" box="[151,275,1399,1423]" pageId="5" pageNumber="271">FIGURE 3.</emphasis>
|
||
<taxonomicName id="82AA4D0AFFD6FF881407FAA618B56B8D" authority="Szawaryn & Czerwiński, 2023" authorityName="Szawaryn & Czerwiński" authorityYear="2023" box="[282,475,1399,1423]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="antakotako" status="sp. nov.">
|
||
<emphasis id="77DEEA9BFFD6FF881407FAA618B56B8D" box="[282,475,1399,1423]" italics="true" pageId="5" pageNumber="271">Merma antakotako</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="77DEEA9BFFD6FF8814FCFAA91B5C6B8D" bold="true" box="[481,562,1399,1423]" pageId="5" pageNumber="271">
|
||
<taxonomicNameLabel id="6CED57E0FFD6FF8814FCFAA91B5C6B8D" box="[481,562,1399,1423]" pageId="5" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male, paratype (MNHN):
|
||
<emphasis id="77DEEA9BFFD6FF88165AFAA91A376B8D" bold="true" box="[839,857,1399,1423]" pageId="5" pageNumber="271">A</emphasis>
|
||
) habitus, ventral;
|
||
<emphasis id="77DEEA9BFFD6FF88110FFAA91D4D6B8D" bold="true" box="[1042,1059,1399,1423]" pageId="5" pageNumber="271">B</emphasis>
|
||
) antenna;
|
||
<emphasis id="77DEEA9BFFD6FF881193FAA91DCE6B8D" bold="true" box="[1166,1184,1399,1423]" pageId="5" pageNumber="271">C</emphasis>
|
||
) mouthparts;
|
||
<emphasis id="77DEEA9BFFD6FF881032FAA91C2F6B8D" bold="true" box="[1327,1345,1399,1423]" pageId="5" pageNumber="271">D</emphasis>
|
||
) apex of hind tibia and tarsus;
|
||
<emphasis id="77DEEA9BFFD6FF88146CFA4518EC6BB1" bold="true" box="[369,386,1435,1459]" pageId="5" pageNumber="271">E</emphasis>
|
||
) pro-, meso- and metathotax, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="45153689FFD6FF8815DAFA3F1995686A" blockId="5.[151,1437,1505,2000]" pageId="5" pageNumber="271">
|
||
Abdomen with five ventrites with ventrite 5 emarginate posteriorly (
|
||
<figureCitation id="DD912A0CFFD6FF8816EDFA3F1D3E6BFE" box="[1008,1104,1505,1532]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4A</figureCitation>
|
||
). Abdominal postcoxal lines descending, sub-parallel to posterior margin of ventrite 1. Apical margin of tergite VIII rounded (
|
||
<figureCitation id="DD912A0CFFD6FF88101BF9DB1C0F6822" box="[1286,1377,1541,1568]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4D</figureCitation>
|
||
) and sternite VIII straight (
|
||
<figureCitation id="DD912A0CFFD6FF88148DF9F718876846" box="[400,489,1577,1604]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4E</figureCitation>
|
||
); apodeme of sternum IX (
|
||
<figureCitation id="DD912A0CFFD6FF881603F9F71A1D6846" box="[798,883,1577,1604]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4F</figureCitation>
|
||
) rod-like. Tergite X transverse, truncate apically (
|
||
<figureCitation id="DD912A0CFFD6FF881582F9931980686A" box="[159,238,1613,1640]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="45153689FFD6FF8815DAF9AF1DA36961" blockId="5.[151,1437,1505,2000]" pageId="5" pageNumber="271">
|
||
Male genitalia (
|
||
<figureCitation id="DD912A0CFFD6FF881466F9AF1B5B688E" box="[379,565,1649,1676]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4B, C, G–I</figureCitation>
|
||
). Tegminal basal piece with a pair of indistinct spines near base of tegminal strut. Penis guide slightly asymmetrical in inner view (
|
||
<figureCitation id="DD912A0CFFD6FF8817E8F94B1A2268B2" box="[757,844,1685,1712]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4C</figureCitation>
|
||
), as long as parameres, distal ¼ of his length strongly tapering into apex; in lateral view (
|
||
<figureCitation id="DD912A0CFFD6FF88173CF9671B1668D6" box="[545,632,1721,1748]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4B</figureCitation>
|
||
) outer edge straight with curved tip and with densely setose field in the median part, inner edge weakly sinuate; in inner view with a pair of small thorns laterally. Parameres well developed, rounded apically. Tegminal strut distinctly shorter than penis guide, stout, strongly expanded distally. Penis (
|
||
<figureCitation id="DD912A0CFFD6FF881055F8DF1CFA691E" box="[1352,1428,1793,1820]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4I</figureCitation>
|
||
) rod-like, sinuate; penis tip (
|
||
<figureCitation id="DD912A0CFFD6FF8814D6F8FB1B4C6942" box="[459,546,1829,1856]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4H</figureCitation>
|
||
) asymmetrical, apex in inner view (
|
||
<figureCitation id="DD912A0CFFD6FF8816A8F8FB1D636942" box="[949,1037,1829,1856]" captionStart="FIGURE 4" captionStartId="6.[151,250,1528,1552]" captionTargetBox="[151,1435,184,1504]" captionTargetId="figure-17@6.[151,1436,181,1504]" captionTargetPageId="6" captionText="FIGURE 4. Merma antakotako sp. nov., male, holotype (MNHN): A) abdomen; B) tegmen, lateral; C) tegmen, inner; D) tergite VIII; E) sternite VIII; F) abdominal segments IX and X; G) penis tip, inner; H) penis tip, lateral; I) penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.8058821" httpUri="https://zenodo.org/record/8058821/files/figure.png" pageId="5" pageNumber="271">Fig. 4G</figureCitation>
|
||
) with wide thorn-like projection on one side and with row of denticles on the other side; penis capsule with arms weakly developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DB06502FFD6FF8815DAF8B31B3E6985" box="[199,592,1901,1927]" pageId="5" pageNumber="271" type="distribution">
|
||
<paragraph id="45153689FFD6FF8815DAF8B31B3E6985" blockId="5.[151,1437,1505,2000]" box="[199,592,1901,1927]" pageId="5" pageNumber="271">
|
||
<emphasis id="77DEEA9BFFD6FF8815DAF8B3180F6985" bold="true" box="[199,353,1901,1927]" pageId="5" pageNumber="271">Distribution.</emphasis>
|
||
<collectingCountry id="3DBD7619FFD6FF88147AF8B318806985" box="[359,494,1901,1927]" name="Madagascar" pageId="5" pageNumber="271">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="DD912A0CFFD6FF8814E3F8B31B2C6985" box="[510,578,1901,1927]" captionStart="FIGURE 6" captionStartId="8.[151,250,1464,1488]" captionTargetBox="[427,1161,351,1435]" captionTargetId="figure-85@8.[420,1166,345,1441]" captionTargetPageId="8" captionText="FIGURE 6. Distribution map of Merma." figureDoi="http://doi.org/10.5281/zenodo.8063852" httpUri="https://zenodo.org/record/8063852/files/figure.png" pageId="5" pageNumber="271">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DB06502FFD6FF8815DAF84C1DDA69D2" pageId="5" pageNumber="271" type="materials_examined">
|
||
<paragraph id="45153689FFD6FF8815DAF84C1C1569AE" blockId="5.[151,1437,1505,2000]" box="[199,1403,1937,1964]" pageId="5" pageNumber="271">
|
||
<emphasis id="77DEEA9BFFD6FF8815DAF84C181B69A9" bold="true" box="[199,373,1937,1964]" pageId="5" pageNumber="271">Type material.</emphasis>
|
||
<materialsCitation id="F5C23CD4FFD6FF881461F84C1C1569AE" ID-GBIF-Occurrence="4133181302" box="[380,1403,1937,1964]" collectingDate="1970-10" collectionCode="MNHN" collectorName="A. Villier" country="Madagascar" location="Baie d'Antongil" municipality="Voloina" pageId="5" pageNumber="271" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="77DEEA9BFFD6FF881461F84C189C69AE" bold="true" box="[380,498,1938,1964]" pageId="5" pageNumber="271">
|
||
<typeStatus id="9A11882BFFD6FF881461F84C188369AE" box="[380,493,1938,1964]" pageId="5" pageNumber="271">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male, “
|
||
<collectingCountry id="3DBD7619FFD6FF88175AF84F1BBD69A9" box="[583,723,1937,1963]" name="Madagascar" pageId="5" pageNumber="271">Madagascar</collectingCountry>
|
||
,
|
||
<location id="40756052FFD6FF8817C3F84F1AF869A9" LSID="urn:lsid:plazi:treatment:CD03879FFFD7FF88158AFD991DDA69D2:40756052FFD6FF8817C3F84F1AF869A9" box="[734,918,1937,1963]" country="Madagascar" municipality="Voloina" name="Baie d'Antongil" pageId="5" pageNumber="271">Baie d’Antongil</location>
|
||
,
|
||
<collectingMunicipality id="A571ACF3FFD6FF8816BCF84F1A9569A9" box="[929,1019,1937,1963]" pageId="5" pageNumber="271">Voloina</collectingMunicipality>
|
||
,
|
||
<date id="31141049FFD6FF881118F84C1D3569AE" box="[1029,1115,1938,1964]" pageId="5" pageNumber="271" value="1970-10">
|
||
<collectingDate id="2150E9A1FFD6FF881118F84C1D3569AE" box="[1029,1115,1938,1964]" pageId="5" pageNumber="271" value="1970-10">X.1970</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="E85F535FFFD6FF881178F84C1DA069A9" box="[1125,1230,1937,1964]" pageId="5" pageNumber="271">A. Villier</collectorName>
|
||
rec” (
|
||
<collectionCode id="23BBAE4CFFD6FF88100EF84C1C0169AE" box="[1299,1391,1938,1964]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="271">MNHN</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="45153689FFD6FF8815DAF8681DDA69D2" blockId="5.[151,1437,1505,2000]" box="[199,1204,1973,2000]" pageId="5" pageNumber="271">
|
||
<materialsCitation id="F5C23CD4FFD6FF8815DAF8681DDF69D2" ID-GBIF-Occurrence="4133181303" box="[199,1201,1973,2000]" collectionCode="MNHN" country="Madagascar" county="Antakotako" location="Ron Maroantsetra" municipality="Vadon" pageId="5" pageNumber="271" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="77DEEA9BFFD6FF8815DAF868185369D2" bold="true" box="[199,317,1974,2000]" pageId="5" pageNumber="271">
|
||
<typeStatus id="9A11882BFFD6FF8815DAF868185669D2" box="[199,312,1974,2000]" pageId="5" pageNumber="271">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male, “
|
||
<collectingCountry id="3DBD7619FFD6FF88148EF86B1B7169CD" box="[403,543,1973,1999]" name="Madagascar" pageId="5" pageNumber="271">Madagascar</collectingCountry>
|
||
,
|
||
<location id="40756052FFD6FF881734F8681B8969D2" LSID="urn:lsid:plazi:treatment:CD03879FFFD7FF88158AFD991DDA69D2:40756052FFD6FF881734F8681B8969D2" box="[553,743,1973,2000]" country="Madagascar" county="Antakotako" municipality="Vadon" name="Ron Maroantsetra" pageId="5" pageNumber="271">
|
||
<superScript id="B2DF9BC1FFD6FF881734F8681B2269C1" attach="left" box="[553,588,1973,2000]" fontSize="6" pageId="5" pageNumber="271">Ron</superScript>
|
||
Maroantsetra
|
||
</location>
|
||
, 11.39,
|
||
<collectingMunicipality id="A571ACF3FFD6FF881623F86B1AEB69CD" box="[830,901,1973,1999]" pageId="5" pageNumber="271">Vadon</collectingMunicipality>
|
||
/
|
||
<collectingCounty id="AC744E05FFD6FF88168FF86B1D7969CD" box="[914,1047,1973,1999]" pageId="5" pageNumber="271">Antakotako</collectingCounty>
|
||
” (1,
|
||
<collectionCode id="23BBAE4CFFD6FF881153F8681DC769D2" box="[1102,1193,1974,2000]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="271">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |