342 lines
61 KiB
XML
342 lines
61 KiB
XML
<document id="BE84AA11CF08DD3B65ADFF180CA7FB6D" ENCODING="UTF-8" ID-GBIF-Dataset="3a2de000-203e-4438-b4ef-77c37c83c5ba" ModsDocID="z00105p001" checkinTime="1246626767654" checkinUser="thomas" docAuthor="David G. Smith & Jeffrey T. Williams" docDate="2002" docId="9C41683B5E9A70F101035390F972B54E" docLanguage="en" docName="2002_Smith_Williams_gg1.xml" docOrigin="Zootaxa 105" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:2EE9E183-4F8E-4125-BEE0-BD1F29307B6C" docTitle="Enneanectes Jordan and Evermann 1895" docType="treatment" docVersion="14" lastPageNumber="7" masterDocId="FE8B598E79ED08AFF0FB8F000DAA22EF" masterDocTitle="History and status of the genera Enneanectes and Axoclinus (Teleostei: Blennioidei: Tripterygiidae)." masterLastPageNumber="10" masterPageNumber="1" pageNumber="2" updateTime="1701313826093" updateUser="plazi">
|
||
<mods:mods id="4328834A213C3D839D4447504E140792" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="13B786FF86F7114F1F66CB6293B49EF3">
|
||
<mods:title id="35CC95111A308E6DF5B394D3B097CE14">History and status of the genera Enneanectes and Axoclinus (Teleostei: Blennioidei: Tripterygiidae).</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="7E7754B47E954D489324542E1D73F3CD" type="personal">
|
||
<mods:role id="B3948DD6D0A8EBE12F8267A1FC93A1DA">
|
||
<mods:roleTerm id="EEA3674FF61474174D8D16460E26A466">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D5742A6AA602C1DA954CDF3B55BB5CBC">David G. Smith</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="997AF38AD6675F200C72B6F6AC1428A6" type="personal">
|
||
<mods:role id="52A026F8199E6CBA5ED8307FFABA8528">
|
||
<mods:roleTerm id="36E19824DDB538585965DB8068E3F8A0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="640310E46D4A116BF1F4934053A19576">Jeffrey T. Williams</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="6D3E4D7807ECC7A8370F1614856413DE">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A2AB915F6641FF544A8CFFCDC3461663" type="host">
|
||
<mods:titleInfo id="14C2B04B847A30B01A582D2D70B82A3B">
|
||
<mods:title id="AA44C5B29BD5B91EE312A16FBC757990">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="3B4FDCD4CA28DDD9A83E426B83DEF351">
|
||
<mods:date id="88937A3704C20159B61C22DDDC18E11F">2002</mods:date>
|
||
<mods:detail id="11CA4EE99A1C95BEF1859DDFFAA48180" type="volume">
|
||
<mods:number id="42F65D0BB41D87E23250399D09344160">105</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="2C077C7302FAF7E1251D22C84AFD9D86" unit="page">
|
||
<mods:start id="FAE87D73F6E8153339901560DBD48993">1</mods:start>
|
||
<mods:end id="BC251F06B52EDDC0F252B2C93BBB77A6">10</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="7A87ACF4435050738469984FCBD0C0BD">
|
||
<mods:url id="B27E61B4395A5D3613BD048C5BA63B6E">http://www.zoobank.org/urn:lsid:zoobank.org:pub:2EE9E183-4F8E-4125-BEE0-BD1F29307B6C</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="3220971CAA897D6702CC7D4BE8814395">journal article</mods:classification>
|
||
<mods:identifier id="EB146817D9E0692EBBFCB2E2BA065307" type="Plazi-Custom">z00105p001</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="9C41683B5E9A70F101035390F972B54E" ID-DOI="http://doi.org/10.5281/zenodo.6277964" ID-GBIF-Taxon="100122983" ID-Zenodo-Dep="6277964" LSID="urn:lsid:plazi:treatment:9C41683B5E9A70F101035390F972B54E" httpUri="http://treatment.plazi.org/id/9C41683B5E9A70F101035390F972B54E" lastPageNumber="7" pageNumber="2">
|
||
<subSubSection id="476302EDDA8C31EAF1658E393F3701BE" type="nomenclature">
|
||
<paragraph id="50059A48A5EA973808F154B26521CE5E">
|
||
[[
|
||
<taxonomicName id="DBDEB276B2CB8ECDF4AA2F53A533F01B" ID-CoL="4BXB" ID-ENA="722435" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Enneanectes Jordan and Evermann, 1895</taxonomicName>
|
||
]]
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9025F0A1BB3DA6AF955DCE54A96CC33" type="discussion">
|
||
<paragraph id="8E3B8934DAF0C592349C8FB02C6158A5" pageNumber="2">History of the problem</paragraph>
|
||
<paragraph id="0C3F64E14C0185B6B58EC3FBA0EC5956" pageNumber="2">The problem involves two distinct biological species, three nominal species, and three nominal genera. The names have been applied to different species at different times. The two biological species are characterized mainly by the configuration of the lateral line (Fig. 1). In the first species, which we will call Species 1, the lateral line is continuous, with an anterior section of tubed scales extending to approximately the origin of the third dorsal fin followed by a few notched scales. The lateral line descends gradually from the upper end of the opercle to the midlateral line, and there are 4-5 scale rows between the lateral line and the first dorsal fin. The dorsal rays are III + XII + 9-10, and the anal rays are II, 16-17 (Allen and Robertson, 1994: 219) In Species 2, the lateral line is discontinuous. The anterior section of tubed scales extends more or less horizontally from the opercle to approximately the posterior end of the second dorsal fin; it is followed by a series of notched scales that begins two scale rows ventral to the anterior section. There are three rows of scales between the lateral line and the first dorsal fin. Dorsal rays are III + XI-XII + 7-8, and anal rays are II, 14-15 (Allen and Robertson, 1994: 221).</paragraph>
|
||
<paragraph id="CCB3B9D924203993E967F4FF36C06EEB" lastPageNumber="3" pageNumber="2">
|
||
The nomenclatural history begins with the description of
|
||
<taxonomicName id="2C02E5CBCED2BCC219B8B54CC015CC53" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="carminale">Tripterygium carminale</taxonomicName>
|
||
by Jordan and Gilbert (1882: 362), based on four specimens collected in a tide pool at Mazatlan, on the Pacific coast of Mexico. The lateral line is described as “extending to opposite last ray of soft dorsal, ascending anteriorly but without convex curve.” Fin ray counts are given as D. III - XI, 9, A. II, 17. Based on the form of the lateral line and the fin-ray counts,
|
||
<taxonomicName id="2B0BBC4E5B91C484F8DCEFA77DE5A808" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="carminale">Tripterygium carminale</taxonomicName>
|
||
agrees with Species 1. The holotype was given as USNM 28118, but neither it nor any of the three paratypes can be found today. It is assumed that they were destroyed in the fire at Indiana University in 1883 (Brittan, 1997: 234), as no mention of them exists after that time. Jordan often received USNM catalog numbers for specimens on which he and his group were working at Indiana; the specimens would be forwarded to the Smithsonian when the work was completed. If these specimens existed today, they would be either at the USNM or at the CAS, where the Indiana University collection was ultimately transferred. We confirm that the specimens are not present in the USNM collection, and they were not mentioned by
|
||
<normalizedToken id="65A4A9D4698DFAB679C2555A7796BE03" originalValue="Böhlke">Boehlke</normalizedToken>
|
||
(1952) in his type catalog of the Stanford collection (which then held the former Indiana University collection and was subsequently moved to the CAS).
|
||
</paragraph>
|
||
<paragraph id="EAEABE99BE7D0C03700F3EF44C347765" pageNumber="3">
|
||
Thirteen years later, Jordan (1895: 501) placed the species epithet in a new genus,
|
||
<taxonomicName id="146F867B7583DE46B58C84820A418CD4" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
, based on a very brief diagnosis of the characters in which
|
||
<taxonomicName id="B934D2CDF326641ED1F1123CDB1B3888" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="carminalis">carminale</taxonomicName>
|
||
differed from
|
||
<taxonomicName id="5979DB1E0B61C2BC61FD3EA08EEF4D15" LSID-ZBK="urn:lsid:zoobank.org:act:C2E64947-48AA-45F3-8054-2A00F93B227A" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygion Risso 1827:241" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Tripterygion Risso</taxonomicName>
|
||
(the correct spelling). Jordan stated that
|
||
<taxonomicName id="206659EF06C7BEDDBAE8270A564AC5DC" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
was “framed...by Jordan and Evermann,” suggesting that it was to be published in the upcoming“Fishes of North and Middle America” (Jordan and Evermann, 1898), which indeed contains a full description of the genus, attributed to Jordan and Evermann, 1895. It is significant that Jordan and Evermann did not use one of the types in their description, but rather a small specimen collected by Jordan and his colleagues during a later trip to Mexico. This is further evidence that the types were missing at least as early as 1895. They also expressed some doubt about the identity of their specimen, stating that it “...differs somewhat in the count of the fin rays; but the very small size of the specimen prevents us from being entirely sure of its correctness.”
|
||
</paragraph>
|
||
<paragraph id="D87D8A6373B2A602872297BA3636A63F" pageNumber="4">
|
||
Brock (1938: 131) collected two specimens that he identified as
|
||
<taxonomicName id="2FEE2BE1F09428CF596B444D03F03C7B" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="carminalis">Enneapterygius carminalis</taxonomicName>
|
||
from Cape San Lucas, Baja California. He also briefly discussed the generic problem, saying that the specimens agreed with
|
||
<taxonomicName id="E9B3F0A55C51DBAC734155AEF52B1D41" LSID-ZBK="urn:lsid:zoobank.org:act:7C622ED0-36C5-494A-B739-8ED707EBB3EC" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias jordani Evermann & Marsh 1899:357" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="jordani">Gillias jordani Evermann and Marsh, 1899</taxonomicName>
|
||
, “in the generic characters they used to separate
|
||
<taxonomicName id="D887FC4E19B1528F3B9B927B11F16CFA" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
from
|
||
<taxonomicName id="C0AC575952E80D0222FCA6D815124BC7" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
....Even if
|
||
<taxonomicName id="7D8E7C926A2EA754F1DC50BF88267930" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
be withdrawn from the synonymy of
|
||
<taxonomicName id="082C657B046B060ABE6175BC28238486" LSID-ZBK="urn:lsid:zoobank.org:act:A085B8E9-4680-4C66-8184-1BB67ADB5A1E" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius Rüppell 1835:2" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneapterygius</taxonomicName>
|
||
,
|
||
<taxonomicName id="3B63BAA55AFD707606716723DF8F1F01" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
must still remain a synonym of
|
||
<taxonomicName id="73030CFAC18CCF91AB0ED9B3E33A6486" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
.” Brock thus framed the nomenclatural question that would persist over the next 60 years. Two years later, Brock (1940: 33) revisited the problem in more detail. He decided that the specimens (Species 1) he had identified as
|
||
<taxonomicName id="62BE3D058CF285FBCA1455B8BE95302B" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="carminalis">E. carminalis</taxonomicName>
|
||
were not the species described by Jordan and Gilbert, and he therefore described them as a new species,
|
||
<taxonomicName id="C1E8751EC83C9638B34124925C97D16A" LSID-ZBK="urn:lsid:zoobank.org:act:F3C7B4C4-7567-4A33-8FB4-743F2E71647D" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius storeyae Brock 1940:33" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="storeyae">Enneapterygius storeyae</taxonomicName>
|
||
(he continued to regard
|
||
<taxonomicName id="A90C5B9E64B80C1F759300C25A3DB70A" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
as a synonym of
|
||
<taxonomicName id="BE5361F4D220DE7AC1FC0D572F96E4BC" LSID-ZBK="urn:lsid:zoobank.org:act:A085B8E9-4680-4C66-8184-1BB67ADB5A1E" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius Rüppell 1835:2" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneapterygius</taxonomicName>
|
||
). He also discussed the confusion between the species called here
|
||
<taxonomicName id="8DC555B92BCEBDDF2067D9E1FC694869" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="carminalis">E. carminalis</taxonomicName>
|
||
and
|
||
<taxonomicName id="AA5698B7680E85B9B41677CE056B4438" LSID-ZBK="urn:lsid:zoobank.org:act:F3C7B4C4-7567-4A33-8FB4-743F2E71647D" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius storeyae Brock 1940:33" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="storeyae">E. storeyae</taxonomicName>
|
||
, asserting that the original description of
|
||
<taxonomicName id="74CBA4E0367E44850C4FE63B2F0B0245" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
could apply to either species. In order to resolve the ambiguity, and in the absence of a holotype, Brock designated a neotype for
|
||
<taxonomicName id="A577C13C7A851047BE5D594002D66712" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
. The specimen (Species 2) he chose was SU 3854, the specimen Jordan and Evermann (in Jordan 1895) described as representative of the type species of
|
||
<taxonomicName id="F5C10CCB8EDA1C2DC5E921A80C65B79E" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
. He selected this specimen on the perfectly rational assumption that Jordan would have been able to recognize the species he himself had described 13 years earlier. Brock also noted that the “American members of the genus
|
||
<taxonomicName id="868C4459BF28C46EE8A73A1E9AE74844" LSID-ZBK="urn:lsid:zoobank.org:act:A085B8E9-4680-4C66-8184-1BB67ADB5A1E" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius Rüppell 1835:2" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneapterygius</taxonomicName>
|
||
” differ from the Indo-West Pacific species in the scalation of the head, and, if this character is agreed to be of generic significance, “
|
||
<taxonomicName id="E065E84AA4B785549A50BED6343EC93A" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
must replace
|
||
<taxonomicName id="AA92A7535338127F1DF7D4A246AE05D6" LSID-ZBK="urn:lsid:zoobank.org:act:A085B8E9-4680-4C66-8184-1BB67ADB5A1E" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius Rüppell 1835:2" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Enneapterygius</taxonomicName>
|
||
, and, of course,
|
||
<taxonomicName id="75EA332F82A0035EE1B0C47CE6202B75" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
, for these American species.”
|
||
</paragraph>
|
||
<paragraph id="388AE443230FC9D7518C5086A1FBED29" lastPageNumber="5" pageNumber="4">
|
||
Fowler (1944: 286) described a new species,
|
||
<taxonomicName id="5755D13807D8E8259EE5877CC08F1908" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="sexmaculatus">Gillias sexmaculatus</taxonomicName>
|
||
, from the Pacific coast of Panama. He described a discontinuous lateral line and three scales between the lateral line and the spinous dorsal-fin origin, characters that align it with Species 2. He emphasized its distinction from the description of
|
||
<taxonomicName id="5A513F8696969AEC39049BA16D16743F" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="carminale">carminalis Jordan and Gilbert</taxonomicName>
|
||
and its resemblance to the Atlantic
|
||
<taxonomicName id="A8D191F8A7516C7DF4138212A546F2D1" LSID-ZBK="urn:lsid:zoobank.org:act:7C622ED0-36C5-494A-B739-8ED707EBB3EC" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias jordani Evermann & Marsh 1899:357" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="jordani">Gillias jordani Evermann and Marsh</taxonomicName>
|
||
. Fowler was thus basing his interpretation on Jordan and Gilbert’s original description of
|
||
<taxonomicName id="EA7A1A2102C466551B6E5F9CF37AA009" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="carminale">carminalis</taxonomicName>
|
||
(Species 1) and not on the specimen from Jordan (1895), which Brock had selected as the neotype (Species 2). Fowler described a new genus,
|
||
<taxonomicName id="EA1171C32BA6D5C3EC3E07FB426DCCA3" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
, based on a new species
|
||
<taxonomicName id="1729412E6015D8D4356C1494578D57AB" LSID-ZBK="urn:lsid:zoobank.org:act:F8918A5A-8FB4-4911-9F41-477B8B5D4642" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus lucillae Fowler 1944:289" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="lucillae">A. lucillae</taxonomicName>
|
||
.
|
||
<taxonomicName id="03A197930011286319E957A0C0D9270B" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
was distinguished mainly by the lateral line, described as “axial along side of body, incomplete, only extends as far as third dorsal.” Although
|
||
<taxonomicName id="79EE3A47C606D4653247B0CA435D24CD" LSID-ZBK="urn:lsid:zoobank.org:act:F8918A5A-8FB4-4911-9F41-477B8B5D4642" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus lucillae Fowler 1944:289" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="lucillae">Axoclinus lucillae</taxonomicName>
|
||
is distinct from Species 1 in its scale and fin-ray counts, the lateral-line character agrees, and
|
||
<taxonomicName id="09488FFB7E6FE7668AAB8842173E561A" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="4" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
is the genus in which Species 1 would be placed.
|
||
</paragraph>
|
||
<paragraph id="249C0CEE2B8D263F3A0B75477E57E0E0" pageNumber="5">
|
||
Fowler’s treatment of the nominal species carminalis and storeyae is confusing and contradictory. On page 288, he accepted
|
||
<taxonomicName id="315D4508C22F60F98FCCD52082701533" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminale">Tripterygium carminale Jordan and Gilbert</taxonomicName>
|
||
as the “genotype” of
|
||
<taxonomicName id="232408F6EC9209E714A3578CE803CF67" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
, listing some characters from the description (i.e., Species 1, not from Brock’s neotype, which represents Species 2) that “are all out of harmony with
|
||
<taxonomicName id="81233330805207F6DBDDC5A5A29AC7D1" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="sexmaculatus">Gillias sexmaculatus</taxonomicName>
|
||
.” Aside from noting its distinction from his new species
|
||
<taxonomicName id="F27009F3946EF932E1EE883857BDA548" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="sexmaculatus">sexmaculatus</taxonomicName>
|
||
, he did not treat it further. Also on page 288, he noted that “the
|
||
<taxonomicName id="AE225B3A64655A757556B290BD78E753" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="storyae">Gillias storyae [sic] (Brock)</taxonomicName>
|
||
” also belongs in
|
||
<taxonomicName id="6C3EB79C6D614957E6DF96B4E16FFD0E" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
, although he further noted that it differed in having “5 rows of scales between the lateral line and the base of the first dorsal, though the course of the lateral line is not described” or illustrated. Without knowing the condition of the lateral line, it is difficult to understand how Fowler determined that
|
||
<taxonomicName id="B48271297279D54BFE29281CA5901DC4" LSID-ZBK="urn:lsid:zoobank.org:act:F3C7B4C4-7567-4A33-8FB4-743F2E71647D" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius storeyae Brock 1940:33" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="storeyae">storeyae</taxonomicName>
|
||
belonged in
|
||
<taxonomicName id="A7F84339550A14F2EF91B1D9FF4DB838" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
. Then on page 289, he stated that his new species
|
||
<taxonomicName id="A266C827EE1E749FD063A52FC6718DA9" LSID-ZBK="urn:lsid:zoobank.org:act:F8918A5A-8FB4-4911-9F41-477B8B5D4642" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus lucillae Fowler 1944:289" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="lucillae">Axoclinus lucillae</taxonomicName>
|
||
“approaches
|
||
<taxonomicName id="508E220E55E9F313C66885A3208BCD82" LSID-ZBK="urn:lsid:zoobank.org:act:F3C7B4C4-7567-4A33-8FB4-743F2E71647D" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius storeyae Brock 1940:33" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="storeyae">Enneapterygius storeyae Brock</taxonomicName>
|
||
but [is] apparently quite different in coloration.” Here he seemed to be saying that
|
||
<taxonomicName id="18427B6C465C416B6FC33780D5E5C6C6" LSID-ZBK="urn:lsid:zoobank.org:act:F3C7B4C4-7567-4A33-8FB4-743F2E71647D" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius storeyae Brock 1940:33" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="storeyae">storeyae</taxonomicName>
|
||
belonged in
|
||
<taxonomicName id="EF4F44C485CEFB44402435CF1D1A042B" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="FE39542A86B06C0DD05174C2AA33E7EB" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
, where he had placed it on the previous page.
|
||
</paragraph>
|
||
<paragraph id="ABC94BD0A95E96965CFFF5910E631C47" pageNumber="5">
|
||
Schultz (1950: 268) added to the confusion by synonymizing the genera
|
||
<taxonomicName id="EE75EC78CD0173CCF1FF50CFB5526DC0" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
and
|
||
<taxonomicName id="009A38E31C849DABFEC6BC330CFDE403" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
with
|
||
<taxonomicName id="D4838F38449FDB65513E50D0A5292E89" LSID-ZBK="urn:lsid:zoobank.org:act:C2E64947-48AA-45F3-8054-2A00F93B227A" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygion Risso 1827:241" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Tripterygion Risso</taxonomicName>
|
||
and
|
||
<taxonomicName id="E5511D507FF243BCB3F496C7536CB9EE" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
with
|
||
<taxonomicName id="41AA5506263AEA93AC969B69A8C602F5" LSID-ZBK="urn:lsid:zoobank.org:act:F9BD3D42-105B-42EB-9BBF-CB059FBD25A9" family="Tripterygiidae" genus="Helcogramma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Helcogramma McCulloch & Waite 1918:51" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Helcogramma McCulloch and Waite</taxonomicName>
|
||
. He based this decision on the form of the lateral line, i.e., continuous in
|
||
<taxonomicName id="ED454574AB954EAA17C916CE45D5FC48" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
and
|
||
<taxonomicName id="F647FE203CDCFC9B9F4865E2DA969BE6" LSID-ZBK="urn:lsid:zoobank.org:act:F9BD3D42-105B-42EB-9BBF-CB059FBD25A9" family="Tripterygiidae" genus="Helcogramma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Helcogramma McCulloch & Waite 1918:51" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Helcogramma</taxonomicName>
|
||
, discontinuous in
|
||
<taxonomicName id="DB70612666131C0047685DE2DEBEE549" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
,
|
||
<taxonomicName id="2026094F6FE6DBD490E03E524FA4053C" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2B48291D6144F303EBEF466100C4546B" LSID-ZBK="urn:lsid:zoobank.org:act:C2E64947-48AA-45F3-8054-2A00F93B227A" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygion Risso 1827:241" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Tripterygion</taxonomicName>
|
||
. The species
|
||
<taxonomicName id="CDDE87D0D206AA3279D2B2C74BCF559F" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminale">carminalis</taxonomicName>
|
||
(i.e., based on the Jordan and Gilbert description, Species 1, not on the Brock neotype, Species 2) was therefore transferred to
|
||
<taxonomicName id="0E9A0158FBAD23BA598F29142D5604B6" LSID-ZBK="urn:lsid:zoobank.org:act:F9BD3D42-105B-42EB-9BBF-CB059FBD25A9" family="Tripterygiidae" genus="Helcogramma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Helcogramma McCulloch & Waite 1918:51" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Helcogramma</taxonomicName>
|
||
, and
|
||
<taxonomicName id="172A2E42B533D6D2FD15D5721B060B47" family="Tripterygiidae" genus="Helcogramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="storeyae">storeyae</taxonomicName>
|
||
and
|
||
<taxonomicName id="6063188F46F572F843ACBFDD2072B005" family="Tripterygiidae" genus="Helcogramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="lucillae">lucillae</taxonomicName>
|
||
were included as synonyms. The species
|
||
<taxonomicName id="7A3F228C43A8A08056BC1DCF0F8095C8" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="sexmaculatus">sexmaculatus</taxonomicName>
|
||
was placed in
|
||
<taxonomicName id="0CAB8495D368536485EEA55FD1599F57" LSID-ZBK="urn:lsid:zoobank.org:act:C2E64947-48AA-45F3-8054-2A00F93B227A" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygion Risso 1827:241" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Tripterygion</taxonomicName>
|
||
. Schultz also claimed to have found the holotype of
|
||
<taxonomicName id="EE2E06CB1DDBA5429F9126D224F98CDC" family="Tripterygiidae" genus="Helcogramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminale">carminalis</taxonomicName>
|
||
, USNM 120946, which he removed from a syntypic lot of
|
||
<taxonomicName id="6D5FDBC8C0676B898264F1A7584E2814" LSID-ZBK="urn:lsid:zoobank.org:act:87EBA367-999C-4C39-862B-CAFFBB5328B4" family="Gobiesocidae" genus="Gobiesox" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobiesox zebra Jordan & Gilbert 1882:359" order="Gobiesociformes" pageNumber="5" phylum="Chordata" rank="species" species="zebra">Gobiesox zebra Jordan and Gilbert</taxonomicName>
|
||
. Although the specimen was collected at the same time and place as the type series, its type status is doubtful. In the first place, Jordan and Gilbert (1882: 363) said that the four types of
|
||
<taxonomicName id="2A36C1387D7BFEE5E5ABCA0EC8EA142C" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminale">carminalis</taxonomicName>
|
||
were “each about 1 1/2 inches long,” USNM 120946 is barely one inch in standard length and 1 1/4 inches (33 mm) in total length. Second, why would Jordan and Gilbert have put the holotype back into a jar full of another species? It seems much more likely that the specimen was just a stray that was mixed in with the type lot of
|
||
<taxonomicName id="1C29CB7791EBE8D7949973D9394E9658" LSID-ZBK="urn:lsid:zoobank.org:act:87EBA367-999C-4C39-862B-CAFFBB5328B4" family="Gobiesocidae" genus="Gobiesox" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobiesox zebra Jordan & Gilbert 1882:359" order="Gobiesociformes" pageNumber="5" phylum="Chordata" rank="species" species="zebra">Gobiesox zebra</taxonomicName>
|
||
; in their description of
|
||
<taxonomicName id="A22481145D656C849A0F17E147C3EC1A" LSID-ZBK="urn:lsid:zoobank.org:act:87EBA367-999C-4C39-862B-CAFFBB5328B4" family="Gobiesocidae" genus="Gobiesox" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobiesox zebra Jordan & Gilbert 1882:359" order="Gobiesociformes" pageNumber="5" phylum="Chordata" rank="species" species="zebra">G. zebra</taxonomicName>
|
||
, Jordan and Gilbert (1882: 359) said only that “about 30 specimens” were collected, suggesting that they did not count them very carefully or even look at them all. If the holotype of
|
||
<taxonomicName id="B94F2B504AF71FB619AF08D893CA043E" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminale">carminalis</taxonomicName>
|
||
really existed, as Schultz clearly believed, then Brock’s neotype designation would be invalid. Schultz followed this logic and stated that Brock’s “neotype ” (quotation marks used by Schultz) “does not belong to this genus and species [i.e.,
|
||
<taxonomicName id="B7AA809126A50913D343AC9C5F24AF8C" family="Tripterygiidae" genus="Helcogramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminale">Helcogramma carminale</taxonomicName>
|
||
as used by Schultz] but is a specimen of
|
||
<taxonomicName id="27840DC64D19F8898D788EEEB7639E2A" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="sexmaculatus">Tripterygion sexmaculatus (Fowler)</taxonomicName>
|
||
.”
|
||
</paragraph>
|
||
<paragraph id="A0B903BF62670EEAE4EDE2AF751BBEB6" lastPageNumber="6" pageNumber="5">
|
||
Rosenblatt (1960: 3) resurrected
|
||
<taxonomicName id="BCCE4D7C7CDFA9BE3C94F67C9D9FD411" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
, pointing out its differences from both
|
||
<taxonomicName id="82DC1470F0F9284CCC6A61D2F45956E2" LSID-ZBK="urn:lsid:zoobank.org:act:C2E64947-48AA-45F3-8054-2A00F93B227A" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygion Risso 1827:241" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Tripterygion</taxonomicName>
|
||
and
|
||
<taxonomicName id="E85417B22B533ED20B84EF9BC8AAF7C6" LSID-ZBK="urn:lsid:zoobank.org:act:A085B8E9-4680-4C66-8184-1BB67ADB5A1E" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius Rüppell 1835:2" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Enneapterygius</taxonomicName>
|
||
(which Schultz had also synonymized with
|
||
<taxonomicName id="43917B2945C1CCF4F2698B304D6F573D" LSID-ZBK="urn:lsid:zoobank.org:act:C2E64947-48AA-45F3-8054-2A00F93B227A" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygion Risso 1827:241" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Tripterygion</taxonomicName>
|
||
). He then summarized the nomenclatural history of
|
||
<taxonomicName id="CD42F3AA44756C5FD2CAB4D0070AC842" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="5" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
, emphasizing again that the specimen of “
|
||
<taxonomicName id="8DA5750828382C9FC186428F1D09329E" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
” used by Jordan and Evermann (in Jordan 1895) represented a different species (Species 2) from the one originally described by Jordan and Gilbert (Species 1) and that Brock’s
|
||
<taxonomicName id="529BC71842B73127D00472B09C5C69ED" LSID-ZBK="urn:lsid:zoobank.org:act:F3C7B4C4-7567-4A33-8FB4-743F2E71647D" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius storeyae Brock 1940:33" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="storeyae">storeyae</taxonomicName>
|
||
represented “the true
|
||
<taxonomicName id="8B8AA6AD79E4269391E3947CF7248E54" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="carminale">T. carminale</taxonomicName>
|
||
.” He outlined two alternative scenarios that would flow from this analysis and gave his reasons for choosing one of these scenarios:
|
||
</paragraph>
|
||
<paragraph id="281957856770C418ABFECC894F0E4BEC" pageNumber="6">
|
||
It would seem that the most logical course is to recognize as the type of
|
||
<taxonomicName id="D6B7CC288150CE2C69C1B974574B0DE9" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
the actual specimens upon which Jordan and Evermann based the generic description, namely
|
||
<taxonomicName id="4003C8BFB85F4F5D95D4FE10DB177A67" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="sexmaculatus">Gillias sexmaculatus Fowler</taxonomicName>
|
||
. The alternative to this [i.e., to recognize the nominal species that Jordan and Evermann had misidentified their specimen as] would be to replace
|
||
<taxonomicName id="1BFCA5DF3A0BFC65131BF60D35D1CBF5" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
with
|
||
<taxonomicName id="3E0A98424220F865F3091A0A49B2DDD4" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
, and refer the species here placed in
|
||
<taxonomicName id="F4DFA27B299D6E610643D824C20AFFE1" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
to
|
||
<taxonomicName id="5B21959C1FA5B4EA24A0E5F4AE1BE742" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
. This would mean that not one of the species of
|
||
<taxonomicName id="75F6C5CF3981F0F8A9755102C18550B5" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
would then agree in any generic character with the original description of the genus. This latter alternative, while admissible nomenclatorially, would create a zoological paradox. It is for this reason that
|
||
<taxonomicName id="3E5757E1C6B46D02E138D2AEC74529A3" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="sexmaculatus">Gillias sexmaculatus</taxonomicName>
|
||
is here recognized as the type species of
|
||
<taxonomicName id="F2414CC61625193D65772C88144754DE" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
. (Rosenblatt, 1960: 5)
|
||
</paragraph>
|
||
<paragraph id="4AB3B00D195EEA21A424922506936014" pageNumber="6">
|
||
Rosenblatt, however, was still under the impression that a holotype existed for
|
||
<taxonomicName id="73838E4992DD0A0B0F32655BDE03BEFC" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="carminale">Tripterygion carminale</taxonomicName>
|
||
and that the name carminale applied to the species actually described by Jordan and Gilbert (1882). In his view, Brock’s (1940) neotype designation was invalid, as evidenced by his use of quotation marks around the word “neotype.” As we have explained above, however, Schultz’s “holotype” is not a holotype, and Brock’s neotype designation stands.
|
||
</paragraph>
|
||
<paragraph id="E20A9870BC79132AB189C9FF7B97612B" pageNumber="6">
|
||
Almost all subsequent authors have followed Rosenblatt’s interpretation and assigned species to
|
||
<taxonomicName id="679C992B92E0C68D264AD60BD457B7AB" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
according to his concept of that genus. We note in particular Allen and Robertson (1994: 218), which has become a standard reference to fishes of the eastern tropical Pacific. Those authors dealt with all the eastern Pacific species of Tripterygiidae and explained the lateral-line feature that characterizes
|
||
<taxonomicName id="1CDE211A331DF5742F70634124C97F3C" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
and
|
||
<taxonomicName id="02E50F12F74291CA68B9C73D3203E44C" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
. A prevailing consensus has thus been achieved on the definition of these two genera and on the species that are assigned to each.
|
||
</paragraph>
|
||
<paragraph id="7735D9453F191AD8D3EBF6FD2542D5A6" lastPageNumber="7" pageNumber="6">
|
||
This consensus has been challenged by Fricke (1997: 564). He recognized Brock’s neotype designation (though incorrectly attributing it to Schultz) for
|
||
<taxonomicName id="D788BC6511EFDB2D3B69DAD9CF259ACF" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
as fixing the name to Jordan and Evermann’s (1895) species (i.e., Species 2, not the one originally described under that name by Jordan and Gilbert, Species 1). However, he incorrectly included
|
||
<taxonomicName id="F27335BA0672ACB7A823D7F85F72031C" LSID-ZBK="urn:lsid:zoobank.org:act:F3C7B4C4-7567-4A33-8FB4-743F2E71647D" family="Tripterygiidae" genus="Enneapterygius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneapterygius storeyae Brock 1940:33" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="storeyae">storeyae</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="21DA6D2D1DA2381167CFD42FDE72C27D" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
instead of treating it as the next available name for the original
|
||
<taxonomicName id="2C3674427DDAE407607D395AF4E1D0F3" LSID-ZBK="urn:lsid:zoobank.org:act:4A31B536-6B8B-43D8-B70C-3737C32D3AA7" family="Tripterygiidae" genus="Tripterygium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tripterygium carminale Jordan & Gilbert 1882:362" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="carminale">carminalis</taxonomicName>
|
||
of Jordan and Gilbert (Species 1). He correctly included
|
||
<taxonomicName id="49B67F0B3AE3EA49AE82BD25CF72E1C4" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="sexmaculatus">sexmaculatus</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="EA2C7C0C982A465AC9AEF2502AE44763" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
of Brock (Species 2). Then he incorrectly synonymized
|
||
<taxonomicName id="18D6B01771FD73ABAEDBE85DD5B5D8FF" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
with
|
||
<taxonomicName id="9AF23B17BADF2FEECAD35EBC3458AEDA" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
and transferred the species currently assigned to
|
||
<taxonomicName id="262FAAD139F9B69BA38650639512CA40" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
to
|
||
<taxonomicName id="4BEB009CF7912AAF7C92DCDE605DFE42" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
, along with
|
||
<taxonomicName id="F648CF07049D23947425D3D472E2AE1A" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
. The remaining species currently in
|
||
<taxonomicName id="9419547A8A6C9247D52E67FC2E0E364A" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
were transferred to
|
||
<taxonomicName id="5097B7C44F5D73AF9BE5CE8A0C32156C" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
. Fricke did not explain why one of the current
|
||
<taxonomicName id="F3FACBA01DAE052569A4D00EC53D29B3" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
species (
|
||
<taxonomicName id="49D9FE32B95C703365188FC55107982A" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="carminalis">carminalis</taxonomicName>
|
||
) remained in his concept of
|
||
<taxonomicName id="2A625972521D2289C3C20E66542E3299" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
while all of its congeners were moved to
|
||
<taxonomicName id="836BB98D9E65471DF9D12D56C4907129" LSID-ZBK="urn:lsid:zoobank.org:act:75DC3F0D-228F-4B30-9ABA-13F4B53F9DF3" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias Evermann & Marsh 1899:357" order="Perciformes" pageNumber="6" phylum="Chordata" rank="genus">Gillias</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FE519E3A0C86158BCD47375DD24E54BA" pageNumber="7">Conclusion</paragraph>
|
||
<paragraph id="1FA93C06B6E1ECA4C97D33C76267C6E6" pageNumber="7">
|
||
The resolution of this case rests upon whether Brock’s designation of a neotype for
|
||
<taxonomicName id="FDD0B28D082F6D2ACFF588FAA7EC4CC4" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="carminale">Tripterygion carminale Jordan and Gilbert</taxonomicName>
|
||
is accepted. If it is, then the specific name carminalis is applied to Species 2 and assigned to the genus
|
||
<taxonomicName id="EB418CB03CD709CDC37EB385A8C32152" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="7" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
. If the neotype is not accepted, then carminalis is applied to Species 1, and
|
||
<taxonomicName id="B1CBEFB82E5BC1518AB2169DBAFA0C9E" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="7" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
goes with it. It is fairly clear from the description that
|
||
<taxonomicName id="3B931E6C15E0F7EEE6C32B0F47B98C9B" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="carminale">Tripterygion carminale</taxonomicName>
|
||
as described by Jordan and Gilbert refers to the taxon we call Species 1. Jordan and Evermann (in Jordan, 1895) misidentified the specimen they chose as the type species of
|
||
<taxonomicName id="C6EC352632BBA4E2DB9D91AA60BF24B0" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="7" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
; it actually belongs to the taxon we call Species 2. Article 70.3 of the ICZN addresses the problem of misidentified type species.
|
||
</paragraph>
|
||
<paragraph id="020843CE82254939A662CE19F574E8F6" pageNumber="7">If an author discovers that a type species was misidentified..., the author may select, and thereby fix as type species, the species that will, in his or her judgment, best serve stability and universality, either 70.3.1. the nominal species previously cited as type species, or 70.3.2. the taxonomic species actually involved in the misidentification. If the latter choice is made, the author must refer to this Article and cite together both the name previously cited as type species and the name of the species selected (ICZN, 1999: 74).</paragraph>
|
||
<paragraph id="AB722AD1F4EE64C37F2CAF32B4959A2F" pageNumber="7">
|
||
We hereby invoke Article 70.3.2 and select “the taxonomic species actually involved in the misidentification,” i.e., the species represented by the specimen designated by Brock (1940) as the neotype of
|
||
<taxonomicName id="946BE00BD0585E48A334A9BAE76CE76C" family="Tripterygiidae" genus="Tripterygion" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="carminale">Tripterygion carminale Jordan and Gilbert, 1882</taxonomicName>
|
||
. This maintains prevailing usage by fixing the genus
|
||
<taxonomicName id="220C00A426F813E56442AA9CD3A2AA68" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="7" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
to the species that have been almost universally assigned to it since 1960.
|
||
<taxonomicName id="AA47D4E196AF88A73A71259670A4339E" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="carminalis">Enneanectes carminalis (Jordan and Gilbert, 1882)</taxonomicName>
|
||
becomes the valid name of the taxon we call Species 2, with
|
||
<taxonomicName id="49B62AEAB8BC6450154D65537D95FB06" LSID-ZBK="urn:lsid:zoobank.org:act:14735E09-CCE6-4914-B77B-50BE63ACDDD0" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gillias sexmaculatus Fowler 1944:286" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="sexmaculatus">Gillias sexmaculatus Fowler, 1944</taxonomicName>
|
||
as a junior synonym. The taxon we call Species 1 takes the next available name and becomes
|
||
<taxonomicName id="0F3EAD82F0DA3A5A6942620644E14A94" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="storeyae">Axoclinus storeyae (Brock, 1940)</taxonomicName>
|
||
. Table 1 summarizes the names applied to the two species by the various authors. Table 2 summarizes the historical and current placement of the species included in the two genera
|
||
<taxonomicName id="9A8D9BB78878CD29B65DA36A9A59B760" LSID-ZBK="urn:lsid:zoobank.org:act:671F7E3C-D3A5-4E77-AA43-F6BF07D334A2" family="Tripterygiidae" genus="Axoclinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Axoclinus Fowler 1944:288" order="Perciformes" pageNumber="7" phylum="Chordata" rank="genus">Axoclinus</taxonomicName>
|
||
and
|
||
<taxonomicName id="8A62067F4301C26D8563F344E0E67E87" LSID-ZBK="urn:lsid:zoobank.org:act:0DC6140F-53B4-4736-B530-A4DF2777144C" family="Tripterygiidae" genus="Enneanectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Enneanectes Jordan & Evermann 1895:501" order="Perciformes" pageNumber="7" phylum="Chordata" rank="genus">Enneanectes</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |