treatments-xml/data/98/24/16/982416653C1C4B39FF17BAEEFC99403F.xml
2024-06-21 12:44:59 +02:00

416 lines
66 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="21842E72A3BDC1271C3F149C61692594" ID-CLB-Dataset="55842" ID-DOI="10.11646/zootaxa.5128.4.1" ID-GBIF-Dataset="6793d5d0-7fcd-415a-8bba-2867cf76501c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6479917" ID-ZooBank="26CA4EF0-61A1-4B60-8C7F-99CA8B8F8700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650722012437" checkinUser="plazi" docAuthor="Pearson, Kaila A. M. &amp; Rouse, Greg W." docDate="2022" docId="982416653C1C4B39FF17BAEEFC99403F" docLanguage="en" docName="zootaxa.5128.4.1.pdf" docOrigin="Zootaxa 5128 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Galapagomystides patricki Pearson &amp; Rouse 2022, n. sp." docType="treatment" docVersion="8" lastPageNumber="480" masterDocId="641D6E1D3C0B4B24FF80BF01FFF34442" masterDocTitle="Vampire Worms; A revision of Galapagomystides (Phyllodocidae, Annelida), with the description of three new species" masterLastPageNumber="485" masterPageNumber="451" pageNumber="474" updateTime="1699329107976" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DB2E7377441CC4E2DB5E7825D4E1AAA0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="620D9E582ED54CC1E5C155264A68E65F">
<mods:title id="69C3AB2D81037C936AD1C4D91769321E">Vampire Worms; A revision of Galapagomystides (Phyllodocidae, Annelida), with the description of three new species</mods:title>
</mods:titleInfo>
<mods:name id="CE7AB97D8DEC654F22128A178E9AF076" type="personal">
<mods:role id="493E2858CA133C3418B3BB1B68E80A71">
<mods:roleTerm id="B63A5035E8E994D88DDC275B1013D90E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B0078A1DA397AD48E029F44F7153876">Pearson, Kaila A. M.</mods:namePart>
<mods:affiliation id="6178911F248C72FDE7F080C6C0D62160">Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA 92093 - 0202, USA.</mods:affiliation>
</mods:name>
<mods:name id="8A2505D2B22FBF4D087090F1D1CF6C76" type="personal">
<mods:role id="EEC9C1C7869FBBE3439B38150DA352C0">
<mods:roleTerm id="CCAA1202FA6DCED01553AB2FEDC8D567">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="33FBA59C0BF02F8DEC6C6A83D2C67652">Rouse, Greg W.</mods:namePart>
<mods:affiliation id="682758980B5EBE522679C2BB6DB59268">Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA 92093 - 0202, USA. &amp; South Australian Museum, North Terrace, Adelaide SA 5000 Australia.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B307D03E69083EAAFC57AAEE63EB3D86">text</mods:typeOfResource>
<mods:relatedItem id="0B39311FB4E48BFA41DB6F53DB0AB76E" type="host">
<mods:titleInfo id="F42C6019EEA9C9DC053F15976FF3703C">
<mods:title id="8C8D50EF1E11257E4D3C4D18372C1A29">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D62CDE5BCB15786B6EE20A6DC9D475A5">
<mods:date id="E08C88A856320070B6A6A30D0E345C17">2022</mods:date>
<mods:detail id="A5A7413761116B746A2C5AF55F760CCC" type="pubDate">
<mods:number id="16AB1A857014A9897ACFDBA2F42BE6D3">2022-04-22</mods:number>
</mods:detail>
<mods:detail id="0CA44D87ADB052869AEAF8BFE79489E3" type="volume">
<mods:number id="8F6886814090EFE0CA5E7C5615D8CF8C">5128</mods:number>
</mods:detail>
<mods:detail id="2B37D4B0F5E226B5D8E471730FAA2D34" type="issue">
<mods:number id="DBDDE94E195D9117CF18896FC600D6BF">4</mods:number>
</mods:detail>
<mods:extent id="6A4E8BEE34C81124A95AC57132DA68D2" unit="page">
<mods:start id="F8024B2319268B7A51EB82C0303CF918">451</mods:start>
<mods:end id="D99D79E903C841FE51E46604E2E029F6">485</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D08D7B5CE9E07B1B42E8668A9A434690">journal article</mods:classification>
<mods:identifier id="29BA379BF568BCE6A5BA682BCC047B17" type="CLB-Dataset">55842</mods:identifier>
<mods:identifier id="457A55F00666A6D36414676CDCC7F4B8" type="DOI">10.11646/zootaxa.5128.4.1</mods:identifier>
<mods:identifier id="934E81959BEE5FDBEE5B38F3BD1013AE" type="GBIF-Dataset">6793d5d0-7fcd-415a-8bba-2867cf76501c</mods:identifier>
<mods:identifier id="1A338FB9E05FA2479824426AFCC7E13E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3C74D68EB4C3A98C90976069DC50AAB5" type="Zenodo-Dep">6479917</mods:identifier>
<mods:identifier id="845F66917F5AA9D3962062060E74F9A1" type="ZooBank">26CA4EF0-61A1-4B60-8C7F-99CA8B8F8700</mods:identifier>
</mods:mods>
<treatment id="982416653C1C4B39FF17BAEEFC99403F" ID-DOI="http://doi.org/10.5281/zenodo.6499258" ID-GBIF-Taxon="194848065" ID-Zenodo-Dep="6499258" LSID="urn:lsid:plazi:treatment:982416653C1C4B39FF17BAEEFC99403F" httpUri="http://treatment.plazi.org/id/982416653C1C4B39FF17BAEEFC99403F" lastPageId="29" lastPageNumber="480" pageId="23" pageNumber="474">
<subSubSection id="5897F4F83C1C4B33FF17BAEEFDD2424B" box="[151,545,1519,1546]" pageId="23" pageNumber="474" type="nomenclature">
<paragraph id="1032A7733C1C4B33FF17BAEEFDD2424B" blockId="23.[151,545,1519,1581]" box="[151,545,1519,1546]" pageId="23" pageNumber="474">
<heading id="4B7A101F3C1C4B33FF17BAEEFDD2424B" bold="true" box="[151,545,1519,1546]" fontSize="11" level="1" pageId="23" pageNumber="474" reason="1">
<emphasis id="22F97B613C1C4B33FF17BAEEFDD2424B" bold="true" box="[151,545,1519,1546]" pageId="23" pageNumber="474">
<taxonomicName id="D78DDCF03C1C4B33FF17BAEEFE2B4248" ID-CoL="8ZF29" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[151,472,1519,1546]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="23" pageNumber="474" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C1C4B33FF17BAEEFE2B4248" bold="true" box="[151,472,1519,1546]" italics="true" pageId="23" pageNumber="474">Galapagomystides patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C1C4B33FE5FBAEEFDD2424B" box="[479,545,1519,1545]" pageId="23" pageNumber="474" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5897F4F83C1C4B33FF17B912FECB426F" box="[151,312,1555,1581]" pageId="23" pageNumber="474" type="description">
<paragraph id="1032A7733C1C4B33FF17B912FECB426F" blockId="23.[151,545,1519,1581]" box="[151,312,1555,1581]" pageId="23" pageNumber="474">
<figureCitation id="88B6BBF63C1C4B33FF17B912FEE3426F" box="[151,272,1555,1581]" captionStart="FIGURE 19" captionStartId="26.[151,250,1802,1826]" captionTargetBox="[177,1411,181,1778]" captionTargetId="figure-25@26.[177,1411,181,1778]" captionTargetPageId="26" captionText="FIGURE 19. Light microscopy (LM) images of Galapagomystides patricki n. sp. A—in situ Galapagomystides inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC A13420; A13420). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419).Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="23" pageNumber="474">Figures 19</figureCitation>
,
<figureCitation id="88B6BBF63C1C4B33FE9CB912FECB426F" box="[284,312,1555,1581]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="23" pageNumber="474">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5897F4F83C1C4B33FF17B95AFF3E42DB" pageId="23" pageNumber="474" type="diagnosis">
<paragraph id="1032A7733C1C4B33FF17B95AFF3E42DB" blockId="23.[151,1437,1627,2014]" pageId="23" pageNumber="474">
<emphasis id="22F97B613C1C4B33FF17B95AFEE14237" bold="true" box="[151,274,1627,1653]" pageId="23" pageNumber="474">Diagnosis.</emphasis>
First segment not fused to prostomium. Elongated dorsal cirri on segments 1 and 2. No elongated ventral cirri.
</paragraph>
</subSubSection>
<subSubSection id="5897F4F83C1C4B33FF47B9A2FDFC4333" pageId="23" pageNumber="474" type="materials_examined">
<paragraph id="1032A7733C1C4B33FF47B9A2FDFC4333" blockId="23.[151,1437,1627,2014]" pageId="23" pageNumber="474">
<emphasis id="22F97B613C1C4B33FF47B9A2FE4942FF" bold="true" box="[199,442,1699,1725]" pageId="23" pageNumber="474">Material Examined.</emphasis>
<materialsCitation id="A0E5AD2E3C1C4B33FE44B9A2FB6942A3" ID-GBIF-Occurrence="3756545323" accessionNumber="A13419, MZ711314" collectingDate="2009-03-01" collectionCode="SIO-BIC, SEM, MZ" country="Costa Rica" elevation="1415" latitude="9.0303" location="Parrita Seep" longLatPrecision="54" longitude="-84.623" pageId="23" pageNumber="474" specimenCount="1" typeStatus="holotype">
<emphasis id="22F97B613C1C4B33FE44B9A2FDC942FF" bold="true" box="[452,570,1699,1725]" pageId="23" pageNumber="474">
<typeStatus id="CF3619D13C1C4B33FE44B9A2FDC642FF" box="[452,565,1699,1725]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="769C3FB63C1C4B33FDC4B9A2FD5A42FF" box="[580,681,1699,1725]" pageId="23" pageNumber="474">SIO-BIC</collectionCode>
A13419* (prepared for
<collectionCode id="769C3FB63C1C4B33FC45B9A2FBF042FF" box="[965,1027,1699,1725]" pageId="23" pageNumber="474">SEM</collectionCode>
),
<location id="1552F1A83C1C4B33FB99B9A2FB5542FF" LSID="urn:lsid:plazi:treatment:982416653C1C4B39FF17BAEEFC99403F:1552F1A83C1C4B33FB99B9A2FB5542FF" box="[1049,1190,1699,1725]" country="Costa Rica" latitude="9.0303" longLatPrecision="54" longitude="-84.623" name="Parrita Seep" pageId="23" pageNumber="474">Parrita Seep</location>
,
<collectingCountry id="689AE7E33C1C4B33FB35B9A2FAC542FF" box="[1205,1334,1699,1725]" name="Costa Rica" pageId="23" pageNumber="474">Costa Rica</collectingCountry>
;
<geoCoordinate id="75B9C1B43C1C4B33FAC4B9A2FF5842A0" degrees="9.0303" direction="north" orientation="latitude" pageId="23" pageNumber="474" precision="5" value="9.0303">9.0303° N</geoCoordinate>
<geoCoordinate id="75B9C1B43C1C4B33FF35B9C6FEC142A0" box="[181,306,1735,1762]" degrees="84.623" direction="west" orientation="longitude" pageId="23" pageNumber="474" precision="55" value="-84.623">84.623° W</geoCoordinate>
, ~
<elevation id="9BA040403C1C4B33FECFB9C6FE0142A3" box="[335,498,1735,1762]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="23" pageNumber="474" unit="m" value="1415.0">
<quantity id="D7750A963C1C4B33FECFB9C6FE5442A0" box="[335,423,1735,1762]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="23" pageNumber="474" unit="m" value="1415.0">1415 m</quantity>
depth
</elevation>
,
<date id="643381B33C1C4B33FD80B9C6FD5D42A3" box="[512,686,1735,1761]" pageId="23" pageNumber="474" value="2009-03-01">
<collectingDate id="7477785B3C1C4B33FD80B9C6FD5D42A3" box="[512,686,1735,1761]" pageId="23" pageNumber="474" value="2009-03-01">March 1, 2009</collectingDate>
</date>
, HOV Alvin. [Genbank COI=
<accessionNumber id="0FDE3A903C1C4B33FB94B9C7FB6742A3" box="[1044,1172,1734,1761]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MZ711314" pageId="23" pageNumber="474">
<collectionCode id="769C3FB63C1C4B33FB94B9C7FBCD42A3" box="[1044,1086,1734,1761]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="23" pageNumber="474">MZ</collectionCode>
711314
</accessionNumber>
]
</materialsCitation>
<materialsCitation id="A0E5AD2E3C1C4B33FB24B9C6FB404347" ID-GBIF-Occurrence="3756545307" accessionNumber="A13425, A13426, A13427, A13420, A13421" collectionCode="SIO-BIC" country="Costa Rica" elevation="1400" location="Parrita Seep" pageId="23" pageNumber="474" specimenCount="1" typeStatus="paratype">
<emphasis id="22F97B613C1C4B33FB24B9C6FAD642A3" bold="true" box="[1188,1317,1735,1761]" pageId="23" pageNumber="474">
<typeStatus id="CF3619D13C1C4B33FB24B9C6FAD342A3" box="[1188,1312,1735,1761]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="769C3FB63C1C4B33FAAFB9C6FA6A42A3" box="[1327,1433,1735,1761]" pageId="23" pageNumber="474">SIO-BIC</collectionCode>
: A13425, A13426, A13427,
<accessionNumber id="0FDE3A903C1C4B33FE5BB9EAFDC94347" box="[475,570,1771,1797]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13420" pageId="23" pageNumber="474" type="EnaNcbi">A13420</accessionNumber>
*, A13421*,
<location id="1552F1A83C1C4B33FD50B9EAFC934347" LSID="urn:lsid:plazi:treatment:982416653C1C4B39FF17BAEEFC99403F:1552F1A83C1C4B33FD50B9EAFC934347" box="[720,864,1771,1797]" country="Costa Rica" name="Parrita Seep" pageId="23" pageNumber="474">Parrita Seep</location>
,
<collectingCountry id="689AE7E33C1C4B33FCF0B9EAFC1C4347" box="[880,1007,1771,1797]" name="Costa Rica" pageId="23" pageNumber="474">Costa Rica</collectingCountry>
~
<elevation id="9BA040403C1C4B33FB8AB9EAFB404347" box="[1034,1203,1771,1798]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="474" unit="m" value="1400.0">
<quantity id="D7750A963C1C4B33FB8AB9EAFB964344" box="[1034,1125,1771,1798]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="474" unit="m" value="1400.0">1400 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="A0E5AD2E3C1C4B33FB43B9EAFC1E436B" ID-GBIF-Occurrence="3756545303" collectionCode="SIO-BIC" country="Costa Rica" elevation="850" location="Parrita Seep" pageId="23" pageNumber="474" specimenCode="A9964" specimenCount="1" typeStatus="paratype">
<collectionCode id="769C3FB63C1C4B33FB43B9EAFADE4347" box="[1219,1325,1771,1797]" pageId="23" pageNumber="474">SIO-BIC</collectionCode>
:
<specimenCode id="402B0F083C1C4B33FABBB9EAFA7F4347" box="[1339,1420,1771,1797]" collectionCode="SIO-BIC" pageId="23" pageNumber="474">A9964</specimenCode>
*, A9875*, A9877*, A9961*, A9934,
<location id="1552F1A83C1C4B33FDADB80EFD4A436B" LSID="urn:lsid:plazi:treatment:982416653C1C4B39FF17BAEEFC99403F:1552F1A83C1C4B33FDADB80EFD4A436B" box="[557,697,1807,1833]" country="Costa Rica" name="Parrita Seep" pageId="23" pageNumber="474">Parrita Seep</location>
,
<collectingCountry id="689AE7E33C1C4B33FD46B80EFCB1436B" box="[710,834,1807,1833]" name="Costa Rica" pageId="23" pageNumber="474">Costa Rica</collectingCountry>
~
<elevation id="9BA040403C1C4B33FCD9B80EFC1E436B" box="[857,1005,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="23" pageNumber="474" unit="m" value="850.0">
<quantity id="D7750A963C1C4B33FCD9B80EFC504368" box="[857,931,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="23" pageNumber="474" unit="m" value="850.0">850 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="A0E5AD2E3C1C4B33FC7AB80EFE75430F" ID-GBIF-Occurrence="3756545311" collectionCode="MZUCR" country="Costa Rica" elevation="1770" location="Jaco Scar" pageId="23" pageNumber="474" specimenCode="MZUCR 1510-01" specimenCount="1" typeStatus="paratype">
<specimenCode id="402B0F083C1C4B33FC7AB80EFB3A4368" box="[1018,1225,1807,1834]" collectionCode="MZUCR" pageId="23" pageNumber="474">MZUCR 1510-01</specimenCode>
*,
<location id="1552F1A83C1C4B33FB61B80EFABC436B" LSID="urn:lsid:plazi:treatment:982416653C1C4B39FF17BAEEFC99403F:1552F1A83C1C4B33FB61B80EFABC436B" box="[1249,1359,1807,1833]" country="Costa Rica" name="Jaco Scar" pageId="23" pageNumber="474">Jacó Scar</location>
,
<collectingCountry id="689AE7E33C1C4B33FADCB80EFF3E430F" name="Costa Rica" pageId="23" pageNumber="474">Costa Rica</collectingCountry>
, ~
<elevation id="9BA040403C1C4B33FF67B832FE75430F" box="[231,390,1843,1870]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="23" pageNumber="474" unit="m" value="1770.0">
<quantity id="D7750A963C1C4B33FF67B832FECE430C" box="[231,317,1843,1870]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="23" pageNumber="474" unit="m" value="1770.0">1770 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="A0E5AD2E3C1C4B33FE11B832FBC7430F" ID-GBIF-Occurrence="3756545301" box="[401,1076,1843,1870]" collectionCode="SIO-BIC" country="Costa Rica" elevation="805" location="Parrita Seep" pageId="23" pageNumber="474" specimenCode="A1424" specimenCount="1" typeStatus="paratype">
<collectionCode id="769C3FB63C1C4B33FE11B832FE08430F" box="[401,507,1843,1869]" pageId="23" pageNumber="474">SIO-BIC</collectionCode>
:
<specimenCode id="402B0F083C1C4B33FD84B832FDA0430F" box="[516,595,1843,1869]" collectionCode="SIO-BIC" pageId="23" pageNumber="474">A1424</specimenCode>
, ~
<quantity id="D7750A963C1C4B33FDECB832FD46430F" box="[620,693,1843,1869]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="474" unit="km" value="20.0">20 km</quantity>
away
<location id="1552F1A83C1C4B33FD7EB832FC7A430F" LSID="urn:lsid:plazi:treatment:982416653C1C4B39FF17BAEEFC99403F:1552F1A83C1C4B33FD7EB832FC7A430F" box="[766,905,1843,1869]" country="Costa Rica" name="Parrita Seep" pageId="23" pageNumber="474">Parrita Seep</location>
, ~
<elevation id="9BA040403C1C4B33FC23B832FBC7430F" box="[931,1076,1843,1870]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="23" pageNumber="474" unit="m" value="805.0">
<quantity id="D7750A963C1C4B33FC23B832FC18430C" box="[931,1003,1843,1870]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="23" pageNumber="474" unit="m" value="805.0">805 m</quantity>
depth
</elevation>
</materialsCitation>
. For locality details see
<tableCitation id="5D0F92C83C1C4B33FAC2B832FA65430F" box="[1346,1430,1843,1869]" captionStart="TABLE 1" captionStartId="2.[127,215,464,490]" captionTargetPageId="2" captionText="TABLE 1. Voucher name, origin, locational coordinate, collection depth, date of collection and GenBank accession numbers of sequenced vouchers and types for COI, 16S rRNA, 18S rRNA, 28S rRNA. Note that Protomystides verenae is referred to as Galapagomystides verenae n. comb.. * indicates holotype." httpUri="http://table.plazi.org/id/44F2F7FB3C094B26FFFFBED1FA0A464C" pageId="23" pageNumber="474" tableUuid="44F2F7FB3C094B26FFFFBED1FA0A464C">Table 1</tableCitation>
. * indicates sequenced specimens.
</paragraph>
</subSubSection>
<subSubSection id="5897F4F83C1C4B39FF47B87AFCCE46B3" lastPageId="29" lastPageNumber="480" pageId="23" pageNumber="474" type="description">
<paragraph id="1032A7733C1C4B39FF47B87AFB11467F" blockId="23.[151,1437,1627,2014]" lastBlockId="29.[151,1437,150,1149]" lastPageId="29" lastPageNumber="480" pageId="23" pageNumber="474">
<emphasis id="22F97B613C1C4B33FF47B87AFEAA43D7" bold="true" box="[199,345,1915,1941]" pageId="23" pageNumber="474">Description.</emphasis>
<typeStatus id="CF3619D13C1C4B33FEDFB87AFE3443D7" box="[351,455,1915,1941]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
body length
<quantity id="D7750A963C1C4B33FDD7B87AFD5443D4" box="[599,679,1915,1942]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="23" pageNumber="474" unit="mm" value="16.0">16 mm</quantity>
,
<quantity id="D7750A963C1C4B33FD32B87AFD1F43D7" box="[690,748,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="474" unit="mm" value="1.0">1mm</quantity>
wide at segment 10 for ~70 segments. Body red or white with red gut in life (
<figureCitation id="88B6BBF63C1C4B33FEC3B89EFE5F43FB" box="[323,428,1951,1977]" captionStart="FIGURE 19" captionStartId="26.[151,250,1802,1826]" captionTargetBox="[177,1411,181,1778]" captionTargetId="figure-25@26.[177,1411,181,1778]" captionTargetPageId="26" captionText="FIGURE 19. Light microscopy (LM) images of Galapagomystides patricki n. sp. A—in situ Galapagomystides inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC A13420; A13420). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419).Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="23" pageNumber="474">Fig. 19A</figureCitation>
). Body brown/orange with numerous dark pigmentation speckles in preserved (formalin/ ethanol) state (
<figureCitation id="88B6BBF63C1C4B33FEBFB8C2FE39439C" box="[319,458,1987,2014]" captionStart="FIGURE 19" captionStartId="26.[151,250,1802,1826]" captionTargetBox="[177,1411,181,1778]" captionTargetId="figure-25@26.[177,1411,181,1778]" captionTargetPageId="26" captionText="FIGURE 19. Light microscopy (LM) images of Galapagomystides patricki n. sp. A—in situ Galapagomystides inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC A13420; A13420). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419).Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="23" pageNumber="474">Fig. 19C, D</figureCitation>
). Lobe-like prostomium; nuchal organs not visible (
<figureCitation id="88B6BBF63C1C4B33FB97B8C2FB72439F" box="[1047,1153,1987,2013]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="23" pageNumber="474">Fig. 20A</figureCitation>
). Anterior dorsal edge of A A prostomium with paired cylindrical antennae (~
<quantity id="D7750A963C164B39FD2ABF97FCE344F3" box="[682,784,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="480" unit="mm" value="0.25">0.25 mm</quantity>
long) (
<figureCitation id="88B6BBF63C164B39FCE0BF96FC3A44F3" box="[864,969,151,177]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20A</figureCitation>
). Paired palps ventral to antennae, similar in shape and length to antennae (
<figureCitation id="88B6BBF63C164B39FD80BFBAFD7B4497" box="[512,648,187,213]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20A, D</figureCitation>
). Segment one not fused to prostomium, distinct dorsally and ventrally, following segments clearly demarcated (
<figureCitation id="88B6BBF63C164B39FDE8BFDEFD0644BB" box="[616,757,223,249]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20A, C</figureCitation>
). Pair of elongated dorsal cirri [tentacular cirri] on each of segments 1 (~
<quantity id="D7750A963C164B39FEB5BE02FE7C455F" box="[309,399,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="480" unit="mm" value="0.4">0.4 mm</quantity>
long), 2 (~
<quantity id="D7750A963C164B39FD8FBE03FD84455F" box="[527,631,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="29" pageNumber="480" unit="mm" value="0.35">0.35 mm</quantity>
long) (
<figureCitation id="88B6BBF63C164B39FD4CBE02FCC5455F" box="[716,822,259,285]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20A</figureCitation>
). All elongated dorsal cirri cirriform, tapering distally (
<figureCitation id="88B6BBF63C164B39FF20BE26FEF94503" box="[160,266,295,321]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20A</figureCitation>
). Dorsal elongated cirri on segment one, right side and dorsal elongated cirri on segment two, left side were broken off the
<typeStatus id="CF3619D13C164B39FE07BE4AFE1E4527" box="[391,493,331,357]" pageId="29" pageNumber="480" type="holotype">holotype</typeStatus>
. No elongated ventral cirri (
<figureCitation id="88B6BBF63C164B39FCB7BE4AFC574527" box="[823,932,331,357]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20D</figureCitation>
). Segment 3 lacking dorsal cirri (
<figureCitation id="88B6BBF63C164B39FAAABE4AFF5F45CB" captionStart="FIGURE 19" captionStartId="26.[151,250,1802,1826]" captionTargetBox="[177,1411,181,1778]" captionTargetId="figure-25@26.[177,1411,181,1778]" captionTargetPageId="26" captionText="FIGURE 19. Light microscopy (LM) images of Galapagomystides patricki n. sp. A—in situ Galapagomystides inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC A13420; A13420). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419).Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="29" pageNumber="480">Figs 19B, C</figureCitation>
,
<figureCitation id="88B6BBF63C164B39FF3BBE6EFF0345CB" box="[187,240,367,393]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">20A</figureCitation>
). Bulbous, rounded dorsal cirri (~
<quantity id="D7750A963C164B39FDFCBE6EFD2445CB" box="[636,727,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="480" unit="mm" value="0.1">0.1 mm</quantity>
long) begin on segment 4 continuing posteriorly (
<figureCitation id="88B6BBF63C164B39FAA5BE6EFA6345CB" box="[1317,1424,367,393]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20C</figureCitation>
). Ventral cirri absent on segment 1 (
<figureCitation id="88B6BBF63C164B39FDAEBE92FD6845EF" box="[558,667,403,429]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20D</figureCitation>
). Bulbous ventral cirri (~
<quantity id="D7750A963C164B39FC42BE92FBEE45EF" box="[962,1053,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="480" unit="mm" value="0.1">0.1 mm</quantity>
long) (
<figureCitation id="88B6BBF63C164B39FBF6BE92FB1045EF" box="[1142,1251,403,429]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20D</figureCitation>
). Chaetae begin on segment 2 continuing posteriorly (
<figureCitation id="88B6BBF63C164B39FDCABEB6FD454593" box="[586,694,439,465]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20A</figureCitation>
). Parapodia uniramous, notopodial chaetae absent; neuropodium with central fascicle containing ~610 compound chaetae; one simple emergent acicula (
<figureCitation id="88B6BBF63C164B39FB06BEDAFAA745B7" box="[1158,1364,475,501]" captionStart="FIGURE 19" captionStartId="26.[151,250,1802,1826]" captionTargetBox="[177,1411,181,1778]" captionTargetId="figure-25@26.[177,1411,181,1778]" captionTargetPageId="26" captionText="FIGURE 19. Light microscopy (LM) images of Galapagomystides patricki n. sp. A—in situ Galapagomystides inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC A13420; A13420). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419).Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="29" pageNumber="480">Figs 19E, F, G, H</figureCitation>
,
<figureCitation id="88B6BBF63C164B39FAE0BEDAFA6345B7" box="[1376,1424,475,501]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">20F</figureCitation>
). Compound chaetal shaft cylindrical; thin, flattened triangular short blade extending from rounded joint (
<figureCitation id="88B6BBF63C164B39FAAFBEFEFA64465B" box="[1327,1431,511,537]" captionStart="FIGURE 19" captionStartId="26.[151,250,1802,1826]" captionTargetBox="[177,1411,181,1778]" captionTargetId="figure-25@26.[177,1411,181,1778]" captionTargetPageId="26" captionText="FIGURE 19. Light microscopy (LM) images of Galapagomystides patricki n. sp. A—in situ Galapagomystides inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC A13420; A13420). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419).Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="29" pageNumber="480">Figs 19E</figureCitation>
,
<figureCitation id="88B6BBF63C164B39FF17BD22FF34467F" box="[151,199,547,573]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">20F</figureCitation>
). Pygidium with one pair of lobed pygidial cirri (~
<quantity id="D7750A963C164B39FD7BBD23FC92467F" box="[763,865,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="480" unit="mm" value="0.15">0.15 mm</quantity>
long) rounded distally (
<figureCitation id="88B6BBF63C164B39FBEEBD22FB26467F" box="[1134,1237,547,573]" captionStart="FIGURE 20" captionStartId="27.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@27.[151,1435,181,1844]" captionTargetPageId="27" captionText="FIGURE 20. Scanning electron microscopy (SEM) of Galapagomystides patricki n. sp. holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="29" pageNumber="480">Fig. 20B</figureCitation>
).
</paragraph>
<caption id="44F2F7FB3C134B3CFF17B84AFBB143B6" ID-DOI="http://doi.org/10.5281/zenodo.6479953" ID-Zenodo-Dep="6479953" httpUri="https://zenodo.org/record/6479953/files/figure.png" pageId="24" pageNumber="475" startId="24.[151,250,1867,1891]" targetBox="[151,1435,181,1844]" targetPageId="24">
<paragraph id="1032A7733C134B3CFF17B84AFBB143B6" blockId="24.[151,1437,1867,2036]" pageId="24" pageNumber="475">
<emphasis id="22F97B613C134B3CFF17B84AFEEC4326" bold="true" box="[151,287,1867,1892]" pageId="24" pageNumber="475">FIGURE 17.</emphasis>
Light microscopy (LM) images of
<taxonomicName id="D78DDCF03C134B3CFDFEB84DFC7C4326" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[638,911,1868,1892]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="24" pageNumber="475" phylum="Annelida" rank="species" species="kathyae" status="sp. nov.">
<emphasis id="22F97B613C134B3CFDFEB84DFC7C4326" box="[638,911,1868,1892]" italics="true" pageId="24" pageNumber="475">Galapagomystides kathyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C134B3CFC14B84DFC3A4321" box="[916,969,1868,1891]" pageId="24" pageNumber="475" rank="species">n. sp.</taxonomicNameLabel>
A—dorsal view, holotype (SIO-BIC
<accessionNumber id="0FDE3A903C134B3CFABCB84DFA624326" box="[1340,1425,1868,1892]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13418" pageId="24" pageNumber="475" type="EnaNcbi">A13418</accessionNumber>
). B—ventral view of holotype, holotype (SIO-BIC
<accessionNumber id="0FDE3A903C134B3CFD0CB871FD1243CA" box="[652,737,1904,1928]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13418" pageId="24" pageNumber="475" type="EnaNcbi">A13418</accessionNumber>
). C—live dorsal view, holotype (SIO-BIC
<accessionNumber id="0FDE3A903C134B3CFB0FB871FB1643CA" box="[1167,1253,1904,1928]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13418" pageId="24" pageNumber="475" type="EnaNcbi">A13418</accessionNumber>
). D—whole body, paratype (SIO-BIC A4651). E—drawing of compound chaetae, holotype (SIO-BIC
<accessionNumber id="0FDE3A903C134B3CFC74B895FBB943EE" box="[1012,1098,1940,1964]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13418" pageId="24" pageNumber="475" type="EnaNcbi">A13418</accessionNumber>
). F—dorsal cirri, holotype (SIO-BIC
<accessionNumber id="0FDE3A903C134B3CFF48B8B9FEEE4392" box="[200,285,1976,2000]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13418" pageId="24" pageNumber="475" type="EnaNcbi">A13418</accessionNumber>
). G—parapodia, holotype (SIO-BIC
<accessionNumber id="0FDE3A903C134B3CFD18B8B9FD1E4392" box="[664,749,1976,2000]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13418" pageId="24" pageNumber="475" type="EnaNcbi">A13418</accessionNumber>
). Abbreviations: a, antennae; dc, dorsal cirri; edc, elongated dorsal cirri; evc, elongated ventral cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium.
</paragraph>
</caption>
<caption id="44F2F7FB3C124B3DFF17B84AFA684392" ID-DOI="http://doi.org/10.5281/zenodo.6479955" ID-Zenodo-Dep="6479955" httpUri="https://zenodo.org/record/6479955/files/figure.png" pageId="25" pageNumber="476" startId="25.[151,250,1867,1891]" targetBox="[151,1435,181,1844]" targetPageId="25">
<paragraph id="1032A7733C124B3DFF17B84AFA684392" blockId="25.[151,1437,1867,2000]" pageId="25" pageNumber="476">
<emphasis id="22F97B613C124B3DFF17B84AFED04326" bold="true" box="[151,291,1867,1892]" pageId="25" pageNumber="476">FIGURE 18.</emphasis>
Scanning electron microscopy (SEM) of
<taxonomicName id="D78DDCF03C124B3DFD5AB84DFC1C4326" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[730,1007,1868,1892]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="25" pageNumber="476" phylum="Annelida" rank="species" species="kathyae" status="sp. nov.">
<emphasis id="22F97B613C124B3DFD5AB84DFC1C4326" box="[730,1007,1868,1892]" italics="true" pageId="25" pageNumber="476">Galapagomystides kathyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C124B3DFC78B84DFBC14321" box="[1016,1074,1868,1891]" pageId="25" pageNumber="476" rank="species">n. sp.</taxonomicNameLabel>
holotype (SIO-BIC
<accessionNumber id="0FDE3A903C124B3DFA88B84DFAAD4326" box="[1288,1374,1868,1892]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13418" pageId="25" pageNumber="476" type="EnaNcbi">A13418</accessionNumber>
). A— close-up dorsal view of anterior and prostomium. B—ventral view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; ci, cilia; dc, dorsal cirri; edc, elongated dorsal cirri; evc, elongated ventral cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium.
</paragraph>
</caption>
<caption id="44F2F7FB3C114B3EFF17B80BFE1A43B8" ID-DOI="http://doi.org/10.5281/zenodo.6479957" ID-Zenodo-Dep="6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="26" pageNumber="477" startId="26.[151,250,1802,1826]" targetBox="[177,1411,181,1778]" targetPageId="26">
<paragraph id="1032A7733C114B3EFF17B80BFE1A43B8" blockId="26.[151,1437,1802,2042]" pageId="26" pageNumber="477">
<emphasis id="22F97B613C114B3EFF17B80BFED54360" bold="true" box="[151,294,1802,1826]" pageId="26" pageNumber="477">FIGURE 19.</emphasis>
Light microscopy (LM) images of
<taxonomicName id="D78DDCF03C114B3EFD30B80BFC3B4360" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[688,968,1802,1826]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="26" pageNumber="477" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C114B3EFD30B80BFC3B4360" box="[688,968,1802,1826]" italics="true" pageId="26" pageNumber="477">Galapagomystides patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C114B3EFC54B80BFBE34363" box="[980,1040,1802,1825]" pageId="26" pageNumber="477" rank="species">n. sp.</taxonomicNameLabel>
A—
<emphasis id="22F97B613C114B3EFBC6B80AFAA74360" box="[1094,1364,1802,1826]" italics="true" pageId="26" pageNumber="477">
in situ
<taxonomicName id="D78DDCF03C114B3EFB17B80BFAA74360" authorityName="Blake" authorityYear="1985" box="[1175,1364,1802,1826]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="26" pageNumber="477" phylum="Annelida" rank="genus">Galapagomystides</taxonomicName>
</emphasis>
inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC
<accessionNumber id="0FDE3A903C114B3EFB2CB82FFAF24304" box="[1196,1281,1838,1862]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13420" pageId="26" pageNumber="477" type="EnaNcbi">A13420</accessionNumber>
;
<accessionNumber id="0FDE3A903C114B3EFA8AB82FFA924304" box="[1290,1377,1838,1862]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A13420" pageId="26" pageNumber="477" type="EnaNcbi">A13420</accessionNumber>
). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419). Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium.
</paragraph>
</caption>
<caption id="44F2F7FB3C104B3FFF17B84AFBC94392" ID-DOI="http://doi.org/10.5281/zenodo.6479959" ID-Zenodo-Dep="6479959" httpUri="https://zenodo.org/record/6479959/files/figure.png" pageId="27" pageNumber="478" startId="27.[151,250,1867,1891]" targetBox="[151,1435,181,1844]" targetPageId="27">
<paragraph id="1032A7733C104B3FFF17B84AFBC94392" blockId="27.[151,1437,1867,2000]" pageId="27" pageNumber="478">
<emphasis id="22F97B613C104B3FFF17B84AFED04326" bold="true" box="[151,291,1867,1892]" pageId="27" pageNumber="478">FIGURE 20.</emphasis>
Scanning electron microscopy (SEM) of
<taxonomicName id="D78DDCF03C104B3FFD5AB84DFC1C4326" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[730,1007,1868,1892]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="27" pageNumber="478" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C104B3FFD5AB84DFC1C4326" box="[730,1007,1868,1892]" italics="true" pageId="27" pageNumber="478">Galapagomystides patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C104B3FFC78B84DFBC14321" box="[1016,1074,1868,1891]" pageId="27" pageNumber="478" rank="species">n. sp.</taxonomicNameLabel>
holotype (SIO-BIC A13419). A— close-up dorsal view of anterior and prostomium. B—dorsal view of pygidium with pygidial cirri. C—dorsal view of anterior. D—ventral view of anterior. E—parapodia. F—compound chaetae. Abbreviations: a, antennae; bo, buccal organ; dc, dorsal cirri; edc, elongated dorsal cirri; vc, ventral cirri; p, palps; pro, prostomium; pyg, pygidium.
</paragraph>
</caption>
<caption id="44F2F7FB3C174B38FF17B84EFEDE43BA" ID-DOI="http://doi.org/10.5281/zenodo.6479961" ID-Zenodo-Dep="6479961" httpUri="https://zenodo.org/record/6479961/files/figure.png" pageId="28" pageNumber="479" startId="28.[151,250,1871,1895]" targetBox="[157,1436,181,1791]" targetPageId="28">
<paragraph id="1032A7733C174B38FF17B84EFEDE43BA" blockId="28.[151,1437,1871,2040]" pageId="28" pageNumber="479">
<emphasis id="22F97B613C174B38FF17B84EFED14325" bold="true" box="[151,290,1871,1895]" pageId="28" pageNumber="479">FIGURE 21.</emphasis>
Maximum Likelihood (ML) tree rendered from the concatenation of genes COI, 18S, 28S and 16S including outgroups (rooted with
<taxonomicName id="D78DDCF03C174B38FE06B875FDA643CE" authorityName="Audouin &amp; Milne Edwards" authorityYear="1834" box="[390,597,1908,1932]" class="Polychaeta" family="Lumbrineridae" genus="Lumbrineris" kingdom="Animalia" order="Eunicida" pageId="28" pageNumber="479" phylum="Annelida" rank="species" species="latreilli">
<emphasis id="22F97B613C174B38FE06B875FDA643CE" box="[390,597,1908,1932]" italics="true" pageId="28" pageNumber="479">Lumbrineris latreilli</emphasis>
</taxonomicName>
), that were not shown in Figure 2. * indicates type species of
<taxonomicName id="D78DDCF03C174B38FB59B875FA6543CE" authorityName="Blake" authorityYear="1985" box="[1241,1430,1908,1932]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="28" pageNumber="479" phylum="Annelida" rank="genus">
<emphasis id="22F97B613C174B38FB59B875FA6543CE" box="[1241,1430,1908,1932]" italics="true" pageId="28" pageNumber="479">Galapagomystides</emphasis>
</taxonomicName>
. Support values are not shown for the parts of the topology seen in Figure 2. Numbers on nodes represent ML bootstrap values (above node) and Bayesian posterior probabilities (below node). Values below 50% for ML and 0.70 for posterior probabilities are not shown.
</paragraph>
</caption>
<paragraph id="1032A7733C164B39FF47BD47FCCE46B3" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="480">
<emphasis id="22F97B613C164B39FF47BD47FECC4622" bold="true" box="[199,319,582,608]" pageId="29" pageNumber="480">Variation.</emphasis>
Most
<taxonomicName id="D78DDCF03C164B39FE03BD46FE094623" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[387,506,583,609]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FE03BD46FE094623" box="[387,506,583,609]" italics="true" pageId="29" pageNumber="480">G. patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FE7FBD46FDC94623" box="[511,570,583,609]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
type material specimens were within
<superScript id="E7F80A3B3C164B39FC57BD46FC134617" attach="right" box="[983,992,583,597]" fontSize="6" pageId="29" pageNumber="480">+</superScript>
/-
<quantity id="D7750A963C164B39FC77BD46FB974623" box="[1015,1124,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="29" pageNumber="480" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 mm</quantity>
from the
<typeStatus id="CF3619D13C164B39FB4CBD46FADD4623" box="[1228,1326,583,609]" pageId="29" pageNumber="480" type="holotype">holotype</typeStatus>
in overall length, though
<specimenCount id="068B6CFA3C164B39FEC6BD6AFE2046C7" box="[326,467,619,645]" pageId="29" pageNumber="480" type="juvenile">one juvenile</specimenCount>
was ~
<quantity id="D7750A963C164B39FDA2BD6AFD8746C7" box="[546,628,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="480" unit="mm" value="10.0">10 mm</quantity>
smaller than the
<typeStatus id="CF3619D13C164B39FCC1BD6AFC5446C7" box="[833,935,619,645]" pageId="29" pageNumber="480" type="holotype">holotype</typeStatus>
. The specimens ranged from ~40 segments to ~70 segments. In life, the
<typeStatus id="CF3619D13C164B39FE69BD8EFDA746EB" box="[489,596,655,681]" pageId="29" pageNumber="480" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="D78DDCF03C164B39FD00BD8EFD0F46EB" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[640,764,655,681]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FD00BD8EFD0F46EB" box="[640,764,655,681]" italics="true" pageId="29" pageNumber="480">G. patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FC86BD8EFCB646EB" box="[774,837,655,681]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
had a body that was semi-translucent white/pink at parapodial lobes, dorsal and ventral cirri, prostomium and pygidium; red in the gut region center with a yellow patch on prostomium, elongated cirri and pygidial cirri (
<figureCitation id="88B6BBF63C164B39FD49BDD6FCDC46B3" box="[713,815,727,753]" captionStart="FIGURE 19" captionStartId="26.[151,250,1802,1826]" captionTargetBox="[177,1411,181,1778]" captionTargetId="figure-25@26.[177,1411,181,1778]" captionTargetPageId="26" captionText="FIGURE 19. Light microscopy (LM) images of Galapagomystides patricki n. sp. A—in situ Galapagomystides inside Vestimentiferan tube, Alvin Dive 4508, Mound Quepos, Parrita Seep, Costa Rica, paratype (SIO-BIC A13420; A13420). B— dorsal view of anterior, paratype (SIO-BIC A1424). C—dorsal view of anterior and prostomium, holotype (SIO-BIC A13419). D—dorsal view of pygidium with pygidial cirri, holotype (SIO-BIC A13419). E—drawing of compound chaetae, holotype (SIO-BIC A13419). F—dorsal cirri, holotype (SIO-BIC A13419). G—aciculum, holotype (SIO-BIC A13419). H—compound cirri, holotype (SIO-BIC A13419).Abbreviations: a, antennae; a, aciculum; dc, dorsal cirri; edc, elongated dorsal cirri; p, palps; pro, prostomium; pyg, pygidium." figureDoi="http://doi.org/10.5281/zenodo.6479957" httpUri="https://zenodo.org/record/6479957/files/figure.png" pageId="29" pageNumber="480">Fig. 19B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5897F4F83C164B39FF47BDFAFDD34074" pageId="29" pageNumber="480" type="discussion">
<paragraph id="1032A7733C164B39FF47BDFAFDD34074" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="480">
<emphasis id="22F97B613C164B39FF47BDFAFEC84757" bold="true" box="[199,315,763,789]" pageId="29" pageNumber="480">Remarks.</emphasis>
Most
<taxonomicName id="D78DDCF03C164B39FE05BDFAFE0C4757" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[389,511,763,789]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FE05BDFAFE0C4757" box="[389,511,763,789]" italics="true" pageId="29" pageNumber="480">G. patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FD87BDFAFDB74757" box="[519,580,763,789]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
samples were collected from Parrita Seep,
<collectingCountry id="689AE7E33C164B39FBB0BDFAFB584757" box="[1072,1195,763,789]" name="Costa Rica" pageId="29" pageNumber="480">Costa Rica</collectingCountry>
(
<quantity id="D7750A963C164B39FB3BBDFAFAE64757" box="[1211,1301,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="480" unit="m" value="1400.0">1400 m</quantity>
), except for
<specimenCount id="068B6CFA3C164B39FF17BC1EFEC0477B" box="[151,307,799,825]" pageId="29" pageNumber="480" type="generic">one specimen</specimenCount>
collected from the deeper Jacó Scar,
<collectingCountry id="689AE7E33C164B39FD64BC1EFCAC477B" box="[740,863,799,825]" name="Costa Rica" pageId="29" pageNumber="480">Costa Rica</collectingCountry>
(
<quantity id="D7750A963C164B39FCF1BC1EFC3E477B" box="[881,973,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="480" unit="m" value="1800.0">1800 m</quantity>
). A single sample, A1424, was collected from a multicore that was deployed offsite in the vicinity of the seep. Some
<taxonomicName id="D78DDCF03C164B39FC62BC42FBA8471F" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[994,1115,835,861]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FC62BC42FBA8471F" box="[994,1115,835,861]" italics="true" pageId="29" pageNumber="480">G. patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FBE1BC42FB6E471F" box="[1121,1181,835,861]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
specimens were found in sediment tubes inside empty vestimentiferan tubes. Morphologically,
<taxonomicName id="D78DDCF03C164B39FC4BBC66FBB547C3" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[971,1094,871,897]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FC4BBC66FBB547C3" box="[971,1094,871,897]" italics="true" pageId="29" pageNumber="480">G. patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FBCFBC66FB7F47C3" box="[1103,1164,871,897]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
is most like
<taxonomicName id="D78DDCF03C164B39FAA1BC66FA6E47C2" authorityName="Pearson &amp; Rouse" authorityYear="2022" baseAuthorityName="Blake and Hilbig" baseAuthorityYear="1990" box="[1313,1437,871,897]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="verenae" status="comb. nov.">
<emphasis id="22F97B613C164B39FAA1BC66FA6E47C2" box="[1313,1437,871,897]" italics="true" pageId="29" pageNumber="480">G. verenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FF17BC8AFEE147E7" box="[151,274,907,933]" pageId="29" pageNumber="480" rank="species">new comb.</taxonomicNameLabel>
as both species show no fusion of anterior segments with the prostomium. However, in the phylogenetic analyses,
<taxonomicName id="D78DDCF03C164B39FE85BCAEFE8C478B" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[261,383,943,969]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FE85BCAEFE8C478B" box="[261,383,943,969]" italics="true" pageId="29" pageNumber="480">G. patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FE08BCAEFE36478B" box="[392,453,943,969]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
was the sister group to the rest of the genus (
<figureCitation id="88B6BBF63C164B39FC4CBCAEFBE1478B" box="[972,1042,943,969]" captionStart="FIGURE 2" captionStartId="7.[151,250,1907,1931]" captionTargetBox="[181,1408,279,1879]" captionTargetId="figure-61@7.[177,1411,273,1884]" captionTargetPageId="7" captionText="FIGURE 2. Maximum Likelihood (ML) tree generated from the concatenation of mitochondrial (COI 16rRNA) and nuclear genes (18S rRNA, 28S rRNA). Numbers on nodes represent ML bootstrap values (above node) and Bayesian posterior probabilities (below node). Values below 50% for ML and 0.70 for posterior probabilities are not shown. * indicates G. aristata as the type species of Galapagomystides." figureDoi="http://doi.org/10.5281/zenodo.6479921" httpUri="https://zenodo.org/record/6479921/files/figure.png" pageId="29" pageNumber="480">Fig. 2</figureCitation>
).
<taxonomicName id="D78DDCF03C164B39FBA7BCAEFAA4478B" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[1063,1367,943,969]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FBA7BCAEFAA4478B" box="[1063,1367,943,969]" italics="true" pageId="29" pageNumber="480">Galapagomystides patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FADFBCAEFA6F478B" box="[1375,1436,943,969]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
lacks dorsal cirri on segment three (all other species have either elongated or regular dorsal cirri on segment three). The chaetae of
<taxonomicName id="D78DDCF03C164B39FEC1BCF6FE494053" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[321,442,1015,1041]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FEC1BCF6FE494053" box="[321,442,1015,1041]" italics="true" pageId="29" pageNumber="480">G. patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FE40BCF6FE0F4053" box="[448,508,1015,1041]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
had the shortest blades compared to other species and are also triangular (while the blades in other species are falcate).
</paragraph>
</subSubSection>
<subSubSection id="5897F4F83C164B39FF47BB3EFC99403F" pageId="29" pageNumber="480" type="etymology">
<paragraph id="1032A7733C164B39FF47BB3EFC99403F" blockId="29.[151,1437,150,1149]" pageId="29" pageNumber="480">
<emphasis id="22F97B613C164B39FF47BB3EFEA1401B" bold="true" box="[199,338,1087,1113]" pageId="29" pageNumber="480">Etymology:</emphasis>
<taxonomicName id="D78DDCF03C164B39FEDCBB3EFD7E401B" authority="Pearson &amp; Rouse, 2022" authorityName="Pearson &amp; Rouse" authorityYear="2022" box="[348,653,1087,1113]" class="Polychaeta" family="Phyllodocidae" genus="Galapagomystides" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="480" phylum="Annelida" rank="species" species="patricki" status="sp. nov.">
<emphasis id="22F97B613C164B39FEDCBB3EFD7E401B" box="[348,653,1087,1113]" italics="true" pageId="29" pageNumber="480">Galapagomystides patricki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39CAC61A3C164B39FD17BB3EFD25401B" box="[663,726,1087,1113]" pageId="29" pageNumber="480" rank="species">n. sp.</taxonomicNameLabel>
is named after Patrick Shaughnessy, whose love and support during this project was instrumental to the lead authors success.
</paragraph>
</subSubSection>
</treatment>
</document>