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<document id="26A81F9D84BF1E9FA0E27AAEFFD8BFD4" ID-CLB-Dataset="56366" ID-DOI="10.11646/zootaxa.5306.2.6" ID-GBIF-Dataset="05c564eb-3640-4ffb-903d-06d5219856e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8058811" ID-ZooBank="7D456B11-BBF6-4816-AB81-5D02ACE02DAD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1687249692376" checkinUser="plazi" docAuthor="Szawaryn, Karol &amp; Czerwiński, Tomasz" docDate="2023" docId="CD03879FFFD2FF89158AF9D41DED6FF7" docLanguage="en" docName="zootaxa.5306.2.6.pdf" docOrigin="Zootaxa 5306 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5306.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Merma ankarana Szawaryn &amp; Czerwiński 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="270" masterDocId="313AFFE7FFD3FF8D151DFFDE196E6E02" masterDocTitle="Three new species of Merma Weise (Coleoptera: Coccinellidae: Epilachnini) from Madagascar" masterLastPageNumber="276" masterPageNumber="266" pageNumber="267" updateTime="1687345819786" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FF5F6B57E8A5E7BB51A3056A9052C3C4">Three new species of Merma Weise (Coleoptera: Coccinellidae: Epilachnini) from Madagascar</mods:title>
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<mods:namePart id="865B2E40BE3F2B379509C9FB69BA648B">Szawaryn, Karol</mods:namePart>
<mods:nameIdentifier id="A024CFECE4730F734723B27C13815958" type="ORCID">0000-0002-9329-4268</mods:nameIdentifier>
<mods:affiliation id="C8BFC726E865FE9707A63F99ABE33BF0">k. szawaryn @ gmail. com; https: // orcid. org / 0000 - 0002 - 9329 - 4268</mods:affiliation>
<mods:nameIdentifier id="22883486CB2E96CEBF9C760556BB5836" type="email">k.szawaryn@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8710D696C08EE517C09404BF91C1CF12">Czerwiński, Tomasz</mods:namePart>
<mods:nameIdentifier id="109B21B42108B8A154D11D92A7E67859" type="ORCID">0000-0002-0598-7858</mods:nameIdentifier>
<mods:affiliation id="7805B9D9E5FFD0530F24115CC9A559C7">tczerwinski @ miiz. waw. pl; https: // orcid. org / 0000 - 0002 - 0598 - 7858</mods:affiliation>
<mods:nameIdentifier id="036514FFAB90EF0EA441267546416A0C" type="email">tczerwinski@miiz.waw.pl</mods:nameIdentifier>
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<mods:title id="809C6BF77A706E42CB428B0684623326">Zootaxa</mods:title>
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<mods:part id="2C1FDA727998FDCC01AEEAE154BE2EBC">
<mods:date id="5FFDC292320DBE1990C0FCF0F1861B37">2023</mods:date>
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<mods:number id="E2C67861EA309B6C1E167575F76466A7">2023-06-20</mods:number>
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<mods:number id="0C3B5B4CDE40B3306E2F3BB9BA34377B">5306</mods:number>
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<mods:number id="6E655799B7ECF00C613AB6BC7BA62EDF">2</mods:number>
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<treatment id="CD03879FFFD2FF89158AF9D41DED6FF7" ID-DOI="http://doi.org/10.5281/zenodo.8058887" ID-GBIF-Taxon="209696208" ID-Zenodo-Dep="8058887" LSID="urn:lsid:plazi:treatment:CD03879FFFD2FF89158AF9D41DED6FF7" httpUri="http://treatment.plazi.org/id/CD03879FFFD2FF89158AF9D41DED6FF7" lastPageId="4" lastPageNumber="270" pageId="1" pageNumber="267">
<subSubSection id="0DB06502FFD2FF8C158AF9D418BF6821" box="[151,465,1545,1572]" pageId="1" pageNumber="267" type="nomenclature">
<paragraph id="45153689FFD2FF8C158AF9D418BF6821" blockId="1.[151,1003,1545,1644]" box="[151,465,1545,1572]" pageId="1" pageNumber="267">
<heading id="1E5D81E5FFD2FF8C158AF9D418BF6821" bold="true" box="[151,465,1545,1572]" fontSize="11" level="1" pageId="1" pageNumber="267" reason="1">
<emphasis id="77DEEA9BFFD2FF8C158AF9D418BF6821" bold="true" box="[151,465,1545,1572]" pageId="1" pageNumber="267">
<taxonomicName id="82AA4D0AFFD2FF8C158AF9D418026826" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[151,364,1545,1572]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
<emphasis id="77DEEA9BFFD2FF8C158AF9D418026826" bold="true" box="[151,364,1545,1572]" italics="true" pageId="1" pageNumber="267">Merma ankarana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD2FF8C146EF9D718BF6821" box="[371,465,1545,1571]" pageId="1" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFD2FF8C158AF9F31A85686E" pageId="1" pageNumber="267" type="description">
<paragraph id="45153689FFD2FF8C158AF9F318436845" blockId="1.[151,1003,1545,1644]" box="[151,301,1581,1607]" pageId="1" pageNumber="267">
<figureCitation id="DD912A0CFFD2FF8C158AF9F3187A6845" box="[151,276,1581,1607]" captionStart="FIGURE 1" captionStartId="2.[151,250,1512,1536]" captionTargetBox="[175,1428,187,1482]" captionTargetId="figure-17@2.[153,1434,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Merma species. A–C) M. ankarana sp. nov., male, holotype (MNHN); A) dorsal; B) lateral; C) frontal; D–F) M. antakotako sp. nov., male, paratype (MNHN); D) dorsal; E) lateral; F) frontal; G–I) M. fisheri sp. nov., male, holotype (CAS); G) dorsal; H) lateral; I) frontal." figureDoi="http://doi.org/10.5281/zenodo.8058813" httpUri="https://zenodo.org/record/8058813/files/figure.png" pageId="1" pageNumber="267">Figs 1A–C</figureCitation>
,
<figureCitation id="DD912A0CFFD2FF8C1402F9F318436845" box="[287,301,1581,1607]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="1" pageNumber="267">2</figureCitation>
</paragraph>
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<uri id="313B3A8BFFD2FF8C158AF98F1A85686E" box="[151,1003,1617,1644]" pageId="1" pageNumber="267">
urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="0DB06502FFD2FF8C158AF9471B9569D1" pageId="1" pageNumber="267" type="diagnosis">
<paragraph id="45153689FFD2FF8C158AF9471B9569D1" blockId="1.[151,1437,1689,2004]" pageId="1" pageNumber="267">
<emphasis id="77DEEA9BFFD2FF8C158AF947187C68B1" bold="true" box="[151,274,1689,1715]" pageId="1" pageNumber="267">Diagnosis.</emphasis>
Coloration of
<taxonomicName id="82AA4D0AFFD2FF8C14B2F9441B5368B1" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[431,573,1689,1716]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="ankarana">
<emphasis id="77DEEA9BFFD2FF8C14B2F94418A868B1" box="[431,454,1690,1715]" italics="true" pageId="1" pageNumber="267">M</emphasis>
.
<emphasis id="77DEEA9BFFD2FF8C14CFF9471B5368B1" box="[466,573,1689,1715]" italics="true" pageId="1" pageNumber="267">ankarana</emphasis>
</taxonomicName>
is identical to all other Malagasy species of the genus
<taxonomicName id="82AA4D0AFFD2FF8C1194F9441DB468B1" authorityName="Weise" authorityYear="1898" box="[1161,1242,1690,1715]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="77DEEA9BFFD2FF8C1194F9441DB468B1" box="[1161,1242,1690,1715]" italics="true" pageId="1" pageNumber="267">Merma</emphasis>
</taxonomicName>
. From
<taxonomicName id="82AA4D0AFFD2FF8C103AF9441CF268B1" baseAuthorityName="Sicard" baseAuthorityYear="1908" box="[1319,1436,1689,1716]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="hovana">
<emphasis id="77DEEA9BFFD2FF8C103AF9441C5068B1" box="[1319,1342,1690,1715]" italics="true" pageId="1" pageNumber="267">M</emphasis>
.
<emphasis id="77DEEA9BFFD2FF8C1057F9471CF268B1" box="[1354,1436,1689,1715]" italics="true" pageId="1" pageNumber="267">hovana</emphasis>
</taxonomicName>
and
<taxonomicName id="82AA4D0AFFD2FF8C15D8F96018ED68D5" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[197,387,1725,1752]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="quadriguttata">
<emphasis id="77DEEA9BFFD2FF8C15D8F96019B268D5" box="[197,220,1726,1751]" italics="true" pageId="1" pageNumber="267">M</emphasis>
.
<emphasis id="77DEEA9BFFD2FF8C15F4F96318ED68D5" box="[233,387,1725,1751]" italics="true" pageId="1" pageNumber="267">quadriguttata</emphasis>
</taxonomicName>
it can be easily separated by its larger body size, tarsal claws with additional subquadrate basal tooth and emarginate apex of ventrite
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male (males of
<taxonomicName id="82AA4D0AFFD2FF8C1620F93C1AD968F9" baseAuthorityName="Sicard" baseAuthorityYear="1908" box="[829,951,1761,1787]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="hovana">
<emphasis id="77DEEA9BFFD2FF8C1620F93C1AD968F9" box="[829,951,1761,1787]" italics="true" pageId="1" pageNumber="267">M. hovana</emphasis>
</taxonomicName>
and
<taxonomicName id="82AA4D0AFFD2FF8C16EEF93C1DDB68F9" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[1011,1205,1761,1787]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="quadriguttata">
<emphasis id="77DEEA9BFFD2FF8C16EEF93C1DDB68F9" box="[1011,1205,1761,1787]" italics="true" pageId="1" pageNumber="267">M. quadriguttata</emphasis>
</taxonomicName>
have arcuate apex). General body outline and less explanate lateral elytral margin
<taxonomicName id="82AA4D0AFFD2FF8C164DF8D81A8E691D" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[848,992,1797,1823]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
<emphasis id="77DEEA9BFFD2FF8C164DF8D81A8E691D" box="[848,992,1797,1823]" italics="true" pageId="1" pageNumber="267">M. ankarana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD2FF8C16F5F8D81D506922" box="[1000,1086,1798,1824]" pageId="1" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
shares with
<taxonomicName id="82AA4D0AFFD2FF8C11D6F8D81C56691D" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[1227,1336,1797,1823]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="77DEEA9BFFD2FF8C11D6F8D81C56691D" box="[1227,1336,1797,1823]" italics="true" pageId="1" pageNumber="267">M. fisheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD2FF8C1022F8D81CF66922" box="[1343,1432,1798,1824]" pageId="1" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
, while
<taxonomicName id="82AA4D0AFFD2FF8C15C1F8F418136941" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[220,381,1833,1859]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="antakotako">
<emphasis id="77DEEA9BFFD2FF8C15C1F8F418136941" box="[220,381,1833,1859]" italics="true" pageId="1" pageNumber="267">M. antakotako</emphasis>
</taxonomicName>
has slightly more explanate elytral margin.
<taxonomicName id="82AA4D0AFFD2FF8C1674F8F41D426941" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[873,1068,1833,1859]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="ankarana">
<emphasis id="77DEEA9BFFD2FF8C1674F8F41D426941" box="[873,1068,1833,1859]" italics="true" pageId="1" pageNumber="267">Merma ankarana</emphasis>
</taxonomicName>
also has the largest elytral spots, however, it is based only on a single
<typeStatus id="9A11882BFFD2FF8C1756F8931BC36965" box="[587,685,1869,1895]" pageId="1" pageNumber="267">holotype</typeStatus>
specimen. With
<taxonomicName id="82AA4D0AFFD2FF8C166FF8901D786965" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[882,1046,1869,1895]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="antakotako">
<emphasis id="77DEEA9BFFD2FF8C166FF8901D786965" box="[882,1046,1869,1895]" italics="true" pageId="1" pageNumber="267">M. antakotako</emphasis>
</taxonomicName>
it shares emarginate ventrite
<quantity id="82529B6CFFD2FF8C1072F8931CF26965" box="[1391,1436,1869,1896]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="267" unit="in" value="5.0">5 in</quantity>
male, while in
<taxonomicName id="82AA4D0AFFD2FF8C1421F8AC18C76989" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[316,425,1905,1931]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="267" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="77DEEA9BFFD2FF8C1421F8AC18C76989" box="[316,425,1905,1931]" italics="true" pageId="1" pageNumber="267">M. fisheri</emphasis>
</taxonomicName>
it is rounded. The most reliable character to separate that species are structures of the male genitalia: penis guide in lateral view with inner margin distinctly sinusoidal in middle; penis with narrow lateral thorn-like projection, and a single row of small thorns.
</paragraph>
</subSubSection>
<caption id="11D56601FFD1FF8F158AFA361B49684A" ID-DOI="http://doi.org/10.5281/zenodo.8058813" ID-Zenodo-Dep="8058813" httpUri="https://zenodo.org/record/8058813/files/figure.png" pageId="2" pageNumber="268" startId="2.[151,250,1512,1536]" targetBox="[175,1428,187,1482]" targetPageId="2" targetType="figure">
<paragraph id="45153689FFD1FF8F158AFA361B49684A" blockId="2.[151,1437,1512,1608]" pageId="2" pageNumber="268">
<emphasis id="77DEEA9BFFD1FF8F158AFA36187D6802" bold="true" box="[151,275,1512,1536]" pageId="2" pageNumber="268">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="82AA4D0AFFD1FF8F1494FA371B516802" box="[393,575,1512,1536]" class="Insecta" family="Coccinellidae" genus="Merma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="268" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="77DEEA9BFFD1FF8F1494FA3718BC6802" box="[393,466,1513,1536]" italics="true" pageId="2" pageNumber="268">Merma</emphasis>
species.
<emphasis id="77DEEA9BFFD1FF8F1730FA361B516802" bold="true" box="[557,575,1512,1536]" pageId="2" pageNumber="268">A</emphasis>
</taxonomicName>
<emphasis id="77DEEA9BFFD1FF8F175DFA361B306802" bold="true" box="[576,606,1512,1536]" pageId="2" pageNumber="268">–C</emphasis>
)
<taxonomicName id="82AA4D0AFFD1FF8F1770FA371B816802" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[621,751,1512,1536]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="268" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
<emphasis id="77DEEA9BFFD1FF8F1770FA371BEC6802" box="[621,642,1513,1536]" italics="true" pageId="2" pageNumber="268">M</emphasis>
.
<emphasis id="77DEEA9BFFD1FF8F1793FA361B816802" box="[654,751,1512,1536]" italics="true" pageId="2" pageNumber="268">ankarana</emphasis>
</taxonomicName>
<emphasis id="77DEEA9BFFD1FF8F17E8FA361A2B6802" bold="true" box="[757,837,1512,1536]" pageId="2" pageNumber="268">
<taxonomicNameLabel id="6CED57E0FFD1FF8F17E8FA361A2B6802" box="[757,837,1512,1536]" pageId="2" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (MNHN);
<emphasis id="77DEEA9BFFD1FF8F1147FA361D026802" bold="true" box="[1114,1132,1512,1536]" pageId="2" pageNumber="268">A</emphasis>
) dorsal;
<emphasis id="77DEEA9BFFD1FF8F11D8FA361DB86802" bold="true" box="[1221,1238,1512,1536]" pageId="2" pageNumber="268">B</emphasis>
) lateral;
<emphasis id="77DEEA9BFFD1FF8F102CFA361C2D6802" bold="true" box="[1329,1347,1512,1536]" pageId="2" pageNumber="268">C</emphasis>
) frontal;
<emphasis id="77DEEA9BFFD1FF8F158AF9D219AB6826" bold="true" box="[151,197,1548,1572]" pageId="2" pageNumber="268">D–F</emphasis>
)
<taxonomicName id="82AA4D0AFFD1FF8F15CEF9D3180B6826" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[211,357,1548,1572]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="268" phylum="Arthropoda" rank="species" species="antakotako" status="sp. nov.">
<emphasis id="77DEEA9BFFD1FF8F15CEF9D319866826" box="[211,232,1549,1572]" italics="true" pageId="2" pageNumber="268">M</emphasis>
.
<emphasis id="77DEEA9BFFD1FF8F15E9F9D2180B6826" box="[244,357,1548,1572]" italics="true" pageId="2" pageNumber="268">antakotako</emphasis>
</taxonomicName>
<emphasis id="77DEEA9BFFD1FF8F1477F9D218D46826" bold="true" box="[362,442,1548,1572]" pageId="2" pageNumber="268">
<taxonomicNameLabel id="6CED57E0FFD1FF8F1477F9D218D46826" box="[362,442,1548,1572]" pageId="2" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, paratype (MNHN);
<emphasis id="77DEEA9BFFD1FF8F17D7F9D21BB26826" bold="true" box="[714,732,1548,1572]" pageId="2" pageNumber="268">D</emphasis>
) dorsal;
<emphasis id="77DEEA9BFFD1FF8F1629F9D21A2B6826" bold="true" box="[820,837,1548,1572]" pageId="2" pageNumber="268">E</emphasis>
) lateral;
<emphasis id="77DEEA9BFFD1FF8F1682F9D21AC06826" bold="true" box="[927,942,1548,1572]" pageId="2" pageNumber="268">F</emphasis>
) frontal;
<emphasis id="77DEEA9BFFD1FF8F1111F9D21D586826" bold="true" box="[1036,1078,1548,1572]" pageId="2" pageNumber="268">G–I</emphasis>
)
<taxonomicName id="82AA4D0AFFD1FF8F1159F9D31DC86826" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[1092,1190,1548,1572]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="268" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="77DEEA9BFFD1FF8F1159F9D31D376826" box="[1092,1113,1549,1572]" italics="true" pageId="2" pageNumber="268">M</emphasis>
.
<emphasis id="77DEEA9BFFD1FF8F1178F9D21DC86826" box="[1125,1190,1548,1572]" italics="true" pageId="2" pageNumber="268">fisheri</emphasis>
</taxonomicName>
<emphasis id="77DEEA9BFFD1FF8F11B6F9D21D946826" bold="true" box="[1195,1274,1548,1572]" pageId="2" pageNumber="268">
<taxonomicNameLabel id="6CED57E0FFD1FF8F11B6F9D21D946826" box="[1195,1274,1548,1572]" pageId="2" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (CAS);
<emphasis id="77DEEA9BFFD1FF8F15FAF9EE1995684A" bold="true" box="[231,251,1584,1608]" pageId="2" pageNumber="268">G</emphasis>
) dorsal;
<emphasis id="77DEEA9BFFD1FF8F1449F9EE1806684A" bold="true" box="[340,360,1584,1608]" pageId="2" pageNumber="268">H</emphasis>
) lateral;
<emphasis id="77DEEA9BFFD1FF8F14D9F9EE18A0684A" bold="true" box="[452,462,1584,1608]" pageId="2" pageNumber="268">I</emphasis>
) frontal.
</paragraph>
</caption>
<subSubSection id="0DB06502FFD1FF8F15DAF9A8186A68B6" pageId="2" pageNumber="268" type="etymology">
<paragraph id="45153689FFD1FF8F15DAF9A8186A68B6" blockId="2.[151,1437,1654,2041]" pageId="2" pageNumber="268">
<emphasis id="77DEEA9BFFD1FF8F15DAF9A818206892" bold="true" box="[199,334,1654,1680]" pageId="2" pageNumber="268">Etymology.</emphasis>
The specific epithet is given after the Réserve spéciale d’Ankarana, where the
<typeStatus id="9A11882BFFD1FF8F101EF9A81C0B6892" box="[1283,1381,1654,1680]" pageId="2" pageNumber="268">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFD1FF8915DAF960187C6FAF" lastPageId="4" lastPageNumber="270" pageId="2" pageNumber="268" type="description">
<paragraph id="45153689FFD1FF8F15DAF9601D426946" blockId="2.[151,1437,1654,2041]" pageId="2" pageNumber="268">
<emphasis id="77DEEA9BFFD1FF8F15DAF960183768DA" bold="true" box="[199,345,1726,1752]" pageId="2" pageNumber="268">Description.</emphasis>
Length
<quantity id="82529B6CFFD1FF8F14AEF9601B6768DB" box="[435,521,1726,1753]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="268" unit="mm" value="5.5">5.5 mm</quantity>
, width
<quantity id="82529B6CFFD1FF8F1744F9601BDE68DB" box="[601,688,1726,1753]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="268" unit="mm" value="4.8">4.8 mm</quantity>
; length/width ratio = 1.14. Body (
<figureCitation id="DD912A0CFFD1FF8F113EF9601DF468DA" box="[1059,1178,1726,1752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1512,1536]" captionTargetBox="[175,1428,187,1482]" captionTargetId="figure-17@2.[153,1434,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Merma species. A–C) M. ankarana sp. nov., male, holotype (MNHN); A) dorsal; B) lateral; C) frontal; D–F) M. antakotako sp. nov., male, paratype (MNHN); D) dorsal; E) lateral; F) frontal; G–I) M. fisheri sp. nov., male, holotype (CAS); G) dorsal; H) lateral; I) frontal." figureDoi="http://doi.org/10.5281/zenodo.8058813" httpUri="https://zenodo.org/record/8058813/files/figure.png" pageId="2" pageNumber="268">Fig. 1A–C</figureCitation>
) roundly oval, strongly convex, dorsum pubescent. Head, mouthparts, legs and abdomen reddish-orange. Pronotum chestnut brown with anterior corners and lateral margins orange. Elytra chestnut brown, with two orange or reddish maculae on each elytron, the first placed below humeral callus, the second in apical part of elytron.
</paragraph>
<paragraph id="45153689FFD1FF8E15DAF890185E6EFB" blockId="2.[151,1437,1654,2041]" lastBlockId="3.[151,1437,151,933]" lastPageId="3" lastPageNumber="269" pageId="2" pageNumber="268">Head transverse, partially concealed in prothorax. Interocular distance about 0.6 times of head width. Inner eye orbits slightly emarginate antero-medially, closest in the middle. Antenna composed of 11 antennomeres, about as long as 0.6 head width; scape large, elongate, slightly swollen; pedicel distinctly narrower than scape, elongate; antennomere 3 elongate, 1.3 times longer than pedicel; antennomeres 4–7 elongate, about 1.5 times as long as its width; antennomere 8 as long as wide; antennomeres 9–11 forming a distinct asymmetrical club. Ventral antennal grooves distinct in anterior part, moderately long and straight, along inner margin of eye. Clypeus short, parallel-sided, its anterior margin emarginate, smooth without groove. Labrum transverse, short, anterior margin emarginate, membranous.</paragraph>
<paragraph id="45153689FFD0FF8E15DAFEDD1A986F8B" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="269">Prothorax. Hypomeron smooth, simply / finely punctate. Prosternal process smooth, without carinae, bordered laterally. Prosternum in front of coxa about as long as coxal longitudinal diameter; anterior margin, uniformly arcuate, bordered; posterior margin with bordering line starting from the base of prosternal process, laterally reaching notosternal suture. Surface of prosternum covered with sparse small punctae.</paragraph>
<paragraph id="45153689FFD0FF8E15DAFE4D1AF26C63" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="269">Pterothorax.Mesoventral process covered with small tubercles along lateral margins, about as wide as mesocoxal diameter; meso-metaventral suture straight. Elytra with puncte of two sizes; lateral margins distinctly explanate, entirely visible from above. Epipleuron complete, broad, with distinct depressions to accommodate mid and hind legs in repose, inner margin with broad bordering line present in median and apical part. Mesoventrite covered with sparse and small punctures, with just a few larger in median part. Metaventral postcoxal lines joined on metaventral process forming straight line, recurved and then descending laterally.</paragraph>
<paragraph id="45153689FFD0FF8E15DAFDB519886D17" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="269">Legs short and stout with apices of mid and hind femora not protruding from outer margin of elytral epipleuron. Trochanters elongate, angulately produced, with bordering line on their outer surfaces, and cavities on their inner surfaces for receiving tip of tibiae in repose. Protibiae slightly widened, all tibiae on outer edge with oblique carina near apex, without distinct apical spurs. Tarsal claws double with additional subquadrate basal tooth, empodium absent.</paragraph>
<paragraph id="45153689FFD0FF8E15DAFCC118AA6DA7" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="269">
Abdomen with five ventrites with ventrite 5 emarginate posteriorly (
<figureCitation id="DD912A0CFFD0FF8E16EDFCC11D3E6D3B" box="[1008,1104,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="3" pageNumber="269">Fig. 2A</figureCitation>
). Abdominal postcoxal lines descending, sub-parallel to posterior margin of ventrite 1, recurved shortly laterally. Apical margin of tergite VIII rounded (
<figureCitation id="DD912A0CFFD0FF8E141DFCB9183A6D83" box="[256,340,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="3" pageNumber="269">Fig. 2B</figureCitation>
) and sternite VIII straight (
<figureCitation id="DD912A0CFFD0FF8E1766FCB91BBC6D83" box="[635,722,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="3" pageNumber="269">Fig. 2C</figureCitation>
); apodeme of sternum IX (
<figureCitation id="DD912A0CFFD0FF8E16E9FCB91D286D83" box="[1012,1094,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="3" pageNumber="269">Fig. 2E</figureCitation>
) rod-like.Tergite X transverse, truncate apically (
<figureCitation id="DD912A0CFFD0FF8E147DFC5518D66DA7" box="[352,440,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="3" pageNumber="269">Fig. 2E</figureCitation>
).
</paragraph>
<caption id="11D56601FFD0FF8E158AF84A1C5B69D2" ID-DOI="http://doi.org/10.5281/zenodo.8058817" ID-Zenodo-Dep="8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="3" pageNumber="269" startId="3.[151,250,1940,1964]" targetBox="[151,1436,957,1917]" targetPageId="3" targetType="figure">
<paragraph id="45153689FFD0FF8E158AF84A1C5B69D2" blockId="3.[151,1436,1940,2001]" pageId="3" pageNumber="269">
<emphasis id="77DEEA9BFFD0FF8E158AF84A187B69AE" bold="true" box="[151,277,1940,1964]" pageId="3" pageNumber="269">FIGURE 2.</emphasis>
<taxonomicName id="82AA4D0AFFD0FF8E1400F84B18A169AE" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[285,463,1940,1964]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
<emphasis id="77DEEA9BFFD0FF8E1400F84B18A169AE" box="[285,463,1940,1964]" italics="true" pageId="3" pageNumber="269">Merma ankarana</emphasis>
</taxonomicName>
<emphasis id="77DEEA9BFFD0FF8E14CAF84A1B4769AE" bold="true" box="[471,553,1940,1964]" pageId="3" pageNumber="269">
<taxonomicNameLabel id="6CED57E0FFD0FF8E14CAF84A1B4769AE" box="[471,553,1940,1964]" pageId="3" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (MNHN):
<emphasis id="77DEEA9BFFD0FF8E165BF84A1A3669AE" bold="true" box="[838,856,1940,1964]" pageId="3" pageNumber="269">A</emphasis>
) abdomen;
<emphasis id="77DEEA9BFFD0FF8E16C8F84A1A8869AE" bold="true" box="[981,998,1940,1964]" pageId="3" pageNumber="269">B</emphasis>
) tergite VIII;
<emphasis id="77DEEA9BFFD0FF8E1164F84A1DE569AE" bold="true" box="[1145,1163,1940,1964]" pageId="3" pageNumber="269">C</emphasis>
) sternite VIII;
<emphasis id="77DEEA9BFFD0FF8E1034F84A1C5569AE" bold="true" box="[1321,1339,1940,1964]" pageId="3" pageNumber="269">D</emphasis>
) tegmen, lateral;
<emphasis id="77DEEA9BFFD0FF8E15F9F866199B69D2" bold="true" box="[228,245,1976,2000]" pageId="3" pageNumber="269">E</emphasis>
) abdominal segments IX and X;
<emphasis id="77DEEA9BFFD0FF8E1758F8661B3A69D2" bold="true" box="[581,596,1976,2000]" pageId="3" pageNumber="269">F</emphasis>
) tegmen, inner;
<emphasis id="77DEEA9BFFD0FF8E17E1F8661A7E69D2" bold="true" box="[764,784,1976,2000]" pageId="3" pageNumber="269">G</emphasis>
) penis, lateral;
<emphasis id="77DEEA9BFFD0FF8E16B0F8661AAF69D2" bold="true" box="[941,961,1976,2000]" pageId="3" pageNumber="269">H</emphasis>
) penis tip, lateral;
<emphasis id="77DEEA9BFFD0FF8E119DF8661DE469D2" bold="true" box="[1152,1162,1976,2000]" pageId="3" pageNumber="269">I</emphasis>
) penis tip, inner.
</paragraph>
</caption>
<paragraph id="45153689FFD7FF8915DAFF49187C6FAF" blockId="4.[151,1437,151,501]" pageId="4" pageNumber="270">
Male genitalia (
<figureCitation id="DD912A0CFFD7FF891467FF491B626EB3" box="[378,524,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="4" pageNumber="270">Fig. 2D, F–I</figureCitation>
). Tegminal basal piece with a pair of indistinct spines on inner margin near base of tegminal strut. Penis guide symmetrical in inner view (
<figureCitation id="DD912A0CFFD7FF891625FF651AFE6ED7" box="[824,912,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="4" pageNumber="270">Fig. 2F</figureCitation>
), as long as parameres, distal ¼ of its length strongly tapering into apex; in lateral view (
<figureCitation id="DD912A0CFFD7FF89179BFF011BB16EFB" box="[646,735,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="4" pageNumber="270">Fig. 2D</figureCitation>
) outer edge straight with strongly curved tip and with densely setose field in the median part, inner edge strongly sinusoidal in median part. Parameres well developed, rounded apically. Tegminal strut distinctly shorter than penis guide, stout, strongly expanded distally. Penis rod-like, sinuate (
<figureCitation id="DD912A0CFFD7FF891582FE9519926F67" box="[159,252,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="4" pageNumber="270">Fig. 2G</figureCitation>
); penis tip (
<figureCitation id="DD912A0CFFD7FF891498FE9518B06F67" box="[389,478,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="4" pageNumber="270">Fig. 2H</figureCitation>
) asymmetrical, with narrow thorn-like projection; apex in inner view (
<figureCitation id="DD912A0CFFD7FF891017FE951C376F67" box="[1290,1369,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1940,1964]" captionTargetBox="[151,1436,957,1917]" captionTargetId="figure-335@3.[151,1436,957,1917]" captionTargetPageId="3" captionText="FIGURE 2. Merma ankarana sp. nov., male, holotype (MNHN): A) abdomen; B) tergite VIII; C) sternite VIII; D) tegmen, lateral; E) abdominal segments IX and X; F) tegmen, inner; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner." figureDoi="http://doi.org/10.5281/zenodo.8058817" httpUri="https://zenodo.org/record/8058817/files/figure.png" pageId="4" pageNumber="270">Fig. 2I</figureCitation>
) with narrow thorn-like projection on one side and with row of denticles on the other side; penis capsule with arms weakly developed.
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFD7FF8915DAFE681B3E6FD3" box="[199,592,438,465]" pageId="4" pageNumber="270" type="distribution">
<paragraph id="45153689FFD7FF8915DAFE681B3E6FD3" blockId="4.[151,1437,151,501]" box="[199,592,438,465]" pageId="4" pageNumber="270">
<emphasis id="77DEEA9BFFD7FF8915DAFE68180F6FD2" bold="true" box="[199,353,438,464]" pageId="4" pageNumber="270">Distribution.</emphasis>
<collectingCountry id="3DBD7619FFD7FF89147AFE6918806FD3" box="[359,494,439,465]" name="Madagascar" pageId="4" pageNumber="270">Madagascar</collectingCountry>
(
<figureCitation id="DD912A0CFFD7FF8914E3FE691B2C6FD3" box="[510,578,439,465]" captionStart="FIGURE 6" captionStartId="8.[151,250,1464,1488]" captionTargetBox="[427,1161,351,1435]" captionTargetId="figure-85@8.[420,1166,345,1441]" captionTargetPageId="8" captionText="FIGURE 6. Distribution map of Merma." figureDoi="http://doi.org/10.5281/zenodo.8063852" httpUri="https://zenodo.org/record/8063852/files/figure.png" pageId="4" pageNumber="270">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFD7FF8915DAFE051DED6FF7" box="[199,1155,474,501]" pageId="4" pageNumber="270" type="materials_examined">
<paragraph id="45153689FFD7FF8915DAFE051DED6FF7" blockId="4.[151,1437,151,501]" box="[199,1155,474,501]" pageId="4" pageNumber="270">
<emphasis id="77DEEA9BFFD7FF8915DAFE05181B6FF6" bold="true" box="[199,373,474,501]" pageId="4" pageNumber="270">Type material.</emphasis>
<materialsCitation id="F5C23CD4FFD7FF891461FE051DED6FF7" ID-GBIF-Occurrence="4133181305" box="[380,1155,474,501]" collectionCode="MNHN" country="Madagascar" location="Ankarana" municipality="Vadon" pageId="4" pageNumber="270" specimenCount="1" typeStatus="holotype">
<emphasis id="77DEEA9BFFD7FF891461FE05189C6FF7" bold="true" box="[380,498,475,501]" pageId="4" pageNumber="270">
<typeStatus id="9A11882BFFD7FF891461FE0518836FF7" box="[380,493,475,501]" pageId="4" pageNumber="270">Holotype</typeStatus>
:
</emphasis>
male, “
<collectingCountry id="3DBD7619FFD7FF89175AFE051BBD6FF7" box="[583,723,475,501]" name="Madagascar" pageId="4" pageNumber="270">Madagascar</collectingCountry>
,
<location id="40756052FFD7FF8917C1FE051A226FF7" LSID="urn:lsid:plazi:treatment:CD03879FFFD2FF89158AF9D41DED6FF7:40756052FFD7FF8917C1FE051A226FF7" box="[732,844,475,501]" country="Madagascar" municipality="Vadon" name="Ankarana" pageId="4" pageNumber="270">Ankarana</location>
, 25 X 45,
<collectingMunicipality id="A571ACF3FFD7FF8916A2FE051D626FF7" box="[959,1036,475,501]" pageId="4" pageNumber="270">Vadon</collectingMunicipality>
”(
<collectionCode id="23BBAE4CFFD7FF891103FE051D166FF7" box="[1054,1144,475,501]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="270">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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