190 lines
23 KiB
XML
190 lines
23 KiB
XML
<document id="690C47A9FC382393CBBAEF210AAD640E" ID-DOI="10.5281/zenodo.157788" ID-GBIF-Dataset="dbb243fc-1d94-4cef-8a13-410bc468103e" ID-ISSN="11755326" ID-Zenodo-Dep="157788" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461017416568" checkinUser="plazi" docAuthor="Merrin, Kelly L." docDate="2004" docId="C43687B62836D004177D394105AD745E" docLanguage="en" docName="zt00513.pdf" docOrigin="Zootaxa 513" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Notopais beddardi Merrin, 2004, n. sp." docType="treatment" docVersion="10" lastPageNumber="13" masterDocId="380FFFCE283FD00816753E4D01467678" masterDocTitle="Review of the deepwater asellote genus Notopais Hodgson, 1910 (Crustacea: Isopoda: Munnopsididae) with description of three new species from the southwestern Pacific" masterLastPageNumber="27" masterPageNumber="1" pageNumber="10" updateTime="1698572651680" updateUser="ExternalLinkService">
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<mods:title id="76DF916A1654E40245FBBF2C76F5D8A8">Review of the deepwater asellote genus Notopais Hodgson, 1910 (Crustacea: Isopoda: Munnopsididae) with description of three new species from the southwestern Pacific</mods:title>
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<mods:namePart id="59E952E8377DE282C51261CF7E21C92B">Merrin, Kelly L.</mods:namePart>
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<mods:typeOfResource id="278D0BA268DFF07649BF2DA2B2486830">text</mods:typeOfResource>
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<mods:title id="8D4793606207E9926538F7329ED54D6E">Zootaxa</mods:title>
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<mods:part id="05DAC8A9ECCB5F8387868875BF7FE5AE">
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<mods:date id="8F956D2B8560CFC5F9E0FCC1E372255D">2004</mods:date>
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<mods:detail id="A9CB7397497E3F223DD9962ACE872FA3" type="volume">
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<mods:number id="79257FF36D7A023075628F1D2FD43933">513</mods:number>
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<mods:identifier id="527384B0E8F3D1BCC9A2C24E535FD780" type="DOI">10.5281/zenodo.157788</mods:identifier>
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<treatment id="C43687B62836D004177D394105AD745E" ID-DOI="http://doi.org/10.5281/zenodo.6269546" ID-GBIF-Taxon="119601806" ID-Zenodo-Dep="6269546" LSID="urn:lsid:plazi:treatment:C43687B62836D004177D394105AD745E" httpUri="http://treatment.plazi.org/id/C43687B62836D004177D394105AD745E" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
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<subSubSection id="0485652B2836D001177D394103D6715E" box="[264,656,1804,1830]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="4C2036A02836D001177D394103D6715E" blockId="9.[264,656,1804,1830]" box="[264,656,1804,1830]" pageId="9" pageNumber="10">
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<heading id="176881CC2836D001177D394103D6715E" bold="true" box="[264,656,1804,1830]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
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<emphasis id="7EEBEAB22836D001177D394103D6715E" bold="true" box="[264,656,1804,1830]" pageId="9" pageNumber="10">
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<taxonomicName id="8B9F4D232836D001177D394100A3715E" ID-CoL="74BF8" box="[264,485,1804,1830]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="beddardi" status="sp. nov.">
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<emphasis id="7EEBEAB22836D001177D394100A3715E" bold="true" box="[264,485,1804,1830]" italics="true" pageId="9" pageNumber="10">Notopais beddardi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="65D857C92836D001179939410369715E" box="[492,559,1804,1830]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="D4A42A252836D001144B394103C1715E" box="[574,647,1804,1830]" captionStart="FIGURE 3" captionStartId="10.[264,367,1808,1832]" captionTargetBox="[287,1310,765,1807]" captionTargetId="figure@10.[264,1323,728,1815]" captionTargetPageId="10" captionText="FIGURE 3. Notopais beddardi n. sp. Female holotype, 4.0 mm. A, dorsal view; B, lateral view; C, left antenna 1 and 2, dorsal view; D, left antenna 2, ventral view; E, right pereopod 2; F, operculum." httpUri="https://zenodo.org/record/157791/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0485652B2836D002177D391A029A7726" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="4C2036A02836D002177D391A029A7726" blockId="9.[264,1322,1879,1905]" lastBlockId="10.[264,1323,284,670]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<emphasis id="7EEBEAB22836D001177D391A00AC7108" box="[264,490,1879,1904]" italics="true" pageId="9" pageNumber="10">Material examined</emphasis>
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.—
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<typeStatus id="932488022836D001146F391A03CD7109" box="[538,651,1879,1905]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
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. Female (4.0 mm), Station 149H, off Cumberland Bay, Kerguelen Island, southern Indian Ocean,
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<geoCoordinate id="29AB50672835D00214833F510215774E" box="[758,851,284,310]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="-48.75">48º45’S</geoCoordinate>
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<geoCoordinate id="29AB50672835D002152E3F5102FA774E" box="[859,956,284,310]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="69.23333">69º14’E</geoCoordinate>
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,
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<date id="382110602835D00215BC3F5105CB774E" box="[969,1165,284,310]" pageId="10" pageNumber="11" value="1874-01-29">29 January 1874</date>
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, 127 fathoms (
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<quantity id="8B679B452835D00217653F0900197726" box="[272,351,324,350]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="10" pageNumber="11" unit="m" value="232.0">232 m</quantity>
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), volcanic mud,
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<collectionCode id="2A8EAE652835D00214543F0903277726" LSID="urn:lsid:biocol.org:col:15108" box="[545,609,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15108" name="Embrapa Gado de Corte" pageId="10" pageNumber="11">HMS</collectionCode>
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<emphasis id="7EEBEAB22835D002141C3F0903AB7725" box="[617,749,324,349]" italics="true" pageId="10" pageNumber="11">Challenger</emphasis>
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(
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<collectionCode id="2A8EAE652835D00214893F0902107726" box="[764,854,324,350]" pageId="10" pageNumber="11">BMNH</collectionCode>
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2004.677).
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</paragraph>
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</subSubSection>
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<subSubSection id="0485652B2835D003174D3F2105CF722E" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
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<paragraph id="4C2036A02835D002174D3F21001F74E6" blockId="10.[264,1323,284,670]" pageId="10" pageNumber="11">
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<emphasis id="7EEBEAB22835D002174D3F21008477FD" box="[312,450,364,389]" italics="true" pageId="10" pageNumber="11">Description</emphasis>
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:
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<emphasis id="7EEBEAB22835D00217A73F21037B77FD" box="[466,573,364,389]" italics="true" pageId="10" pageNumber="11">
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<typeStatus id="932488022835D00217A73F21037B77FD" box="[466,573,364,389]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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</emphasis>
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. Body 2.4 times as long as greatest width (from spine to spine) of pereonite 2; widest at pereonite 4; cuticle not highly calcified, lightly setose. Cephalon lightly setose with 2 dorsal spines; posterior margins rounded, but almost square; ridge encompassing antennae with no extension. Pereonites 1–4 anterior margins each with 4 well developed spines; pereonite 5 anterior margin with evenly spaced stiff simple setae; pereonite 5 with anterolateral margin rounded; pereonites 1–4 anterolateral margins with spines, 5–7 rounded and smooth; pereonites 1–4 each with pair of lateral, submarginal spines.
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</paragraph>
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<caption id="18E066282835D002177D395D04637132" httpUri="https://zenodo.org/record/157791/files/figure.png" pageId="10" pageNumber="11" targetBox="[287,1310,765,1807]" targetPageId="10">
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<paragraph id="4C2036A02835D002177D395D04637132" blockId="10.[264,1323,1808,1866]" pageId="10" pageNumber="11">
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<emphasis id="7EEBEAB22835D002177D395D00CC7150" bold="true" box="[264,394,1808,1832]" pageId="10" pageNumber="11">FIGURE 3.</emphasis>
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<taxonomicName id="8B9F4D232835D00217E4395C031E7150" box="[401,600,1809,1832]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="beddardi" status="sp. nov.">
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<emphasis id="7EEBEAB22835D00217E4395C031E7150" box="[401,600,1809,1832]" italics="true" pageId="10" pageNumber="11">Notopais beddardi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="65D857C92835D002142B395C03DE7150" box="[606,664,1809,1832]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
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Female holotype, 4.0 mm. A, dorsal view; B, lateral view; C, left antenna 1 and 2, dorsal view; D, left antenna 2, ventral view; E, right pereopod 2; F, operculum.
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</paragraph>
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</caption>
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<paragraph id="4C2036A02834D003174D3F51056A740E" blockId="11.[264,1325,284,1870]" pageId="11" pageNumber="12">Antenna 1 article 1 rectangular and elongate, 1.7 times as long as wide, dorsal surface concave and forms shallow depression, distal margin with 5 robust setae; article 2 elongate, 0.4 times as long as article 1, 2.4 times as long as wide, with 2 robust setae (distal). Antenna 2 articles 1–3 more or less triangular; article 1 spine on distolateral angle with 3 additional robust setae on ventral side; article 2 inferior margin 1.2 times as long as article 1, distolateral angle with spine and 4 terminal robust setae, distoinferior margin with 2 robust setae; article 3 inferior margin 1.3 times as long as article 1, with 5 long robust and 3 short robust setae on distosuperior rim of article, distoinferior margin with 1 robust seta; article 4 small, 0.6 times as long as article 1, with no ornamentation.</paragraph>
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<paragraph id="4C2036A02834D003174D3CC902EF75CE" blockId="11.[264,1325,284,1870]" pageId="11" pageNumber="12">Pereopod 2 basis 2.7 times as long as wide, inferior margin with 6 setae (all SS); ischium 2.1 times as long as wide, inferior margin with 7 setae (1 SS, 6 submarginal SS), lateral face with 3 setae (all SS), superior margin with 2 setae (both SS, in distal half); merus 1.7 times as long as wide, inferior margin with 4 setae (all SS), distosuperior margin with 4 setae (all SS); carpus 6.0 times as long as wide, inferior margin with 13 setae (all RS, evenly spaced), superior margin with 4 setae (all SS, in distal half); propodus 8.2 times as long as wide, inferior margin with 6 setae (all SS), superior margin with 3 setae (all SS, distal half); dactylus 9.7 times as long as proximal width.</paragraph>
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<paragraph id="4C2036A02834D003174D3D8905CF722E" blockId="11.[264,1325,284,1870]" pageId="11" pageNumber="12">
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Pleon 0.6 times as long as proximal width, lateral sides rounded and posterior end coming to a rounded point, with scattered simple setae. Operculum 1.9 times as long as proximal width, medial keel with evenly spaced downward facing robust setae, lateral margins and distal surface with few setae (unable to determine actual
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<typeStatus id="932488022834D003124C3A71053A722E" box="[1081,1148,1084,1110]" pageId="11" pageNumber="12">types</typeStatus>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="0485652B2834D004174D3A2903F377FE" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="discussion">
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<paragraph id="4C2036A02834D004174D3A2903F377FE" blockId="11.[264,1325,284,1870]" lastBlockId="12.[264,1323,284,550]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<emphasis id="7EEBEAB22834D003174D3A2900D87205" box="[312,414,1124,1149]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
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:
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<taxonomicName id="8B9F4D232834D00317D83A2903C17205" box="[429,647,1124,1149]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beddardi" status="sp. nov.">
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<emphasis id="7EEBEAB22834D00317D83A2903C17205" box="[429,647,1124,1149]" italics="true" pageId="11" pageNumber="12">Notopais beddardi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="65D857C92834D00314F93A29038D7206" box="[652,715,1124,1150]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
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can be identified by the combination of having only two dorsal spines on the cephalon; anterior facing marginal spines on only the first four pereonites, with robust setae on the corresponding margin of the fifth; the unique setal combinations of the first two articles in antenna 1 and the first three articles in antenna 2; the rounded anterior margin of pereonite 5; and the anterolateral margins of pereonites 5 and 6 rounded and smooth, and not coming to a point. The spines on the anterior margin of article 4 on
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<taxonomicName id="8B9F4D232834D00317E23B1903627315" box="[407,548,1364,1389]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beddardi">
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<emphasis id="7EEBEAB22834D00317E23B1903627315" box="[407,548,1364,1389]" italics="true" pageId="11" pageNumber="12">N. beddardi</emphasis>
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</taxonomicName>
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are set on the absolute anterior margin, similar to that seen in
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<taxonomicName id="8B9F4D232834D00313653B19001A73ED" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="spicata">
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<emphasis id="7EEBEAB22834D00313653B19001A73ED" italics="true" pageId="11" pageNumber="12">N. spicata</emphasis>
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</taxonomicName>
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(although they are not as pronounced or as acutely angled) instead of almost being dorsal as is in
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<taxonomicName id="8B9F4D232834D00317C83BE903CC73C5" box="[445,650,1444,1469]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quadrispinosa">
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<emphasis id="7EEBEAB22834D00317C83BE903CC73C5" box="[445,650,1444,1469]" italics="true" pageId="11" pageNumber="12">N. quadrispinosa</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="8B9F4D232834D00314E93BE9026D73C5" box="[668,811,1444,1469]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beddardi">
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<emphasis id="7EEBEAB22834D00314E93BE9026D73C5" box="[668,811,1444,1469]" italics="true" pageId="11" pageNumber="12">N. beddardi</emphasis>
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</taxonomicName>
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does not have anterior spines on the fifth pereonite as in
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<taxonomicName id="8B9F4D232834D00317B63B8103C9739D" box="[451,655,1484,1509]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quadrispinosa">
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<emphasis id="7EEBEAB22834D00317B63B8103C9739D" box="[451,655,1484,1509]" italics="true" pageId="11" pageNumber="12">N. quadrispinosa</emphasis>
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</taxonomicName>
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, instead it has a row of short stout setae. The shape of pereonites 5–7 also differs between the two species. In
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<taxonomicName id="8B9F4D232834D00315D23BB905737075" box="[935,1077,1524,1549]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beddardi">
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<emphasis id="7EEBEAB22834D00315D23BB905737075" box="[935,1077,1524,1549]" italics="true" pageId="11" pageNumber="12">N. beddardi</emphasis>
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</taxonomicName>
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these pereonites are broader than those in
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<taxonomicName id="8B9F4D232834D003147E38510391704D" box="[523,727,1564,1589]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quadrispinosa">
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<emphasis id="7EEBEAB22834D003147E38510391704D" box="[523,727,1564,1589]" italics="true" pageId="11" pageNumber="12">N. quadrispinosa</emphasis>
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</taxonomicName>
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. Pereonite
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<quantity id="8B679B452834D003152B385102CA704E" box="[862,908,1564,1590]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="11" pageNumber="12" unit="in" value="5.0">5 in</quantity>
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<taxonomicName id="8B9F4D232834D00315E138510567704D" box="[916,1057,1564,1589]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beddardi">
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<emphasis id="7EEBEAB22834D00315E138510567704D" box="[916,1057,1564,1589]" italics="true" pageId="11" pageNumber="12">N. beddardi</emphasis>
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</taxonomicName>
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has a rounded anterolateral margin, and the corresponding margins of pereonites 6 and 7 are not as acute as those in
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<taxonomicName id="8B9F4D232834D003171A3821037D70FD" box="[367,571,1644,1669]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quadrispinosa">
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<emphasis id="7EEBEAB22834D003171A3821037D70FD" box="[367,571,1644,1669]" italics="true" pageId="11" pageNumber="12">N. quadrispinosa</emphasis>
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</taxonomicName>
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. The spination of both antennae are also different between the two species. The first article of antenna
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<quantity id="8B679B452834D003149638D9025770D6" box="[739,785,1684,1710]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="12" unit="in" value="1.0">1 in</quantity>
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<taxonomicName id="8B9F4D232834D003156C38D902E370D5" box="[793,933,1684,1709]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beddardi">
|
||
<emphasis id="7EEBEAB22834D003156C38D902E370D5" box="[793,933,1684,1709]" italics="true" pageId="11" pageNumber="12">N. beddardi</emphasis>
|
||
</taxonomicName>
|
||
has long robust setae around the distal margin, while
|
||
<taxonomicName id="8B9F4D232834D003147338F1039370AD" box="[518,725,1724,1749]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quadrispinosa">
|
||
<emphasis id="7EEBEAB22834D003147338F1039370AD" box="[518,725,1724,1749]" italics="true" pageId="11" pageNumber="12">N. quadrispinosa</emphasis>
|
||
</taxonomicName>
|
||
has robust setae along the superior margin. The number of robust setae on the spines of the first two antennal articles of antenna 2 also differ, with
|
||
<taxonomicName id="8B9F4D232834D0031706394100B9715D" box="[371,511,1804,1829]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beddardi">
|
||
<emphasis id="7EEBEAB22834D0031706394100B9715D" box="[371,511,1804,1829]" italics="true" pageId="11" pageNumber="12">N. beddardi</emphasis>
|
||
</taxonomicName>
|
||
having many more robust setae on the distosuperior margins than in
|
||
<taxonomicName id="8B9F4D232834D003177D397900927135" box="[264,468,1844,1869]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quadrispinosa">
|
||
<emphasis id="7EEBEAB22834D003177D397900927135" box="[264,468,1844,1869]" italics="true" pageId="11" pageNumber="12">N. quadrispinosa</emphasis>
|
||
</taxonomicName>
|
||
which has a single robust seta terminating each spine. The position of robust setae on the distal margin of the article 3 of antenna 2 also differs between the two species, with those of
|
||
<taxonomicName id="8B9F4D232833D00414603F0903E57725" box="[533,675,324,349]" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beddardi">
|
||
<emphasis id="7EEBEAB22833D00414603F0903E57725" box="[533,675,324,349]" italics="true" pageId="12" pageNumber="13">N. beddardi</emphasis>
|
||
</taxonomicName>
|
||
being on the superior margin, while on
|
||
<taxonomicName id="8B9F4D232833D00412F93F09007977FD" class="Malacostraca" family="Munnopsidae" genus="Notopais" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quadrispinosa">
|
||
<emphasis id="7EEBEAB22833D00412F93F09007977FD" italics="true" pageId="12" pageNumber="13">N. quadrispinosa</emphasis>
|
||
</taxonomicName>
|
||
they are on the inferior margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0485652B2833D004174D3FD9001D77AE" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="4C2036A02833D004174D3FD9001D77AE" blockId="12.[264,1323,284,550]" pageId="12" pageNumber="13">
|
||
<emphasis id="7EEBEAB22833D004174D3FD9008177D5" box="[312,455,404,429]" italics="true" pageId="12" pageNumber="13">Distribution</emphasis>
|
||
: Known only from
|
||
<typeStatus id="932488022833D00414CB3FD903B777D6" box="[702,753,404,430]" pageId="12" pageNumber="13">type</typeStatus>
|
||
locality, off Kerguelen Island, southern Indian Ocean.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0485652B2833D004174D3FA905AD745E" pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph id="4C2036A02833D004174D3FA905AD745E" blockId="12.[264,1323,284,550]" pageId="12" pageNumber="13">
|
||
<emphasis id="7EEBEAB22833D004174D3FA900F37785" box="[312,437,484,509]" italics="true" pageId="12" pageNumber="13">Etymology</emphasis>
|
||
: For Frank E. Beddard, the distinguished biologist, who described the many isopods which were collected during the
|
||
<emphasis id="7EEBEAB22833D00415443C4102F3745D" box="[817,949,524,549]" italics="true" pageId="12" pageNumber="13">Challenger</emphasis>
|
||
expedition of
|
||
<date id="382110602833D004122A3C4105A0745E" box="[1119,1254,524,550]" pageId="12" pageNumber="13" value="1873" valueMax="1876">1873–1876</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |